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<document id="562F41D6B9A1B2A0BDC130095D21D120" ID-CLB-Dataset="27000" ID-DOI="10.11646/zootaxa.4586.3.8" ID-GBIF-Dataset="42d8c42b-cc7c-4c9e-9828-1a73f9eedc4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="2647350" ID-ZooBank="3CFD2CE3-6E93-4DFB-81CB-5406DB34BF2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555665942860" checkinUser="plazi" docAuthor="Hendrich, Lars, Manuel, Michael &amp; Balke, Michael" docDate="2019" docId="082F1226FFBBFFEEC1B0696DFD374ED3" docLanguage="en" docName="zootaxa.4586.3.8.pdf" docOrigin="Zootaxa 4586 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megadytes (Bifurcitus) magnus Tremouilles &amp; Bachmann 1980" docType="treatment" docVersion="8" lastPageNumber="529" masterDocId="F4166A5EFFB3FFE2C1276E07FFE14D1B" masterDocTitle="The return of the Duke - locality data for Megadytes ducalis Sharp, 1882, the world's largest diving beetle, with notes on related species (Coleoptera: Dytiscidae)" masterLastPageNumber="535" masterPageNumber="517" pageNumber="525" updateTime="1698723126771" updateUser="ExternalLinkService">
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<mods:title id="67705545CB90A0D23898C13A7A0DF90A">The return of the Duke - locality data for Megadytes ducalis Sharp, 1882, the world's largest diving beetle, with notes on related species (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="132A9E12EFA9FF59158DDC0F8EE38438">Hendrich, Lars</mods:namePart>
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<treatment id="082F1226FFBBFFEEC1B0696DFD374ED3" ID-DOI="http://doi.org/10.5281/zenodo.5943242" ID-GBIF-Taxon="156827858" ID-Zenodo-Dep="5943242" LSID="urn:lsid:plazi:treatment:082F1226FFBBFFEEC1B0696DFD374ED3" httpUri="http://treatment.plazi.org/id/082F1226FFBBFFEEC1B0696DFD374ED3" lastPageId="12" lastPageNumber="529" pageId="8" pageNumber="525">
<subSubSection id="C89CF0BBFFBBFFEAC1B0696DFC474A9F" box="[151,934,1898,1924]" pageId="8" pageNumber="525" type="nomenclature">
<paragraph id="8039A330FFBBFFEAC1B0696DFC474A9F" blockId="8.[151,934,1898,1924]" box="[151,934,1898,1924]" pageId="8" pageNumber="525">
<heading id="DB71145CFFBBFFEAC1B0696DFC474A9F" bold="true" box="[151,934,1898,1924]" fontSize="11" level="1" pageId="8" pageNumber="525" reason="1">
<taxonomicName id="4786D8B3FFBBFFEAC1B0696DFC474A9F" ID-CoL="7VSTN" authority="Tremouilles &amp; Bachmann, 1980" authorityName="Tremouilles &amp; Bachmann" authorityYear="1980" box="[151,934,1898,1924]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="magnus" subGenus="Bifurcitus">
<emphasis id="B2F27F22FFBBFFEAC1B0696DFC474A9F" bold="true" box="[151,934,1898,1924]" pageId="8" pageNumber="525">
<emphasis id="B2F27F22FFBBFFEAC1B0696DFEF64A9F" bold="true" box="[151,279,1898,1924]" italics="true" pageId="8" pageNumber="525">Megadytes</emphasis>
(
<emphasis id="B2F27F22FFBBFFEAC00F696DFE434A9F" bold="true" box="[296,418,1898,1924]" italics="true" pageId="8" pageNumber="525">Bifurcitus</emphasis>
)
<emphasis id="B2F27F22FFBBFFEAC093696DFDF54A9F" bold="true" box="[436,532,1898,1924]" italics="true" pageId="8" pageNumber="525">magnus</emphasis>
<bibRefCitation id="E417DEC1FFBBFFEAC33C696DFC474A9F" author="Tremouilles, E. R. &amp; Bachmann, A. O." box="[539,934,1898,1924]" pageId="8" pageNumber="525" pagination="101 - 125" refId="ref7687" refString="Tremouilles, E. R. &amp; Bachmann, A. O. (1980) La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae). Revista de la Sociedad Entomologica Argentina, 39, 101 - 125." type="journal article" year="1980">Trémouilles &amp; Bachmann, 1980</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C89CF0BBFFBBFFEAC1B069B6FDAA4AFD" pageId="8" pageNumber="525" type="reference_group">
<paragraph id="8039A330FFBBFFEAC1B069B6FDAA4AFD" blockId="8.[151,1436,1969,2022]" pageId="8" pageNumber="525">
<treatmentCitationGroup id="A096841EFFBBFFEAC1B069B6FDAA4AFD" pageId="8" pageNumber="525">
<taxonomicName id="4786D8B3FFBBFFEAC1B069B6FCB14ADC" ID-CoL="7VSTN" authority="Tremouilles &amp; Bachmann, 1980: 118" authorityName="Tremouilles &amp; Bachmann" authorityPageNumber="118" authorityYear="1980" box="[151,848,1969,1991]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="525" phylum="Arthropoda" rank="species" species="magnus" subGenus="Bifurcitus">
<emphasis id="B2F27F22FFBBFFEAC1B069B6FEE44ADC" box="[151,261,1969,1991]" italics="true" pageId="8" pageNumber="525">Megadytes</emphasis>
(
<emphasis id="B2F27F22FFBBFFEAC03469B6FE974ADC" box="[275,374,1969,1991]" italics="true" pageId="8" pageNumber="525">Bifurcitus</emphasis>
)
<emphasis id="B2F27F22FFBBFFEAC0A369B5FE324ADC" box="[388,467,1970,1991]" italics="true" pageId="8" pageNumber="525">magnus</emphasis>
<treatmentCitation id="01278521FFBBFFEAC0FE69B6FCB14ADC" author="Tremouilles, E. R. &amp; Bachmann, A. O." box="[473,848,1969,1991]" page="118" pageId="8" pageNumber="525" year="1980">
<bibRefCitation id="E417DEC1FFBBFFEAC0FE69B6FCC04ADC" author="Tremouilles, E. R. &amp; Bachmann, A. O." box="[473,801,1969,1991]" pageId="8" pageNumber="525" pagination="101 - 125" refId="ref7687" refString="Tremouilles, E. R. &amp; Bachmann, A. O. (1980) La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae). Revista de la Sociedad Entomologica Argentina, 39, 101 - 125." type="journal article" year="1980">Trémouilles &amp; Bachmann, 1980</bibRefCitation>
: 118
</treatmentCitation>
</taxonomicName>
(orig. descr.); Trémouilles 1989b: 161;
<treatmentCitation id="01278521FFBBFFEAC5C469B6FED14AFD" author="Miller, K. B. &amp; Bergsten, J." page="108" pageId="8" pageNumber="525" year="2016">
<bibRefCitation id="E417DEC1FFBBFFEAC5C469B6FF1C4AFD" author="Miller, K. B. &amp; Bergsten, J." pageId="8" pageNumber="525" refId="ref7159" refString="Miller, K. B. &amp; Bergsten, J. (2016) Diving beetles of the World. Systematics and biology of the Dytiscidae. Johns Hopkins University Press, Baltimore, 320 pp." type="book" year="2016">Miller &amp; Bergsten 2016</bibRefCitation>
: 108
</treatmentCitation>
;
<treatmentCitation id="01278521FFBBFFEAC01D69D7FDA64AFD" author="Nilsson, A. N. &amp; Hajek, J." box="[314,583,2000,2022]" page="81" pageId="8" pageNumber="525" year="2019">
<bibRefCitation id="E417DEC1FFBBFFEAC01D69D7FDC04AFD" author="Nilsson, A. N. &amp; Hajek, J." box="[314,545,2000,2022]" pageId="8" pageNumber="525" refId="ref7262" refString="Nilsson, A. N. &amp; Hajek, J. (2019) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Fersion 1. I. 2019. Distributed as a PDF file via Internet. Available from: http: // www. waterbeetles. eu (accessed 8 February 2019)" type="url" year="2019">Nilsson &amp; Hájek 2019</bibRefCitation>
: 81
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="D4F9F3B8FFBAFFEBC1B069BDFBE74AF4" httpUri="https://zenodo.org/record/2647362/files/figure.png" pageId="9" pageNumber="526" startId="9.[151,249,1978,2000]" targetBox="[199,1381,195,1953]" targetPageId="9">
<paragraph id="8039A330FFBAFFEBC1B069BDFBE74AF4" blockId="9.[151,1436,1978,2031]" pageId="9" pageNumber="526">
<emphasis id="B2F27F22FFBAFFEBC1B069BDFEF54ACB" bold="true" box="[151,276,1978,2000]" pageId="9" pageNumber="526">FIGURE 6.</emphasis>
<taxonomicName id="4786D8B3FFBAFFEBC03969BDFE024ACB" authorityName="Tremouilles &amp; Bachmann" authorityYear="1980" box="[286,483,1978,2000]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="526" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="B2F27F22FFBAFFEBC03969BDFE024ACB" box="[286,483,1978,2000]" italics="true" pageId="9" pageNumber="526">Megadytes magnus</emphasis>
</taxonomicName>
: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) median lobe in ventro-apical view to show the long nose-like tip. Scale bars = 1.5 mm.
</paragraph>
</caption>
<caption id="D4F9F3B8FFB9FFE8C1B06933FEFF4A9C" httpUri="https://zenodo.org/record/2647364/files/figure.png" pageId="10" pageNumber="527" startId="10.[151,249,1844,1866]" targetBox="[166,1427,204,1822]" targetPageId="10">
<paragraph id="8039A330FFB9FFE8C1B06933FEFF4A9C" blockId="10.[151,1436,1844,1927]" pageId="10" pageNumber="527">
<emphasis id="B2F27F22FFB9FFE8C1B06933FEF24A51" bold="true" box="[151,275,1844,1866]" pageId="10" pageNumber="527">FIGURE 7.</emphasis>
<taxonomicName id="4786D8B3FFB9FFE8C03D6933FE394A51" box="[282,472,1844,1866]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="527" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB9FFE8C03D6933FE394A51" box="[282,472,1844,1866]" italics="true" pageId="10" pageNumber="527">Megadytes ducalis</emphasis>
</taxonomicName>
, male from collection Guignot: A) frontal view, B) ventral view of head and protarsus, C) left middle leg, D) dorso-lateral view of left side of head, pronotum and elytral base, E) ventro-lateral view of left hind leg. Scale bars = 5 mm.
</paragraph>
</caption>
<caption id="D4F9F3B8FFB8FFE9C1B0697BFEFC4AF5" httpUri="https://zenodo.org/record/2647366/files/figure.png" pageId="11" pageNumber="528" startId="11.[151,249,1916,1938]" targetBox="[213,1368,207,1893]" targetPageId="11">
<paragraph id="8039A330FFB8FFE9C1B0697BFEFC4AF5" blockId="11.[151,1436,1916,2030]" pageId="11" pageNumber="528">
<emphasis id="B2F27F22FFB8FFE9C1B0697BFEF34A89" bold="true" box="[151,274,1916,1938]" pageId="11" pageNumber="528">FIGURE 8.</emphasis>
<taxonomicName id="4786D8B3FFB8FFE9C03E697BFE1E4A89" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1829" box="[281,511,1916,1938]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="528" phylum="Arthropoda" rank="species" species="lherminieri">
<emphasis id="B2F27F22FFB8FFE9C03E697BFE1E4A89" box="[281,511,1916,1938]" italics="true" pageId="11" pageNumber="528">Megadytes lherminieri</emphasis>
</taxonomicName>
, male, Ecuador, Esmeraldas: A) ventral view of head, left protarsus and left middle leg, B) dorsal view of left protarsus, C) ventro-lateral view of left hind leg;
<taxonomicName id="4786D8B3FFB8FFE9C246699BFC324AAA" authorityName="Tremouilles &amp; Bachmann" authorityYear="1980" box="[865,979,1948,1969]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="528" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="B2F27F22FFB8FFE9C246699BFC324AAA" box="[865,979,1948,1969]" italics="true" pageId="11" pageNumber="528">M. magnus</emphasis>
</taxonomicName>
, male, Paraguay, cerro Lambaré: D) ventral view of head, right protarsus and right middle leg, E) dorsal view of left protarsus, F) ventro-lateral view of left hind leg. Scale bars = 5 mm.
</paragraph>
</caption>
<subSubSection id="C89CF0BBFFBFFFEEC1B06E90FEAC4FEB" pageId="12" pageNumber="529" type="materials_examined">
<paragraph id="8039A330FFBFFFEEC1B06E90FDC34DB4" blockId="12.[151,1436,151,968]" box="[151,546,151,176]" pageId="12" pageNumber="529">
<materialsCitation id="30EEA96DFFBFFFEEC1B06E90FDFC4DB4" ID-GBIF-Occurrence="2236620154" box="[151,541,151,176]" country="Argentina" location="Santa Fe" pageId="12" pageNumber="529" specimenCount="1" stateProvince="Santa Fe" typeStatus="holotype">
<emphasis id="B2F27F22FFBFFFEEC1B06E90FECC4DAB" bold="true" box="[151,301,151,176]" pageId="12" pageNumber="529">
<typeStatus id="5F3D1D92FFBFFFEEC1B06E90FF334DAB" box="[151,210,151,176]" pageId="12" pageNumber="529">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F891E3A0FFBFFFEEC01B6E90FE4E4DAB" box="[316,431,151,176]" name="Argentina" pageId="12" pageNumber="529">Argentina</collectingCountry>
,
<collectingRegion id="42426DD2FFBFFFEEC09E6E9FFDFC4DB4" box="[441,541,152,176]" country="Argentina" name="Santa Fe" pageId="12" pageNumber="529">Santa Fe</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8039A330FFBFFFEEC1E06EBBFEE44C5B" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="529">
<emphasis id="B2F27F22FFBFFFEEC1E06EBBFE104DCE" bold="true" box="[199,497,188,213]" pageId="12" pageNumber="529">Type material examined</emphasis>
.
<materialsCitation id="30EEA96DFFBFFFEEC3246EBBFEE04C5B" ID-GBIF-Occurrence="2236620169" collectingDate="1943-03" collectionCode="BMNH" collectorName="S. Fe Ceres &amp; Polmer &amp; Col. A. O. Bachmann" country="Argentina" location="Argentina" pageId="12" pageNumber="529" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B2F27F22FFBFFFEEC3246EBBFD914DCE" bold="true" box="[515,624,188,213]" pageId="12" pageNumber="529">
<typeStatus id="5F3D1D92FFBFFFEEC3246EBBFD914DCE" box="[515,624,188,213]" pageId="12" pageNumber="529" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="968068B9FFBFFFEEC3A46EBBFD374DCE" box="[643,726,188,213]" pageId="12" pageNumber="529" type="male">1 male</specimenCount>
, &quot;
<collectingCountry id="F891E3A0FFBFFFEEC3C96EBBFC844DCE" box="[750,869,188,213]" name="Argentina" pageId="12" pageNumber="529">Argentina</collectingCountry>
:
<collectorName id="2D73C6E6FFBFFFEEC2536EBBFC1F4DCF" box="[884,1022,188,212]" pageId="12" pageNumber="529">S. Fe Ceres</collectorName>
<date id="F43885F0FFBFFFEEC52E6EBBFB854DCF" box="[1033,1124,188,212]" pageId="12" pageNumber="529" value="1943-03">
<collectingDate id="E47C7C18FFBFFFEEC52E6EBBFB854DCF" box="[1033,1124,188,212]" pageId="12" pageNumber="529" value="1943-03">III.1943</collectingDate>
</date>
leg.
<collectorName id="2D73C6E6FFBFFFEEC5826EBBFB184DCE" box="[1189,1273,188,213]" pageId="12" pageNumber="529">Polmer</collectorName>
&quot; [handwritten label], &quot;
<typeStatus id="5F3D1D92FFBFFFEEC1C96EE7FE734DE3" box="[238,402,224,248]" pageId="12" pageNumber="529" type="paratype">PARATYPUS</typeStatus>
&quot; [red printed label], &quot;Brit. Mus. 1989-185&quot; [printed label], &quot;
<collectorName id="2D73C6E6FFBFFFEEC5186ED8FAD24DE3" box="[1087,1331,223,248]" pageId="12" pageNumber="529">Col. A.O. Bachmann</collectorName>
&quot; [printed label], &quot;
<taxonomicName id="4786D8B3FFBFFFEEC1D66F03FBE94C06" authority="E. R. Tremouilles y A. O. Bachmann" authorityName="E. R. Tremouilles y A. O. Bachmann" box="[241,1032,260,285]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="species" species="magnus" subGenus="Bifurcitus">Megadytes (Bifurcitus) magnus E.R. Tremouilles y A.O. Bachmann</taxonomicName>
ano 1979&quot; [red handwritten label] (
<collectionCode id="E6973BF5FFBFFFEEC1B86F2FFF184C5B" box="[159,249,296,320]" pageId="12" pageNumber="529">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8039A330FFBFFFEEC1E06F4BFDB04F03" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="529">
<emphasis id="B2F27F22FFBFFFEEC1E06F4BFD6B4C7E" bold="true" box="[199,650,332,357]" pageId="12" pageNumber="529">
Other material examined.
<materialsCitation id="30EEA96DFFBFFFEEC3306F4BFD6B4C7E" ID-GBIF-Occurrence="2236620184" box="[535,650,332,357]" collectingDate="1990-02-02" collectionCode="CLH" collectorName="U. Drechsel" country="Paraguay" location="Chaco Dep." pageId="12" pageNumber="529" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Asuncion">
<collectingCountry id="F891E3A0FFBFFFEEC3306F4BFD6B4C7E" box="[535,650,332,357]" name="Paraguay" pageId="12" pageNumber="529">Paraguay</collectingCountry>
</materialsCitation>
</emphasis>
:
<specimenCount id="968068B9FFBFFFEEC3876F4BFD144C7E" box="[672,757,332,357]" pageId="12" pageNumber="529" type="male">1 male</specimenCount>
,
<specimenCount id="968068B9FFBFFFEEC2206F4BFC9C4C7E" box="[775,893,332,357]" pageId="12" pageNumber="529" type="female">2 females</specimenCount>
, &quot;
<collectingRegion id="42426DD2FFBFFFEEC2B06F4BFBE04C7E" box="[919,1025,332,357]" country="Paraguay" name="Asuncion" pageId="12" pageNumber="529">Paraguay</collectingRegion>
/
<location id="8559F5EBFFBFFFEEC52D6F4BFB774C7F" LSID="urn:lsid:plazi:treatment:082F1226FFBBFFEEC1B0696DFD374ED3:8559F5EBFFBFFFEEC52D6F4BFB774C7F" box="[1034,1174,332,357]" country="Paraguay" name="Chaco Dep." pageId="12" pageNumber="529" stateProvince="Asuncion">Chaco Dep.</location>
<collectingRegion id="42426DD2FFBFFFEEC5846F4BFAF04C7F" box="[1187,1297,332,356]" country="Paraguay" name="Boqueron" pageId="12" pageNumber="529">Boquéron</collectingRegion>
<locationDeviation id="918BBA12FFBFFFEEC4076F4BFE674C93" location="Mariscal Estigarribia" pageId="12" pageNumber="529">
<quantity id="477E0ED5FFBFFFEEC4076F4BFA8F4C7E" box="[1312,1390,332,357]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="529" unit="km" value="10.0">10 km</quantity>
SE Mariscal Estigarribia
</locationDeviation>
,
<date id="F43885F0FFBFFFEEC0B66F77FE144C93" box="[401,501,368,392]" pageId="12" pageNumber="529" value="1990-02-02">
<collectingDate id="E47C7C18FFBFFFEEC0B66F77FE144C93" box="[401,501,368,392]" pageId="12" pageNumber="529" value="1990-02-02">2.2.1990</collectingDate>
</date>
, leg.
<collectorName id="2D73C6E6FFBFFFEEC3156F77FD5B4C93" box="[562,698,367,392]" pageId="12" pageNumber="529">U. Drechsel</collectorName>
&quot; (CLH);
<materialsCitation id="30EEA96DFFBFFFEEC2036F77FC124CB7" ID-GBIF-Occurrence="2236620185" collectingDate="1990-10-02" collectionCode="CLH, ZSM" collectorName="U. Drechsel" country="Paraguay" location="Chaco Dep." pageId="12" pageNumber="529" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Asuncion">
<specimenCount id="968068B9FFBFFFEEC2036F77FC9C4C93" box="[804,893,367,392]" pageId="12" pageNumber="529" type="male">4 males</specimenCount>
,
<specimenCount id="968068B9FFBFFFEEC2AE6F68FC194C93" box="[905,1016,367,392]" pageId="12" pageNumber="529" type="female">6 females</specimenCount>
, &quot;
<collectingRegion id="42426DD2FFBFFFEEC52B6F68FB974C93" box="[1036,1142,367,392]" country="Paraguay" name="Asuncion" pageId="12" pageNumber="529">Paraguay</collectingRegion>
/
<location id="8559F5EBFFBFFFEEC5596F68FAE54C9C" LSID="urn:lsid:plazi:treatment:082F1226FFBBFFEEC1B0696DFD374ED3:8559F5EBFFBFFFEEC5596F68FAE54C9C" box="[1150,1284,367,392]" country="Paraguay" name="Chaco Dep." pageId="12" pageNumber="529" stateProvince="Asuncion">Chaco Dep.</location>
<collectingRegion id="42426DD2FFBFFFEEC42C6F77FA984C93" box="[1291,1401,368,392]" country="Paraguay" name="Boqueron" pageId="12" pageNumber="529">Boquéron</collectingRegion>
<locationDeviation id="918BBA12FFBFFFEEC4A76F77FE0B4CB6" location="Mariscal Estigarribia" pageId="12" pageNumber="529">
<quantity id="477E0ED5FFBFFFEEC4A76F77FF5A4CB6" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="529" unit="km" value="10.0">10 km</quantity>
SE Mariscal Estigarribia
</locationDeviation>
,
<date id="F43885F0FFBFFFEEC0DD6F93FD8D4CB7" box="[506,620,404,428]" pageId="12" pageNumber="529" value="1990-10-02">
<collectingDate id="E47C7C18FFBFFFEEC0DD6F93FD8D4CB7" box="[506,620,404,428]" pageId="12" pageNumber="529" value="1990-10-02">10.2.1990</collectingDate>
</date>
, leg.
<collectorName id="2D73C6E6FFBFFFEEC3936F92FCA24CB6" box="[692,835,404,429]" pageId="12" pageNumber="529">U. Drechsel</collectorName>
&quot; (
<collectionCode id="E6973BF5FFBFFFEEC2476F93FC7D4CB7" box="[864,924,404,428]" pageId="12" pageNumber="529">CLH</collectionCode>
,
<collectionCode id="E6973BF5FFBFFFEEC28A6F93FC0A4CB7" box="[941,1003,404,428]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="529" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="30EEA96DFFBFFFEEC5226F93FD134CCB" ID-GBIF-Occurrence="2236620180" collectingDate="1992-06-05" collectionCode="CLH" collectorName="U. Drechsel &amp; Coll. Hendrich" country="Paraguay" location="Cerro Lambare" pageId="12" pageNumber="529" specimenCount="1" specimenCount-male="1" stateProvince="Asuncion">
<specimenCount id="968068B9FFBFFFEEC5226F93FBB94CB6" box="[1029,1112,404,429]" pageId="12" pageNumber="529" type="male">1 male</specimenCount>
, &quot;
<collectingRegion id="42426DD2FFBFFFEEC5556F93FB3A4CB7" box="[1138,1243,404,428]" country="Paraguay" name="Asuncion" pageId="12" pageNumber="529">Paraguay</collectingRegion>
/
<location id="8559F5EBFFBFFFEEC5C36F93FA794CB6" LSID="urn:lsid:plazi:treatment:082F1226FFBBFFEEC1B0696DFD374ED3:8559F5EBFFBFFFEEC5C36F93FA794CB6" box="[1252,1432,404,429]" country="Paraguay" name="Cerro Lambare" pageId="12" pageNumber="529" stateProvince="Asuncion">Cerro Lambaré</location>
,
<date id="F43885F0FFBFFFEEC1B06FB0FF1A4CCB" box="[151,251,439,464]" pageId="12" pageNumber="529" value="1992-06-05">
<collectingDate id="E47C7C18FFBFFFEEC1B06FB0FF1A4CCB" box="[151,251,439,464]" pageId="12" pageNumber="529" value="1992-06-05">5.6.1992</collectingDate>
</date>
, leg.
<collectorName id="2D73C6E6FFBFFFEEC0616FBFFE364CCB" box="[326,471,439,464]" pageId="12" pageNumber="529">U. Drechsel</collectorName>
/
<collectorName id="2D73C6E6FFBFFFEEC0C76FB0FD6C4CCB" box="[480,653,439,464]" pageId="12" pageNumber="529">Coll. Hendrich</collectorName>
&quot; (
<collectionCode id="E6973BF5FFBFFFEEC38B6FBFFD0B4CCB" box="[684,746,440,464]" pageId="12" pageNumber="529">CLH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="30EEA96DFFBFFFEEC2226FBFFD114CEF" ID-GBIF-Occurrence="2236620192" collectingDate="31.10. 1991, 5.6. 1992" collectionCode="CLH" collectorName="U. Drechsel &amp; Coll. Hendrich" country="Paraguay" location="Dep. Central" pageId="12" pageNumber="529" specimenCount="1" specimenCount-male="1" stateProvince="Asuncion">
<specimenCount id="968068B9FFBFFFEEC2226FBFFCBA4CCB" box="[773,859,439,464]" pageId="12" pageNumber="529" type="male">1 male</specimenCount>
, &quot;
<collectingRegion id="42426DD2FFBFFFEEC2506FBFFC024CCB" box="[887,995,440,464]" country="Paraguay" name="Asuncion" pageId="12" pageNumber="529">Paraguay</collectingRegion>
/
<location id="8559F5EBFFBFFFEEC2CB6FBFFB624CCB" LSID="urn:lsid:plazi:treatment:082F1226FFBBFFEEC1B0696DFD374ED3:8559F5EBFFBFFFEEC2CB6FBFFB624CCB" box="[1004,1155,439,464]" country="Paraguay" name="Dep. Central" pageId="12" pageNumber="529" stateProvince="Asuncion">Dep. Central</location>
,
<collectingRegion id="42426DD2FFBFFFEEC5B16FB0FAE54CCB" box="[1174,1284,439,464]" country="Paraguay" name="Asuncion" pageId="12" pageNumber="529">Asunción</collectingRegion>
,
<collectingDate id="E47C7C18FFBFFFEEC4306FBFFF1A4CEE" isEnumeration="true" pageId="12" pageNumber="529">
<date id="F43885F0FFBFFFEEC4306FBFFA794CCB" box="[1303,1432,440,464]" pageId="12" pageNumber="529" value="1991-10-31">31.10.1991</date>
,
<date id="F43885F0FFBFFFEEC1B06FDBFF1A4CEE" box="[151,251,476,501]" pageId="12" pageNumber="529" value="1992-06-05">5.6.1992</date>
</collectingDate>
, leg.
<collectorName id="2D73C6E6FFBFFFEEC0616FDAFE374CEE" box="[326,470,476,501]" pageId="12" pageNumber="529">U. Drechsel</collectorName>
/
<collectorName id="2D73C6E6FFBFFFEEC0F96FDBFD6A4CEE" box="[478,651,476,501]" pageId="12" pageNumber="529">Coll. Hendrich</collectorName>
&quot; (
<collectionCode id="E6973BF5FFBFFFEEC38D6FDBFD094CEF" box="[682,744,476,500]" pageId="12" pageNumber="529">CLH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="30EEA96DFFBFFFEEC2246FDBFDAC4F03" ID-GBIF-Occurrence="2236620172" collectingDate="1984-10-12" collectionCode="CLH" collectorName="Gottwald" country="Brazil" location="Brasilien" pageId="12" pageNumber="529" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo">
<emphasis id="B2F27F22FFBFFFEEC2246FDBFCAD4CEE" bold="true" box="[771,844,476,501]" pageId="12" pageNumber="529">
<collectingCountry id="F891E3A0FFBFFFEEC2246FDBFCAD4CEE" box="[771,844,476,501]" name="Brazil" pageId="12" pageNumber="529">Brazil</collectingCountry>
</emphasis>
:
<specimenCount id="968068B9FFBFFFEEC2456FDBFC2C4CEE" box="[866,973,476,501]" pageId="12" pageNumber="529" type="female">1 female</specimenCount>
, &quot;
<location id="8559F5EBFFBFFFEEC2CF6FDBFBB34CEE" LSID="urn:lsid:plazi:treatment:082F1226FFBBFFEEC1B0696DFD374ED3:8559F5EBFFBFFFEEC2CF6FDBFBB34CEE" box="[1000,1106,476,501]" country="Brazil" name="Brasilien" pageId="12" pageNumber="529" stateProvince="Sao Paulo">Brasilien</location>
,
<collectingRegion id="42426DD2FFBFFFEEC5426FDBFB004CEE" box="[1125,1249,476,501]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="529">Sao Paulo</collectingRegion>
,
<date id="F43885F0FFBFFFEEC5D46FDBFA954CEF" box="[1267,1396,476,500]" pageId="12" pageNumber="529" value="1984-10-12">
<collectingDate id="E47C7C18FFBFFFEEC5D46FDBFA954CEF" box="[1267,1396,476,500]" pageId="12" pageNumber="529" value="1984-10-12">10.12.1984</collectingDate>
</date>
, in swimming pool,
<collectorName id="2D73C6E6FFBFFFEEC0736FF8FE5D4F03" box="[340,444,511,536]" pageId="12" pageNumber="529">Gottwald</collectorName>
leg.&quot; (
<collectionCode id="E6973BF5FFBFFFEEC3206C07FDA44F03" box="[519,581,512,536]" pageId="12" pageNumber="529">CLH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8039A330FFBFFFEEC1E06C23FEAC4FEB" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="529">
<emphasis id="B2F27F22FFBFFFEEC1E06C23FE764F26" bold="true" box="[199,407,548,573]" pageId="12" pageNumber="529">Descriptive notes</emphasis>
. A description of this species was given by
<bibRefCitation id="E417DEC1FFBFFFEEC28F6C23FACA4F27" author="Tremouilles, E. R. &amp; Bachmann, A. O." box="[936,1323,548,573]" pageId="12" pageNumber="529" pagination="101 - 125" refId="ref7687" refString="Tremouilles, E. R. &amp; Bachmann, A. O. (1980) La tribu Cybisterini en la Argentina (Coleoptera, Dytiscidae). Revista de la Sociedad Entomologica Argentina, 39, 101 - 125." type="journal article" year="1980">Trémouilles &amp; Bachmann (1980)</bibRefCitation>
. Here we provide photos of the habitus (
<figureCitation id="18BDBFB5FFBFFFEEC0CD6C40FD804F44" box="[490,609,583,608]" captionStart="FIGURE 1" captionStartId="2.[151,249,1676,1698]" captionTargetBox="[171,1414,205,1629]" captionTargetId="figure@2.[166,1421,193,1654]" captionTargetPageId="2" captionText="FIGURE 1. Habitus in dorsal view of: A) M. ducalis, B) M. lherminieri, C) M. magnus; habitus in ventral view of: D) M. ducalis, E) M. lherminieri (Ecuador, Esmeraldas), F) M. magnus (Paraguay, Cerro Lambaré)." httpUri="https://zenodo.org/record/2647352/files/figure.png" pageId="12" pageNumber="529">Figs 1C, F</figureCitation>
,
<figureCitation id="18BDBFB5FFBFFFEEC34B6C4FFD6C4F7B" box="[620,653,584,608]" captionStart="FIGURE 2" captionStartId="3.[151,249,1959,1981]" captionTargetBox="[193,1409,484,1938]" captionTargetId="figure@3.[151,1436,481,1938]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in lateral view of: A) M. ducalis, B) M. lherminieri, C) M. magnus." httpUri="https://zenodo.org/record/2647354/files/figure.png" pageId="12" pageNumber="529">2C</figureCitation>
) and focus on morphological characters of the male in different views (
<figureCitation id="18BDBFB5FFBFFFEEC1B86C6BFF094F9F" box="[159,232,620,645]" captionStart="FIGURE 8" captionStartId="11.[151,249,1916,1938]" captionTargetBox="[213,1368,207,1893]" captionTargetId="figure@11.[204,1372,193,1893]" captionTargetPageId="11" captionText="FIGURE 8. Megadytes lherminieri, male, Ecuador, Esmeraldas: A) ventral view of head, left protarsus and left middle leg, B) dorsal view of left protarsus, C) ventro-lateral view of left hind leg; M. magnus, male, Paraguay, cerro Lambaré: D) ventral view of head, right protarsus and right middle leg, E) dorsal view of left protarsus, F) ventro-lateral view of left hind leg. Scale bars = 5 mm." httpUri="https://zenodo.org/record/2647366/files/figure.png" pageId="12" pageNumber="529">Figs 8</figureCitation>
D–F). The median lobe with both parameres of a specimen from Mariscal Estigarribia in
<collectingCountry id="F891E3A0FFBFFFEEC4376C6AFA984F9E" box="[1296,1401,621,645]" name="Paraguay" pageId="12" pageNumber="529">Paraguay</collectingCountry>
, is figured in ventral view (
<figureCitation id="18BDBFB5FFBFFFEEC0806C88FDE04FB3" box="[423,513,655,680]" captionStart="FIGURE 6" captionStartId="9.[151,249,1978,2000]" captionTargetBox="[199,1381,195,1953]" captionTargetId="figure@9.[199,1381,195,1953]" captionTargetPageId="9" captionText="FIGURE 6. Megadytes magnus: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) median lobe in ventro-apical view to show the long nose-like tip. Scale bars = 1.5 mm." httpUri="https://zenodo.org/record/2647362/files/figure.png" pageId="12" pageNumber="529">Fig. 6A</figureCitation>
), the tip of the median lobe in lateral view (
<figureCitation id="18BDBFB5FFBFFFEEC2CB6C88FBA54FB3" box="[1004,1092,655,680]" captionStart="FIGURE 6" captionStartId="9.[151,249,1978,2000]" captionTargetBox="[199,1381,195,1953]" captionTargetId="figure@9.[199,1381,195,1953]" captionTargetPageId="9" captionText="FIGURE 6. Megadytes magnus: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) median lobe in ventro-apical view to show the long nose-like tip. Scale bars = 1.5 mm." httpUri="https://zenodo.org/record/2647362/files/figure.png" pageId="12" pageNumber="529">Fig. 6B</figureCitation>
), and the tip of median lobe in ventro-apical view (
<figureCitation id="18BDBFB5FFBFFFEEC0586CB3FE3A4FD6" box="[383,475,692,717]" captionStart="FIGURE 6" captionStartId="9.[151,249,1978,2000]" captionTargetBox="[199,1381,195,1953]" captionTargetId="figure@9.[199,1381,195,1953]" captionTargetPageId="9" captionText="FIGURE 6. Megadytes magnus: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) median lobe in ventro-apical view to show the long nose-like tip. Scale bars = 1.5 mm." httpUri="https://zenodo.org/record/2647362/files/figure.png" pageId="12" pageNumber="529">Fig. 6C</figureCitation>
). Note that
<figureCitation id="18BDBFB5FFBFFFEEC34E6CB3FCF64FD7" box="[617,791,692,717]" captionStart="FIGURE 6" captionStartId="9.[151,249,1978,2000]" captionTargetBox="[199,1381,195,1953]" captionTargetId="figure@9.[199,1381,195,1953]" captionTargetPageId="9" captionText="FIGURE 6. Megadytes magnus: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) median lobe in ventro-apical view to show the long nose-like tip. Scale bars = 1.5 mm." httpUri="https://zenodo.org/record/2647362/files/figure.png" pageId="12" pageNumber="529">Figs 6A and C</figureCitation>
show the median lobe of the same individual, tilted at different angles.
</paragraph>
</subSubSection>
<subSubSection id="C89CF0BBFFBFFFEEC1E06CFBFD374ED3" pageId="12" pageNumber="529" type="distribution">
<paragraph id="8039A330FFBFFFEEC1E06CFBFB314E23" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="529">
<emphasis id="B2F27F22FFBFFFEEC1E06CFBFEB84E0E" bold="true" box="[199,345,764,789]" pageId="12" pageNumber="529">Distribution</emphasis>
. Southern part of tropical and subtropical South America. The species was described from
<collectingCountry id="F891E3A0FFBFFFEEC1B06D18FEE84E23" box="[151,265,799,824]" name="Argentina" pageId="12" pageNumber="529">Argentina</collectingCountry>
,
<collectingCountry id="F891E3A0FFBFFFEEC0336D27FE9D4E23" box="[276,380,800,824]" name="Paraguay" pageId="12" pageNumber="529">Paraguay</collectingCountry>
and
<collectingCountry id="F891E3A0FFBFFFEEC0946D27FDF74E23" box="[435,534,800,824]" name="Uruguay" pageId="12" pageNumber="529">Uruguay</collectingCountry>
, and later recorded from
<collectingCountry id="F891E3A0FFBFFFEEC2086D18FC924E23" box="[815,883,799,824]" name="Brazil" pageId="12" pageNumber="529">Brazil</collectingCountry>
(Trémouilles 1989b) (
<figureCitation id="18BDBFB5FFBFFFEEC5576D18FB234E23" box="[1136,1218,799,824]" captionStart="FIGURE 11" captionStartId="16.[151,249,1904,1926]" captionTargetBox="[151,1436,193,1883]" captionTargetId="figure@16.[151,1436,193,1883]" captionTargetPageId="16" captionText="FIGURE 11. Distribution of the subgenus Bifurcitus in Central and South America: White line: distribution of M. lherminieri; black line: distribution of M. magnus; red spot: only known locality for M. ducalis." httpUri="https://zenodo.org/record/2647372/files/figure.png" pageId="12" pageNumber="529">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8039A330FFBFFFEEC1E06D43FD374ED3" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="529">
<emphasis id="B2F27F22FFBFFFEEC1E06D43FEC24E46" bold="true" box="[199,291,836,861]" pageId="12" pageNumber="529">Habitat</emphasis>
. Unknown, probably similar to
<taxonomicName id="4786D8B3FFBFFFEEC3B76D42FCD54E46" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1829" box="[656,820,836,861]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="species" species="lherminieri">
<emphasis id="B2F27F22FFBFFFEEC3B76D42FCD54E46" box="[656,820,836,861]" italics="true" pageId="12" pageNumber="529">M. lherminieri</emphasis>
</taxonomicName>
. Most of the specimens in
<collectingCountry id="F891E3A0FFBFFFEEC54C6D42FB324E46" box="[1131,1235,837,861]" name="Paraguay" pageId="12" pageNumber="529">Paraguay</collectingCountry>
were collected at light in a savannah region with temporary swamps and permanent farm dams. The swimming pool in Sao Paulo, were the single female has been collected, was not far away from a big reservoir or dam, overgrown with water lilies and water hyacinths (S. Gottwald pers. com.).
</paragraph>
</subSubSection>
</treatment>
</document>
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