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<document ID-DOI="10.24349/esot-nc22" ID-ISSN="2107-7207" ID-Zenodo-Dep="7160296" approvalRequired="22" approvalRequired_for_textStreams="22" checkinTime="1665196356350" checkinUser="felipe" docAuthor="Gerecke, Reinhard &amp; Smit, Harry" docDate="2022" docId="038687E4FFA7182DFE7CD81E7742FC84" docLanguage="en" docName="Acarologia.62.2.302-316.pdf.pdf" docOrigin="Acarologia 62 (2)" docSource="https://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4509" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.4:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="4" docTitle="Lebertia (Lebertia) khatrii Gerecke &amp; Smit 2022, n. sp." docType="treatment" docVersion="2" lastPageNumber="307" masterDocId="FFBFFF9CFFA3182BFF97DB327407FFA5" masterDocTitle="Water mites of the genusLebertia Neuman, 1880 from the eastern Himalayas (Acari: Hydrachnidia: Lebertiidae)" masterLastPageNumber="316" masterPageNumber="302" pageNumber="305" updateTime="1665197544334" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Water mites of the genusLebertia Neuman, 1880 from the eastern Himalayas (Acari: Hydrachnidia: Lebertiidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gerecke, Reinhard</mods:namePart>
<mods:affiliation>Biesingerstrasse 11, DE 72070 Tübingen, Germany. &amp; Arachnology, Senckenberg, Senckenberganlage 25, D- 60325 Frankfurt, Germany.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smit, Harry</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands. &amp; Biesingerstrasse 11, DE 72070 Tübingen, Germany.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acarologia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>62</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>302</mods:start>
<mods:end>316</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>https://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4509</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.24349/esot-nc22</mods:identifier>
<mods:identifier type="ISSN">2107-7207</mods:identifier>
<mods:identifier type="Zenodo-Dep">7160296</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:038687E4FFA7182DFE7CD81E7742FC84" httpUri="http://treatment.plazi.org/id/038687E4FFA7182DFE7CD81E7742FC84" lastPageId="6" lastPageNumber="307" pageId="4" pageNumber="305">
<subSubSection pageId="4" pageNumber="305" type="nomenclature">
<paragraph blockId="4.[491,894,812,838]" box="[491,894,812,838]" pageId="4" pageNumber="305">
<heading bold="true" box="[491,894,812,838]" fontSize="11" level="2" pageId="4" pageNumber="305" reason="3">
<emphasis bold="true" box="[491,894,812,838]" pageId="4" pageNumber="305">
<taxonomicName authority="Gerecke &amp; Smit, 2022" authorityName="Gerecke &amp; Smit" authorityYear="2022" box="[491,819,812,838]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="khatrii" status="n. sp." subGenus="Lebertia">
<emphasis bold="true" box="[491,597,812,838]" italics="true" pageId="4" pageNumber="305">Lebertia</emphasis>
(
<emphasis bold="true" box="[613,719,812,838]" italics="true" pageId="4" pageNumber="305">Lebertia</emphasis>
)
<emphasis bold="true" box="[735,819,812,838]" italics="true" pageId="4" pageNumber="305">khatrii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[827,894,812,838]" pageId="4" pageNumber="305" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="4.[490,1539,868,2028]" box="[491,1003,868,887]" pageId="4" pageNumber="305">
Zoobank:
<uuid box="[580,1003,868,887]" pageId="4" pageNumber="305">9FB20554-994E-4E0C-8D7C-D423BD7BFCFB</uuid>
</paragraph>
</subSubSection>
<subSubSection box="[531,739,899,923]" pageId="4" pageNumber="305" type="description">
<paragraph blockId="4.[490,1539,868,2028]" box="[531,739,899,923]" pageId="4" pageNumber="305">
(
<figureCitation box="[539,644,899,923]" captionStart="Figure 2" captionStartId="5.[119,186,1470,1491]" captionTargetBox="[127,1481,230,1403]" captionTargetId="figure-2@5.[103,1494,221,1414]" captionTargetPageId="5" captionText="Figure 2 Lebertia(Lebertia) khatriin. sp., type series. A-D, ♂; A – I-L-4-5; B – IV-L-4-6; C – right palp medial; D – coxal and genital field; E – ♀ genital. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7160300" httpUri="https://zenodo.org/record/7160300/files/figure.png" pageId="4" pageNumber="305">Figures 2</figureCitation>
A-E, 6B)
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="305" type="materials_examined">
<paragraph blockId="4.[490,1539,868,2028]" box="[531,1487,933,958]" pageId="4" pageNumber="305">
<emphasis bold="true" box="[531,655,934,958]" pageId="4" pageNumber="305">Type series</emphasis>
—
<materialsCitation box="[691,1487,933,958]" country="Bhutan" location="Bhutan" municipality="Royal Botanical Park Lampelri" pageId="4" pageNumber="305" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[691,790,933,957]" pageId="4" pageNumber="305">Holotype</typeStatus>
<specimenCount box="[795,817,934,957]" pageId="4" pageNumber="305" type="male">♂</specimenCount>
,
<emphasis bold="true" box="[827,943,934,958]" pageId="4" pageNumber="305">
<collectingCountry box="[827,917,934,958]" name="Bhutan" pageId="4" pageNumber="305">Bhutan</collectingCountry>
01
</emphasis>
,
<location LSID="urn:lsid:plazi:treatment:038687E4FFA7182DFE7CD81E7742FC84:8EF06029FFA7182FFC2DD8977008FC18" box="[954,1039,933,957]" country="Bhutan" municipality="Royal Botanical Park Lampelri" name="Bhutan" pageId="4" pageNumber="305">Bhutan</location>
,
<collectingMunicipality box="[1049,1377,933,957]" pageId="4" pageNumber="305">Royal Botanical Park Lampelri</collectingMunicipality>
, 27°30.450′
</materialsCitation>
</paragraph>
<paragraph blockId="4.[490,1539,868,2028]" pageId="4" pageNumber="305">
N,
<geoCoordinate box="[525,651,968,992]" degrees="89" direction="east" minutes="45.162" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="89.7527">89°45.162′ E</geoCoordinate>
, alt
<quantity box="[697,770,968,992]" metricMagnitude="3" metricUnit="m" metricValue="2.667" pageId="4" pageNumber="305" unit="m" value="2667.0">2667 m</quantity>
,
<date box="[782,934,968,992]" pageId="4" pageNumber="305" value="2016-08-10">10 Aug. 2016</date>
, leg. W. Klein, slide mounted (
<collectionCode box="[1290,1350,968,992]" pageId="4" pageNumber="305">NBC</collectionCode>
).
<typeStatus box="[1368,1476,969,993]" pageId="4" pageNumber="305">Paratypes</typeStatus>
: Same site and date,
<specimenCount box="[716,774,1003,1027]" pageId="4" pageNumber="305" type="male">2 ♂♂</specimenCount>
,
<specimenCount box="[785,858,1003,1027]" pageId="4" pageNumber="305" type="female">10 ♀♀</specimenCount>
(
<specimenCount box="[872,932,1003,1027]" pageId="4" pageNumber="305" type="male">2 ♂♂</specimenCount>
,
<specimenCount box="[943,1003,1003,1027]" pageId="4" pageNumber="305" type="female">4 ♀♀</specimenCount>
slide mounted,
<specimenCount box="[1185,1225,1003,1027]" pageId="4" pageNumber="305" type="female">6 ♀</specimenCount>
in liquid) (
<collectionCode box="[1348,1399,1003,1027]" pageId="4" pageNumber="305">NBC</collectionCode>
);
<emphasis bold="true" pageId="4" pageNumber="305">
<collectingCountry box="[1409,1499,1003,1027]" name="Bhutan" pageId="4" pageNumber="305">Bhutan</collectingCountry>
02
</emphasis>
, E of Takchu, springs in meadow,
<geoCoordinate box="[905,1023,1037,1061]" degrees="27" direction="north" minutes="21.492" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="27.3582">27°21.492′ N</geoCoordinate>
,
<geoCoordinate box="[1034,1146,1037,1061]" degrees="89" direction="east" minutes="17.173" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="89.28622">89°17.173′ E</geoCoordinate>
,
<quantity box="[1158,1221,1037,1061]" metricMagnitude="3" metricUnit="m" metricValue="2.907" pageId="4" pageNumber="305" unit="m" value="2907.0">2907 m</quantity>
a.s.l.,
<date box="[1290,1434,1037,1061]" pageId="4" pageNumber="305" value="2016-08-07">07 Aug. 2016</date>
,
</paragraph>
<paragraph blockId="4.[490,1539,868,2028]" pageId="4" pageNumber="305">
leg. W. Klein,
<specimenCount box="[653,695,1072,1096]" pageId="4" pageNumber="305" type="male">1 ♂</specimenCount>
,
<specimenCount box="[706,746,1072,1096]" pageId="4" pageNumber="305" type="female">1 ♀</specimenCount>
slide mounted,
<specimenCount box="[928,983,1072,1096]" pageId="4" pageNumber="305" type="female">2 ♀♀</specimenCount>
in liquid;
<emphasis bold="true" box="[1079,1197,1072,1096]" pageId="4" pageNumber="305">
<collectingCountry box="[1079,1169,1072,1096]" name="Bhutan" pageId="4" pageNumber="305">Bhutan</collectingCountry>
03
</emphasis>
, Dumcho,
<collectingRegion box="[1320,1363,1072,1095]" country="Bhutan" name="Ha" pageId="4" pageNumber="305">Haa</collectingRegion>
river, pool 3,
<geoCoordinate box="[491,607,1106,1130]" degrees="27" direction="north" minutes="21.883" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="27.364717">27°21.883′ N</geoCoordinate>
,
<geoCoordinate box="[618,725,1106,1130]" degrees="89" direction="east" minutes="17.738" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="89.29563">89°17.738′ E</geoCoordinate>
,
<quantity box="[737,807,1106,1130]" metricMagnitude="3" metricUnit="m" metricValue="2.675" pageId="4" pageNumber="305" unit="m" value="2675.0">2675 m</quantity>
a.s.l.,
<date box="[876,1019,1106,1130]" pageId="4" pageNumber="305" value="2016-08-07">07 Aug. 2016</date>
, leg. W. Klein,
<specimenCount box="[1191,1231,1106,1130]" pageId="4" pageNumber="305" type="male">1 ♂</specimenCount>
slide mounted.
</paragraph>
<paragraph blockId="4.[490,1539,868,2028]" box="[531,1425,1141,1165]" pageId="4" pageNumber="305">
Other material –
<emphasis bold="true" box="[711,829,1141,1165]" pageId="4" pageNumber="305">
<collectingCountry box="[711,801,1141,1165]" name="Bhutan" pageId="4" pageNumber="305">Bhutan</collectingCountry>
03
</emphasis>
, Dumcho,
<collectingRegion box="[952,995,1141,1164]" country="Bhutan" name="Ha" pageId="4" pageNumber="305">Haa</collectingRegion>
river, pool 3,
<geoCoordinate box="[1139,1254,1141,1165]" degrees="27" direction="north" minutes="21.883" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="27.364717">27°21.883′ N</geoCoordinate>
,
<geoCoordinate box="[1265,1372,1141,1165]" degrees="89" direction="east" minutes="17.738" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="89.29563">89°17.738′ E</geoCoordinate>
, 2675
</paragraph>
<paragraph blockId="4.[490,1539,868,2028]" pageId="4" pageNumber="305">
m a.s.l.,
<date box="[582,718,1175,1199]" pageId="4" pageNumber="305" value="2016-08-07">07 Aug.2016</date>
, leg. W. Klein (
<figureCitation box="[893,1012,1175,1199]" captionStart="Figure 6" captionStartId="12.[119,186,1883,1904]" captionTargetBox="[103,1494,277,1829]" captionTargetId="figure-2@12.[103,1494,277,1829]" captionTargetPageId="12" captionText="Figure 6 Collection sites. A – Haa River (Bhutan 03); B – Sertu Khola, Nunthala (Nepal 06); C – Garden stream, Nunthala (Nepal 05); D – Surke Khola (Nepal 04); E – stream S of Gokyo (Nepal 01); F – stream Dudh Kosi Nadi (Nepal 02). Authors: W. Klein (A), P.V. Veel (B-F)." figureDoi="http://doi.org/10.5281/zenodo.7160312" httpUri="https://zenodo.org/record/7160312/files/figure.png" pageId="4" pageNumber="305">Figure 6A</figureCitation>
), 1 dn slide mounted;
<emphasis bold="true" box="[1243,1343,1176,1200]" pageId="4" pageNumber="305">
<collectingCountry box="[1243,1315,1176,1200]" name="Nepal" pageId="4" pageNumber="305">Nepal</collectingCountry>
04
</emphasis>
, Surke Khola, stream,
<geoCoordinate box="[576,695,1210,1234]" degrees="27" direction="north" minutes="40.327" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="27.672117">27°40.327′ N</geoCoordinate>
,
<geoCoordinate box="[706,821,1210,1234]" degrees="86" direction="east" minutes="43.297" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="86.72162">86°43.297′ E</geoCoordinate>
,
<quantity box="[833,899,1210,1234]" metricMagnitude="3" metricUnit="m" metricValue="2.29" pageId="4" pageNumber="305" unit="m" value="2290.0">2290 m</quantity>
a.s.l.,
<date box="[969,1105,1210,1234]" pageId="4" pageNumber="305" value="2016-04-11">11 Apr. 2016</date>
leg P.W. Veel (
<figureCitation box="[1274,1393,1210,1234]" captionStart="Figure 6" captionStartId="12.[119,186,1883,1904]" captionTargetBox="[103,1494,277,1829]" captionTargetId="figure-2@12.[103,1494,277,1829]" captionTargetPageId="12" captionText="Figure 6 Collection sites. A – Haa River (Bhutan 03); B – Sertu Khola, Nunthala (Nepal 06); C – Garden stream, Nunthala (Nepal 05); D – Surke Khola (Nepal 04); E – stream S of Gokyo (Nepal 01); F – stream Dudh Kosi Nadi (Nepal 02). Authors: W. Klein (A), P.V. Veel (B-F)." figureDoi="http://doi.org/10.5281/zenodo.7160312" httpUri="https://zenodo.org/record/7160312/files/figure.png" pageId="4" pageNumber="305">Figure 6D</figureCitation>
), 1
</paragraph>
<paragraph blockId="4.[490,1539,868,2028]" pageId="4" pageNumber="305">
<specimenCount box="[491,511,1245,1268]" pageId="4" pageNumber="305" type="female">♀</specimenCount>
slide mounted (juvenile, grading
<emphasis box="[896,914,1245,1268]" italics="true" pageId="4" pageNumber="305">L</emphasis>
to.
<taxonomicName authority="Gerecke &amp; Smit, 2022" authorityName="Gerecke &amp; Smit" authorityYear="2022" box="[924,974,1244,1268]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="veeli" status="n. sp.">
<emphasis box="[924,974,1244,1268]" italics="true" pageId="4" pageNumber="305">veeli</emphasis>
</taxonomicName>
<emphasis bold="true" box="[980,1040,1245,1269]" pageId="4" pageNumber="305">
<taxonomicNameLabel box="[980,1040,1245,1269]" pageId="4" pageNumber="305" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
possibly due to incomplete sclerotization);
<emphasis bold="true" box="[491,591,1279,1303]" pageId="4" pageNumber="305">
<collectingCountry box="[491,563,1279,1303]" name="Nepal" pageId="4" pageNumber="305">Nepal</collectingCountry>
06
</emphasis>
, Sertu Khola stream,
<geoCoordinate box="[825,932,1279,1303]" degrees="27" direction="north" minutes="33.899" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="27.564983">27°33.899′ N</geoCoordinate>
,
<geoCoordinate box="[941,1061,1279,1303]" degrees="86" direction="east" minutes="35.752" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="86.59586">86°35.752′ E</geoCoordinate>
,
<quantity box="[1072,1143,1279,1303]" metricMagnitude="3" metricUnit="m" metricValue="2.563" pageId="4" pageNumber="305" unit="m" value="2563.0">2563 m</quantity>
a.s.l.,
<date box="[1211,1345,1279,1303]" pageId="4" pageNumber="305" value="2016-04-14">14 Apr. 2016</date>
leg P.W.
</paragraph>
<paragraph blockId="4.[490,1539,868,2028]" box="[491,1468,1313,1338]" pageId="4" pageNumber="305">
Veel (
<figureCitation box="[551,667,1313,1337]" captionStart="Figure 6" captionStartId="12.[119,186,1883,1904]" captionTargetBox="[103,1494,277,1829]" captionTargetId="figure-2@12.[103,1494,277,1829]" captionTargetPageId="12" captionText="Figure 6 Collection sites. A – Haa River (Bhutan 03); B – Sertu Khola, Nunthala (Nepal 06); C – Garden stream, Nunthala (Nepal 05); D – Surke Khola (Nepal 04); E – stream S of Gokyo (Nepal 01); F – stream Dudh Kosi Nadi (Nepal 02). Authors: W. Klein (A), P.V. Veel (B-F)." figureDoi="http://doi.org/10.5281/zenodo.7160312" httpUri="https://zenodo.org/record/7160312/files/figure.png" pageId="4" pageNumber="305">Figure 6B</figureCitation>
),
<specimenCount box="[685,724,1313,1337]" pageId="4" pageNumber="305" type="female">1 ♀</specimenCount>
slide mounted;
<emphasis bold="true" box="[886,984,1314,1338]" pageId="4" pageNumber="305">
<collectingCountry box="[886,958,1314,1338]" name="Nepal" pageId="4" pageNumber="305">Nepal</collectingCountry>
09
</emphasis>
, Bhujekhung, Stream,
<geoCoordinate box="[1245,1362,1313,1337]" degrees="28" direction="north" minutes="31.709" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="28.528482">28°31.709′ N</geoCoordinate>
, 83°01.598′
</paragraph>
<paragraph blockId="4.[490,1539,868,2028]" pageId="4" pageNumber="305">
E,
<quantity box="[520,596,1348,1372]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="305" unit="m" value="3100.0">3100 m</quantity>
a.s.l.,
<date box="[663,795,1348,1372]" pageId="4" pageNumber="305" value="2016-10-10">10 Oct. 2016</date>
leg P.W. Veel
<specimenCount box="[951,992,1348,1372]" pageId="4" pageNumber="305" type="male">1 ♂</specimenCount>
,
<specimenCount box="[1003,1057,1348,1372]" pageId="4" pageNumber="305" type="female">2 ♀♀</specimenCount>
slide mounted;
<emphasis bold="true" box="[1233,1332,1348,1372]" pageId="4" pageNumber="305">
<collectingCountry box="[1233,1305,1348,1372]" name="Nepal" pageId="4" pageNumber="305">Nepal</collectingCountry>
10
</emphasis>
, Rupgad, Thuli Beri River,
<geoCoordinate box="[615,747,1382,1406]" degrees="28" direction="north" minutes="58.460" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="28.974333">28°58.460′ N</geoCoordinate>
,
<geoCoordinate box="[757,876,1382,1406]" degrees="82" direction="east" minutes="51.991" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="82.866516">82°51.991′ E</geoCoordinate>
,
<quantity box="[888,958,1382,1406]" metricMagnitude="3" metricUnit="m" metricValue="2.0669999999999997" pageId="4" pageNumber="305" unit="m" value="2067.0">2067 m</quantity>
a.s.l.,
<date box="[1027,1167,1382,1406]" pageId="4" pageNumber="305" value="2016-11-01">01 Nov. 2016</date>
leg P.W. Veel,
<specimenCount box="[1333,1389,1382,1406]" pageId="4" pageNumber="305" type="female">2 ♀♀</specimenCount>
slide mounted.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="305" type="diagnosis">
<paragraph blockId="4.[490,1539,868,2028]" pageId="4" pageNumber="305">
<emphasis bold="true" box="[531,646,1452,1476]" pageId="4" pageNumber="305">Diagnosis</emphasis>
— Integument lineated, with lines in parts of the dorsum shortened and fusing to form a reticulate pattern; legs without swimming setae; P-3 basally relatively narrow (ratio maximum H/basal H 1.3-1.4), with 5 long setae, distal setae nearly equidistant or centrodistal seta closer to dorsodistal seta (A/B 1.3-2.1); P-4 maximum H in central part, insertions of setae dividing ventral margin in general into a short proximal, and subequal central and distal sector(s) (17-27/33-48/26-50 %).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305" type="description">
<paragraph blockId="4.[490,1539,868,2028]" lastBlockId="5.[491,1551,1635,2039]" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305">
<emphasis bold="true" box="[531,662,1659,1683]" pageId="4" pageNumber="305">Description</emphasis>
— Integument lines surrounding longish fields of fine porosity, in parts of the dorsum shortened and fusing to form a characteristic reticulation, extension of this reticulation highly variable, in most specimens covering most of the dorsum, in others restricted to small dorsocentral fields. Cx-I medially distinctly longer than Cx-II (ratio 1.1-1.4), posterior margin W 25-35. Posterolateral margin of Cx-IV equally rounded or with a nearly straight sector on the level of glandularia V3. These and setae V1 not fused to coxal shield, but V3 often very close to Cx-IV margin (
<figureCitation box="[746,863,1866,1890]" captionStart="Figure 2" captionStartId="5.[119,186,1470,1491]" captionTargetBox="[127,1481,230,1403]" captionTargetId="figure-2@5.[103,1494,221,1414]" captionTargetPageId="5" captionText="Figure 2 Lebertia(Lebertia) khatriin. sp., type series. A-D, ♂; A – I-L-4-5; B – IV-L-4-6; C – right palp medial; D – coxal and genital field; E – ♀ genital. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7160300" httpUri="https://zenodo.org/record/7160300/files/figure.png" pageId="4" pageNumber="305">Figure 2D</figureCitation>
). Proportions and setation of distal segments of I- and IV-L as in
<figureCitation box="[548,642,1901,1925]" captionStart="Figure 2" captionStartId="5.[119,186,1470,1491]" captionTargetBox="[127,1481,230,1403]" captionTargetId="figure-2@5.[103,1494,221,1414]" captionTargetPageId="5" captionText="Figure 2 Lebertia(Lebertia) khatriin. sp., type series. A-D, ♂; A – I-L-4-5; B – IV-L-4-6; C – right palp medial; D – coxal and genital field; E – ♀ genital. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7160300" httpUri="https://zenodo.org/record/7160300/files/figure.png" pageId="4" pageNumber="305">Figure 2</figureCitation>
A-B; distal 2/3 of the ventral surface of IV-L-5 with more than 10 setae inserting close to each other, IV-L-6 slightly bowed, with 2-3 fine and short ventral setae; IV-L- claw L 55-67. Genital field (
<figureCitation box="[750,861,1970,1994]" captionStart="Figure 2" captionStartId="5.[119,186,1470,1491]" captionTargetBox="[127,1481,230,1403]" captionTargetId="figure-2@5.[103,1494,221,1414]" captionTargetPageId="5" captionText="Figure 2 Lebertia(Lebertia) khatriin. sp., type series. A-D, ♂; A – I-L-4-5; B – IV-L-4-6; C – right palp medial; D – coxal and genital field; E – ♀ genital. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7160300" httpUri="https://zenodo.org/record/7160300/files/figure.png" pageId="4" pageNumber="305">Figure 2E</figureCitation>
) with lateral margins of genital flaps strongly diverging from anterior to posterior end, medial setae (
<specimenCount box="[989,1012,2004,2028]" pageId="4" pageNumber="305" type="male">♂</specimenCount>
:&gt; 30,
<specimenCount box="[1087,1109,2005,2028]" pageId="4" pageNumber="305" type="female">♀</specimenCount>
: ca 15), in general in a single row (posteriorly doubled in a few
<specimenCount box="[826,869,1635,1659]" pageId="5" pageNumber="306" type="male">♂♂</specimenCount>
). Excretory pore smooth. Gnathosoma ventral margin in distal part nearly straight. P-2 relatively robust (L/H 1.37-1.54), ventral seta inserted 7-10 from distoventral edge, basally strong, distinctly shorter than L of segment (
<figureCitation box="[1291,1405,1704,1728]" captionStart="Figure 2" captionStartId="5.[119,186,1470,1491]" captionTargetBox="[127,1481,230,1403]" captionTargetId="figure-2@5.[103,1494,221,1414]" captionTargetPageId="5" captionText="Figure 2 Lebertia(Lebertia) khatriin. sp., type series. A-D, ♂; A – I-L-4-5; B – IV-L-4-6; C – right palp medial; D – coxal and genital field; E – ♀ genital. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7160300" httpUri="https://zenodo.org/record/7160300/files/figure.png" pageId="5" pageNumber="306">Figure 2C</figureCitation>
); P-3 slender, in basal part narrower than distally, dorsal and ventral diverging distally, 2 of the 5 long setae arranged near the centre of the dorsum (close to, or clearly away from each other), centrodistal seta closer to dorsodistal one, distoventral seta near segment edge; P-4 with maximum H in centre or proximal part, mediodistal peg seta strong, pointed; proximoventral seta rather close to proximal segment edge, distoventral seta halfway between proximoventral seta and distal segment edge (in one exceptional case, the central sector is enlarged (26/48/26 %).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160300" ID-Zenodo-Dep="7160300" httpUri="https://zenodo.org/record/7160300/files/figure.png" pageId="5" pageNumber="306" startId="5.[119,186,1470,1491]" targetBox="[127,1481,230,1403]" targetPageId="5">
<paragraph blockId="5.[119,1507,1470,1517]" box="[119,1507,1470,1492]" pageId="5" pageNumber="306">
<emphasis bold="true" box="[119,204,1470,1491]" pageId="5" pageNumber="306">Figure 2</emphasis>
<taxonomicName authority="Gerecke &amp; Smit, 2022" authorityName="Gerecke &amp; Smit" authorityYear="2022" box="[216,476,1470,1492]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="khatrii" status="n. sp." subGenus="Lebertia">
<emphasis box="[216,303,1470,1492]" italics="true" pageId="5" pageNumber="306">Lebertia</emphasis>
(
<emphasis box="[311,392,1470,1492]" italics="true" pageId="5" pageNumber="306">Lebertia</emphasis>
)
<emphasis box="[406,476,1470,1492]" italics="true" pageId="5" pageNumber="306">khatrii</emphasis>
</taxonomicName>
<emphasis bold="true" box="[475,530,1470,1491]" pageId="5" pageNumber="306">
<taxonomicNameLabel box="[475,530,1470,1491]" pageId="5" pageNumber="306" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, type series. A-D, ♂; A – I-L-4-5; B – IV-L-4-6; C – right palp medial; D – coxal and genital field;
</paragraph>
<paragraph blockId="5.[119,1507,1470,1517]" box="[119,460,1495,1517]" pageId="5" pageNumber="306">E – ♀ genital. Scale bars: 100 µm.</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="306" type="etymology">
<paragraph blockId="5.[491,1551,1635,2039]" pageId="5" pageNumber="306">
<emphasis bold="true" box="[531,654,1946,1970]" pageId="5" pageNumber="306">Etymology</emphasis>
— Named after Rajan Khatri (Kathmandu), for his assistance with the collecting of the material.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306" type="discussion">
<paragraph blockId="5.[491,1551,1635,2039]" lastBlockId="6.[491,1538,229,801]" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
<emphasis bold="true" box="[531,651,2015,2039]" pageId="5" pageNumber="306">Discussion</emphasis>
— From the Asian fauna the only
<taxonomicName authorityName="Neuman" authorityYear="1880" box="[1029,1127,2015,2039]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis box="[1029,1127,2015,2039]" italics="true" pageId="5" pageNumber="306">Lebertia</emphasis>
</taxonomicName>
(s.str.) species reported to have a reticulated integument is
<taxonomicName authority="Sokolow, 1934" authorityName="Sokolow" authorityYear="1934" box="[762,1009,229,253]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="acuta">
<emphasis box="[762,857,230,253]" italics="true" pageId="6" pageNumber="307">L. acuta</emphasis>
Sokolow, 1934
</taxonomicName>
, described from the Ussuri catchment. This species differs from
<taxonomicName authority="Gerecke &amp; Smit, 2022" authorityName="Gerecke &amp; Smit" authorityYear="2022" box="[705,808,264,288]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="khatrii" status="n. sp.">
<emphasis box="[705,808,264,288]" italics="true" pageId="6" pageNumber="307">L. khatrii</emphasis>
</taxonomicName>
<emphasis bold="true" box="[808,867,264,288]" pageId="6" pageNumber="307">
<taxonomicNameLabel box="[808,867,264,288]" pageId="6" pageNumber="307" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in the position of the centrodistal of P-3 which is very close to the dorsodistal one (A/B 3.0). The same is true for
<taxonomicName authority="Sokolow, 1948" authorityName="Sokolow" authorityYear="1948" box="[1076,1362,298,322]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="hissarica">
<emphasis box="[1076,1210,298,322]" italics="true" pageId="6" pageNumber="307">L. hissarica</emphasis>
Sokolow, 1948
</taxonomicName>
, a species from
<collectingCountry box="[551,675,332,356]" name="Tajikistan" pageId="6" pageNumber="307">Tadjikistan</collectingCountry>
that might have a similar integument structure (but not described explicitly, ratio A/B&gt; 3.0). In the European fauna, two
<taxonomicName authorityName="Neuman" authorityYear="1880" box="[995,1094,366,390]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis box="[995,1094,366,390]" italics="true" pageId="6" pageNumber="307">Lebertia</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1093,1142,367,390]" pageId="6" pageNumber="307" sensu="stricto">s.str.</taxonomicNameLabel>
species are known to have a reticulated integument, both differing from the new species, among others, in the position of ventral setae on P-4:
<taxonomicName authority="Koenike, 1908" authorityName="Koenike" authorityYear="1908" box="[722,1078,435,459]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="salebrosa">
<emphasis box="[722,929,435,459]" italics="true" pageId="6" pageNumber="307">Lebertia salebrosa</emphasis>
Koenike, 1908
</taxonomicName>
, in comparison with the new species with a distinctly finer integument lineation locally producing a rather irregular reticulation, is generally minor in dimensions (e.g., P-4 L
<specimenCount box="[970,990,504,527]" pageId="6" pageNumber="307" type="male">♂</specimenCount>
&lt;115,
<specimenCount box="[1072,1092,504,527]" pageId="6" pageNumber="307" type="female">♀</specimenCount>
&lt;130), a farther distance between P-3 ventro- and centrodistal setae (ratio A/B 2.2-4.0), and the two proximal ventral sectors of P-4 shorter than the distal one (25/25/50 %), rarely equal in size (33/33/33 %). Specimens of
<taxonomicName authority="Koenike, 1919" authorityName="Koenike" authorityYear="1919" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis italics="true" pageId="6" pageNumber="307">L. reticulata</emphasis>
Koenike, 1919
</taxonomicName>
differ in presence of one swimming seta each on III-/IV-L-5, and a row of strong ventral setae on IV-L-6. Furthermore, their P-4 is club-shaped (maximum H in distal part) and both ventral setae insert in the distal segment sector.
</paragraph>
<paragraph blockId="6.[491,1538,229,801]" pageId="6" pageNumber="307">
Several specimens from
<collectingCountry box="[793,856,708,732]" name="Nepal" pageId="6" pageNumber="307">Nepal</collectingCountry>
do not show a distinct integument reticulation, but a pattern of longer or shorter lines similar to the situation in
<taxonomicName authority="Gerecke &amp; Smit, 2022" authorityName="Gerecke &amp; Smit" authorityYear="2022" box="[1051,1131,742,766]" class="Arachnida" family="Lebertiidae" genus="Lebertia" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="veeli" status="n. sp.">
<emphasis box="[1051,1131,742,766]" italics="true" pageId="6" pageNumber="307">L. veeli</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1140,1201,743,767]" pageId="6" pageNumber="307">
<taxonomicNameLabel box="[1140,1201,743,767]" pageId="6" pageNumber="307" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but they agree with the diagnosis in palp morphology.
</paragraph>
</subSubSection>
</treatment>
</document>
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