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<document id="1F4EE04A2D7C815DFAB0FDD15458C0E7" ID-CLB-Dataset="25281" ID-DOI="10.5852/ejt.2019.563" ID-GBIF-Dataset="4de0fe4e-219b-432f-9d36-43d2f4a41835" ID-Zenodo-Dep="3476748" ID-ZooBank="6F6E082D-0675-49C1-A603-F7BABB546C46" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570549751342" checkinUser="plazi" docAuthor="Onuferko, Thomas M." docDate="2019" docId="201E87ADFF87FFCE1868FBD741828C98" docLanguage="en" docName="ejt-563_onuferko.pdf.imf" docOrigin="European Journal of Taxonomy 563" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Epeolus fumipennis Say 1837" docType="treatment" docVersion="13" lastPageNumber="28" masterDocId="DC27FFD5FF90FFD51A0BFFD24347886C" masterDocTitle="A review of the cleptoparasitic bee genus Epeolus Latreille, 1802 (Hymenoptera: Apidae) in the Caribbean, Central America and Mexico" masterLastPageNumber="69" masterPageNumber="1" pageNumber="24" updateTime="1698758470402" updateUser="ExternalLinkService">
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<mods:title id="6CB9B57C523A6D9CB2843E51EC31D174">A review of the cleptoparasitic bee genus Epeolus Latreille, 1802 (Hymenoptera: Apidae) in the Caribbean, Central America and Mexico</mods:title>
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<mods:namePart id="D9DBB0ABD425626AB17C784C20F36B70">Onuferko, Thomas M.</mods:namePart>
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<treatment id="201E87ADFF87FFCE1868FBD741828C98" ID-DOI="http://doi.org/10.5281/zenodo.3477513" ID-GBIF-Taxon="159660838" ID-Zenodo-Dep="3477513" LSID="urn:lsid:plazi:treatment:201E87ADFF87FFCE1868FBD741828C98" httpUri="http://treatment.plazi.org/id/201E87ADFF87FFCE1868FBD741828C98" lastPageId="27" lastPageNumber="28" pageId="23" pageNumber="24">
<subSubSection id="E0AD6530FF87FFC21868FBD740978C4C" box="[611,976,1029,1056]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="A80836BBFF87FFC21868FBD740978C4C" blockId="23.[611,976,1029,1092]" box="[611,976,1029,1056]" pageId="23" pageNumber="24">
<heading id="F34081D7FF87FFC21868FBD740978C4C" box="[611,976,1029,1056]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="6FB74D38FF87FFC21868FBD740978C4C" ID-CoL="6FNQV" authority="Say, 1837" authorityName="Say" authorityYear="1837" box="[611,976,1029,1056]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF87FFC21868FBD740128C73" bold="true" box="[611,853,1029,1056]" italics="true" pageId="23" pageNumber="24">Epeolus fumipennis</emphasis>
<bibRefCitation id="CC264B4AFF87FFC21956FBD440978C4C" author="Say" box="[861,976,1030,1056]" firstAuthor="Say" pageId="23" pageNumber="24" pagination="361 - 416" refId="ref37138" refString="Say T. 1837. Descriptions of new species of North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1: 361 - 416." type="journal article" year="1837">Say, 1837</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF87FFC21866FBFB40808C2F" box="[621,967,1065,1092]" pageId="23" pageNumber="24" type="description">
<paragraph id="A80836BBFF87FFC21866FBFB40808C2F" blockId="23.[611,976,1029,1092]" box="[621,967,1065,1092]" pageId="23" pageNumber="24">
<figureCitation id="308C2A3EFF87FFC21866FBFB418B8C2F" box="[621,716,1065,1091]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[194,1398,502,1710]" captionTargetId="figure@6.[189,1398,501,1710]" captionTargetPageId="6" captionText="Fig. 1. Male genitalia of Epeolus spp. A. E. boliviensis Friese, 1908, in which the penis lacks lobes. B. E. claripennis Friese, 1908, in which the penis lacks lobes. C. E. fulvopilosus Cameron, 1902, in which the penis lacks lobes. D. E. fumipennis Say, 1837, in which the penis lacks lobes. E. E. hanusiae sp. nov., paratype (CNC 754086), showing the penis with a short, fleshy lateral lobe (blue arrow) on each side. F. E. luteipennis Friese, 1916, showing the penis with a long, fleshy lateral lobe (blue arrow) on each side. G. E. niger (Michener, 1954), in which the penis lacks lobes. H. E. obscuripes Cockerell, 1917 stat. nov., in which the penis lacks lobes. I. E. pulchellus Cresson, 1865, in which the penis lacks lobes. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3476750" httpUri="https://zenodo.org/record/3476750/files/figure.png" pageId="23" pageNumber="24">Figs 1D</figureCitation>
,
<figureCitation id="308C2A3EFF87FFC218D2FBFB41B98C2F" box="[729,766,1065,1091]" captionStart="Fig" captionStartId="7.[189,232,1895,1921]" captionTargetBox="[189,1398,1441,1838]" captionTargetId="figure@7.[189,1398,1441,1847]" captionTargetPageId="7" captionText="Fig. 2. Pronotal collar of female, dorsal view. A. E. boliviensis Friese, 1908, which is straight along its anterior margin. B. E. fumipennis Say, 1837, which is relatively straight along its anterior margin. C. E. hanusiae sp. nov., paratype (UCBME), which is convex along its anterior margin. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3476752" httpUri="https://zenodo.org/record/3476752/files/figure.png" pageId="23" pageNumber="24">2B</figureCitation>
,
<figureCitation id="308C2A3EFF87FFC21901FBFB40688C2F" box="[778,815,1065,1091]" captionStart="Fig" captionStartId="8.[189,232,1537,1563]" captionTargetBox="[189,1398,266,1493]" captionTargetId="figure@8.[189,1398,266,1500]" captionTargetPageId="8" captionText="Fig. 3. Mesosoma of female, posterior view. A. E. fulvopilosus Cameron, 1902, showing mesoscutellum with a distinct ridge (blue arrow) overhanging its depressed posterior margin. B. E. fumipennis Say, 1837, showing mesoscutellum with a faint ridge (blue arrow) overhanging its depressed posterior margin. C. E. pulchellus Cresson, 1865, showing mesoscutellum without a distinct ridge delineating its dorsal and posterior portions. D. E. odyneroides sp. nov., paratype (KUNHM SEMC1248301), showing mesoscutellum without a distinct ridge delineating its dorsal and posterior portions. E. E. axillaris Onuferko, 2018, paratype, showing metanotum with a distinct posteromedial depression (blue arrow). F. E. rufulus Cockerell, 1941, holotype, showing flat metanotum. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3476754" httpUri="https://zenodo.org/record/3476754/files/figure.png" pageId="23" pageNumber="24">3B</figureCitation>
,
<figureCitation id="308C2A3EFF87FFC21930FBF840198C28" box="[827,862,1066,1092]" captionStart="Fig" captionStartId="12.[189,232,1824,1850]" captionTargetId="figure@13.[241,1373,265,2019]" captionTargetPageId="13" captionText="Fig. 7 (opposite page). Occurrence records (yellow circles except where indicated otherwise) of: A. Epeolus boliviensis Friese, 1908. B. E. claripennis Friese, 1908. C. E. danieli (Genaro, 2014) comb. nov. (orange square) and E. luteipennis Friese, 1916. D. E. fulvopilosus Cameron, 1902. E. E. fumipennis Say, 1837. F. E. hanusiae sp. nov. G. E. niger (Michener, 1954). H. E. nomadiformis sp. nov. I. E. obscuripes Cockerell, 1917 stat. nov. J. E. odyneroides sp. nov. K. E. pulchellus Cresson, 1865." figureDoi="http://doi.org/10.5281/zenodo.3476765" httpUri="https://zenodo.org/record/3476765/files/figure.png" pageId="23" pageNumber="24">7E</figureCitation>
,
<figureCitation id="308C2A3EFF87FFC21960FBFB40CE8C2F" box="[875,905,1065,1091]" captionStart="Fig" captionStartId="24.[189,232,1895,1921]" captionTargetBox="[194,1398,1040,1852]" captionTargetId="figure@24.[189,1398,1040,1860]" captionTargetPageId="24" captionText="Fig. 11. Epeolus fumipennis Say, 1837. A. Female, habitus, lateral view. B. Female, habitus, dorsal view. C. Male, habitus, lateral view. D. Female axillae and mesoscutellum, dorsal view (blue lines indicate the posterior extent of the axilla relative to the length of the horizontal dorsal portion of the mesoscutellum). Scale bars: A–C = 3 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3476773" httpUri="https://zenodo.org/record/3476773/files/figure.png" pageId="23" pageNumber="24">11</figureCitation>
,
<figureCitation id="308C2A3EFF87FFC2199EFBFB40F68C2F" box="[917,945,1065,1091]" captionStart="Fig" captionStartId="25.[189,232,1789,1815]" captionTargetBox="[189,1398,1351,1748]" captionTargetId="figure@25.[189,1398,1351,1753]" captionTargetPageId="25" captionText="Fig. 12. Pseudopygidial area of female, dorsal view. A. E. fumipennis Say, 1837, which consists of a basal impressed triangular or semicircular portion covered in silvery short, appressed setae differentiated from a transverse band of coppery short, appressed setae along the posterior-facing apical margin of T5. B. E. bifasciatus Cresson, 1864, which is more than twice as wide as long. C. E. obscuripes Cockerell, 1917 stat. nov., which is not more than twice as wide as long. The pseudopygidial area is the apical portion of T5 that changes slope from the rest of the tergum and is covered in short, coppery or silvery hairs uniform in length (posteromesad the light blue lines). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3476775" httpUri="https://zenodo.org/record/3476775/files/figure.png" pageId="23" pageNumber="24">12</figureCitation>
A
</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF87FFC21AB6FBB942B78CAB" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="A80836BBFF87FFC21AB6FBB9405B8CEA" blockId="23.[189,796,1131,1158]" box="[189,796,1131,1158]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="88A71195FF87FFC21AB6FBB9405B8CEA" box="[189,796,1131,1158]" pageId="23" pageNumber="24">
<taxonomicName id="6FB74D38FF87FFC21AB6FBB9411D8CE9" ID-CoL="6FNQV" authority="Say, 1837: 403" authorityName="Say" authorityPageNumber="403" authorityYear="1837" box="[189,602,1131,1157]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF87FFC21AB6FBB942E38CE9" box="[189,420,1131,1157]" italics="true" pageId="23" pageNumber="24">Epeolus fumipénnis</emphasis>
<treatmentCitation id="291610AAFF87FFC21BA0FBB9411D8CE9" author="Say T." box="[427,602,1131,1157]" page="403" pageId="23" pageNumber="24" year="1837">
<bibRefCitation id="CC264B4AFF87FFC21BA0FBB9411D8CE9" author="Say" box="[427,602,1131,1157]" firstAuthor="Say" pageId="23" pageNumber="24" pagination="361 - 416" refId="ref37138" refString="Say T. 1837. Descriptions of new species of North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1: 361 - 416." type="journal article" year="1837">Say, 1837: 403</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(sex not stated).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A80836BBFF87FFC21AB6FB7F42B78CAB" blockId="23.[189,496,1197,1223]" box="[189,496,1197,1223]" pageId="23" pageNumber="24">
<heading id="F34081D7FF87FFC21AB6FB7F42B78CAB" bold="true" box="[189,496,1197,1223]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="9AC3EAA9FF87FFC21AB6FB7F42B78CAB" bold="true" box="[189,496,1197,1223]" pageId="23" pageNumber="24">Proposed common name</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF87FFC21AB6FB0E42958C9A" box="[189,466,1244,1270]" pageId="23" pageNumber="24" type="description">
<paragraph id="A80836BBFF87FFC21AB6FB0E42958C9A" blockId="23.[189,466,1244,1270]" box="[189,466,1244,1270]" pageId="23" pageNumber="24">Dusky-winged epeolus.</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF87FFCD1AB6FACC47D08997" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="A80836BBFF87FFC21AB6FACC42718D54" blockId="23.[189,310,1310,1336]" box="[189,310,1310,1336]" pageId="23" pageNumber="24">
<heading id="F34081D7FF87FFC21AB6FACC42718D54" bold="true" box="[189,310,1310,1336]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="9AC3EAA9FF87FFC21AB6FACC42718D54" bold="true" box="[189,310,1310,1336]" pageId="23" pageNumber="24">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A80836BBFF87FFCD1AB6FA9F47D08997" blockId="23.[189,1399,1357,2027]" lastBlockId="24.[189,1399,267,508]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Together with most of the morphological features that are diagnostic for the ‘
<taxonomicName id="6FB74D38FF87FFC21E66FA9F46678D0B" authorityName="Holmberg" authorityYear="1886" box="[1133,1312,1357,1383]" class="Insecta" family="Apidae" genus="Trophocleptria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Trophocleptria</taxonomicName>
group’ (see exception below), the following in combination can be used to tell
<taxonomicName id="6FB74D38FF87FFC21E2FFAA0478F8DE7" authorityName="Say" authorityYear="1837" box="[1060,1224,1393,1419]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF87FFC21E2FFAA0478F8DE7" box="[1060,1224,1393,1419]" italics="true" pageId="23" pageNumber="24">E. fumipennis</emphasis>
</taxonomicName>
apart from all other
<taxonomicName id="6FB74D38FF87FFC21B08FA4742258DC3" authorityName="Latreille" authorityYear="1802" box="[259,354,1429,1455]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="9AC3EAA9FF87FFC21B08FA4742258DC3" box="[259,354,1429,1455]" italics="true" pageId="23" pageNumber="24">Epeolus</emphasis>
</taxonomicName>
: the mesoscutum has a pair of well-defined paramedian bands (
<figureCitation id="308C2A3EFF87FFC21E69FA4747828DC3" box="[1122,1221,1429,1455]" captionStart="Fig" captionStartId="7.[189,232,1895,1921]" captionTargetBox="[189,1398,1441,1838]" captionTargetId="figure@7.[189,1398,1441,1847]" captionTargetPageId="7" captionText="Fig. 2. Pronotal collar of female, dorsal view. A. E. boliviensis Friese, 1908, which is straight along its anterior margin. B. E. fumipennis Say, 1837, which is relatively straight along its anterior margin. C. E. hanusiae sp. nov., paratype (UCBME), which is convex along its anterior margin. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3476752" httpUri="https://zenodo.org/record/3476752/files/figure.png" pageId="23" pageNumber="24">Figs 2B</figureCitation>
,
<figureCitation id="308C2A3EFF87FFC21ED8FA4746408DC3" box="[1235,1287,1429,1455]" captionStart="Fig" captionStartId="24.[189,232,1895,1921]" captionTargetBox="[194,1398,1040,1852]" captionTargetId="figure@24.[189,1398,1040,1860]" captionTargetPageId="24" captionText="Fig. 11. Epeolus fumipennis Say, 1837. A. Female, habitus, lateral view. B. Female, habitus, dorsal view. C. Male, habitus, lateral view. D. Female axillae and mesoscutellum, dorsal view (blue lines indicate the posterior extent of the axilla relative to the length of the horizontal dorsal portion of the mesoscutellum). Scale bars: A–C = 3 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3476773" httpUri="https://zenodo.org/record/3476773/files/figure.png" pageId="23" pageNumber="24">11B</figureCitation>
); there is no large tooth laterally near the base of each axilla (
<figureCitation id="308C2A3EFF87FFC21928FA6A40C88DBF" box="[803,911,1464,1491]" captionStart="Fig" captionStartId="24.[189,232,1895,1921]" captionTargetBox="[194,1398,1040,1852]" captionTargetId="figure@24.[189,1398,1040,1860]" captionTargetPageId="24" captionText="Fig. 11. Epeolus fumipennis Say, 1837. A. Female, habitus, lateral view. B. Female, habitus, dorsal view. C. Male, habitus, lateral view. D. Female axillae and mesoscutellum, dorsal view (blue lines indicate the posterior extent of the axilla relative to the length of the horizontal dorsal portion of the mesoscutellum). Scale bars: A–C = 3 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3476773" httpUri="https://zenodo.org/record/3476773/files/figure.png" pageId="23" pageNumber="24">Fig. 11D</figureCitation>
); the mesoscutellum does not have a pair of posteriorly directed teeth (
<figureCitation id="308C2A3EFF87FFC21813FA0E41C38D9A" box="[536,644,1500,1526]" captionStart="Fig" captionStartId="24.[189,232,1895,1921]" captionTargetBox="[194,1398,1040,1852]" captionTargetId="figure@24.[189,1398,1040,1860]" captionTargetPageId="24" captionText="Fig. 11. Epeolus fumipennis Say, 1837. A. Female, habitus, lateral view. B. Female, habitus, dorsal view. C. Male, habitus, lateral view. D. Female axillae and mesoscutellum, dorsal view (blue lines indicate the posterior extent of the axilla relative to the length of the horizontal dorsal portion of the mesoscutellum). Scale bars: A–C = 3 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3476773" httpUri="https://zenodo.org/record/3476773/files/figure.png" pageId="23" pageNumber="24">Fig. 11D</figureCitation>
); the mesopleura are finely punctate, each with punctures more or less equally dense throughout (few i≥2d); the fore wings are deeply infuscate apically (
<figureCitation id="308C2A3EFF87FFC21ED1F9D2466B8E76" box="[1242,1324,1536,1562]" captionStart="Fig" captionStartId="24.[189,232,1895,1921]" captionTargetBox="[194,1398,1040,1852]" captionTargetId="figure@24.[189,1398,1040,1860]" captionTargetPageId="24" captionText="Fig. 11. Epeolus fumipennis Say, 1837. A. Female, habitus, lateral view. B. Female, habitus, dorsal view. C. Male, habitus, lateral view. D. Female axillae and mesoscutellum, dorsal view (blue lines indicate the posterior extent of the axilla relative to the length of the horizontal dorsal portion of the mesoscutellum). Scale bars: A–C = 3 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3476773" httpUri="https://zenodo.org/record/3476773/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
A–B); T1 has a broad, medially narrowed, bright to pale yellow submedial fascia (
<figureCitation id="308C2A3EFF87FFC21E3DF9F647E78E52" box="[1078,1184,1572,1598]" captionStart="Fig" captionStartId="24.[189,232,1895,1921]" captionTargetBox="[194,1398,1040,1852]" captionTargetId="figure@24.[189,1398,1040,1860]" captionTargetPageId="24" captionText="Fig. 11. Epeolus fumipennis Say, 1837. A. Female, habitus, lateral view. B. Female, habitus, dorsal view. C. Male, habitus, lateral view. D. Female axillae and mesoscutellum, dorsal view (blue lines indicate the posterior extent of the axilla relative to the length of the horizontal dorsal portion of the mesoscutellum). Scale bars: A–C = 3 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3476773" httpUri="https://zenodo.org/record/3476773/files/figure.png" pageId="23" pageNumber="24">Fig. 11B</figureCitation>
); and T2–T4 have complete bright to pale yellow apical fasciae (
<figureCitation id="308C2A3EFF87FFC218D8F995407A8E0E" box="[723,829,1607,1634]" captionStart="Fig" captionStartId="24.[189,232,1895,1921]" captionTargetBox="[194,1398,1040,1852]" captionTargetId="figure@24.[189,1398,1040,1860]" captionTargetPageId="24" captionText="Fig. 11. Epeolus fumipennis Say, 1837. A. Female, habitus, lateral view. B. Female, habitus, dorsal view. C. Male, habitus, lateral view. D. Female axillae and mesoscutellum, dorsal view (blue lines indicate the posterior extent of the axilla relative to the length of the horizontal dorsal portion of the mesoscutellum). Scale bars: A–C = 3 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3476773" httpUri="https://zenodo.org/record/3476773/files/figure.png" pageId="23" pageNumber="24">Fig. 11B</figureCitation>
). Whereas in other species in the ‘
<taxonomicName id="6FB74D38FF87FFC21ECFF99546308E0D" authorityName="Holmberg" authorityYear="1886" box="[1220,1399,1607,1633]" class="Insecta" family="Apidae" genus="Trophocleptria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Trophocleptria</taxonomicName>
group’ each mesopleuron has a carina delineating its anterior and lateral surfaces, in
<taxonomicName id="6FB74D38FF87FFC21EDAF9BE46308EE9" authorityName="Say" authorityYear="1837" box="[1233,1399,1643,1669]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF87FFC21EDAF9BE46308EE9" box="[1233,1399,1643,1669]" italics="true" pageId="23" pageNumber="24">E. fumipennis</emphasis>
</taxonomicName>
the anterior and lateral surfaces of each mesopleuron are not clearly differentiated from one another.
<taxonomicName id="6FB74D38FF87FFC21AB6F96142E48EA1" authorityName="Say" authorityYear="1837" box="[189,419,1715,1741]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF87FFC21AB6F96142E48EA1" box="[189,419,1715,1741]" italics="true" pageId="23" pageNumber="24">Epeolus fumipennis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="6FB74D38FF87FFC21832F96641A08EA1" authorityName="Cameron" authorityYear="1902" box="[569,743,1715,1741]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fulvopilosus">
<emphasis id="9AC3EAA9FF87FFC21832F96641A08EA1" box="[569,743,1715,1741]" italics="true" pageId="23" pageNumber="24">E. fulvopilosus</emphasis>
</taxonomicName>
in terms of the patterns of pubescence on the mesosoma and metasoma. However, in
<taxonomicName id="6FB74D38FF87FFC21815F90541968E9C" authorityName="Cameron" authorityYear="1902" box="[542,721,1750,1776]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fulvopilosus">
<emphasis id="9AC3EAA9FF87FFC21815F90541968E9C" box="[542,721,1750,1776]" italics="true" pageId="23" pageNumber="24">E. fulvopilosus</emphasis>
</taxonomicName>
the mesoscutum lacks paramedian bands, although the mesoscutum of the female anteriorly has faint lines of bright yellow tomentum along the midline and between the midline and parapsidal lines; there is a large tooth laterally near the base of each axilla; the mesoscutellum has a pair of posteriorly directed teeth; each mesopleuron has a carina delineating its anterior and lateral surfaces; the mesopleura are more coarsely punctate, each with sparser punctures ventrolaterally (many i≥2d) than in the upper half, with interspaces shining; and T1 has a broad, medially narrowed or interrupted bright to pale yellow basal fascia. Additionally, females may be separated on the basis of the pseudopygidial area, which uniquely in
<taxonomicName id="6FB74D38FF87FFC21904F80040E88F87" authorityName="Say" authorityYear="1837" box="[783,943,2001,2027]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF87FFC21904F80040E88F87" box="[783,943,2001,2027]" italics="true" pageId="23" pageNumber="24">E. fumipennis</emphasis>
</taxonomicName>
consists of a basal impressed triangular portion covered in silvery short, appressed setae differentiated from a transverse band of coppery short, appressed setae along the posterior-facing apical margin of T5 (
<figureCitation id="308C2A3EFF88FFCD19BEFEE44763893C" box="[949,1060,310,336]" captionStart="Fig" captionStartId="25.[189,232,1789,1815]" captionTargetBox="[189,1398,1351,1748]" captionTargetId="figure@25.[189,1398,1351,1753]" captionTargetPageId="25" captionText="Fig. 12. Pseudopygidial area of female, dorsal view. A. E. fumipennis Say, 1837, which consists of a basal impressed triangular or semicircular portion covered in silvery short, appressed setae differentiated from a transverse band of coppery short, appressed setae along the posterior-facing apical margin of T5. B. E. bifasciatus Cresson, 1864, which is more than twice as wide as long. C. E. obscuripes Cockerell, 1917 stat. nov., which is not more than twice as wide as long. The pseudopygidial area is the apical portion of T5 that changes slope from the rest of the tergum and is covered in short, coppery or silvery hairs uniform in length (posteromesad the light blue lines). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3476775" httpUri="https://zenodo.org/record/3476775/files/figure.png" pageId="24" pageNumber="25">Fig. 12A</figureCitation>
).
<taxonomicName id="6FB74D38FF88FFCD1E31FEE44665893C" authorityName="Say" authorityYear="1837" box="[1082,1314,310,336]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF88FFCD1E31FEE44665893C" box="[1082,1314,310,336]" italics="true" pageId="24" pageNumber="25">Epeolus fumipennis</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="6FB74D38FF88FFCD1B3DFEB041DC8917" authority="Cockerell, 1917" authorityName="Onuferko" authorityYear="2019" baseAuthorityName="Cockerell" baseAuthorityYear="1917" box="[310,667,353,379]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="obscuripes" status="stat. nov.">
<emphasis id="9AC3EAA9FF88FFCD1B3DFEB042918917" box="[310,470,353,379]" italics="true" pageId="24" pageNumber="25">E. obscuripes</emphasis>
<bibRefCitation id="CC264B4AFF88FFCD1BD5FEB341DC8917" author="Cockerell" box="[478,667,353,379]" firstAuthor="Cockerell" pageId="24" pageNumber="25" pagination="298 - 304" refId="ref35268" refString="Cockerell T. D. A. 1917. XXXIII. - Descriptions and records of bees. - LXXVII. Annals and Magazine of Natural History 20: 298 - 304. https: // doi. org / 10.1080 / 00222931709487008" type="journal article" year="1917">Cockerell, 1917</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="81F057D2FF88FFCD18A9FEB0404B8910" box="[674,780,354,380]" pageId="24" pageNumber="25">stat. nov.</taxonomicNameLabel>
in that in males of both species T1–T6 are typically fasciate. However, in
<taxonomicName id="6FB74D38FF88FFCD1BB1FE5F411E89CA" authorityName="Onuferko" authorityYear="2019" baseAuthorityName="Cockerell" baseAuthorityYear="1917" box="[442,601,396,422]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="obscuripes">
<emphasis id="9AC3EAA9FF88FFCD1BB1FE5F411E89CA" box="[442,601,396,422]" italics="true" pageId="24" pageNumber="25">E. obscuripes</emphasis>
</taxonomicName>
the mesoscutum lacks paramedian bands, the axillae are shorter, not extending as far back as the ridge overhanging the depressed posterior margin of the mesoscutellum, and T1 has a broad, medially narrowed or interrupted, bright to pale yellow basal fascia.
</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF88FFCC1AB6FDF841FA8B60" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="A80836BBFF88FFCD1AB6FDF842ED8A28" blockId="24.[189,426,554,580]" box="[189,426,554,580]" pageId="24" pageNumber="25">
<heading id="F34081D7FF88FFCD1AB6FDF842ED8A28" bold="true" box="[189,426,554,580]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="9AC3EAA9FF88FFCD1AB6FDF842ED8A28" bold="true" box="[189,426,554,580]" pageId="24" pageNumber="25">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A80836BBFF88FFCD1AE1FDB341448A17" blockId="24.[189,789,609,679]" box="[234,515,609,635]" pageId="24" pageNumber="25">
<emphasis id="9AC3EAA9FF88FFCD1AE1FDB341448A17" bold="true" box="[234,515,609,635]" pageId="24" pageNumber="25">
Primary
<typeStatus id="770C8819FF88FFCD1B57FDB342D58A17" box="[348,402,609,635]" pageId="24" pageNumber="25">type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="A80836BBFF88FFCD1AB6FD5E40528ACB" blockId="24.[189,789,609,679]" box="[189,789,652,679]" pageId="24" pageNumber="25">
<collectingCountry id="D0A0762BFF88FFCD1AB6FD5E42768ACA" box="[189,305,652,678]" name="Mexico" pageId="24" pageNumber="25">MEXICO</collectingCountry>
• (not examined, presumably destroyed).
</paragraph>
<paragraph id="A80836BBFF88FFCD1AE1FD07408B8B76" blockId="24.[189,972,725,794]" pageId="24" pageNumber="25">
<emphasis id="9AC3EAA9FF88FFCD1AE1FD0740DD8A83" bold="true" box="[234,922,725,751]" pageId="24" pageNumber="25">DNA barcoded material with BIN-compliant sequences</emphasis>
Available. BOLD:ACZ0714. Specimens examined and sequenced:
</paragraph>
<paragraph id="A80836BBFF88FFCD1AB6FC9B41728B8F" blockId="24.[189,1399,841,995]" pageId="24" pageNumber="25">
<materialsCitation id="18DF3CE6FF88FFCD1AB6FC9B47358BE1" ID-GBIF-Occurrence="2425042242" collectingDate="2012-01-14" collectingDateMax="2012-01-20" collectingDateMin="2012-01-14" collectionCode="BBSL" collectorName="F. D. Parker &amp; T. D. McIntyre" country="Panama" latitude="8.6986" location="Boquete" longLatPrecision="7" longitude="-82.4505" pageId="24" pageNumber="25" specimenCode="FDP119767" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui">
<collectingCountry id="D0A0762BFF88FFCD1AB6FC9B427F8B0F" box="[189,312,841,867]" name="Panama" pageId="24" pageNumber="25">PANAMA</collectingCountry>
•
<specimenCount id="BEB1FD32FF88FFCD1B58FC9B42C48B0F" box="[339,387,841,867]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
;
<collectingRegion id="6A73F859FF88FFCD1B9AFC9B42B18B0F" box="[401,502,841,867]" country="Panama" name="Chiriqui" pageId="24" pageNumber="25">Chiriquí</collectingRegion>
,
<quantity id="6F4F9B5EFF88FFCD1809FC9B41798B0F" box="[514,574,841,867]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="25" unit="km" value="8.0">8 km</quantity>
S of
<location id="AD686060FF88FFCD188AFC9B41A08B0F" LSID="urn:lsid:plazi:treatment:201E87ADFF87FFCE1868FBD741828C98:AD686060FF88FFCD188AFC9B41A08B0F" box="[641,743,841,867]" country="Panama" latitude="8.6986" longLatPrecision="7" longitude="-82.4505" name="Boquete" pageId="24" pageNumber="25" stateProvince="Chiriqui">Boquete</location>
;
<geoCoordinate id="CD83507CFF88FFCD18FFFC9B40378B0F" box="[756,880,841,867]" degrees="8.6986" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="8.6986">8.6986° N</geoCoordinate>
,
<geoCoordinate id="CD83507CFF88FFCD1976FC9B47498B0F" box="[893,1038,841,868]" degrees="82.4505" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="-82.4505">82.4505° W</geoCoordinate>
;
<date id="DC09107BFF88FFCD1E17FC9B47A58B0F" box="[1052,1250,841,867]" pageId="24" pageNumber="25" value="2012-01-14" valueMax="2012-01-20" valueMin="2012-01-14">
<collectingDate id="CC4DE993FF88FFCD1E17FC9B47A58B0F" box="[1052,1250,841,867]" pageId="24" pageNumber="25" value="2012-01-14" valueMax="2012-01-20" valueMin="2012-01-14">14–20 Jan. 2012</collectingDate>
</date>
;
<collectorName id="0542536DFF88FFCD1EFBFC9B46308B0F" box="[1264,1399,841,867]" pageId="24" pageNumber="25">F.D. Parker</collectorName>
and
<collectorName id="0542536DFF88FFCD1AE6FCA642D48BE2" box="[237,403,884,910]" pageId="24" pageNumber="25">T.D. McIntyre</collectorName>
leg.; BOLD sample ID: CCDB-28239 H01;
<collectionCode id="CEA6AE7EFF88FFCD199CFCA640988BE2" box="[919,991,884,910]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="24" pageNumber="25" type="Museum">BBSL</collectionCode>
<specimenCode id="F8119EC0FF88FFCD19EEFCA147358BE1" box="[997,1138,883,909]" collectionCode="BBSL" country="USA" httpUri="http://grbio.org/cool/q49p-qw3v" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="24" pageNumber="25" type="Museum">FDP119767</specimenCode>
</materialsCitation>
•
<materialsCitation id="18DF3CE6FF88FFCD1E83FCA641768B8F" ID-GBIF-Occurrence="2425042150" collectingDate="2012-01-21" collectingDateMax="61-12-02" collectingDateMin="2012-01-21" collectionCode="BBSL" collectorName="F. D. Parker &amp; T. D. McIntyre" country="Panama" latitude="8.6986" location="Boquete" longLatPrecision="7" longitude="-82.4505" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui">
<specimenCount id="BEB1FD32FF88FFCD1E83FCA647F38BE2" box="[1160,1204,884,910]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<date id="DC09107BFF88FFCD1BC9FC4D41C08BD5" box="[450,647,927,953]" pageId="24" pageNumber="25" value="2012-01-21" valueMax="2012-01-31" valueMin="2012-01-21">
<collectingDate id="CC4DE993FF88FFCD1BC9FC4D41C08BD5" box="[450,647,927,953]" pageId="24" pageNumber="25" value="2012-01-21" valueMax="2012-01-31" valueMin="2012-01-21">21–31 Jan. 2012</collectingDate>
</date>
;
<collectorName id="0542536DFF88FFCD1898FC4D405D8BD4" box="[659,794,926,953]" pageId="24" pageNumber="25">F.D. Parker</collectorName>
and
<collectorName id="0542536DFF88FFCD1959FC4D40BD8BD5" box="[850,1018,927,953]" pageId="24" pageNumber="25">T.D. McIntyre</collectorName>
leg.; BOLD sample ID: CCDB- 28239 G12;
<collectionCode id="CEA6AE7EFF88FFCD1B5BFC1B42DF8B8F" box="[336,408,969,995]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="24" pageNumber="25" type="Museum">BBSL</collectionCode>
FDP
<date id="DC09107BFF88FFCD1BD2FC1B41768B8F" box="[473,561,969,995]" pageId="24" pageNumber="25" value="61-12-02">
<collectingDate id="CC4DE993FF88FFCD1BD2FC1B41768B8F" box="[473,561,969,995]" pageId="24" pageNumber="25" value="61-12-02">120261</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="FCC86633FF88FFCD1AB6F8B541C58F87" ID-DOI="http://doi.org/10.5281/zenodo.3476773" ID-Zenodo-Dep="3476773" httpUri="https://zenodo.org/record/3476773/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1895,1921]" targetBox="[194,1398,1040,1852]" targetPageId="24">
<paragraph id="A80836BBFF88FFCD1AB6F8B541C58F87" blockId="24.[189,1399,1895,2027]" pageId="24" pageNumber="25">
<emphasis id="9AC3EAA9FF88FFCD1AB6F8B542518FED" bold="true" box="[189,278,1895,1921]" pageId="24" pageNumber="25">Fig. 11.</emphasis>
<taxonomicName id="6FB74D38FF88FFCD1B17F8B5413A8FED" authority="Say, 1837" authorityName="Say" authorityYear="1837" box="[284,637,1895,1921]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF88FFCD1B17F8B541458FED" box="[284,514,1895,1921]" italics="true" pageId="24" pageNumber="25">Epeolus fumipennis</emphasis>
<bibRefCitation id="CC264B4AFF88FFCD1803F8B5413A8FED" author="Say" box="[520,637,1895,1921]" firstAuthor="Say" pageId="24" pageNumber="25" pagination="361 - 416" refId="ref37138" refString="Say T. 1837. Descriptions of new species of North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1: 361 - 416." type="journal article" year="1837">Say, 1837</bibRefCitation>
</taxonomicName>
.
<emphasis id="9AC3EAA9FF88FFCD188CF8B541DB8FED" bold="true" box="[647,668,1895,1921]" pageId="24" pageNumber="25">A</emphasis>
. Female, habitus, lateral view.
<emphasis id="9AC3EAA9FF88FFCD1E08F8B547508FED" bold="true" box="[1027,1047,1895,1921]" pageId="24" pageNumber="25">B</emphasis>
. Female, habitus, dorsal view.
<emphasis id="9AC3EAA9FF88FFCD1AB6F85843958FC8" bold="true" box="[189,210,1930,1956]" pageId="24" pageNumber="25">C</emphasis>
. Male, habitus, lateral view.
<emphasis id="9AC3EAA9FF88FFCD182BF85841728FC8" bold="true" box="[544,565,1930,1956]" pageId="24" pageNumber="25">D</emphasis>
. Female axillae and mesoscutellum, dorsal view (blue lines indicate the posterior extent of the axilla relative to the length of the horizontal dorsal portion of the mesoscutellum). Scale bars: A–C = 3 mm; D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="A80836BBFF89FFCC1AE1FED941498949" blockId="25.[189,1398,267,364]" box="[234,526,267,293]" pageId="25" pageNumber="26">
<emphasis id="9AC3EAA9FF89FFCC1AE1FED941498949" bold="true" box="[234,526,267,293]" pageId="25" pageNumber="26">Non-barcoded material</emphasis>
</paragraph>
<paragraph id="A80836BBFF89FFCC1AB6FEFC416B8900" blockId="25.[189,1398,267,364]" pageId="25" pageNumber="26">
<materialsCitation id="18DF3CE6FF89FFCC1AB6FEFC416E8900" ID-GBIF-Occurrence="2425042126" collectingDate="2001-02-01" collectingDateMax="2001-04-12" collectingDateMin="2001-02-01" collectionCode="KUNHM" collectorName="T. H. Ricketts" country="Costa Rica" location="Perez Zeledon" pageId="25" pageNumber="26" specimenCode="SEMC1248326" specimenCount="1" specimenCount-male="1" stateProvince="San Jose">
<collectingCountry id="D0A0762BFF89FFCC1AB6FEFC42228924" box="[189,357,302,328]" name="Costa Rica" pageId="25" pageNumber="26">COSTA RICA</collectingCountry>
<emphasis id="9AC3EAA9FF89FFCC1B66FEFD42308925" bold="true" box="[365,375,303,329]" pageId="25" pageNumber="26">•</emphasis>
<specimenCount id="BEB1FD32FF89FFCC1B75FEFC42EA8925" box="[382,429,302,329]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
;
<collectingRegion id="6A73F859FF89FFCC1BB1FEFC41648924" box="[442,547,302,328]" country="Costa Rica" name="San Jose" pageId="25" pageNumber="26">San José</collectingRegion>
,
<location id="AD686060FF89FFCC1824FEFC419F8924" LSID="urn:lsid:plazi:treatment:201E87ADFF87FFCE1868FBD741828C98:AD686060FF89FFCC1824FEFC419F8924" box="[559,728,302,328]" country="Costa Rica" name="Perez Zeledon" pageId="25" pageNumber="26" stateProvince="San Jose">Pérez Zeledón</location>
(San Isidro vicinity);
<date id="DC09107BFF89FFCC19D0FEFC47888924" box="[987,1231,302,329]" pageId="25" pageNumber="26" value="2001-02-01" valueMax="2001-04-12" valueMin="2001-02-01">
<collectingDate id="CC4DE993FF89FFCC19D0FEFC47888924" box="[987,1231,302,329]" pageId="25" pageNumber="26" value="2001-02-01" valueMax="2001-04-12" valueMin="2001-02-01">1 Feb.–12 Apr. 2001</collectingDate>
</date>
;
<collectorName id="0542536DFF89FFCC1ED0FEFD46318924" box="[1243,1398,302,329]" pageId="25" pageNumber="26">T.H. Ricketts</collectorName>
leg.;
<collectionCode id="CEA6AE7EFF89FFCC1AF3FE8042208900" box="[248,359,338,364]" pageId="25" pageNumber="26">KUNHM</collectionCode>
<specimenCode id="F8119EC0FF89FFCC1B65FE80416E8900" box="[366,553,338,364]" collectionCode="KUNHM" pageId="25" pageNumber="26">SEMC1248326</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="A80836BBFF89FFCC1AB6FE46426A8ACA" blockId="25.[189,1399,403,678]" pageId="25" pageNumber="26">
<materialsCitation id="18DF3CE6FF89FFCC1AB6FE46416189C2" ID-GBIF-Occurrence="2425042104" box="[189,550,404,430]" collectionCode="ANSP" country="Mexico" location="MEXICO" pageId="25" pageNumber="26" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectingCountry id="D0A0762BFF89FFCC1AB6FE46427689C2" box="[189,305,404,430]" name="Mexico" pageId="25" pageNumber="26">MEXICO</collectingCountry>
<emphasis id="9AC3EAA9FF89FFCC1B32FE46420489C2" bold="true" box="[313,323,404,430]" pageId="25" pageNumber="26">•</emphasis>
<specimenCount id="BEB1FD32FF89FFCC1B47FE4642D689C2" box="[332,401,404,430]" pageId="25" pageNumber="26" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="BEB1FD32FF89FFCC1BABFE46429789C2" box="[416,464,404,430]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
;
<collectionCode id="CEA6AE7EFF89FFCC1BD7FE46416189C2" box="[476,550,404,430]" country="USA" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">ANSP</collectionCode>
</materialsCitation>
<emphasis id="9AC3EAA9FF89FFCC1824FE46417E89C2" bold="true" box="[559,569,404,430]" pageId="25" pageNumber="26">•</emphasis>
<materialsCitation id="18DF3CE6FF89FFCC1849FE4642008999" ID-GBIF-Occurrence="2425042166" collectingDate="2004-12-17" collectionCode="ECOSUR" collectorName="M. Rincon &amp; R. Ayala &amp; M. Guzman &amp; J. Esponda &amp; C. Balboa" country="Mexico" latitude="15.5665" location="Acacoyagua" longLatPrecision="7" longitude="-92.7902" municipality="Ei. La Palma" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Chiapas">
<specimenCount id="BEB1FD32FF89FFCC1849FE46413789C2" box="[578,624,404,430]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
;
<collectingRegion id="6A73F859FF89FFCC188BFE4141A689C1" box="[640,737,403,429]" country="Mexico" name="Chiapas" pageId="25" pageNumber="26">Chiapas</collectingRegion>
,
<collectingMunicipality id="486CACC1FF89FFCC18E5FE4140CE89C1" box="[750,905,403,430]" pageId="25" pageNumber="26">Ei. La Palma</collectingMunicipality>
(
<location id="AD686060FF89FFCC1991FE46477589C2" LSID="urn:lsid:plazi:treatment:201E87ADFF87FFCE1868FBD741828C98:AD686060FF89FFCC1991FE46477589C2" box="[922,1074,404,430]" country="Mexico" latitude="15.5665" longLatPrecision="7" longitude="-92.7902" municipality="Ei. La Palma" name="Acacoyagua" pageId="25" pageNumber="26" stateProvince="Chiapas">Acacoyagua</location>
);
<geoCoordinate id="CD83507CFF89FFCC1E43FE41479489C2" box="[1096,1235,403,430]" degrees="15.5665" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="5" value="15.5665">15.5665° N</geoCoordinate>
,
<geoCoordinate id="CD83507CFF89FFCC1EEBFE46463589C2" box="[1248,1394,404,430]" degrees="92.7902" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="5" value="-92.7902">92.7902° W</geoCoordinate>
;
<date id="DC09107BFF89FFCC1AB6FE65422189BD" box="[189,358,439,465]" pageId="25" pageNumber="26" value="2004-12-17">
<collectingDate id="CC4DE993FF89FFCC1AB6FE65422189BD" box="[189,358,439,465]" pageId="25" pageNumber="26" value="2004-12-17">17 Dec. 2004</collectingDate>
</date>
;
<collectorName id="0542536DFF89FFCC1B7DFE6542BB89BD" box="[374,508,439,465]" pageId="25" pageNumber="26">M. Rincón</collectorName>
,
<collectorName id="0542536DFF89FFCC1800FE65413189BD" box="[523,630,439,465]" pageId="25" pageNumber="26">R. Ayala</collectorName>
,
<collectorName id="0542536DFF89FFCC188DFE65405E89BD" box="[646,793,439,465]" pageId="25" pageNumber="26">M. Guzmán</collectorName>
,
<collectorName id="0542536DFF89FFCC1921FE6540EA89BD" box="[810,941,439,465]" pageId="25" pageNumber="26">J. Esponda</collectorName>
and
<collectorName id="0542536DFF89FFCC19E4FE65472F89BD" box="[1007,1128,439,465]" pageId="25" pageNumber="26">C. Balboa</collectorName>
leg.;
<collectionCode id="CEA6AE7EFF89FFCC1EB8FE65466089BD" box="[1203,1319,439,465]" country="Mexico" name="El Colegio de la Frontera Sur (Mexico)" pageId="25" pageNumber="26" type="Museum">ECOSUR</collectionCode>
ECO- TAE-42220
</materialsCitation>
<emphasis id="9AC3EAA9FF89FFCC1B5AFE09421C8999" bold="true" box="[337,347,475,501]" pageId="25" pageNumber="26">•</emphasis>
<materialsCitation id="18DF3CE6FF89FFCC1B6EFE0946248A75" ID-GBIF-Occurrence="2425042112" collectingDate="2006-11-17" collectionCode="ECOSUR" collectorName="C. Balboa &amp; J. Merida &amp; M. Guzman &amp; M. Cigarroa &amp; J. Toto" country="Mexico" latitude="15.2783" location="Acacoyagua" longLatPrecision="7" longitude="-92.3983" municipality="Ei. Rosario Zacatonal" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas">
<specimenCount id="BEB1FD32FF89FFCC1B6EFE0942D08999" box="[357,407,475,501]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
;
<collectingRegion id="6A73F859FF89FFCC1BADFE0841408998" box="[422,519,474,500]" country="Mexico" name="Chiapas" pageId="25" pageNumber="26">Chiapas</collectingRegion>
,
<collectingMunicipality id="486CACC1FF89FFCC181DFE08405D8998" box="[534,794,474,500]" pageId="25" pageNumber="26">Ei. Rosario Zacatonal</collectingMunicipality>
(
<location id="AD686060FF89FFCC1927FE0940838999" LSID="urn:lsid:plazi:treatment:201E87ADFF87FFCE1868FBD741828C98:AD686060FF89FFCC1927FE0940838999" box="[812,964,475,501]" country="Mexico" latitude="15.2783" longLatPrecision="7" longitude="-92.3983" municipality="Ei. Rosario Zacatonal" name="Acacoyagua" pageId="25" pageNumber="26" stateProvince="Chiapas">Acacoyagua</location>
);
<geoCoordinate id="CD83507CFF89FFCC19D1FE0847208999" box="[986,1127,474,501]" degrees="15.2783" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="5" value="15.2783">15.2783° N</geoCoordinate>
,
<geoCoordinate id="CD83507CFF89FFCC1E7DFE09464F8999" box="[1142,1288,475,501]" degrees="92.3983" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="5" value="-92.3983">92.3983° W</geoCoordinate>
;
<date id="DC09107BFF89FFCC1F1CFE0943BC8A74" pageId="25" pageNumber="26" value="2006-11-17">
<collectingDate id="CC4DE993FF89FFCC1F1CFE0943BC8A74" pageId="25" pageNumber="26" value="2006-11-17">17 Nov. 2006</collectingDate>
</date>
;
<collectorName id="0542536DFF89FFCC1B02FE2C42C48A74" box="[265,387,510,536]" pageId="25" pageNumber="26">C. Balboa</collectorName>
,
<collectorName id="0542536DFF89FFCC1B9AFE2C41428A74" box="[401,517,510,536]" pageId="25" pageNumber="26">J. Mérida</collectorName>
,
<collectorName id="0542536DFF89FFCC1818FE2C41E38A74" box="[531,676,510,536]" pageId="25" pageNumber="26">M. Guzmán</collectorName>
,
<collectorName id="0542536DFF89FFCC18B9FE2C40038A74" box="[690,836,510,536]" pageId="25" pageNumber="26">M. Cigarroa</collectorName>
and
<collectorName id="0542536DFF89FFCC198BFE2C40968A74" box="[896,977,510,536]" pageId="25" pageNumber="26">J. Toto</collectorName>
leg.;
<collectionCode id="CEA6AE7EFF89FFCC1E1CFE2C47CC8A74" box="[1047,1163,510,536]" country="Mexico" name="El Colegio de la Frontera Sur (Mexico)" pageId="25" pageNumber="26" type="Museum">ECOSUR</collectionCode>
ECO-TAE-50698
</materialsCitation>
<emphasis id="9AC3EAA9FF89FFCC1F67FE2C46318A74" bold="true" box="[1388,1398,510,536]" pageId="25" pageNumber="26">•</emphasis>
<materialsCitation id="18DF3CE6FF89FFCC1AB6FDF3419C8A0C" ID-GBIF-Occurrence="2425042200" collectingDate="2006-11-29" collectionCode="ECOSUR" collectorName="C. Balboa &amp; J. Merida &amp; M. Guzman &amp; M. Cigarroa &amp; J. Toto" country="Mexico" latitude="15.2783" location="Acacoyagua" longLatPrecision="7" longitude="-92.3983" municipality="Ei. Rosario Zacatonal" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Chiapas">
<specimenCount id="BEB1FD32FF89FFCC1AB6FDF343AF8A50" box="[189,232,545,572]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<date id="DC09107BFF89FFCC18A5FDF3400B8A57" box="[686,844,545,572]" pageId="25" pageNumber="26" value="2006-11-29">
<collectingDate id="CC4DE993FF89FFCC18A5FDF3400B8A57" box="[686,844,545,572]" pageId="25" pageNumber="26" value="2006-11-29">29 Nov. 2006</collectingDate>
</date>
;
<collectorName id="0542536DFF89FFCC195CFDF3408A8A57" box="[855,973,545,571]" pageId="25" pageNumber="26">C. Balboa</collectorName>
,
<collectorName id="0542536DFF89FFCC19DCFDF0470F8A57" box="[983,1096,545,572]" pageId="25" pageNumber="26">J. Mérida</collectorName>
,
<collectorName id="0542536DFF89FFCC1E5AFDF047998A57" box="[1105,1246,545,572]" pageId="25" pageNumber="26">M. Guzmán</collectorName>
,
<collectorName id="0542536DFF89FFCC1EE2FDF046308A57" box="[1257,1399,545,572]" pageId="25" pageNumber="26">M. Cigarroa</collectorName>
and
<collectorName id="0542536DFF89FFCC1AF9FD9742038A33" box="[242,324,581,607]" pageId="25" pageNumber="26">J. Toto</collectorName>
leg.;
<collectionCode id="CEA6AE7EFF89FFCC1B85FD9741458A33" box="[398,514,581,607]" country="Mexico" name="El Colegio de la Frontera Sur (Mexico)" pageId="25" pageNumber="26" type="Museum">ECOSUR</collectionCode>
ECO-TAE-51519
</materialsCitation>
<emphasis id="9AC3EAA9FF89FFCC18EEFD9741A88A33" bold="true" box="[741,751,581,607]" pageId="25" pageNumber="26">•</emphasis>
<materialsCitation id="18DF3CE6FF89FFCC18F0FD9742888AEE" ID-GBIF-Occurrence="2425042157" collectingDate="1962-03-19" collectionCode="BBSL" collectorName="F. D. Parker" country="Mexico" location="9 mi E of Chupaderos" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Sinaloa">
<specimenCount id="BEB1FD32FF89FFCC18F0FD97406C8A33" box="[763,811,581,607]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
;
<collectingRegion id="6A73F859FF89FFCC1936FD9740DF8A33" box="[829,920,581,607]" country="Mexico" name="Sinaloa" pageId="25" pageNumber="26">Sinaloa</collectingRegion>
,
<location id="AD686060FF89FFCC19ACFD9747FC8A33" LSID="urn:lsid:plazi:treatment:201E87ADFF87FFCE1868FBD741828C98:AD686060FF89FFCC19ACFD9747FC8A33" box="[935,1211,581,607]" country="Mexico" name="9 mi E of Chupaderos" pageId="25" pageNumber="26" stateProvince="Sinaloa">
<quantity id="6F4F9B5EFF89FFCC19ACFD9740A78A33" box="[935,992,581,607]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="25" pageNumber="26" unit="mi" value="9.0">9 mi</quantity>
E of Chupaderos
</location>
;
<date id="DC09107BFF89FFCC1EC1FD9746358A33" box="[1226,1394,581,607]" pageId="25" pageNumber="26" value="1962-03-19">
<collectingDate id="CC4DE993FF89FFCC1EC1FD9746358A33" box="[1226,1394,581,607]" pageId="25" pageNumber="26" value="1962-03-19">19 Mar. 1962</collectingDate>
</date>
;
<collectorName id="0542536DFF89FFCC1AB6FDBB42038AEE" box="[189,324,616,643]" pageId="25" pageNumber="26">F.D. Parker</collectorName>
leg.;
<collectionCode id="CEA6AE7EFF89FFCC1B8CFDBA42888AEE" box="[391,463,616,642]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="25" pageNumber="26" type="Museum">BBSL</collectionCode>
</materialsCitation>
<emphasis id="9AC3EAA9FF89FFCC1BDCFDBA42A68AEE" bold="true" box="[471,481,616,642]" pageId="25" pageNumber="26">•</emphasis>
<materialsCitation id="18DF3CE6FF89FFCC1BE1FDBA426E8ACA" ID-GBIF-Occurrence="2425042174" collectingDate="2018-04" collectionCode="UCBME" collectorName="F. D. Parker &amp; L. A. Stange" country="Mexico" location="Chichen Itza" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Yucatan">
<specimenCount id="BEB1FD32FF89FFCC1BE1FDBA415F8AEF" box="[490,536,616,643]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
;
<collectingRegion id="6A73F859FF89FFCC182DFDBA41CF8AEE" box="[550,648,616,642]" country="Mexico" name="Yucatan" pageId="25" pageNumber="26">Yucatán</collectingRegion>
,
<location id="AD686060FF89FFCC189FFDBA40698AEE" LSID="urn:lsid:plazi:treatment:201E87ADFF87FFCE1868FBD741828C98:AD686060FF89FFCC189FFDBA40698AEE" box="[660,814,616,642]" country="Mexico" name="Chichen Itza" pageId="25" pageNumber="26" stateProvince="Yucatan">Chichén Itzá</location>
;
<date id="DC09107BFF89FFCC1931FDBA40D28AEE" box="[826,917,616,642]" pageId="25" pageNumber="26" value="2018-04">
<collectingDate id="CC4DE993FF89FFCC1931FDBA40D28AEE" box="[826,917,616,642]" pageId="25" pageNumber="26" value="2018-04">18 Apr.</collectingDate>
</date>
; 1962;
<collectorName id="0542536DFF89FFCC19E7FDBB47348AEE" box="[1004,1139,616,643]" pageId="25" pageNumber="26">F.D. Parker</collectorName>
and
<collectorName id="0542536DFF89FFCC1EA6FDBA467C8AEE" box="[1197,1339,616,642]" pageId="25" pageNumber="26">L.A. Stange</collectorName>
leg.;
<collectionCode id="CEA6AE7EFF89FFCC1AB6FD5E426E8ACA" box="[189,297,652,678]" pageId="25" pageNumber="26">UCBME</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="A80836BBFF89FFCC1AB6FD1C41FA8B60" blockId="25.[189,1399,717,780]" pageId="25" pageNumber="26">
<materialsCitation id="18DF3CE6FF89FFCC1AB6FD1C41FD8B60" ID-GBIF-Occurrence="2425042224" collectingDate="2012-02-15" collectingDateMax="2012-02-29" collectingDateMin="2012-02-15" collectionCode="BBSL" collectorName="F. D. Parker &amp; T. D. McIntyre" country="Panama" latitude="8.6986" location="Boquete" longLatPrecision="7" longitude="-82.4505" pageId="25" pageNumber="26" specimenCode="FDP126465" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui">
<collectingCountry id="D0A0762BFF89FFCC1AB6FD1C427F8A84" box="[189,312,718,744]" name="Panama" pageId="25" pageNumber="26">PANAMA</collectingCountry>
<emphasis id="9AC3EAA9FF89FFCC1B4BFD1C420D8A84" bold="true" box="[320,330,718,744]" pageId="25" pageNumber="26">•</emphasis>
<specimenCount id="BEB1FD32FF89FFCC1B59FD1C42C78A84" box="[338,384,718,744]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
;
<collectingRegion id="6A73F859FF89FFCC1B9BFD1F42B28A8B" box="[400,501,717,743]" country="Panama" name="Chiriqui" pageId="25" pageNumber="26">Chiriquí</collectingRegion>
,
<quantity id="6F4F9B5EFF89FFCC180AFD1C417A8A8B" box="[513,573,717,744]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="km" value="8.0">8 km</quantity>
S of
<location id="AD686060FF89FFCC1874FD1C41A28A84" LSID="urn:lsid:plazi:treatment:201E87ADFF87FFCE1868FBD741828C98:AD686060FF89FFCC1874FD1C41A28A84" box="[639,741,718,744]" country="Panama" latitude="8.6986" longLatPrecision="7" longitude="-82.4505" name="Boquete" pageId="25" pageNumber="26" stateProvince="Chiriqui">Boquete</location>
;
<geoCoordinate id="CD83507CFF89FFCC18FAFD1F402A8A84" box="[753,877,717,744]" degrees="8.6986" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="5" value="8.6986">8.6986° N</geoCoordinate>
,
<geoCoordinate id="CD83507CFF89FFCC1971FD1F474C8A84" box="[890,1035,717,744]" degrees="82.4505" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="5" value="-82.4505">82.4505° W</geoCoordinate>
;
<date id="DC09107BFF89FFCC1E13FD1F47A48A84" box="[1048,1251,717,744]" pageId="25" pageNumber="26" value="2012-02-15" valueMax="2012-02-29" valueMin="2012-02-15">
<collectingDate id="CC4DE993FF89FFCC1E13FD1F47A48A84" box="[1048,1251,717,744]" pageId="25" pageNumber="26" value="2012-02-15" valueMax="2012-02-29" valueMin="2012-02-15">15–29 Feb. 2012</collectingDate>
</date>
;
<collectorName id="0542536DFF89FFCC1EFBFD1C46308A8B" box="[1264,1399,717,744]" pageId="25" pageNumber="26">F.D. Parker</collectorName>
and
<collectorName id="0542536DFF89FFCC1AE5FD2342D18B67" box="[238,406,753,779]" pageId="25" pageNumber="26">T.D. McIntyre</collectorName>
leg.;
<collectionCode id="CEA6AE7EFF89FFCC1BD3FD2341678B67" box="[472,544,753,779]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="25" pageNumber="26" type="Museum">BBSL</collectionCode>
<specimenCode id="F8119EC0FF89FFCC1823FD2341FD8B60" box="[552,698,753,780]" collectionCode="BBSL" country="USA" httpUri="http://grbio.org/cool/q49p-qw3v" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="25" pageNumber="26" type="Museum">FDP126465</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF89FFCF1AB6FCE042528EB2" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="A80836BBFF89FFCC1AB6FCE0422C8B20" blockId="25.[189,363,818,844]" box="[189,363,818,844]" pageId="25" pageNumber="26">
<heading id="F34081D7FF89FFCC1AB6FCE0422C8B20" bold="true" box="[189,363,818,844]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="9AC3EAA9FF89FFCC1AB6FCE0422C8B20" bold="true" box="[189,363,818,844]" pageId="25" pageNumber="26">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="A80836BBFF89FFCC1AE1FCB042028B10" blockId="25.[189,1398,866,963]" box="[234,325,866,892]" pageId="25" pageNumber="26">
<emphasis id="9AC3EAA9FF89FFCC1AE1FCB042028B10" bold="true" box="[234,325,866,892]" pageId="25" pageNumber="26">Female</emphasis>
</paragraph>
<paragraph id="A80836BBFF89FFCC1AB6FC5742158BAF" blockId="25.[189,1398,866,963]" pageId="25" pageNumber="26">
MEASUREMENTS. Length
<quantity id="6F4F9B5EFF89FFCC1BD8FC5741248BF3" box="[467,611,901,927]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.2" metricValueMin="6.8" pageId="25" pageNumber="26" unit="mm" value="8.0" valueMax="9.2" valueMin="6.8">6.8–9.2 mm</quantity>
; head length
<quantity id="6F4F9B5EFF89FFCC18F0FC5740CD8BF3" box="[763,906,901,927]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="25" pageNumber="26" unit="mm" value="1.95" valueMax="2.2" valueMin="1.7">1.7–2.2 mm</quantity>
; head width
<quantity id="6F4F9B5EFF89FFCC1E17FC5747EB8BF3" box="[1052,1196,901,927]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="25" pageNumber="26" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2–2.8 mm</quantity>
; fore wing length
<quantity id="6F4F9B5EFF89FFCC1AB6FC7A420A8BAF" box="[189,333,936,963]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.2" metricValueMin="5.9" pageId="25" pageNumber="26" unit="mm" value="6.55" valueMax="7.2" valueMin="5.9">5.9–7.2 mm</quantity>
.
</paragraph>
<paragraph id="A80836BBFF89FFCC1AB6FC3841B58CD9" blockId="25.[189,1398,1002,1206]" pageId="25" pageNumber="26">INTEGUMENT COLORATION. Mostly black; notable exceptions as follows: at least partially ferruginous on mandible, labrum, antenna, pronotal collar, pronotal lobe, tegula, axilla, mesoscutellum, metanotum, mesopleuron and legs. Mandible with apex darker than all but extreme base; preapical tooth slightly lighter than mandibular apex. Antenna brown and orange in part. Pronotal collar, pronotal lobe and tegula pale ferruginous to amber. Wing membrane dusky subhyaline, slightly darker at apex. Legs more extensively reddish orange than brown or black.</paragraph>
<paragraph id="A80836BBFF89FFCF1AB6FB0F41C78AD2" blockId="25.[189,1399,1245,1307]" lastBlockId="26.[189,1399,267,702]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">PUBESCENCE. Face with tomentum densest around antennal socket, slightly sparser on clypeus, upper paraocular and frontal areas, and vertexal area. Dorsum of mesosoma and metasoma with bands of bright to pale yellow short, appressed setae. Pronotal collar with tomentum uniformly pale yellow. Mesoscutum with paramedian band. Mesopleuron sparsely hairy, but tomentum dense ventrally as well as between two sparsely hairy patches (one beneath base of fore wing (hypoepimeral area), a larger circular patch occupying much of ventrolateral half of mesopleuron). Metanotum with tomentum uninterrupted, uniformly off-white. T1 with broad, medially narrowed bright yellow submedial fascia. T2 with complete bright yellow apical fascia, broadest medially and without anterolateral extensions. T3 and T4 with complete, pale yellow apical fasciae of slightly sparser tomentum. T5 with large, continuous patch of pale yellow tomentum bordering and contacting pseudopygidial area. T5 with pseudopygidial area lunate, its apex less than twice as wide as medial length, with basal impressed triangular portion covered in silvery short, appressed setae differentiated from transverse band of coppery short, appressed setae along posterior-facing apical margin. S5 with apical fimbria of coppery to silvery hairs extending beyond apex of sternum by ~⅓ MOD.</paragraph>
<caption id="FCC86633FF89FFCC1AB6F92F471E8F87" ID-DOI="http://doi.org/10.5281/zenodo.3476775" ID-Zenodo-Dep="3476775" httpUri="https://zenodo.org/record/3476775/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1789,1815]" targetBox="[189,1398,1351,1748]" targetPageId="25">
<paragraph id="A80836BBFF89FFCC1AB6F92F471E8F87" blockId="25.[189,1399,1789,2028]" pageId="25" pageNumber="26">
<emphasis id="9AC3EAA9FF89FFCC1AB6F92F425C8F7B" bold="true" box="[189,283,1789,1815]" pageId="25" pageNumber="26">Fig. 12.</emphasis>
Pseudopygidial area of female, dorsal view.
<emphasis id="9AC3EAA9FF89FFCC1933F92F400A8F7B" bold="true" box="[824,845,1789,1815]" pageId="25" pageNumber="26">A</emphasis>
.
<taxonomicName id="6FB74D38FF89FFCC1955F92C47C58F74" authority="Say, 1837" authorityName="Say" authorityYear="1837" box="[862,1154,1790,1816]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF89FFCC1955F92C47468F74" box="[862,1025,1790,1816]" italics="true" pageId="25" pageNumber="26">E. fumipennis</emphasis>
<bibRefCitation id="CC264B4AFF89FFCC1E01F92C47C58F74" author="Say" box="[1034,1154,1790,1816]" firstAuthor="Say" pageId="25" pageNumber="26" pagination="361 - 416" refId="ref37138" refString="Say T. 1837. Descriptions of new species of North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1: 361 - 416." type="journal article" year="1837">Say, 1837</bibRefCitation>
</taxonomicName>
, which consists of a basal impressed triangular or semicircular portion covered in silvery short, appressed setae differentiated from a transverse band of coppery short, appressed setae along the posterior-facing apical margin of T5.
<emphasis id="9AC3EAA9FF89FFCC1AB6F8B543968FED" bold="true" box="[189,209,1895,1921]" pageId="25" pageNumber="26">B</emphasis>
.
<taxonomicName id="6FB74D38FF89FFCC1AD4F8BA41768FED" authority="Cresson, 1864" authorityName="Cresson" authorityYear="1864" box="[223,561,1895,1921]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="bifasciatus">
<emphasis id="9AC3EAA9FF89FFCC1AD4F8BA42388FED" box="[223,383,1895,1921]" italics="true" pageId="25" pageNumber="26">E. bifasciatus</emphasis>
Cresson, 1864
</taxonomicName>
, which is more than twice as wide as long.
<emphasis id="9AC3EAA9FF89FFCC1E3AF8B547018FED" bold="true" box="[1073,1094,1895,1921]" pageId="25" pageNumber="26">C</emphasis>
.
<taxonomicName id="6FB74D38FF89FFCC1E5DF8BA43BF8FC8" authority="Cockerell, 1917" authorityName="Onuferko" authorityYear="2019" baseAuthorityName="Cockerell" baseAuthorityYear="1917" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="obscuripes" status="stat. nov.">
<emphasis id="9AC3EAA9FF89FFCC1E5DF8BA47B28FED" box="[1110,1269,1895,1921]" italics="true" pageId="25" pageNumber="26">E. obscuripes</emphasis>
<bibRefCitation id="CC264B4AFF89FFCC1EF6F8B543BF8FC8" author="Cockerell" firstAuthor="Cockerell" pageId="25" pageNumber="26" pagination="298 - 304" refId="ref35268" refString="Cockerell T. D. A. 1917. XXXIII. - Descriptions and records of bees. - LXXVII. Annals and Magazine of Natural History 20: 298 - 304. https: // doi. org / 10.1080 / 00222931709487008" type="journal article" year="1917">Cockerell, 1917</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="81F057D2FF89FFCC1B09F85942378FC9" box="[258,368,1931,1957]" pageId="25" pageNumber="26">stat. nov.</taxonomicNameLabel>
, which is not more than twice as wide as long. The pseudopygidial area is the apical portion of T5 that changes slope from the rest of the tergum and is covered in short, coppery or silvery hairs uniform in length (posteromesad the light blue lines). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="A80836BBFF8AFFCF1AB6FD3547CC8BD7" blockId="26.[189,1399,743,956]" pageId="26" pageNumber="27">SURFACE SCULPTURE. Punctures dense. Labrum and clypeus with punctures equally dense (i&lt;1d). Small impunctate shiny spot lateral to lateral ocellus. Mesoscutum, mesoscutellum and axilla very coarsely and densely rugose-punctate. Tegula densely punctate anteriorly and mesally (i≤1d), sparsely punctate (i&gt;2d) to impunctate posteriorly and along margins. Mesopleuron with ventrolateral half densely punctate (i≤1d), interspaces shining; mesopleuron with punctures more or less equally dense throughout. Metasomal terga with punctures very fine, dense (i≈1d), evenly distributed on disc.</paragraph>
<paragraph id="A80836BBFF8AFFCF1AB6FC36428F8D8D" blockId="26.[189,1399,996,1505]" pageId="26" pageNumber="27">
STRUCTURE. Preapical tooth forming right-angled triangle. Labral apex with three small denticles, lateral two each preceded by small discrete longitudinal ridge. Frontal keel strongly raised. Frontal area with pair of very discrete densely punctate protrusions, each located near upper mesal margin of compound eye. Head dorsally with pair of protrusions, each located where upper genal area meets vertexal area. Vertexal area strongly convex in frontal view. Scape with greatest length 1.5 × greatest width. F2 noticeably longer than wide (L/W ratio = 1.4–1.5). Preoccipital ridge separated from hypostomal carina by no less than 1 MOD. Pronotal collar rather short (medial length ~⅔ MOD), expanded laterally to about 2 × medial length in dorsal view, and relatively straight along anterior margin. Mesoscutellum moderately bigibbous, depressed along posterior margin beneath overhanging ridge. Axilla large, its lateral margin more than half as long as mesoscutellar width (AL/MSCW ratio = 0.6) and tip extending as far back as apex of horizontal dorsal portion of mesoscutellum; axilla with tip conspicuously diverging from side of mesoscutellum, distinctly hooked, and axilla with free portion
<superScript id="5FC29BF3FF8AFFCF1E49FAAB470E8DEB" attach="right" box="[1090,1097,1401,1415]" fontSize="6" pageId="26" pageNumber="27">2</superScript>
∕5 its medial length; axilla with lateral margin relatively straight and carinate. Fore wing with three submarginal cells. Pygidial plate apically truncate.
</paragraph>
<paragraph id="A80836BBFF8AFFCF1AE1F9D8426D8E48" blockId="26.[189,1399,1546,1758]" box="[234,298,1546,1572]" pageId="26" pageNumber="27">
<emphasis id="9AC3EAA9FF8AFFCF1AE1F9D8426D8E48" bold="true" box="[234,298,1546,1572]" pageId="26" pageNumber="27">Male</emphasis>
</paragraph>
<paragraph id="A80836BBFF8AFFCF1AB6F9FD42528EB2" blockId="26.[189,1399,1546,1758]" pageId="26" pageNumber="27">Description as for female except for usual secondary sexual characters and as follows: F2 shorter, not noticeably longer than wide (L/W ratio = 1.1); pygidial plate apically rounded, with larger and deeper punctures, closely punctate basomedially and sparsely punctate apically and laterally, with interspaces shining; S3–S5 with much longer (&gt;1 MOD), coppery to silvery subapical hairs, those of S4 and S5 curved.</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF8AFFCF1AB6F8D541CE8F3E" box="[189,649,1799,1874]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="A80836BBFF8AFFCF1AB6F8D542118F4D" blockId="26.[189,342,1799,1825]" box="[189,342,1799,1825]" pageId="26" pageNumber="27">
<heading id="F34081D7FF8AFFCF1AB6F8D542118F4D" bold="true" box="[189,342,1799,1825]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="9AC3EAA9FF8AFFCF1AB6F8D542118F4D" bold="true" box="[189,342,1799,1825]" pageId="26" pageNumber="27">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="A80836BBFF8AFFCF1AB6F8EA41CE8F3E" blockId="26.[189,649,1848,1874]" box="[189,649,1848,1874]" pageId="26" pageNumber="27">
Central America and
<collectingCountry id="D0A0762BFF8AFFCF1BBCF8EA41568F3E" box="[439,529,1848,1874]" name="Mexico" pageId="26" pageNumber="27">Mexico</collectingCountry>
(
<figureCitation id="308C2A3EFF8AFFCF182AF8EA413B8F3E" box="[545,636,1848,1874]" captionStart="Fig" captionStartId="12.[189,232,1824,1850]" captionTargetId="figure@13.[241,1373,265,2019]" captionTargetPageId="13" captionText="Fig. 7 (opposite page). Occurrence records (yellow circles except where indicated otherwise) of: A. Epeolus boliviensis Friese, 1908. B. E. claripennis Friese, 1908. C. E. danieli (Genaro, 2014) comb. nov. (orange square) and E. luteipennis Friese, 1916. D. E. fulvopilosus Cameron, 1902. E. E. fumipennis Say, 1837. F. E. hanusiae sp. nov. G. E. niger (Michener, 1954). H. E. nomadiformis sp. nov. I. E. obscuripes Cockerell, 1917 stat. nov. J. E. odyneroides sp. nov. K. E. pulchellus Cresson, 1865." figureDoi="http://doi.org/10.5281/zenodo.3476765" httpUri="https://zenodo.org/record/3476765/files/figure.png" pageId="26" pageNumber="27">Fig. 7E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF8AFFCE1AB6F8A9427E8925" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="biology_ecology">
<paragraph id="A80836BBFF8AFFCF1AB6F8A942678FF9" blockId="26.[189,288,1915,1941]" box="[189,288,1915,1941]" pageId="26" pageNumber="27">
<heading id="F34081D7FF8AFFCF1AB6F8A942678FF9" bold="true" box="[189,288,1915,1941]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="9AC3EAA9FF8AFFCF1AB6F8A942678FF9" bold="true" box="[189,288,1915,1941]" pageId="26" pageNumber="27">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="A80836BBFF8AFFCF1AE1F87E42CD8FAA" blockId="26.[189,394,1964,2027]" box="[234,394,1964,1990]" pageId="26" pageNumber="27">
<emphasis id="9AC3EAA9FF8AFFCF1AE1F87E42CD8FAA" bold="true" box="[234,394,1964,1990]" pageId="26" pageNumber="27">Host records</emphasis>
</paragraph>
<paragraph id="A80836BBFF8AFFCF1AB6F803427E8F87" blockId="26.[189,394,1964,2027]" box="[189,313,2001,2027]" pageId="26" pageNumber="27">Unknown.</paragraph>
<paragraph id="A80836BBFF8BFFCE1AE1FED942DB8949" blockId="27.[189,412,267,329]" box="[234,412,267,293]" pageId="27" pageNumber="28">
<emphasis id="9AC3EAA9FF8BFFCE1AE1FED942DB8949" bold="true" box="[234,412,267,293]" pageId="27" pageNumber="28">Floral records</emphasis>
</paragraph>
<paragraph id="A80836BBFF8BFFCE1AB6FEFD427E8925" blockId="27.[189,412,267,329]" box="[189,313,303,329]" pageId="27" pageNumber="28">Unknown.</paragraph>
</subSubSection>
<subSubSection id="E0AD6530FF8BFFCE1AB6FEA341828C98" pageId="27" pageNumber="28" type="discussion">
<paragraph id="A80836BBFF8BFFCE1AB6FEA3426889E7" blockId="27.[189,303,369,395]" box="[189,303,369,395]" pageId="27" pageNumber="28">
<heading id="F34081D7FF8BFFCE1AB6FEA3426889E7" bold="true" box="[189,303,369,395]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="9AC3EAA9FF8BFFCE1AB6FEA3426889E7" bold="true" box="[189,303,369,395]" pageId="27" pageNumber="28">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="A80836BBFF8BFFCE1AB6FE7241828C98" blockId="27.[189,1399,416,1268]" pageId="27" pageNumber="28">
The specimen upon which
<bibRefCitation id="CC264B4AFF8BFFCE1BF5FE7241D389D6" author="Say's" box="[510,660,416,442]" firstAuthor="Say's" pageId="27" pageNumber="28" pagination="361 - 416" refId="ref37138" refString="Say T. 1837. Descriptions of new species of North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1: 361 - 416." type="journal article" year="1837">Say’s (1837)</bibRefCitation>
description is based was probably destroyed, along with most of Thomas Say’s insect collection (
<bibRefCitation id="CC264B4AFF8BFFCE186CFE16405089B3" author="LeConte" box="[615,791,452,479]" firstAuthor="LeConte" pageId="27" pageNumber="28" refId="ref36046" refString="LeConte J. L. (ed.) 1859. The complete writings of Thomas Say on the entomology of North America. Vol. I. Bailliere Brothers, New York. https: // doi. org / 10.5962 / bhl. title. 13257" type="book" year="1859">LeConte 1859</bibRefCitation>
: v–vi, xix (footnote)).
<bibRefCitation id="CC264B4AFF8BFFCE1E27FE1647B889B2" author="Mawdsley" box="[1068,1279,452,478]" firstAuthor="Mawdsley" pageId="27" pageNumber="28" pagination="163 - 171" refId="ref36094" refString="Mawdsley J. R. 1993. The entomological collection of Thomas Say. Psyche 100: 163 - 171. https: // doi. org / 10.1155 / 1993 / 59616" type="journal article" year="1993">Mawdsley (1993)</bibRefCitation>
lists only four
<taxonomicName id="6FB74D38FF8BFFCE1AFEFE3A42D28A6E" authorityName="Linnaeus" authorityYear="1758" box="[245,405,488,514]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
specimens from Say’s collection that are presently housed in the Harvard University Museum of Comparative Zoology in Cambridge, MA,
<collectingCountry id="D0A0762BFF8BFFCE193FFDDE40358A4A" box="[820,882,524,550]" name="United States of America" pageId="27" pageNumber="28">USA</collectingCountry>
, and all are ichneumonids. Personal searches through the collections of most (20/28) of the entomological institutions listed in the Materials and Methods of
<bibRefCitation id="CC264B4AFF8BFFCE1B41FD81415B8A02" author="Onuferko" box="[330,540,595,622]" firstAuthor="Onuferko" pageId="27" pageNumber="28" pagination="1 - 185" refId="ref36367" refString="Onuferko T. M. 2018 a. A revision of the cleptoparasitic bee genus Epeolus Latreille for Nearctic species, north of Mexico (Hymenoptera, Apidae). ZooKeys 755: 1 - 185. https: // doi. org / 10.3897 / zookeys. 755.23939" type="journal article" year="2018" yearSuffix="a">Onuferko (2018a)</bibRefCitation>
did not turn up any specimens authoritatively identified as
<taxonomicName id="6FB74D38FF8BFFCE1EC5FD8646288A02" authorityName="Say" authorityYear="1837" box="[1230,1391,596,622]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF8BFFCE1EC5FD8646288A02" box="[1230,1391,596,622]" italics="true" pageId="27" pageNumber="28">E. fumipennis</emphasis>
</taxonomicName>
. Nevertheless, the original description suggests a species of
<taxonomicName id="6FB74D38FF8BFFCE1984FDA540A98AFD" authorityName="Latreille" authorityYear="1802" box="[911,1006,631,657]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="9AC3EAA9FF8BFFCE1984FDA540A98AFD" box="[911,1006,631,657]" italics="true" pageId="27" pageNumber="28">Epeolus</emphasis>
</taxonomicName>
in the ‘
<taxonomicName id="6FB74D38FF8BFFCE1E47FDA547B88AFD" authorityName="Holmberg" authorityYear="1886" box="[1100,1279,631,657]" class="Insecta" family="Apidae" genus="Trophocleptria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Trophocleptria</taxonomicName>
group’ in which the mesoscutum has paramedian bands of pale tomentum, and among the examined Neotropical
<taxonomicName id="6FB74D38FF8BFFCE1AB6FD6D425B8AB5" authorityName="Latreille" authorityYear="1802" box="[189,284,703,729]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="9AC3EAA9FF8BFFCE1AB6FD6D425B8AB5" box="[189,284,703,729]" italics="true" pageId="27" pageNumber="28">Epeolus</emphasis>
</taxonomicName>
specimens are several fitting such a description, and all appear to be conspecific. Morphological features suggesting a species within the ‘
<taxonomicName id="6FB74D38FF8BFFCE189CFD31400D8A91" authorityName="Holmberg" authorityYear="1886" box="[663,842,739,765]" class="Insecta" family="Apidae" genus="Trophocleptria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Trophocleptria</taxonomicName>
group’ are as follows, quoted verbatim: “
<emphasis id="9AC3EAA9FF8BFFCE1F3BFD3146318A91" box="[1328,1398,739,765]" italics="true" pageId="27" pageNumber="28">collar</emphasis>
with a ferruginous disk, contracted in the middle”, “
<emphasis id="9AC3EAA9FF8BFFCE1939FCDA40C38B4D" box="[818,900,776,801]" italics="true" pageId="27" pageNumber="28">tergum</emphasis>
, first and second segments with a yellow band, the first broader and widely interrupted; remaining segments with a whitish band” and “the last segment with the addition of an obscure rufous terminal margin” (
<bibRefCitation id="CC264B4AFF8BFFCE19B3FC9D476D8B05" author="Say" box="[952,1066,847,873]" firstAuthor="Say" pageId="27" pageNumber="28" pagination="361 - 416" refId="ref37138" refString="Say T. 1837. Descriptions of new species of North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1: 361 - 416." type="journal article" year="1837">Say 1837</bibRefCitation>
). The last of these appears to describe the apical portion of the female pseudopygidial area, which is covered in coppery (rather than silvery) setae in the examined specimens that appear to be representatives of this species (
<figureCitation id="308C2A3EFF8BFFCE1EF1FC44462E8BDD" box="[1274,1385,918,945]" captionStart="Fig" captionStartId="25.[189,232,1789,1815]" captionTargetBox="[189,1398,1351,1748]" captionTargetId="figure@25.[189,1398,1351,1753]" captionTargetPageId="25" captionText="Fig. 12. Pseudopygidial area of female, dorsal view. A. E. fumipennis Say, 1837, which consists of a basal impressed triangular or semicircular portion covered in silvery short, appressed setae differentiated from a transverse band of coppery short, appressed setae along the posterior-facing apical margin of T5. B. E. bifasciatus Cresson, 1864, which is more than twice as wide as long. C. E. obscuripes Cockerell, 1917 stat. nov., which is not more than twice as wide as long. The pseudopygidial area is the apical portion of T5 that changes slope from the rest of the tergum and is covered in short, coppery or silvery hairs uniform in length (posteromesad the light blue lines). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3476775" httpUri="https://zenodo.org/record/3476775/files/figure.png" pageId="27" pageNumber="28">Fig. 12A</figureCitation>
). According to
<bibRefCitation id="CC264B4AFF8BFFCE1B6AFC6841668BB9" author="Brumley" box="[353,545,954,981]" firstAuthor="Brumley" pageId="27" pageNumber="28" refId="ref35207" refString="Brumley R. L. 1965. A Revision of the Bee Genus Epeolus Latreille of Western America North of Mexico. M. Sc. thesis, Utah State University, Logan." type="book" year="1965">Brumley (1965)</bibRefCitation>
, who considered
<taxonomicName id="6FB74D38FF8BFFCE18F9FC6940D28BB8" authorityName="Say" authorityYear="1837" box="[754,917,954,980]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF8BFFCE18F9FC6940D28BB8" box="[754,917,954,980]" italics="true" pageId="27" pageNumber="28">E. fumipennis</emphasis>
</taxonomicName>
to be within the ‘
<taxonomicName id="6FB74D38FF8BFFCE1E6EFC68465F8BB8" authorityName="Holmberg" authorityYear="1886" box="[1125,1304,954,980]" class="Insecta" family="Apidae" genus="Trophocleptria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Trophocleptria</taxonomicName>
group’, specimens at the Academy of Natural Sciences of Drexel University in Philadelphia, PA,
<collectingCountry id="D0A0762BFF8BFFCE1F03FC0C46068B94" box="[1288,1345,990,1016]" name="United States of America" pageId="27" pageNumber="28">USA</collectingCountry>
and KUNHM that were collected from the Midwestern and Southeastern
<collectingCountry id="D0A0762BFF8BFFCE1E0EFBD047E28C70" box="[1029,1189,1026,1052]" name="United States of America" pageId="27" pageNumber="28">United States</collectingCountry>
and identified as
<taxonomicName id="6FB74D38FF8BFFCE1AB6FBF542268C2C" authorityName="Say" authorityYear="1837" box="[189,353,1062,1088]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="9AC3EAA9FF8BFFCE1AB6FBF542268C2C" box="[189,353,1062,1088]" italics="true" pageId="27" pageNumber="28">E. fumipennis</emphasis>
</taxonomicName>
are actually
<taxonomicName id="6FB74D38FF8BFFCE1BF5FBF541E78C2C" authorityName="Cresson" authorityYear="1864" box="[510,672,1062,1088]" class="Insecta" family="Apidae" genus="Epeolus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bifasciatus">
<emphasis id="9AC3EAA9FF8BFFCE1BF5FBF541E78C2C" box="[510,672,1062,1088]" italics="true" pageId="27" pageNumber="28">E. bifasciatus</emphasis>
</taxonomicName>
. Given that the taxonomic understanding of this species has been problematic, a re-description is warranted, and is based on all available representatives of this species. Although the sex upon which
<bibRefCitation id="CC264B4AFF8BFFCE1898FBBC406B8CE4" author="Say's" box="[659,812,1134,1160]" firstAuthor="Say's" pageId="27" pageNumber="28" pagination="361 - 416" refId="ref37138" refString="Say T. 1837. Descriptions of new species of North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1: 361 - 416." type="journal article" year="1837">Say’s (1837)</bibRefCitation>
original description is based was not stated, the description of the last metasomal segment suggests a female. Therefore, in the present redescription of the species (
<emphasis id="9AC3EAA9FF8BFFCE1B47FB6742818CA3" box="[332,454,1205,1231]" italics="true" pageId="27" pageNumber="28">vide supra</emphasis>
) a detailed description of the female is given, whereas the description of the male lists only key differences between the sexes.
</paragraph>
</subSubSection>
</treatment>
</document>
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