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<mods:title id="F73195D085AD7A59A2E52674B7B38607">Toward a Natural Classification of Tritomini: Are There Hidden Tribes within the Genus Tritoma Fabricius (Coleoptera: Erotylidae)?</mods:title>
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<paragraph id="8BF8B07597403027FF3DF9A6E236F992" blockId="6.[194,596,1542,1565]" box="[194,596,1542,1565]" pageId="6" pageNumber="634">
<heading id="D0B0071997403027FF3DF9A6E236F992" box="[194,596,1542,1565]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="634" reason="2">
<emphasis id="B9336C6797403027FF3DF9A6E236F992" bold="true" box="[194,596,1542,1565]" pageId="6" pageNumber="634">
<taxonomicName id="4C47CBF697403027FF3DF9A6E1F1F992" authority="Chujo, 1969" authorityName="Chujo" authorityYear="1969" box="[194,403,1542,1564]" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FF3DF9A6E173F992" bold="true" box="[194,273,1543,1564]" italics="true" pageId="6" pageNumber="634">Rotitma</emphasis>
<bibRefCitation id="EFD6CD8497403027FEE9F9A7E1F1F992" author="Chujo, M." box="[278,403,1542,1564]" pageId="6" pageNumber="634" refId="ref7666" refString="Chujo, M. 1969. Erotylidae (Insecta: Coleoptera). Fauna Japonica. Academic Press of Japan, Tokyo, xii + 316 pp., 18 figs., 23 pls." type="book" year="1969">Chûjô, 1969</bibRefCitation>
</taxonomicName>
, full generic status
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35DE3FE97403027FF67F993E12EF9EC" pageId="6" pageNumber="634" type="reference_group">
<paragraph id="8BF8B07597403027FF67F993E12EF9EC" blockId="6.[152,637,1586,1634]" pageId="6" pageNumber="634">
<treatmentCitationGroup id="AB57975B97403027FF67F993E12EF9EC" pageId="6" pageNumber="634">
<treatmentCitation id="0AE6966497403027FF67F993E276F9C6" author="Chujo, M." box="[152,532,1586,1608]" page="226" pageId="6" pageNumber="634" year="1969">
<taxonomicName id="4C47CBF697403027FF67F993E276F9C6" authority="Chujo 1969: 226" authorityName="Chujo" authorityPageNumber="226" authorityYear="1969" box="[152,532,1586,1608]" class="Insecta" family="Erotylidae" genus="Cyrtotriplax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="subGenus" subGenus="Rotitma">
<emphasis id="B9336C6797403027FF67F993E16CF9C9" box="[152,270,1586,1607]" italics="true" pageId="6" pageNumber="634">Cyrtotriplax</emphasis>
(
<emphasis id="B9336C6797403027FEE4F993E105F9C9" box="[283,359,1586,1607]" italics="true" pageId="6" pageNumber="634">Rotitma</emphasis>
)
<bibRefCitation id="EFD6CD8497403027FE8CF993E276F9C6" author="Chujo, M." box="[371,532,1586,1608]" pageId="6" pageNumber="634" refId="ref7666" refString="Chujo, M. 1969. Erotylidae (Insecta: Coleoptera). Fauna Japonica. Academic Press of Japan, Tokyo, xii + 316 pp., 18 figs., 23 pls." type="book" year="1969">Chûjô 1969: 226</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="0AE6966497403027FDE3F993E12AF9EC" author="Chujo, M." page="46" pageId="6" pageNumber="634" year="1990">
<bibRefCitation id="EFD6CD8497403027FDE3F993E12AF9EC" author="Chujo, M." pageId="6" pageNumber="634" pagination="1 - 67" refId="ref7742" refString="Chujo, M., and M. Chujo. 1990. A catalog of the Erotylidae (Insecta, Coleoptera) from the Old World (excl. the Ethiopian Region) III. Esakia 29: 1 - 67." type="journal article" year="1990">Chûjô and Chûjô 1990: 46</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35DE3FE97403027FF4FF9D6E099F94C" pageId="6" pageNumber="634" type="type_taxon">
<paragraph id="8BF8B07597403027FF4FF9D6E099F94C" blockId="6.[152,638,1655,1730]" pageId="6" pageNumber="634">
<emphasis id="B9336C6797403027FF4FF9D6E155F903" bold="true" box="[176,311,1655,1677]" pageId="6" pageNumber="634">
<typeStatus id="54FC0ED797403027FF4FF9D6E080F902" box="[176,226,1655,1676]" pageId="6" pageNumber="634">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="4C47CBF697403027FEC3F9D6E0A9F929" authority="Lewis, 1887" authorityName="Lewis" authorityYear="1887" class="Insecta" family="Erotylidae" genus="Cyrtotriplax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="pallidiventris">
<emphasis id="B9336C6797403027FEC3F9D6E25AF902" box="[316,568,1655,1676]" italics="true" pageId="6" pageNumber="634">Cyrtotriplax pallidiventris</emphasis>
<bibRefCitation id="EFD6CD8497403027FDC2F9D6E0A9F929" author="Lewis, G." pageId="6" pageNumber="634" pagination="53 - 73" refId="ref8802" refString="Lewis, G. 1887. A list of fifty Erotylidae from Japan, including thirty-five new species and four new genera. Annals and Magazine of Natural History (Series 5) 20 (115): 53 - 73. doi. org / 10.1080 / 00222938709460010." type="journal article" year="1887">Lewis, 1887</bibRefCitation>
</taxonomicName>
; by subsequent designation (
<bibRefCitation id="EFD6CD8497403027FE25F933E092F94C" author="Chujo, M." pageId="6" pageNumber="634" pagination="1 - 67" refId="ref7742" refString="Chujo, M., and M. Chujo. 1990. A catalog of the Erotylidae (Insecta, Coleoptera) from the Old World (excl. the Ethiopian Region) III. Esakia 29: 1 - 67." type="journal article" year="1990">Chûjô and Chûjô 1990: 46</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35DE3FE97403027FD3AFF60E295FE73" pageId="6" pageNumber="634" type="diagnosis">
<paragraph id="8BF8B07597403027FD3AFF60E295FE73" blockId="6.[685,1172,193,1576]" pageId="6" pageNumber="634">
<emphasis id="B9336C6797403027FD3AFF60E34DFF59" bold="true" box="[709,815,193,215]" pageId="6" pageNumber="634">Diagnosis.</emphasis>
Body generally ovoid in shape. Clypeus anterior margin with shallow V-shaped emargination; complete marginal bead present. Eyes finely facetted. Antenna with club of 3 antennomeres; antennomere XI transversely oval to nearly circular. Mouthparts with lacinia lacking teeth; maxillary terminal palpomere triangularly widened, not more than 3 times wider than long; mentum pentagonal. Prosternal lines widely spaced, not connecting anteriorly. Elytra lacking distinct basal marginal bead. Tibiae narrow, not triangularly dilated.
</paragraph>
</subSubSection>
<subSubSection id="C35DE3FE97403027FD3AFDA0E35EFD0C" pageId="6" pageNumber="634" type="distribution">
<paragraph id="8BF8B07597403027FD3AFDA0E365FDBC" blockId="6.[685,1172,193,1576]" pageId="6" pageNumber="634">
<emphasis id="B9336C6797403027FD3AFDA0E32AFD99" bold="true" box="[709,840,513,535]" pageId="6" pageNumber="634">Distribution.</emphasis>
<collectingCountry id="F350F0E597403027FCB2FDA3E3E4FD99" box="[845,902,514,535]" name="Japan" pageId="6" pageNumber="634">Japan</collectingCountry>
,
<collectingCountry id="F350F0E597403027FC71FDA3E381FD96" box="[910,995,514,536]" name="Malaysia" pageId="6" pageNumber="634">Sarawak</collectingCountry>
, and eastern North America.
</paragraph>
<paragraph id="8BF8B07597403027FD3AFD96E35EFD0C" blockId="6.[685,1172,193,1576]" pageId="6" pageNumber="634">
<emphasis id="B9336C6797403027FD3AFD96E3E6FDC3" bold="true" box="[709,900,567,589]" pageId="6" pageNumber="634">Number of Species.</emphasis>
6 [5 (
<bibRefCitation id="EFD6CD8497403027FC47FD96E4EEFDC2" author="Chujo, M." box="[952,1164,567,589]" pageId="6" pageNumber="634" pagination="1 - 67" refId="ref7742" refString="Chujo, M., and M. Chujo. 1990. A catalog of the Erotylidae (Insecta, Coleoptera) from the Old World (excl. the Ethiopian Region) III. Esakia 29: 1 - 67." type="journal article" year="1990">Chûjô and Chûjô 1990</bibRefCitation>
) +
<taxonomicName id="4C47CBF697403027FD30FDF3E430FDE9" authority="(Say, 1825)" authorityName="Pecci-Maddalena &amp; Skelley" authorityYear="2021" baseAuthorityName="Pecci-Maddalena &amp; Skelley" baseAuthorityYear="2021" box="[719,1106,593,615]" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="sanguinipennis" status="comb. nov.">
<emphasis id="B9336C6797403027FD30FDF3E3DDFDE9" box="[719,959,594,615]" italics="true" pageId="6" pageNumber="634">Rotitma sanguinipennis</emphasis>
(
<bibRefCitation id="EFD6CD8497403027FC25FDF3E429FDE9" author="Say, T." box="[986,1099,593,615]" pageId="6" pageNumber="634" pagination="237 - 284" refId="ref9542" refString="Say, T. 1825 - 1826. Descriptions of new species of coleopterous insects inhabiting the United States. Journal of the Academy of Natural Sciences, Philadelphia 5 [1825]: 160 - 204; [1826]: 237 - 284, 293 - 304." type="journal article" year="1825">Say, 1825</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B9336C6797403027FB96FDF3E34FFD0C" bold="true" pageId="6" pageNumber="634">
<taxonomicNameLabel id="A200D11C97403027FB96FDF3E34FFD0C" pageId="6" pageNumber="634" rank="species">new combination</taxonomicNameLabel>
</emphasis>
].
</paragraph>
</subSubSection>
<subSubSection id="C35DE3FE97403027FD3AFD26E376F9A6" pageId="6" pageNumber="634" type="discussion">
<paragraph id="8BF8B07597403027FD3AFD26E3E7FCB2" blockId="6.[685,1172,193,1576]" pageId="6" pageNumber="634">
<emphasis id="B9336C6797403027FD3AFD26E34AFD12" bold="true" box="[709,808,647,668]" pageId="6" pageNumber="634">Remarks.</emphasis>
<taxonomicName id="4C47CBF697403027FCD0FD26E44BFD12" authority="Crotch, 1873" authorityName="Crotch" authorityYear="1873" box="[815,1065,647,669]" class="Insecta" family="Erotylidae" genus="Cyrtotriplax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FCD0FD26E3C4FD12" box="[815,934,647,668]" italics="true" pageId="6" pageNumber="634">Cyrtotriplax</emphasis>
<bibRefCitation id="EFD6CD8497403027FC54FD26E44BFD12" author="Crotch, G. R." box="[939,1065,647,669]" pageId="6" pageNumber="634" pagination="349 - 358" refId="ref7818" refString="Crotch, G. R. 1873. Synopsis of the Erotylidae of boreal America. Transactions of the American Entomological Society, Philadelphia 4: 349 - 358. doi. org / 10.2307 / 25076276." type="journal article" year="1873">Crotch, 1873</bibRefCitation>
</taxonomicName>
as used by
<bibRefCitation id="EFD6CD8497403027FD52FD03E328FD36" author="Chujo, M." box="[685,842,674,696]" pageId="6" pageNumber="634" refId="ref7666" refString="Chujo, M. 1969. Erotylidae (Insecta: Coleoptera). Fauna Japonica. Academic Press of Japan, Tokyo, xii + 316 pp., 18 figs., 23 pls." type="book" year="1969">Chûjô (1969: 6)</bibRefCitation>
and
<bibRefCitation id="EFD6CD8497403027FC82FD03E4F1FD39" author="Chujo, M." box="[893,1171,674,696]" pageId="6" pageNumber="634" pagination="1 - 67" refId="ref7742" refString="Chujo, M., and M. Chujo. 1990. A catalog of the Erotylidae (Insecta, Coleoptera) from the Old World (excl. the Ethiopian Region) III. Esakia 29: 1 - 67." type="journal article" year="1990">Chûjô and Chûjô (1990: 30)</bibRefCitation>
(not
<taxonomicName id="4C47CBF697403027FD26FD1DE46AFD5C" authority="Gorham 1896: 286" authorityName="Gorham" authorityPageNumber="286" authorityYear="1896" box="[729,1032,700,722]" class="Insecta" family="Erotylidae" genus="Cyrtotriplax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FD26FD1DE332FD5F" box="[729,848,700,721]" italics="true" pageId="6" pageNumber="634">Cyrtotriplax</emphasis>
<bibRefCitation id="EFD6CD8497403027FCABFD1DE46AFD5C" author="Gorham, H. S." box="[852,1032,700,722]" pageId="6" pageNumber="634" pagination="257 - 302" refId="ref8314" refString="Gorham, H. S. 1896. Viaggio di Leonardo Fea in Birmania e regioni vicine. LXIX. Languriidae, Erotylidae and Endomychidae. Annali del Museo Civico di Storia Naturale di Genova 16 (2): 257 - 302. doi. org / 10.5962 / bhl. part. 25038." type="journal article" year="1896">Gorham 1896: 286</bibRefCitation>
</taxonomicName>
) has a complicated history.
<taxonomicName id="4C47CBF697403027FCCBFD76E390FD63" authority="Crotch" authorityName="Crotch" authorityYear="1873" box="[820,1010,727,749]" class="Insecta" family="Erotylidae" genus="Cyrtotriplax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FCCBFD76E3C9FD62" box="[820,939,727,748]" italics="true" pageId="6" pageNumber="634">Cyrtotriplax</emphasis>
Crotch
</taxonomicName>
is a synonym of
<taxonomicName id="4C47CBF697403027FD52FD53E338FC86" authority="Fabricius" authorityName="Fabricius" box="[685,858,754,776]" class="Insecta" family="Erotylidae" genus="Triplax" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FD52FD53E290FC89" box="[685,754,754,775]" italics="true" pageId="6" pageNumber="634">Triplax</emphasis>
Fabricius
</taxonomicName>
(
<bibRefCitation id="EFD6CD8497403027FC90FD53E4EDFC89" author="Bouchard, P. &amp; Bousquet, A. E. &amp; Davies, M. A. &amp; Lawrence, C. H. C. &amp; Lyal, A. F. &amp; Newton, C. A. M. &amp; Reid, M. &amp; Schmitt, S. A." box="[879,1167,753,776]" pageId="6" pageNumber="634" pagination="1 - 972" refId="ref7460" refString="Bouchard, P., Y. Bousquet, A. E. Davies, M. A. Alon- so-Zarazaga, J. F. Lawrence, C. H. C. Lyal, A. F. Newton, C. A. M. Reid, M. Schmitt, S. A. Slipinski, and A. B. T. Smith. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1 - 972. doi. org / 10.3897 / zookeys. 88.807." type="journal article" year="2011">
Bouchard
<emphasis id="B9336C6797403027FC22FD53E472FC89" box="[989,1040,754,775]" italics="true" pageId="6" pageNumber="634">et al</emphasis>
. 2011: 358
</bibRefCitation>
;
<bibRefCitation id="EFD6CD8497403027FD52FCADE317FCAC" author="Kerzhner, I. M." box="[685,885,780,802]" pageId="6" pageNumber="634" pagination="107 - 134" refId="ref8490" refString="Kerzhner, I. M. 1991. Case 2292. Historie abregee des insectes qui se trouvent aux environs de Paris (Geoffroy, 1762): Proposed conservation of some generic names (Crustacea and Insecta). Bulletin of Zoological Nomenclature 48 (2): 107 - 134. [ruling in ICZN 1994] doi. org / 10.5962 / bhl. part. 695." type="journal article" year="1991">Kerzhner 1991: 126</bibRefCitation>
), while
<taxonomicName id="4C47CBF697403027FC3AFCADE4F1FCAC" authority="Gorham" authorityName="Gorham" authorityYear="1896" box="[965,1171,780,802]" class="Insecta" family="Erotylidae" genus="Cyrtotriplax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FC3AFCADE45EFCAF" box="[965,1084,780,801]" italics="true" pageId="6" pageNumber="634">Cyrtotriplax</emphasis>
Gorham
</taxonomicName>
remains a valid genus.
</paragraph>
<paragraph id="8BF8B07597403027FD3AFCE3E47CFB9C" blockId="6.[685,1172,193,1576]" pageId="6" pageNumber="634">
<taxonomicName id="4C47CBF697403027FD3AFCE3E316FCD6" authority="Fabricius" authorityName="Fabricius" box="[709,884,834,856]" class="Insecta" family="Erotylidae" genus="Tritoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FD3AFCE3E373FCD9" box="[709,785,834,855]" italics="true" pageId="6" pageNumber="634">Tritoma</emphasis>
Fabricius
</taxonomicName>
is a diverse genus with 136 species (
<bibRefCitation id="EFD6CD8497403027FCFEFCFDE3C0FCFF" author="Alvarenga, M." box="[769,930,860,882]" pageId="6" pageNumber="634" pagination="1 - 175" refId="ref7341" refString="Alvarenga, M. 1994. Catalogo dos Erotylidae (Coleoptera) Neotropical. Revista Brasiliera de Zoologia 11 (1): 1 - 175. doi. org / 10.1590 / S 0101 - 81751994000100001." type="journal article" year="1994">Alvarenga 1994</bibRefCitation>
;
<bibRefCitation id="EFD6CD8497403027FC52FCFDE440FCFC" author="Boyle, W. W." box="[941,1058,860,882]" pageId="6" pageNumber="634" pagination="61 - 172" refId="ref7596" refString="Boyle, W. W. 1956. A revision of the Erotylidae of Amer - ica north of Mexico (Coleoptera). Bulletin of the American Museum of Natural History 110 (2): 61 - 172." type="journal article" year="1956">Boyle 1956</bibRefCitation>
;
<bibRefCitation id="EFD6CD8497403027FBD2FCFDE34BFC02" author="Chujo, M." pageId="6" pageNumber="634" pagination="1 - 67" refId="ref7742" refString="Chujo, M., and M. Chujo. 1990. A catalog of the Erotylidae (Insecta, Coleoptera) from the Old World (excl. the Ethiopian Region) III. Esakia 29: 1 - 67." type="journal article" year="1990">Chûjô and Chûjô 1990</bibRefCitation>
;
<bibRefCitation id="EFD6CD8497403027FCC4FCD6E397FC02" author="Delkeskamp, K." box="[827,1013,887,909]" pageId="6" pageNumber="634" pagination="1 - 65" refId="ref7977" refString="Delkeskamp, K. 1981. Erotylidae von Africa und Madagascar. Coleopterorum Catalogus Supplementum 34: 1 - 65." type="journal article" year="1981">Delkeskamp 1981</bibRefCitation>
;
<bibRefCitation id="EFD6CD8497403027FBF8FCD6E34BFC29" author="Goodrich, M. A." pageId="6" pageNumber="634" pagination="31 - 36" refId="ref8099" refString="Goodrich, M. A., and P. E. Skelley. 1991. New synonymy in the genus Tritoma (Coleoptera: Erotylidae). The Coleopterists Bulletin 45 (1): 31 - 36." type="journal article" year="1991">Goodrich and Skelley 1991</bibRefCitation>
) that are widely distributed (Holarctic, Central America, Africa).
<taxonomicName id="4C47CBF697403027FC4EFC0CE39FFC4C" box="[945,1021,941,962]" class="Insecta" family="Erotylidae" genus="Tritoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FC4EFC0CE39FFC4C" box="[945,1021,941,962]" italics="true" pageId="6" pageNumber="634">Tritoma</emphasis>
</taxonomicName>
(
<emphasis id="B9336C6797403027FBF1FC0CE413FC4F" box="[1038,1137,940,962]" italics="true" pageId="6" pageNumber="634">sensu lato</emphasis>
) is readily distinguished from
<taxonomicName id="4C47CBF697403027FC4AFC69E463FC53" authorityName="Chujo" authorityYear="1969" box="[949,1025,968,989]" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FC4AFC69E463FC53" box="[949,1025,968,989]" italics="true" pageId="6" pageNumber="634">Rotitma</emphasis>
</taxonomicName>
by lacking the V-shaped concavity of the epistomal apex and in having a distinctly triangular mentum.
</paragraph>
<paragraph id="8BF8B07597403027FD3AFBB6E3A9FA93" blockId="6.[685,1172,193,1576]" pageId="6" pageNumber="634">
Raising
<taxonomicName id="4C47CBF697403027FCE4FBB9E305FBA3" authorityName="Chujo" authorityYear="1969" box="[795,871,1048,1069]" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FCE4FBB9E305FBA3" box="[795,871,1048,1069]" italics="true" pageId="6" pageNumber="634">Rotitma</emphasis>
</taxonomicName>
to full generic status makes
<taxonomicName id="4C47CBF697403027FD52FB93E29BFBC9" box="[685,761,1074,1095]" class="Insecta" family="Erotylidae" genus="Tritoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FD52FB93E29BFBC9" box="[685,761,1074,1095]" italics="true" pageId="6" pageNumber="634">Tritoma</emphasis>
</taxonomicName>
a more homogeneous genus. However, the morphological diversity and wide geographical distributions of the remaining
<taxonomicName id="4C47CBF697403027FC44FBC9E436FBF3" box="[955,1108,1127,1149]" pageId="6" pageNumber="634">
<emphasis id="B9336C6797403027FC44FBC9E465FBF3" box="[955,1031,1128,1149]" italics="true" pageId="6" pageNumber="634">Tritoma</emphasis>
species
</taxonomicName>
raises questions about the monophyly of this and other related genera. Similarly, the wide geographical distribution of
<taxonomicName id="4C47CBF697403027FCC4FB19E3E5FB43" authorityName="Chujo" authorityYear="1969" box="[827,903,1208,1229]" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FCC4FB19E3E5FB43" box="[827,903,1208,1229]" italics="true" pageId="6" pageNumber="634">Rotitma</emphasis>
</taxonomicName>
indicates additional species may still reside in Asian
<taxonomicName id="4C47CBF697403027FC42FB73E46BFB69" box="[957,1033,1234,1255]" class="Insecta" family="Erotylidae" genus="Tritoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FC42FB73E46BFB69" box="[957,1033,1234,1255]" italics="true" pageId="6" pageNumber="634">Tritoma</emphasis>
</taxonomicName>
(
<emphasis id="B9336C6797403027FBE2FB73E4E6FB69" box="[1053,1156,1234,1255]" italics="true" pageId="6" pageNumber="634">sensu lato</emphasis>
). Robust revisionary and phylogenetic works on the worlds
<taxonomicName id="4C47CBF697403027FD04FAA6E334FA93" authorityName="Curtis" authorityYear="1834" box="[763,854,1287,1309]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="tribe" tribe="Tritomini">Tritomini</taxonomicName>
are needed.
</paragraph>
<paragraph id="8BF8B07597403027FD3AFA83E376F9A6" blockId="6.[685,1172,193,1576]" pageId="6" pageNumber="634">
With the raising of
<taxonomicName id="4C47CBF697403027FC84FA83E3A5FAB9" authorityName="Chujo" authorityYear="1969" box="[891,967,1314,1335]" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FC84FA83E3A5FAB9" box="[891,967,1314,1335]" italics="true" pageId="6" pageNumber="634">Rotitma</emphasis>
</taxonomicName>
to full generic status, the following
<emphasis id="B9336C6797403027FCC6FA9DE396FADC" bold="true" box="[825,1012,1340,1362]" pageId="6" pageNumber="634">new combinations</emphasis>
are recognized:
<taxonomicName id="4C47CBF697403027FD52FAF9E458FAE3" authority="(Lewis 1887: 66)" baseAuthorityName="Lewis" baseAuthorityPageNumber="66" baseAuthorityYear="1887" box="[685,1082,1367,1389]" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="circumcinctus">
<emphasis id="B9336C6797403027FD52FAF9E3E5FAE3" box="[685,903,1368,1389]" italics="true" pageId="6" pageNumber="634">Rotitma circumcinctus</emphasis>
(
<bibRefCitation id="EFD6CD8497403027FC69FAF6E456FAE3" author="Lewis, G." box="[918,1076,1367,1389]" pageId="6" pageNumber="634" pagination="53 - 73" refId="ref8802" refString="Lewis, G. 1887. A list of fifty Erotylidae from Japan, including thirty-five new species and four new genera. Annals and Magazine of Natural History (Series 5) 20 (115): 53 - 73. doi. org / 10.1080 / 00222938709460010." type="journal article" year="1887">Lewis 1887: 66</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C47CBF697403027FBB8FAF9E3CDFA09" authority="(Lewis 1887: 65)" baseAuthorityName="Lewis" baseAuthorityPageNumber="65" baseAuthorityYear="1887" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="discalis">
<emphasis id="B9336C6797403027FBB8FAF9E294FA09" italics="true" pageId="6" pageNumber="634">Rotitma discalis</emphasis>
(
<bibRefCitation id="EFD6CD8497403027FCF8FAD3E3C5FA09" author="Lewis, G." box="[775,935,1393,1416]" pageId="6" pageNumber="634" pagination="53 - 73" refId="ref8802" refString="Lewis, G. 1887. A list of fifty Erotylidae from Japan, including thirty-five new species and four new genera. Annals and Magazine of Natural History (Series 5) 20 (115): 53 - 73. doi. org / 10.1080 / 00222938709460010." type="journal article" year="1887">Lewis 1887: 65</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C47CBF697403027FC42FAD3E338FA2C" authority="(Lewis 1887: 64)" baseAuthorityName="Lewis" baseAuthorityPageNumber="64" baseAuthorityYear="1887" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="pallidiventris">
<emphasis id="B9336C6797403027FC42FAD3E4F1FA09" box="[957,1171,1394,1415]" italics="true" pageId="6" pageNumber="634">Rotitma pallidiventris</emphasis>
(
<bibRefCitation id="EFD6CD8497403027FD4BFA2DE331FA2C" author="Lewis, G." box="[692,851,1420,1442]" pageId="6" pageNumber="634" pagination="53 - 73" refId="ref8802" refString="Lewis, G. 1887. A list of fifty Erotylidae from Japan, including thirty-five new species and four new genera. Annals and Magazine of Natural History (Series 5) 20 (115): 53 - 73. doi. org / 10.1080 / 00222938709460010." type="journal article" year="1887">Lewis 1887: 64</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C47CBF697403027FC6DFA2CE36AFA33" authority="(Lewis 1887: 65)" baseAuthorityName="Lewis" baseAuthorityPageNumber="65" baseAuthorityYear="1887" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="rufipennis">
<emphasis id="B9336C6797403027FC6DFA2CE425FA2F" box="[914,1095,1420,1442]" italics="true" pageId="6" pageNumber="634">Rotitma rufipennis</emphasis>
(
<bibRefCitation id="EFD6CD8497403027FBA8FA2DE363FA33" author="Lewis, G." pageId="6" pageNumber="634" pagination="53 - 73" refId="ref8802" refString="Lewis, G. 1887. A list of fifty Erotylidae from Japan, including thirty-five new species and four new genera. Annals and Magazine of Natural History (Series 5) 20 (115): 53 - 73. doi. org / 10.1080 / 00222938709460010." type="journal article" year="1887">Lewis 1887: 65</bibRefCitation>
)
</taxonomicName>
, all from
<collectingCountry id="F350F0E597403027FCA0FA06E3F5FA32" box="[863,919,1447,1468]" name="Japan" pageId="6" pageNumber="634">Japan</collectingCountry>
; and
<taxonomicName id="4C47CBF697403027FC39FA09E330FA59" authority="Chujo 1974: 13" authorityName="Chujo" authorityPageNumber="13" authorityYear="1974" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="B9336C6797403027FC39FA09E4F1FA32" box="[966,1171,1447,1469]" italics="true" pageId="6" pageNumber="634">Rotitma sarawakensis</emphasis>
<bibRefCitation id="EFD6CD8497403027FD52FA63E330FA59" author="Chujo, M." box="[685,850,1474,1496]" pageId="6" pageNumber="634" pagination="1 - 19" refId="ref7702" refString="Chujo, M. 1974. Erotylid beetles from Thailand, Malaysia, Sarawak and Philippines. Studies on the erotylid-beetles (22). Bulletin of the Japan Entomological Academy 9 (1): 1 - 19." type="journal article" year="1974">Chûjô 1974: 13</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F350F0E597403027FC61FA63E396FA56" box="[926,1012,1474,1496]" name="Malaysia" pageId="6" pageNumber="634">Sarawak</collectingCountry>
. The following North American species is here transferred to the genus
<taxonomicName id="4C47CBF697403027FD12FA59E35BF983" authorityName="Chujo" authorityYear="1969" box="[749,825,1528,1549]" class="Insecta" family="Erotylidae" genus="Rotitma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="634" phylum="Arthropoda" rank="genus">
<emphasis id="B9336C6797403027FD12FA59E35BF983" box="[749,825,1528,1549]" italics="true" pageId="6" pageNumber="634">Rotitma</emphasis>
</taxonomicName>
, bringing the total of included species to six.
</paragraph>
</subSubSection>
</treatment>
</document>