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<mods:title id="C4E8E0E5222BA409F38F312F203AFB74">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<mods:namePart id="47347DEC5A2DF6C446E7BA5E570096A7">Schwarzhans, Werner</mods:namePart>
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<treatment id="039D7D5BFE43FFA8FBC8FBB0FD30F892" LSID="urn:lsid:plazi:treatment:039D7D5BFE43FFA8FBC8FBB0FD30F892" httpUri="http://treatment.plazi.org/id/039D7D5BFE43FFA8FBC8FBB0FD30F892" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection id="C32E9FC6FE43FFA8FBC8FBB0FF60FB52" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8B8BCC4DFE43FFA8FBC8FBB0FCBDFB72" blockId="10.[151,775,1131,1187]" box="[151,775,1131,1155]" pageId="10" pageNumber="11">
<taxonomicName id="4C34B7CEFE43FFA8FBC8FBB0FCBAFB72" authority="Schwarzhans et al., 2020 a" authorityName="Schwarzhans" authorityYear="2020" box="[151,768,1131,1155]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="agiadiae">
<emphasis id="B940105FFE43FFA8FBC8FBB0FD0BFB72" bold="true" box="[151,689,1131,1155]" pageId="10" pageNumber="11">
<emphasis id="B940105FFE43FFA8FBC8FBB0FE0DFB72" bold="true" box="[151,439,1131,1155]" italics="true" pageId="10" pageNumber="11">Odondebuenia agiadiae</emphasis>
<bibRefCitation id="EFA5B1BCFE43FFA8FA9BFBB0FD0BFB72" author="Schwarzhans, W. &amp; Brzobohaty, R. &amp; Radwanska, U." box="[452,689,1131,1155]" pageId="10" pageNumber="11" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., &amp; Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al.,</bibRefCitation>
</emphasis>
2020a
</taxonomicName>
,
</paragraph>
<paragraph id="8B8BCC4DFE43FFA8FBC8FB50FF60FB52" blockId="10.[151,775,1131,1187]" box="[151,218,1163,1187]" pageId="10" pageNumber="11">
<heading id="D0C37B21FE43FFA8FBC8FB50FF60FB52" box="[151,218,1163,1187]" fontSize="10" level="2" pageId="10" pageNumber="11" reason="7">
<bibRefCitation id="EFA5B1BCFE43FFA8FBC8FB50FF60FB52" author="Schwarzhans, W. &amp; Agiadi, K. &amp; Carnevale, G." box="[151,218,1163,1187]" pageId="10" pageNumber="11" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., &amp; Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
</heading>
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<subSubSection id="C32E9FC6FE43FFA8FBC8FB10FD30F892" pageId="10" pageNumber="11" type="description">
<paragraph id="8B8BCC4DFE43FFA8FBC8FB10FEA3FB12" blockId="10.[151,281,1227,1251]" box="[151,281,1227,1251]" pageId="10" pageNumber="11">
<figureCitation id="130FD0C8FE43FFA8FBC8FB10FEA3FB12" box="[151,281,1227,1251]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: ad Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.eh Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (16161622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22 N 87° 52 W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="10" pageNumber="11">Figure 5ad</figureCitation>
</paragraph>
<paragraph id="8B8BCC4DFE43FFA8FBC8FAD0FD9EFAB2" blockId="10.[151,775,1291,1347]" pageId="10" pageNumber="11">
2020a
<taxonomicName id="4C34B7CEFE43FFA8FBB9FAD0FE49FAD2" box="[230,499,1291,1315]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="agiadiae">
<emphasis id="B940105FFE43FFA8FBB9FAD0FE49FAD2" box="[230,499,1291,1315]" italics="true" pageId="10" pageNumber="11">Odondebuenia agiadiae</emphasis>
</taxonomicName>
—Schwarzhans, Brzobohatý &amp; Radwańska: pl. 6, figs. 1222.
</paragraph>
<paragraph id="8B8BCC4DFE43FFA8FBC8FAB0FEF9FA32" blockId="10.[151,774,1387,1475]" pageId="10" pageNumber="11">
<emphasis id="B940105FFE43FFA8FBC8FAB0FF42FA72" box="[151,248,1387,1411]" italics="true" pageId="10" pageNumber="11">Material</emphasis>
7 otoliths:
<specimenCount id="9D3207C4FE43FFA8FADDFAB0FDABFA72" box="[386,529,1387,1411]" count="2" pageId="10" pageNumber="11" type="generic">2 specimens</specimenCount>
, Mlyntsi,
<collectionCode id="ED255488FE43FFA8F9D9FAB7FCBCFA72" box="[646,774,1388,1411]" pageId="10" pageNumber="11">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE43FFA8FBC8FA57FF15FA52" box="[151,175,1420,1443]" country="Russia" name="Paleontological Institute" pageId="10" pageNumber="11">PI</collectionCode>
2570;
<specimenCount id="9D3207C4FE43FFA8FBA8FA50FE3BFA52" box="[247,385,1419,1443]" count="5" pageId="10" pageNumber="11" type="generic">5 specimens</specimenCount>
, Kozatskyi Yar,
<collectionCode id="ED255488FE43FFA8F973FA57FD16FA52" box="[556,684,1420,1443]" pageId="10" pageNumber="11">NMNHU-P</collectionCode>
<collectionCode id="ED255488FE43FFA8F9EBFA57FD76FA52" box="[692,716,1420,1443]" country="Russia" name="Paleontological Institute" pageId="10" pageNumber="11">PI</collectionCode>
2569 and
<collectionCode id="ED255488FE43FFA8FB99FA77FEB8FA32" box="[198,258,1452,1475]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="10" pageNumber="11">NMB</collectionCode>
1219.
</paragraph>
<paragraph id="8B8BCC4DFE43FFA8FBC8FA37FD30F892" blockId="10.[151,775,1515,1891]" pageId="10" pageNumber="11">
<emphasis id="B940105FFE43FFA8FBC8FA37FEB0F9F2" box="[151,266,1516,1539]" italics="true" pageId="10" pageNumber="11">Discussion</emphasis>
Otoliths of
<taxonomicName id="4C34B7CEFE43FFA8FAC7FA37FDA3F9F2" box="[408,537,1515,1539]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="agiadiae">
<emphasis id="B940105FFE43FFA8FAC7FA37FDA3F9F2" box="[408,537,1515,1539]" italics="true" pageId="10" pageNumber="11">O. agiadiae</emphasis>
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are characterized by the preventral projection being stronger than the postdorsal one, a depressed predorsal region, and a small, rather narrow sulcus with reduced sole shape and without a subcaudal iugum. Outline and proportions were found to vary considerably in the original description by Schwarzhans et. al. (2020a); this variability is confirmed here.
<taxonomicName id="4C34B7CEFE43FFA8FB8BF910FE60F912" box="[212,474,1739,1763]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="agiadiae">
<emphasis id="B940105FFE43FFA8FB8BF910FE60F912" box="[212,474,1739,1763]" italics="true" pageId="10" pageNumber="11">Odondebuenia agiadiae</emphasis>
</taxonomicName>
is apparently a species that preferred near-reef environments, such as Lakšárská Nová Ves (late Badenian of
<collectingCountry id="F3238CDDFE43FFA8FA90F8D0FD91F8D2" box="[463,555,1803,1827]" name="Slovakia" pageId="10" pageNumber="11">Slovakia</collectingCountry>
) or Węglinek (early Badenian of
<collectingCountry id="F3238CDDFE43FFA8FA77F8F0FECDF8B2" box="[296,375,1835,1859]" name="Poland" pageId="10" pageNumber="11">Poland</collectingCountry>
), and is now also confirmed in the back-reef environment of the Medobory Reef.
</paragraph>
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