166 lines
18 KiB
XML
166 lines
18 KiB
XML
<document id="6273F47B58DCE875F051E376ABE9BD5C" ID-DOI="10.1093/zoolinnean/zlad071" ID-ISSN="0024-4082" ID-Zenodo-Dep="10470445" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1704719350260" checkinUser="plazi" docAuthor="Costa Santos, Mateus A., Paes Neto, Voltaire D., Schultz, Cesar L., Cisneros, Juan & Pinheiro, Stephanie E. Pierce and Felipe L." docDate="2023" docId="038687C0AF1CFFBA999AF8BD0D2FFF06" docLanguage="en" docName="zlad071.pdf" docOrigin="Zoological Journal of the Linnean Society 199" docSource="https://www.mendeley.com/catalogue/f8822c77-50b8-3b8e-b73f-a2be58219f19/" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Pampaphoneus biccai Cisneros 2012" docType="treatment" docVersion="1" lastPageNumber="1039" masterDocId="FFBFFFB8AF18FFBC9945FFDF0C68FFE0" masterDocTitle="Cranial osteology of the Brazilian dinocephalian Pampaphoneus biccai (Anteosauridae: Syodontinae)" masterLastPageNumber="1058" masterPageNumber="1034" pageNumber="1038" updateTime="1705068750349" updateUser="guilherme">
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<mods:title id="4BA595DCCD53BA8A4B7F9F25F09AA4EA">Cranial osteology of the Brazilian dinocephalian Pampaphoneus biccai (Anteosauridae: Syodontinae)</mods:title>
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<mods:namePart id="CF5680DF3E1404AC3C9E834A217AF943">Costa Santos, Mateus A.</mods:namePart>
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<mods:affiliation id="9E672D22134EC2340C4E1EF25FD3696A">Programa de Pós-Graduação em Geociências, Instituto de Geociências, Universidade Federal do Rio Grande do Sul, Porto Alegre, Rio Grande do Sul, Brazil & Laboratório de Paleobiologia, Universidade Federal do Pampa, Campus São Gabriel, São Gabriel, Rio Grande do Sul, Brazil</mods:affiliation>
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<mods:nameIdentifier id="C636EDBA5C371BDA65D9065F4ABD9662" type="email">santosmac@outlook.com.br</mods:nameIdentifier>
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<mods:roleTerm id="A23F06182BAC46E7D7A46DDCC89E228D">Author</mods:roleTerm>
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<mods:namePart id="1646A0808594AF1326448356D21C0191">Paes Neto, Voltaire D.</mods:namePart>
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<mods:affiliation id="C1323502B53D1CA266D491827452B7FA">Laboratório de Paleobiologia, Universidade Federal do Pampa, Campus São Gabriel, São Gabriel, Rio Grande do Sul, Brazil & Museum of Comparative Zoology and Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachuseưs, USA</mods:affiliation>
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<mods:namePart id="DF7EC711F398B8976A240F77CE1F5DEB">Schultz, Cesar L.</mods:namePart>
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<mods:affiliation id="475B79787492A13C20BD5A33536C4774">Laboratório de Paleovertebrados, Instituto de Geociências, Universidade Federal do Rio Grande do Sul, Porto Alegre, Rio Grande do Sul, Brazil</mods:affiliation>
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<mods:namePart id="13BE8459BB724019B519600CCD281035">Cisneros, Juan</mods:namePart>
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<mods:affiliation id="4C5853A86A80A573C5F394ABEF4AE56E">Museu de Arqueologia e Paleontologia, Universidade Federal do Piauí, Teresina</mods:affiliation>
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<mods:namePart id="4210838C1E6D2FDDF98BDD8AA7807ED0">Pinheiro, Stephanie E. Pierce and Felipe L.</mods:namePart>
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<mods:title id="9B8FAB70C3212483B8509CFBBF45EF2D">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="39FE03298C5B802F087A3FA5648B014D">2023</mods:date>
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<mods:number id="4F3F703F1EEE98A1990F2FC06013F7CD">2023-09-10</mods:number>
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<mods:number id="C4F6E15C5C057A5A321D2EAF46A5A3C7">199</mods:number>
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<mods:start id="C836EC8605A997C847F8C67BB55170BD">1034</mods:start>
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<mods:end id="A9FD184BD490A1D20E946F6DB50D513D">1058</mods:end>
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<mods:classification id="D648141B8408A89825CB19F32BDA1F2A">journal article</mods:classification>
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<mods:identifier id="0D16FB7888A26DA74A9220A911B28108" type="DOI">10.1093/zoolinnean/zlad071</mods:identifier>
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<mods:identifier id="7B7B80C1320D42D94CDFAF367F64804A" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="A32FBE90EE470207463974D31973963B" type="Zenodo-Dep">10470445</mods:identifier>
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<treatment id="038687C0AF1CFFBA999AF8BD0D2FFF06" LSID="urn:lsid:plazi:treatment:038687C0AF1CFFBA999AF8BD0D2FFF06" httpUri="http://treatment.plazi.org/id/038687C0AF1CFFBA999AF8BD0D2FFF06" lastPageId="6" lastPageNumber="1039" pageId="4" pageNumber="1038">
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<subSubSection id="C335655DAF1CFFB8999AF8BD0EE5F89C" box="[223,653,1890,1917]" pageId="4" pageNumber="1038" type="nomenclature">
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<paragraph id="8B9036D6AF1CFFB8999AF8BD0EE5F89C" blockId="4.[223,653,1890,1917]" box="[223,653,1890,1917]" pageId="4" pageNumber="1038">
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<taxonomicName id="4C2F4D55AF1CFFB8999AF8BD0EE5F89C" authority="Cisneros et al. 2012" authorityName="Cisneros" authorityYear="2012" box="[223,653,1890,1917]" family="Anteosauridae" genus="Pampaphoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinocephalia" pageId="4" pageNumber="1038" phylum="Chordata" rank="species" species="biccai">
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<emphasis id="B95BEAC4AF1CFFB8999AF8BD0E74F89C" bold="true" box="[223,540,1890,1917]" pageId="4" pageNumber="1038">
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<emphasis id="B95BEAC4AF1CFFB8999AF8BD0DD0F89D" bold="true" box="[223,440,1890,1917]" italics="true" pageId="4" pageNumber="1038">Pampaphoneus biccai</emphasis>
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<bibRefCitation id="EFBE4B27AF1CFFB898F8F8BD0E74F89C" author="Cisneros JC & Abdala F & Atayman-Guven S" box="[445,540,1890,1916]" pageId="4" pageNumber="1038" pagination="1584 - 8" refId="ref14921" refString="Cisneros JC, Abdala F, Atayman-Guven S et al. Carnivorous dinocephalian from the Middle Late Permiano of Brazil and tetrapod dispersal in Pangaea. PNAS 2012; 109: 1584 - 8." type="journal article" year="2012">Cisneros</bibRefCitation>
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</emphasis>
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||
<emphasis id="B95BEAC4AF1CFFB89B65F8BC0E3AF89C" box="[544,594,1890,1916]" italics="true" pageId="4" pageNumber="1038">et al.</emphasis>
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<emphasis id="B95BEAC4AF1CFFB89B13F8BD0EE5F89C" bold="true" box="[598,653,1890,1916]" pageId="4" pageNumber="1038">2012</emphasis>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C335655DAF1CFFB89934F87C0F12F858" pageId="4" pageNumber="1038" type="materials_examined">
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<paragraph id="8B9036D6AF1CFFB89934F87C0E94F85B" blockId="4.[113,764,1955,1979]" box="[113,764,1955,1979]" pageId="4" pageNumber="1038">
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<emphasis id="B95BEAC4AF1CFFB89934F87C0CA5F85B" box="[113,205,1955,1979]" italics="true" pageId="4" pageNumber="1038">
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<typeStatus id="54948874AF1CFFB89934F87C0CA0F85B" box="[113,200,1955,1979]" pageId="4" pageNumber="1038" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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UFRGS–PV-0386-P, a near complete skull with lower jaw.
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</paragraph>
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<paragraph id="8B9036D6AF1CFFB89A6FF9930FE5F8E1" blockId="4.[810,1459,1611,1793]" pageId="4" pageNumber="1038">
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<emphasis id="B95BEAC4AF1CFFB89A6FF9930876F984" box="[810,1054,1612,1636]" italics="true" pageId="4" pageNumber="1038">Locality and horizon:</emphasis>
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<materialsCitation id="3B473C8BAF1CFFB89D68F9930FE5F8E1" country="Brazil" county="Rio do Rasto Formation" latitude="-30.00316" location="Catucaba District" longLatPrecision="1" longitude="-54.088177" municipality="Sao Gabriel" pageId="4" pageNumber="1038" specimenCount="1" stateProvince="Rio Grande do Sul State">
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Boqueirão Farm (
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||
<geoCoordinate id="EE1B5011AF1CFFB89DBBF9920F51F964" degrees="30" direction="south" minutes="00" orientation="latitude" pageId="4" pageNumber="1038" precision="1" seconds="11.38" value="-30.00316">30° 00ʹ 11.38ʹʹ S</geoCoordinate>
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,
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<geoCoordinate id="EE1B5011AF1CFFB89A08F9B30844F964" box="[845,1068,1642,1668]" degrees="54" direction="west" minutes="05" orientation="longitude" pageId="4" pageNumber="1038" precision="1" seconds="17.44" value="-54.088177">54° 05ʹ 17.44ʹʹ W</geoCoordinate>
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),
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||
<location id="8EF0600DAF1CFFB89D0CF9B3097FF964" LSID="urn:lsid:plazi:treatment:038687C0AF1CFFBA999AF8BD0D2FFF06:8EF0600DAF1CFFB89D0CF9B3097FF964" box="[1097,1303,1644,1668]" country="Brazil" county="Rio do Rasto Formation" latitude="-30.00316" longLatPrecision="1" longitude="-54.088177" municipality="Sao Gabriel" name="Catucaba District" pageId="4" pageNumber="1038" stateProvince="Rio Grande do Sul State">Catuçaba District</location>
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||
,
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||
<collectingMunicipality id="6BF4ACACAF1CFFB89C6FF9B309DAF964" box="[1322,1458,1644,1668]" pageId="4" pageNumber="1038">São Gabriel</collectingMunicipality>
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||
Municipality,
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||
<collectingRegion id="49EBF834AF1CFFB89A82F954088AF943" box="[967,1250,1675,1699]" country="Brazil" name="Rio Grande do Sul" pageId="4" pageNumber="1038">Rio Grande do Sul State</collectingRegion>
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,
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<collectingCountry id="F3387646AF1CFFB89DB5F954095BF943" box="[1264,1331,1675,1699]" name="Brazil" pageId="4" pageNumber="1038">Brazil</collectingCountry>
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. The rocks exposed at this site belongs to the Morro Pelado Member of the Rio do Rasto Formation (Guadalupian) (
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<bibRefCitation id="EFBE4B27AF1CFFB89C7CF9150FEAF8E1" author="Cisneros JC & Abdala F & Atayman-Guven S" pageId="4" pageNumber="1038" pagination="1584 - 8" refId="ref14921" refString="Cisneros JC, Abdala F, Atayman-Guven S et al. Carnivorous dinocephalian from the Middle Late Permiano of Brazil and tetrapod dispersal in Pangaea. PNAS 2012; 109: 1584 - 8." type="journal article" year="2012">
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Cisneros
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<emphasis id="B95BEAC4AF1CFFB89CE4F9140F2CF8E1" italics="true" pageId="4" pageNumber="1038">et al.</emphasis>
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2012
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</bibRefCitation>
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||
)
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||
</materialsCitation>
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</paragraph>
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||
<paragraph id="8B9036D6AF1CFFB89A6FF8FD0F12F858" blockId="4.[810,1459,1826,1976]" pageId="4" pageNumber="1038">
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<emphasis id="B95BEAC4AF1CFFB89A6FF8FD0F87F8DA" box="[810,1007,1826,1850]" italics="true" pageId="4" pageNumber="1038">Referred specimens:</emphasis>
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UNIPAMPA 759, a near complete skull with lower jaw and postcranial remains also collected from the type-locality; UFRGS-PV-0249-P, a fragmentary right dentary, collected from the ‘Fazenda Fagundes’ locality (
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<geoCoordinate id="EE1B5011AF1CFFB89C65F85E09ECF879" box="[1312,1412,1919,1945]" degrees="30" direction="south" minutes="01" orientation="latitude" pageId="4" pageNumber="1038" precision="925" value="-30.016666">30° 01ʹ S</geoCoordinate>
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,
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<geoCoordinate id="EE1B5011AF1CFFB89CD4F85F0F07F858" degrees="54" direction="west" minutes="09" orientation="longitude" pageId="4" pageNumber="1038" precision="925" value="-54.15">54° 09ʹ W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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||
<caption id="DF50665EAF1DFFB999C4F9570F75F95D" ID-DOI="http://doi.org/10.5281/zenodo.10470453" ID-Zenodo-Dep="10470453" httpUri="https://zenodo.org/record/10470453/files/figure.png" pageId="5" pageNumber="1039" startId="5.[129,194,1672,1696]" targetBox="[129,1469,147,1642]" targetPageId="5" targetType="figure">
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<paragraph id="8B9036D6AF1DFFB999C4F9570F75F95D" blockId="5.[129,1443,1672,1725]" pageId="5" pageNumber="1039">
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<emphasis id="B95BEAC4AF1DFFB999C4F9570CB0F941" bold="true" box="[129,216,1672,1697]" pageId="5" pageNumber="1039">Figure 4.</emphasis>
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UFRGS-PV-0386-P, the holotype of
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<taxonomicName id="4C2F4D55AF1DFFB99B73F9560E90F941" box="[566,760,1673,1697]" family="Anteosauridae" genus="Pampaphoneus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1039" phylum="Chordata" rank="species" species="biccai">
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<emphasis id="B95BEAC4AF1DFFB99B73F9560E90F941" box="[566,760,1673,1697]" italics="true" pageId="5" pageNumber="1039">Pampaphoneus biccai</emphasis>
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</taxonomicName>
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. A, right lateral view. B, less lateral view. C, palatal view. D, dorsal view. E, dorsal view of UFRGS-PV-249-P. F, lateral view of UFRGS-PV-0249-P.
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</paragraph>
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||
</caption>
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||
<subSubSection id="C335655DAF1DFFBA99C4F9320D2FFF06" lastPageId="6" lastPageNumber="1040" pageId="5" pageNumber="1039" type="diagnosis">
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||
<paragraph id="8B9036D6AF1DFFBA99C4F9320D2FFF06" blockId="5.[129,778,1773,1984]" lastBlockId="6.[113,763,144,231]" lastPageId="6" lastPageNumber="1040" pageId="5" pageNumber="1039">
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||
<emphasis id="B95BEAC4AF1DFFB999C4F9320E75F8E5" box="[129,541,1773,1797]" italics="true" pageId="5" pageNumber="1039">
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||
Revised diagnosis:
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<taxonomicName id="4C2F4D55AF1DFFB99801F9320E75F8E5" box="[324,541,1773,1797]" family="Anteosauridae" genus="Pampaphoneus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1039" phylum="Chordata" rank="species" species="biccai">Pampaphoneus biccai</taxonomicName>
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</emphasis>
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||
can be distinguished from all other anteosaurids by a squamosal (posterior) process of the jugal extending beyond the anteriormost margin of the temporal fenestra and by the presence of a poorly developed, elliptical boss at the angular. It can be distinguished from all other anteosaurids, except
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<taxonomicName id="4C2F4D55AF1DFFB998D5F8560E2FF841" authorityName="Kutorga" authorityYear="1838" box="[400,583,1929,1953]" family="Anteosauridae" genus="Syodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1039" phylum="Chordata" rank="species" species="biarmicum">
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<emphasis id="B95BEAC4AF1DFFB998D5F8560E2FF841" box="[400,583,1929,1953]" italics="true" pageId="5" pageNumber="1039">Syodon biarmicum</emphasis>
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</taxonomicName>
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, by the presence of nine upper postcanines, which bear mesial and distal serrations and have a bulbous shape. It can be distinguished from
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<taxonomicName id="4C2F4D55AF1DFFB99C3EF9330FCBF8C4" authorityName="Kutorga" authorityYear="1838" family="Anteosauridae" genus="Syodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1039" phylum="Chordata" rank="species" species="biarmicum">
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||
<emphasis id="B95BEAC4AF1DFFB99C3EF9330FCBF8C4" italics="true" pageId="5" pageNumber="1039">Syodon biarmicum</emphasis>
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</taxonomicName>
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by a crest that extends from the pineal boss until the orbital rim; the presence of a diastema separating the upper canine from the first postcanines; the presence of a marked depression on the posterior portion of the maxilla, postfrontal and the entire lacrimal (limited to the lacrimal in
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<taxonomicName id="4C2F4D55AF1DFFB99DB0F85609D8F841" authorityName="Kutorga" authorityYear="1838" box="[1269,1456,1929,1953]" family="Anteosauridae" genus="Syodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1039" phylum="Chordata" rank="species" species="biarmicum">
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||
<emphasis id="B95BEAC4AF1DFFB99DB0F85609D8F841" box="[1269,1456,1929,1953]" italics="true" pageId="5" pageNumber="1039">Syodon biarmicum</emphasis>
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</taxonomicName>
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||
); and in
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<taxonomicName id="4C2F4D55AF1DFFB99AC8F877080FF820" box="[909,1127,1960,1984]" family="Anteosauridae" genus="Pampaphoneus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="1039" phylum="Chordata" rank="species" species="biccai">
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<emphasis id="B95BEAC4AF1DFFB99AC8F877080FF820" box="[909,1127,1960,1984]" italics="true" pageId="5" pageNumber="1039">Pampaphoneus biccai</emphasis>
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</taxonomicName>
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the frontals do not divide the parietals. In addition,
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<taxonomicName id="4C2F4D55AF1EFFBA9815FF4F0E4AFF48" box="[336,546,144,168]" family="Anteosauridae" genus="Pampaphoneus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="1040" phylum="Chordata" rank="species" species="biccai">
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<emphasis id="B95BEAC4AF1EFFBA9815FF4F0E4AFF48" box="[336,546,144,168]" italics="true" pageId="6" pageNumber="1040">Pampaphoneus biccai</emphasis>
|
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</taxonomicName>
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, has a frontal depression anterior to the pineal boss that is deeper than that present in
|
||
<taxonomicName id="4C2F4D55AF1EFFBA99CEFF100D29FF07" authorityName="Kutorga" authorityYear="1838" box="[139,321,207,231]" family="Anteosauridae" genus="Syodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="1040" phylum="Chordata" rank="species" species="biarmicum">
|
||
<emphasis id="B95BEAC4AF1EFFBA99CEFF100D29FF07" box="[139,321,207,231]" italics="true" pageId="6" pageNumber="1040">Syodon biarmicum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |