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<document id="C619767DC83703B891467F8F439780DE" ID-CLB-Dataset="6611" ID-DOI="10.5281/zenodo.4502132" ID-GBIF-Dataset="60740210-999d-4526-88e3-464fd93143ac" ID-ISSN="2345-7600" ID-Zenodo-Dep="4502132" ID-ZooBank="4D5F3585-04B2-45AA-BA00-8795AEF68A83" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612458956132" checkinUser="carolina" docAuthor="Köhler, Frank" docDate="2014" docId="03F3175FEF6B5E41FC663ED5FBD0F9D3" docLanguage="en" docName="RafflesBZool.62.115-123.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Rhagada marghitae Falconieri 1995" docType="treatment" docVersion="12" lastPageNumber="119" masterDocId="FFCA6F27EF6F5E45FFDE3C2BFFD2FFFE" masterDocTitle="On the land snail Rhagada Albers, 1860 (Gastropoda: Camaenidae) from across the Lesser Sunda Islands" masterLastPageNumber="123" masterPageNumber="115" pageNumber="119" updateTime="1702001460782" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8A96981EC310A37FE54DEA0AE7467C29">On the land snail Rhagada Albers, 1860 (Gastropoda: Camaenidae) from across the Lesser Sunda Islands</mods:title>
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<mods:namePart id="9AE087655CBB02A562594BD3A7D93B48">Köhler, Frank</mods:namePart>
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<treatment id="03F3175FEF6B5E41FC663ED5FBD0F9D3" ID-DOI="http://doi.org/10.5281/zenodo.4815063" ID-GBIF-Taxon="182237996" ID-Zenodo-Dep="4815063" LSID="urn:lsid:plazi:treatment:03F3175FEF6B5E41FC663ED5FBD0F9D3" httpUri="http://treatment.plazi.org/id/03F3175FEF6B5E41FC663ED5FBD0F9D3" lastPageNumber="119" pageId="4" pageNumber="119">
<subSubSection id="C340F5C2EF6B5E41FC663ED5FA9FFCE8" box="[952,1357,766,790]" pageId="4" pageNumber="119" type="nomenclature">
<paragraph id="8BE5A649EF6B5E41FC663ED5FA9FFCE8" blockId="4.[952,1357,766,790]" box="[952,1357,766,790]" pageId="4" pageNumber="119">
<heading id="D0AD1125EF6B5E41FC663ED5FA9FFCE8" box="[952,1357,766,790]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="119" reason="3">
<taxonomicName id="4C5ADDCAEF6B5E41FC663ED5FA9FFCE8" ID-CoL="4RYC9" authority="Falconieri, 1995" authorityName="Falconieri" authorityYear="1995" box="[952,1357,766,790]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="marghitae">
<emphasis id="B92E7A5BEF6B5E41FC663ED5FB47FCE8" bold="true" box="[952,1173,766,790]" italics="true" pageId="4" pageNumber="119">Rhagada marghitae</emphasis>
<bibRefCitation id="EFCBDBB8EF6B5E41FB433ED4FA9FFCE8" author="Falconieri A" box="[1181,1357,767,790]" pageId="4" pageNumber="119" pagination="81 - 84" refId="ref5764" refString="Falconieri A (1995) Rhagada marghitae species nova. World Shells, 14: 81 - 84." type="journal article" year="1995">Falconieri, 1995</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340F5C2EF6B5E41FC9D3F15FA4EFC38" pageId="4" pageNumber="119" type="reference_group">
<paragraph id="8BE5A649EF6B5E41FC9D3F15FA10FC8E" blockId="4.[835,1474,830,966]" pageId="4" pageNumber="119">
<treatmentCitation id="0AFB8058EF6B5E41FC9D3F15FB3DFCAD" author="Falconieri A" box="[835,1263,830,851]" page="84" pageId="4" pageNumber="119" year="1995">
<taxonomicName id="4C5ADDCAEF6B5E41FC9D3F15FB3DFCAD" ID-CoL="4RYC9" authority="Falconieri, 1995: 81 - 84" authorityName="Falconieri" authorityPageNumber="81 - 84" authorityYear="1995" box="[835,1263,830,851]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="marghitae">
<emphasis id="B92E7A5BEF6B5E41FC9D3F15FBD0FCAD" box="[835,1026,830,851]" italics="true" pageId="4" pageNumber="119">Rhagada marghitae</emphasis>
<bibRefCitation id="EFCBDBB8EF6B5E41FBD63F15FB3DFCAD" author="Falconieri A" box="[1032,1263,830,851]" pageId="4" pageNumber="119" pagination="81 - 84" refId="ref5764" refString="Falconieri A (1995) Rhagada marghitae species nova. World Shells, 14: 81 - 84." type="journal article" year="1995">Falconieri, 1995: 8184</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1, 2 (from
<materialsCitation id="3B32AC14EF6B5E41FA523F15FA69FC8E" ID-GBIF-Occurrence="3119073301" collectionCode="MSNM" pageId="4" pageNumber="119" specimenCode="MSNM 22635" typeStatus="holotype">
Nusa Penida, Pura Batu Medau;
<typeStatus id="54E118EBEF6B5E41FBA83F70FB18FC8E" box="[1142,1226,859,880]" pageId="4" pageNumber="119" type="holotype">holotype</typeStatus>
<specimenCode id="DBFC0E32EF6B5E41FB0F3F70FA8DFC8E" box="[1233,1375,859,880]" collectionCode="MSNM" country="Italy" lsid="urn:lsid:biocol.org:col:15546" name="Museo Civico di Storia Naturale di Milano" pageId="4" pageNumber="119" type="Herbarium">MSNM 22635</specimenCode>
, not seen
</materialsCitation>
)
</paragraph>
<paragraph id="8BE5A649EF6B5E41FC9D3F53FC69FC57" blockId="4.[835,1474,830,966]" pageId="4" pageNumber="119">
<taxonomicName id="4C5ADDCAEF6B5E41FC9D3F53FBD0FC73" ID-CoL="4RYCR" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[835,1026,888,909]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
<emphasis id="B92E7A5BEF6B5E41FC9D3F53FBD0FC73" box="[835,1026,888,909]" italics="true" pageId="4" pageNumber="119">Rhagada solorensis</emphasis>
</taxonomicName>
(partim)
<treatmentCitation id="0AFB8058EF6B5E41FBAA3F53FA10FC73" box="[1140,1474,888,909]" pageId="4" pageNumber="119">
<bibRefCitation id="EFCBDBB8EF6B5E41FBAA3F53FA6CFC73" author="Vermeulen JJ &amp; Whitten AJ" box="[1140,1470,888,909]" pageId="4" pageNumber="119" pagination="1 - 164" refId="ref6974" refString="Vermeulen JJ &amp; Whitten AJ (1998) Fauna Malesiana. Guide to the Land Snails of Bali. Backhuys Publishers, Leiden. Pp. 1 - 164." type="book chapter" year="1998">Vermeulen &amp; Whitten, 1998: 127</bibRefCitation>
,
</treatmentCitation>
fi g. 129.
</paragraph>
<paragraph id="8BE5A649EF6B5E41FC9D3F9AFA4EFC38" blockId="4.[835,1474,830,966]" box="[835,1436,945,966]" pageId="4" pageNumber="119">
<taxonomicName id="4C5ADDCAEF6B5E41FC9D3F9AFBB8FC38" ID-CoL="4RYCR" authority="solorensis" box="[835,1130,945,966]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="subSpecies" species="solorensis" subSpecies="solorensis">
<emphasis id="B92E7A5BEF6B5E41FC9D3F9AFBB8FC38" box="[835,1130,945,966]" italics="true" pageId="4" pageNumber="119">Rhagada solorensis solorensis</emphasis>
</taxonomicName>
<treatmentCitation id="0AFB8058EF6B5E41FB583F9AFAECFC38" box="[1158,1342,945,966]" pageId="4" pageNumber="119">
<bibRefCitation id="EFCBDBB8EF6B5E41FB583F9AFAECFC38" author="Maassen WJM" box="[1158,1342,945,966]" pageId="4" pageNumber="119" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen, 2009: 66</bibRefCitation>
</treatmentCitation>
(partim).
</paragraph>
</subSubSection>
<subSubSection id="C340F5C2EF6B5E41FC9D3FC0FB1BFBE3" pageId="4" pageNumber="119" type="materials_examined">
<paragraph id="8BE5A649EF6B5E41FC9D3FC0FB1BFBE3" blockId="4.[835,1475,1003,1197]" pageId="4" pageNumber="119">
<emphasis id="B92E7A5BEF6B5E41FC9D3FC0FBDEFBFE" bold="true" box="[835,1036,1003,1024]" pageId="4" pageNumber="119">Material examined.</emphasis>
<materialsCitation id="3B32AC14EF6B5E41FBCF3FC0FB17FBE3" ID-GBIF-Occurrence="3114366310" collectionCode="AM" country="Indonesia" latitude="8.75" location="Nusa Penida" longLatPrecision="1300" longitude="115.5" municipality="Goa Giri Putri" pageId="4" pageNumber="119" specimenCount="1">
<collectionCode id="ED4B3E8CEF6B5E41FBCF3FC0FB9CFBFE" box="[1041,1102,1003,1024]" country="Australia" name="Australian Museum" pageId="4" pageNumber="119" type="Museum">AM C</collectionCode>
.327851 (
<collectingCountry id="F34DE6D9EF6B5E41FB773FC0FAD8FBFE" box="[1193,1290,1003,1024]" name="Indonesia" pageId="4" pageNumber="119">Indonesia</collectingCountry>
,
<location id="8E85F092EF6B5E41FACA3FC0FA42FBFE" LSID="urn:lsid:plazi:treatment:03F3175FEF6B5E41FC663ED5FBD0F9D3:8E85F092EF6B5E41FACA3FC0FA42FBFE" box="[1300,1424,1003,1024]" country="Indonesia" latitude="8.75" longLatPrecision="1300" longitude="115.5" municipality="Goa Giri Putri" name="Nusa Penida" pageId="4" pageNumber="119">Nusa Penida</location>
,
<collectingMunicipality id="6B813C33EF6B5E41FA443FC0FC71FBE3" pageId="4" pageNumber="119">Goa Giri Putri</collectingMunicipality>
,
<geoCoordinate id="EE6EC08EEF6B5E41FC703823FBD1FBE3" box="[942,1027,1032,1053]" degrees="8" direction="north" minutes="45" orientation="latitude" pageId="4" pageNumber="119" precision="925" value="8.75">8° 45' N</geoCoordinate>
,
<geoCoordinate id="EE6EC08EEF6B5E41FBD13823FBAAFBE3" box="[1039,1144,1032,1053]" degrees="115" direction="east" minutes="30" orientation="longitude" pageId="4" pageNumber="119" precision="925" value="115.5">115° 30' E</geoCoordinate>
, 5 wet)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340F5C2EF6B5E41FC9D380FFA5BFB53" pageId="4" pageNumber="119" type="discussion">
<paragraph id="8BE5A649EF6B5E41FC9D380FFA5BFB53" blockId="4.[835,1475,1003,1197]" pageId="4" pageNumber="119">
Taxonomic history.Apparently not aware of Falconieris description, this species was subsumed under
<taxonomicName id="4C5ADDCAEF6B5E41FB48386AFAC6FBA8" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[1174,1300,1089,1110]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
<emphasis id="B92E7A5BEF6B5E41FB48386AFAC6FBA8" box="[1174,1300,1089,1110]" italics="true" pageId="4" pageNumber="119">R. solorensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCBDBB8EF6B5E41FAE0386AFC0DFB8D" author="Vermeulen JJ &amp; Whitten AJ" pageId="4" pageNumber="119" pagination="1 - 164" refId="ref6974" refString="Vermeulen JJ &amp; Whitten AJ (1998) Fauna Malesiana. Guide to the Land Snails of Bali. Backhuys Publishers, Leiden. Pp. 1 - 164." type="book chapter" year="1998">Vermeulen &amp; Whitten (1998)</bibRefCitation>
and
<bibRefCitation id="EFCBDBB8EF6B5E41FBC93875FB6FFB8D" author="Maassen WJM" box="[1047,1213,1118,1139]" pageId="4" pageNumber="119" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen (2009)</bibRefCitation>
. The original description focuses on characters exhibited by several species but provides little information permitting the recognition of this species.
</paragraph>
</subSubSection>
<subSubSection id="C340F5C2EF6B5E41FC9D38FDFA50FA93" pageId="4" pageNumber="119" type="diagnosis">
<paragraph id="8BE5A649EF6B5E41FC9D38FDFA50FA93" blockId="4.[835,1474,1238,1389]" pageId="4" pageNumber="119">
<emphasis id="B92E7A5BEF6B5E41FC9D38FDFC6DFB13" bold="true" box="[835,959,1238,1261]" pageId="4" pageNumber="119">Diagnosis.</emphasis>
Shell moderately large (D =
<quantity id="4CA20BACEF6B5E41FAFE38FDFA64FB13" box="[1312,1462,1238,1261]" metricMagnitude="-2" metricUnit="m" metricValue="1.475" metricValueMax="1.55" metricValueMin="1.4" pageId="4" pageNumber="119" unit="mm" value="14.75" valueMax="15.5" valueMin="14.0">1415.5 mm</quantity>
), with fine axial growth lines, umbilicus forming a chink or narrowly open. Penial verge smooth with pointed tip and well-developed sperm groove, inner penial wall smooth, epiphallus with moderately long flagellum (
<figureCitation id="1361BACCEF6B5E41FAFE397DFAA7FA93" box="[1312,1397,1366,1389]" captionStart="Fig" captionStartId="2.[136,171,2006,2027]" captionTargetBox="[143,1467,293,1913]" captionTargetId="figure-6@2.[129,1480,291,1924]" captionTargetPageId="2" captionText="Fig. 2. Penial anatomy, with penial chamber opened showing internal anatomy. A, Rhagada solorensis, Solor Island, AM C.471154; B, Rhagada marghitae, Nusa Penida AM C.327851; C, Rhagada setzeri atauroensis n. subsp., Atauro Island, holotype AM C.478282; D, Rhagada abbasi n. sp., Lembata Island, holotype AM C.478280. Abbreviations: ep = epiphallus; fl = epiphallic flagellum; gp = genital pore; lwp = longitudinal wall pilaster; pv = penial verge; rm = penial retractor muscle; va = vagina; vd = vas deferens. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4502138" httpUri="https://zenodo.org/record/4502138/files/figure.png" pageId="4" pageNumber="119">Fig. 2B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C340F5C2EF6B5E41FC9D39BDFBD0F9D3" pageId="4" pageNumber="119" type="discussion">
<paragraph id="8BE5A649EF6B5E41FC9D39BDFBD0F9D3" blockId="4.[835,1475,1430,1581]" pageId="4" pageNumber="119">
<emphasis id="B92E7A5BEF6B5E41FC9D39BDFB83FA53" bold="true" box="[835,1105,1430,1453]" pageId="4" pageNumber="119">Comparative remarks.</emphasis>
Confused with
<taxonomicName id="4C5ADDCAEF6B5E41FAD539BDFA4EFA53" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[1291,1436,1430,1453]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
<emphasis id="B92E7A5BEF6B5E41FAD539BDFA4EFA53" box="[1291,1436,1430,1453]" italics="true" pageId="4" pageNumber="119">R. solorensis</emphasis>
</taxonomicName>
by Vermeulen &amp; Whitters (1998) and
<bibRefCitation id="EFCBDBB8EF6B5E41FB0E399DFA57FA33" author="Maassen WJM" box="[1232,1413,1462,1485]" pageId="4" pageNumber="119" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen (2009)</bibRefCitation>
, who depicted shells of this species as
<taxonomicName id="4C5ADDCAEF6B5E41FB7B39FDFAFCFA13" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[1189,1326,1494,1517]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
<emphasis id="B92E7A5BEF6B5E41FB7B39FDFAFCFA13" box="[1189,1326,1494,1517]" italics="true" pageId="4" pageNumber="119">R. solorensis</emphasis>
</taxonomicName>
. Most typical characteristics are smooth, pointed penial verge and smooth inner penial wall.
</paragraph>
</subSubSection>
</treatment>
</document>