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<document id="3EDABBEC4C2AF22E6BA63C85D962CA42" ID-DOI="10.2478/aemnp-2018-0013" ID-GBIF-Dataset="7b94dd56-d3a7-4462-8da2-72ceb277835f" ID-ISSN="1804-6487" ID-Zenodo-Dep="3699163" ID-ZooBank="BF2CDC12-C555-4187-BFAB-274C9C674A5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583496890333" checkinUser="tatiana" docAuthor="Löbl, Ivan" docDate="2018" docId="03F94B1B2173FF866584E4D707A3821F" docLanguage="en" docName="ActaEntMusNatPra.58.1.151-165.pdf.imf" docOrigin="Acta Entomologica Musei Nationalis Pragae 58 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Scaphisoma Leach 1815" docType="key" docVersion="12" lastPageNumber="164" masterDocId="FFC033632178FF8B656AE114055B8309" masterDocTitle="A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="165" masterPageNumber="151" pageNumber="162" updateTime="1698725294610" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="1562D2CD40BB04D39D2D1E4889009A9F">A review of Scaphisomatini from Sulawesi, with descriptions of ten new species (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
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<mods:namePart id="91EE312B97CBA852989DED13ADAFAF92">Löbl, Ivan</mods:namePart>
<mods:affiliation id="9FE604A40D869FB7C32EE20A89FFB58E">Muséum d’histoire naturelle, Route de Malagnou 1, CH- 1208 Geneva, Switzerland</mods:affiliation>
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<mods:title id="8199E9CEB3F99D8C6BB18E772CDE4EE1">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="7D1A7F184838EF88DE911FE3BFBA0F7B">2018</mods:date>
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<mods:number id="3A3E5AF5EE580EC49712F91E77EB82F7">2018-05-30</mods:number>
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<treatment id="03F94B1B2173FF866584E4D707A3821F" ID-DOI="http://doi.org/10.5281/zenodo.3706105" ID-GBIF-Taxon="162467437" ID-Zenodo-Dep="3706105" LSID="urn:lsid:plazi:treatment:03F94B1B2173FF866584E4D707A3821F" httpUri="http://treatment.plazi.org/id/03F94B1B2173FF866584E4D707A3821F" lastPageId="13" lastPageNumber="164" pageId="11" pageNumber="162">
<subSubSection id="C34AA9862173FF806584E4D707FB86D3" box="[238,672,1475,1498]" pageId="11" pageNumber="162" type="nomenclature">
<paragraph id="8BEFFA0D2173FF806584E4D707FB86D3" blockId="11.[238,672,1475,1498]" box="[238,672,1475,1498]" pageId="11" pageNumber="162">
<heading id="D0A74D612173FF806584E4D707FB86D3" box="[238,672,1475,1498]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="162" reason="3">
<emphasis id="B924261F2173FF806584E4D707FB86D3" bold="true" box="[238,672,1475,1498]" pageId="11" pageNumber="162">
Key to
<collectingRegion id="499434EF2173FF806455E4D704F986D3" box="[319,418,1475,1498]" country="Indonesia" name="Sulawesi Utara" pageId="11" pageNumber="162">Sulawesi</collectingRegion>
species of
<taxonomicName id="4C50818E2173FF806771E4D707FB86D3" ID-CoL="8VS35" authorityName="Leach" authorityYear="1815" box="[539,672,1475,1498]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="B924261F2173FF806771E4D707FB86D3" bold="true" box="[539,672,1475,1498]" italics="true" pageId="11" pageNumber="162">Scaphisoma</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34AA9862173FF8665FDE4E607A3821F" lastPageId="13" lastPageNumber="164" pageId="11" pageNumber="162" type="key">
<keyStep id="30A4E2A82173FF8065FDE4E607A38583" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF8065FDE4E607A38543" blockId="11.[151,760,1522,2058]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF8065FDE4E607A38543" pageId="11" pageNumber="162">
1 Pronotum and elytra both with distinct bicoloured colour pattern. Elytra with sutural striae not curved along pronotal lobe.
<emphasis id="B924261F2173FF8064F2E72607A38543" bold="true" box="[408,760,1586,1610]" pageId="11" pageNumber="162">
............
<taxonomicName id="4C50818E2173FF806486E72607C48543" ID-CoL="79T6S" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[492,671,1586,1610]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="versicoloreum" status="sp. nov.">
<emphasis id="B924261F2173FF806486E72607C48543" bold="true" box="[492,671,1586,1610]" italics="true" pageId="11" pageNumber="162">S. versicoloreum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642173FF8067CFE72607A38543" box="[677,760,1586,1610]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF8065FDE74007A38583" blockId="11.[151,760,1522,2058]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF8065FDE74007A38583" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF8065FDE74005FF8562" box="[151,164,1620,1643]" italics="true" pageId="11" pageNumber="162">‒</emphasis>
Pronotum unicoloured, elytra about as unicoloured as pronotum, or with distinctive colour pattern. ........... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF8065FDE78607A38443" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF8065FDE78607A385E3" blockId="11.[151,760,1522,2058]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF8065FDE78607A385E3" pageId="11" pageNumber="162">
2 Elytra bicoloured, mostly dark, each with yellowish, well delimited, subapical transverse fascia. ................... ................................................
<emphasis id="B924261F2173FF806487E7C607A385E3" bold="true" box="[493,760,1746,1770]" pageId="11" pageNumber="162">
<taxonomicName id="4C50818E2173FF806487E7C607C585E3" ID-CoL="6XTCS" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[493,670,1746,1770]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="flavolineatum" status="sp. nov.">
<emphasis id="B924261F2173FF806487E7C607C585E3" bold="true" box="[493,670,1746,1770]" italics="true" pageId="11" pageNumber="162">S. flavolineatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642173FF8067CFE7C607A385E3" box="[677,760,1746,1770]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF8065FDE7E007A38443" blockId="11.[151,760,1522,2058]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF8065FDE7E007A38443" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF8065FDE7E005FF8402" box="[151,164,1780,1803]" italics="true" pageId="11" pageNumber="162">‒</emphasis>
Elytra either unicoloured or if bicoloured, elytral pattern consist of reddish spots; never with light subapical fasciae. ................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF8065FDE64607A38B03" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF8065FDE64607A384A3" blockId="11.[151,760,1522,2058]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF8065FDE64607A384A3" pageId="11" pageNumber="162">3 Elytra with sutural striae extending along bases to form basal striae. Male protarsomere I strongly enlarged, as wide as or wider than tibial apices. ................ 4</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF8065FDE6A007A38B03" blockId="11.[151,760,1522,2058]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF8065FDE6A007A38B03" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF8065FDE6A005FF84C2" box="[151,164,1972,1995]" italics="true" pageId="11" pageNumber="162">‒</emphasis>
Elytra with sutural striae ending near pronotal lobe, not extended laterad to form basal striae. Male protarsomere I usually narrower than tibial apices. ............ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E1AB00C782BF" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E1AB00C7823E" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E1AB00C7823E" pageId="11" pageNumber="162">
4 Length
<quantity id="4CA857E82173FF8066DFE1AA013783DF" box="[949,1132,190,214]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.45" metricValueMin="2.15" pageId="11" pageNumber="162" unit="mm" value="2.3" valueMax="2.45" valueMin="2.15">2.15 to 2.45 mm</quantity>
. Body dark brown to black, mesepimera about as long as third of intervals between them and mesocoxae. Abdominal ventrite I microsculptured. ......................
<taxonomicName id="4C50818E2173FF8061D9E00A00C7823E" ID-CoL="79TC9" authority="Lobl, 2012" authorityName="Lobl" authorityYear="2012" box="[1203,1436,286,311]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="latitarse">
<emphasis id="B924261F2173FF8061D9E00A007C823F" bold="true" box="[1203,1319,286,310]" italics="true" pageId="11" pageNumber="162">S. latitarse</emphasis>
<bibRefCitation id="EFC187FC2173FF806047E00B00C7823E" author="LOBL I." box="[1325,1436,287,311]" firstAuthor="Lobl" journalOrPublisher="Vernate" pageId="11" pageNumber="162" pagination="309 - 317" part="31" refId="ref12955" refString="LOBL I. 2012 a: Irianscapha dimorpha, an unusual new Scaphisomatini from Western New Guinea, and two new Indonesian species of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae). Vernate 31: 309 - 317." title="Irianscapha dimorpha, an unusual new Scaphisomatini from Western New Guinea, and two new Indonesian species of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae)" type="journal article" year="2012">Löbl, 2012</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E05400C782BF" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E05400C782BF" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF806656E0540612825E" box="[828,841,320,343]" italics="true" pageId="11" pageNumber="162">‒</emphasis>
Length
<quantity id="4CA857E82173FF8066D2E02B0128825F" box="[952,1139,319,343]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" metricValueMax="2.24" metricValueMin="1.9" pageId="11" pageNumber="162" unit="mm" value="2.07" valueMax="2.24" valueMin="1.9">1.90 to 2.24 mm</quantity>
long. Body light, reddish- -brown, mesepimera shorter than intervals between them and mesocoxae. Abdominal ventrite I not microsculptured. ................................
<emphasis id="B924261F2173FF80619AE08A00C782BF" bold="true" box="[1264,1436,414,438]" pageId="11" pageNumber="162">
<taxonomicName id="4C50818E2173FF80619AE08A001882BF" ID-CoL="6XTDD" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1264,1347,414,438]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="hulai" status="sp. nov.">
<emphasis id="B924261F2173FF80619AE08A001882BF" bold="true" box="[1264,1347,414,438]" italics="true" pageId="11" pageNumber="162">S. hulai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642173FF806023E08A00C782BF" box="[1353,1436,414,438]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E0AA00C7813E" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E0AA00C782FE" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E0AA00C782FE" pageId="11" pageNumber="162">5 Elytra with sutural striae shortened, starting behind level of scutellum. .......................................................... 6</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E31400C7813E" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E31400C7813E" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF806656E3140612811E" box="[828,841,512,535]" italics="true" pageId="11" pageNumber="162">‒</emphasis>
Elytra with sutural striae not shortened, starting at elytral bases. ................................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E32B00F9801F" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E32B00C781BE" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E32B00C781BE" pageId="11" pageNumber="162">
6 Length
<quantity id="4CA857E82173FF8066D9E32A0138815F" box="[947,1123,574,599]" metricMagnitude="-3" metricUnit="m" metricValue="1.1749999999999998" metricValueMax="1.2" metricValueMin="1.15" pageId="11" pageNumber="162" unit="mm" value="1.1749999999999998" valueMax="1.2" valueMin="1.15">1.15 to 1.20 mm</quantity>
, pronotum and elytra uniformly reddish-brown. Pronotum with punctation coarser near lateral margins than on centre of disc. ........... ...........................................................
<taxonomicName id="4C50818E2173FF8061BFE38A00C781BE" ID-CoL="6XTFK" authority="Lobl, 1983" authorityName="Lobl" authorityYear="1983" box="[1237,1436,670,695]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="napu">
<emphasis id="B924261F2173FF8061BFE38A007C81BF" bold="true" box="[1237,1319,670,694]" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF8061BFE38A01B981BF" bold="true" box="[1237,1250,670,694]" italics="true" pageId="11" pageNumber="162">S</emphasis>
.
<emphasis id="B924261F2173FF806185E38A007C81BF" bold="true" box="[1263,1319,670,694]" italics="true" pageId="11" pageNumber="162">napu</emphasis>
</emphasis>
<bibRefCitation id="EFC187FC2173FF806047E38B00C781BE" author="LOBL I." box="[1325,1436,671,695]" firstAuthor="Lobl" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pageId="11" pageNumber="162" pagination="285 - 293" part="56" refId="ref12872" refString="LOBL I. 1983: Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 56: 285 - 293." title="Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien" type="journal article" year="1983">Löbl, 1983</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E3D400F9801F" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E3D400F9801F" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF806656E3D4061281DE" box="[828,841,704,727]" italics="true" pageId="11" pageNumber="162">‒</emphasis>
Length
<quantity id="4CA857E82173FF8066D9E3AB014E81DF" box="[947,1045,703,727]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="162" unit="mm" value="2.2">2.20 mm</quantity>
, elytra on apical fourth notably lighter than on remaining surface and pronotum. Pronotal punctation even throughout. .................................. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E20B00C780BF" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E20B00C7805F" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E20B00C7805F" pageId="11" pageNumber="162">
7 Metaventrite with mesal carina. Abdominal ventrite I entirely very finely punctate. ....
<emphasis id="B924261F2173FF8061DDE22A00C7805F" bold="true" box="[1207,1436,830,854]" pageId="11" pageNumber="162">
<taxonomicName id="4C50818E2173FF8061DDE22A0018805F" ID-CoL="6Y5MK" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1207,1347,830,854]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="carinatum" status="sp. nov.">
<emphasis id="B924261F2173FF8061DDE22A0018805F" bold="true" box="[1207,1347,830,854]" italics="true" pageId="11" pageNumber="162">S. carinatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642173FF806023E22A00C7805F" box="[1353,1436,830,854]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E27400C780BF" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E27400C780BF" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF806656E2740612807E" box="[828,841,864,887]" italics="true" pageId="11" pageNumber="162">‒</emphasis>
Metaventrite lacking mesal carina. Punctation on middle of abdominal ventrite I much coarser than on lateral parts. ..........................................
<emphasis id="B924261F2173FF80619EE28A00C780BF" bold="true" box="[1268,1436,926,950]" pageId="11" pageNumber="162">
<taxonomicName id="4C50818E2173FF80619EE28A002C80BF" ID-CoL="8VS35" authorityName="Leach" authorityYear="1815" box="[1268,1399,926,950]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="B924261F2173FF80619EE28A002C80BF" bold="true" box="[1268,1399,926,950]" italics="true" pageId="11" pageNumber="162">Scaphisoma</emphasis>
</taxonomicName>
sp.
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E2AB00C7873E" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E2AB00C780DE" blockId="11.[828,1442,190,2039]" box="[828,1436,959,983]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E2AB00C780DE" box="[828,1436,959,983]" pageId="11" pageNumber="162">8 Elytra with distinct bicolorous pattern. ...................... 9</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E2F400C7873E" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E2F400C7873E" pageId="11" pageNumber="162">
<emphasis id="B924261F2173FF806656E2F4061280FE" box="[828,841,992,1015]" italics="true" pageId="11" pageNumber="162">‒</emphasis>
Elytra entirely light reddish-brown to black, or with narrowly lighter apical margins, or somewhat darkened near apical margins. ............................................. 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E52B00C687DE" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E52B00C6879E" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E52B00C6879E" pageId="11" pageNumber="162">
9 Pronotum and elytra dark, blackish-brown to black, elytra each with one or two reddish spots, body length
<quantity id="4CA857E82173FF80660BE56B014B879F" box="[865,1040,1151,1175]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="11" pageNumber="162" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.20 to 1.40 mm</quantity>
. ......................................................... 10
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E58B00C687DE" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E58B00C687DE" pageId="11" pageNumber="162">– Elytral colour pattern different, body length 1.7 to 2.0 mm. .............................................................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E5CB00C786DE" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E5CB00C7865E" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E5CB00C7865E" pageId="11" pageNumber="162">
10 Elytra each with single reddish spot and with reddish apical area. Aedeagus with trifid apical process, internal sac with sclerotized, teeth-like structures. .............. ....................
<taxonomicName id="4C50818E2173FF806680E42A00C7865E" ID-CoL="79TFP" authority="Motschulsky, 1863" authorityName="Motschulsky" authorityYear="1863" box="[1002,1436,1342,1367]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="obliquemaculatum">
<emphasis id="B924261F2173FF806680E42A018A865F" bold="true" box="[1002,1233,1342,1366]" italics="true" pageId="11" pageNumber="162">S. obliquemaculatum</emphasis>
Motschulsky, 1863
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E44B00C786DE" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E44B00C786DE" pageId="11" pageNumber="162">
– Elytra each with two reddish spots, lacking light apical area. Aedeagus with apical process simple, internal sac membranous, lacking sclerotized structures. ............... ............................................................
<taxonomicName id="4C50818E2173FF8061B7E4AA00C786DE" ID-CoL="79TMM" authority="Lobl, 1983" authorityName="Lobl" authorityYear="1983" box="[1245,1436,1470,1495]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="bugi">
<emphasis id="B924261F2173FF8061B7E4AA007C86DF" bold="true" box="[1245,1319,1470,1494]" italics="true" pageId="11" pageNumber="162">S. bugi</emphasis>
<bibRefCitation id="EFC187FC2173FF806047E4AB00C786DE" author="LOBL I." box="[1325,1436,1471,1495]" firstAuthor="Lobl" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pageId="11" pageNumber="162" pagination="285 - 293" part="56" refId="ref12872" refString="LOBL I. 1983: Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 56: 285 - 293." title="Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien" type="journal article" year="1983">Löbl, 1983</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E4CB00C7845F" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E4CB00C7859E" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E4CB00C7859E" pageId="11" pageNumber="162">
11 Elytral punctation very dense and coarse, much coarser than pronotal punctation, puncture diameters in part larger than puncture intervals. Metaventrite with antecoxal puncture row. Aedeagus symmetrical, with apically widened, lobed parameres. ............................... ............................................................
<taxonomicName id="4C50818E2173FF8061B6E76A00C7859E" ID-CoL="79TDY" authority="Lobl, 1983" authorityName="Lobl" authorityYear="1983" box="[1244,1436,1662,1687]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="palu">
<emphasis id="B924261F2173FF8061B6E76A007C859F" bold="true" box="[1244,1319,1662,1686]" italics="true" pageId="11" pageNumber="162">S. palu</emphasis>
<bibRefCitation id="EFC187FC2173FF806047E76B00C7859E" author="LOBL I." box="[1325,1436,1663,1687]" firstAuthor="Lobl" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pageId="11" pageNumber="162" pagination="285 - 293" part="56" refId="ref12872" refString="LOBL I. 1983: Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 56: 285 - 293." title="Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien" type="journal article" year="1983">Löbl, 1983</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E78B00C7845F" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E78B00C7845F" pageId="11" pageNumber="162">
– Elytral punctation fine and sparse, slightly coarser than that on pronotum; puncture diameters much smaller than puncture intervals. Metaventrite without antecoxal puncture rows. Aedeagus asymmetrical, with parameres not lobed, narrowed apically. .............. ........................................................
<taxonomicName id="4C50818E2173FF8061ACE62A001A845F" ID-CoL="6XSWB" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1222,1345,1854,1878]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="sumpichi" status="sp. nov.">
<emphasis id="B924261F2173FF8061ACE62A0188845F" bold="true" box="[1222,1235,1854,1878]" italics="true" pageId="11" pageNumber="162">S</emphasis>
.
<emphasis id="B924261F2173FF8061B5E62A001A845F" bold="true" box="[1247,1345,1854,1878]" italics="true" pageId="11" pageNumber="162">sumpichi</emphasis>
</taxonomicName>
<emphasis id="B924261F2173FF80602DE62A00C7845F" bold="true" box="[1351,1436,1854,1878]" pageId="11" pageNumber="162">
<taxonomicNameLabel id="A2179B642173FF80602DE62A00C7845F" box="[1351,1436,1854,1878]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82173FF806656E64B00C684FE" pageId="11" pageNumber="162">
<paragraph id="8BEFFA0D2173FF806656E64B00C784BE" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E64B00C784BE" pageId="11" pageNumber="162">
12 Body length
<quantity id="4CA857E82173FF80669FE64B0101847F" box="[1013,1114,1887,1911]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="162" unit="mm" value="0.9">0.90 mm</quantity>
long. Elytral punctation becoming coarser apically. Abdomen lacking obvious microsculpture. ...........................
<taxonomicName id="4C50818E2173FF8061D5E68A00C784BE" ID-CoL="79SVR" authority="Lobl, 1983" authorityName="Lobl" authorityYear="1983" box="[1215,1436,1950,1975]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="sadang">
<emphasis id="B924261F2173FF8061D5E68A007D84BF" bold="true" box="[1215,1318,1950,1974]" italics="true" pageId="11" pageNumber="162">S. sadang</emphasis>
<bibRefCitation id="EFC187FC2173FF806047E68B00C784BE" author="LOBL I." box="[1325,1436,1951,1975]" firstAuthor="Lobl" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pageId="11" pageNumber="162" pagination="285 - 293" part="56" refId="ref12872" refString="LOBL I. 1983: Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 56: 285 - 293." title="Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien" type="journal article" year="1983">Löbl, 1983</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2173FF806656E6AB00C684FE" blockId="11.[828,1442,190,2039]" pageId="11" pageNumber="162">
<keyLead id="30A159382173FF806656E6AB00C684FE" pageId="11" pageNumber="162">
– Body length
<quantity id="4CA857E82173FF806682E6AA01CE84DF" box="[1000,1173,1982,2007]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="1.8" metricValueMin="1.55" pageId="11" pageNumber="162" unit="mm" value="1.675" valueMax="1.8" valueMin="1.55">1.55 to 1.80 mm</quantity>
. Elytral punctation even. Abdomen microsculptured. ...................................... 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82174FF8765FDE1AB00C7823F" pageId="12" pageNumber="163">
<paragraph id="8BEFFA0D2174FF8765FDE1AB07A3825E" blockId="12.[151,760,191,439]" pageId="12" pageNumber="163">
<keyLead id="30A159382174FF8765FDE1AB07A3825E" pageId="12" pageNumber="163">13 Head and pronotum blackish, elytra with well visible, sharply delimited discal punctures, elytral apices not sexually dimorphic, with apical margins slightly rounded or truncate. Metaventrite with mesal stria, abdominal microsculpture strigulate. .............................. 14</keyLead>
</paragraph>
<paragraph id="8BEFFA0D2174FF8765FDE04B00C7823F" blockId="12.[151,760,191,439]" lastBlockId="12.[828,1437,191,406]" pageId="12" pageNumber="163">
<keyLead id="30A159382174FF8765FDE04B00C7823F" pageId="12" pageNumber="163">
– Pronotum and elytra reddish-brown, elytral punctures hardly visible, not clearly delimited. Elytral apices sexually dimorphic, in male truncate, in female expanded and strongly oblique. Metaventrite lacking mesal stria, abdominal ventrite I not microsculptured, following ventrite with punctulate microsculpture. ............. ............................................................
<emphasis id="B924261F2174FF8761B4E00A00C7823F" bold="true" box="[1246,1436,286,310]" pageId="12" pageNumber="163">
<taxonomicName id="4C50818E2174FF8761B4E00A001F823F" ID-CoL="6Y5L7" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1246,1348,286,310]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="ancora" status="sp. nov.">
<emphasis id="B924261F2174FF8761B4E00A001F823F" bold="true" box="[1246,1348,286,310]" italics="true" pageId="12" pageNumber="163">S. ancora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642174FF876023E00A00C7823F" box="[1353,1436,286,310]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A4E2A82174FF866656E02B07A3821F" lastPageId="13" lastPageNumber="164" pageId="12" pageNumber="163">
<paragraph id="8BEFFA0D2174FF876656E02B00C7829F" blockId="12.[828,1437,191,406]" pageId="12" pageNumber="163">
<keyLead id="30A159382174FF876656E02B00C7829F" pageId="12" pageNumber="163">
14 Metaventrite bearing strigulate microsculpture. Aedeagus with internal sac lacking distinct spine-like structures. .........................................
<emphasis id="B924261F2174FF8761B1E06A00C7829F" bold="true" box="[1243,1436,382,406]" pageId="12" pageNumber="163">
<taxonomicName id="4C50818E2174FF8761B1E06A0018829F" ID-CoL="79TFQ" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1243,1347,382,406]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="ogawai" status="sp. nov.">
<emphasis id="B924261F2174FF8761B1E06A0018829F" bold="true" box="[1243,1347,382,406]" italics="true" pageId="12" pageNumber="163">S. ogawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642174FF876023E06A00C7829F" box="[1353,1436,382,406]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<caption id="DF2FAA852174FF8765FDE6FF05AD8B11" ID-DOI="http://doi.org/10.5281/zenodo.3699171" ID-Zenodo-Dep="3699171" httpUri="https://zenodo.org/record/3699171/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,188,2027,2046]" targetBox="[206,1382,457,2010]" targetPageId="12">
<paragraph id="8BEFFA0D2174FF8765FDE6FF05AD8B11" blockId="12.[151,1437,2027,2072]" pageId="12" pageNumber="163">
Figs 50–53. Habitus of
<taxonomicName id="4C50818E2174FF87640EE6FF049084F7" ID-CoL="8VS35" authorityName="Leach" authorityYear="1815" box="[356,459,2027,2046]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B924261F2174FF87640EE6FF049084F7" box="[356,459,2027,2046]" italics="true" pageId="12" pageNumber="163">Scaphisoma</emphasis>
</taxonomicName>
. 50, 51 –
<taxonomicName id="4C50818E2174FF87674EE6FF072D84F7" ID-CoL="6Y5L7" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[548,630,2027,2046]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="ancora" status="sp. nov.">
<emphasis id="B924261F2174FF87674EE6FF072D84F7" box="[548,630,2027,2046]" italics="true" pageId="12" pageNumber="163">S. ancora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642174FF876717E6FF079A84F7" box="[637,705,2027,2046]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
(50 – male, 51 – female); 52 –
<taxonomicName id="4C50818E2174FF8766BEE6FF016384F7" ID-CoL="6XSWB" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[980,1080,2027,2046]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="sumpichi" status="sp. nov.">
<emphasis id="B924261F2174FF8766BEE6FF016384F7" box="[980,1080,2027,2046]" italics="true" pageId="12" pageNumber="163">S. sumpichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642174FF876154E6FF01DF84F7" box="[1086,1156,2027,2046]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
, male; 53 –
<taxonomicName id="4C50818E2174FF876184E6FF002684F7" ID-CoL="79T6S" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[1262,1405,2027,2046]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="versicoloreum" status="sp. nov.">
<emphasis id="B924261F2174FF876184E6FF002684F7" box="[1262,1405,2027,2046]" italics="true" pageId="12" pageNumber="163">S. versicoloreum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642174FF8760EEE6FF05E58B11" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
, male.
</paragraph>
</caption>
<paragraph id="8BEFFA0D2175FF8665FDE1AB07A3821F" blockId="13.[151,760,191,278]" pageId="13" pageNumber="164">
<keyLead id="30A159382175FF8665FDE1AB07A3821F" pageId="13" pageNumber="164">
– Metaventrite lacking microsculpture. Aedeagus with internal sac bearing distinct spine-like structures. ...... ..............................................................
<emphasis id="B924261F2175FF866729E1EA07A3821F" bold="true" box="[579,760,254,278]" pageId="13" pageNumber="164">
<taxonomicName id="4C50818E2175FF866729E1EA07C4821F" ID-CoL="6Y5F2" authority="Löbl, 2018" authorityName="Löbl" authorityYear="2018" box="[579,671,254,278]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pellax" status="sp. nov.">
<emphasis id="B924261F2175FF866729E1EA070B821F" bold="true" box="[579,592,254,278]" italics="true" pageId="13" pageNumber="164">S</emphasis>
.
<emphasis id="B924261F2175FF866737E1EA07C4821F" bold="true" box="[605,671,254,278]" italics="true" pageId="13" pageNumber="164">pellax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179B642175FF8667CFE1EA07A3821F" box="[677,760,254,278]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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