171 lines
20 KiB
XML
171 lines
20 KiB
XML
<document id="F2B7DB66CAB52AA9FA60E9A958D1195D" ID-DOI="10.5281/zenodo.280969" ID-GBIF-Dataset="117fb243-8aa0-42f5-bbb0-3c6ceb076004" ID-ISSN="1175-5326" ID-Zenodo-Dep="280969" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460480130987" checkinUser="plazi" docAuthor="Bragagnolo, Cibele & Pinto-Da-Rocha, Ricardo" docDate="2012" docId="038687A2FFD4C26CFF3C5AABFC14FB42" docLanguage="en" docName="zt03308p048.pdf" docOrigin="Zootaxa 3308" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Promitobates trapista Bragagnolo & Pinto-Da-Rocha, 2012, sp. n." docType="treatment" docVersion="8" lastPageNumber="39" masterDocId="FFBFFFDAFFF1C24AFFAB5F78FFCEFF86" masterDocTitle="Systematic review of Promitobates Roewer, 1913 and cladistic analysis of Mitobatinae Simon, 1879 (Arachnida: Opiliones: Gonyleptidae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="38" updateTime="1698307495848" updateUser="plazi">
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<mods:titleInfo id="0F48C78389BA6B99F9445BE31AE20D57">
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<mods:title id="A4E9A818B87F692BD62E7F87ADB3CAF5">Systematic review of Promitobates Roewer, 1913 and cladistic analysis of Mitobatinae Simon, 1879 (Arachnida: Opiliones: Gonyleptidae)</mods:title>
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<mods:name id="ADDEEF3B94D4CFE63548A0D7090F9028" type="personal">
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<mods:roleTerm id="0B47F76986C7E2F48D86C94CBE977527">Author</mods:roleTerm>
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<mods:namePart id="01C4166BCC3DE698F09DC884882EC6FC">Bragagnolo, Cibele</mods:namePart>
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<mods:namePart id="ADA80ABDAB45B3E44F026FE73B9E7272">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:typeOfResource id="894574C5D0DB1FCAE566D4CA952E21B5">text</mods:typeOfResource>
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<mods:title id="919489ECD1C19773CE215DC1802D7D8A">Zootaxa</mods:title>
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<mods:part id="FD4957D4F92FDFA6EF41ECB65CC62290">
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<mods:date id="7009BC01C80CF54CD9393F5B778A4EA4">2012</mods:date>
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<mods:number id="91FF702216173D3A2CB1C8DC82D7B3EF">3308</mods:number>
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<mods:classification id="BAB4F8AC29551FDFDACC29B4C2741604">journal article</mods:classification>
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<mods:identifier id="3B50F050D78B653F0A05FB58899E2502" type="DOI">10.5281/zenodo.280969</mods:identifier>
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<mods:identifier id="9548EC93BB39C24ACFE5AEC2A7A2A4A4" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5FA896CB9498AAE00E738E5F39C9BB27" type="Zenodo-Dep">280969</mods:identifier>
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<treatment id="038687A2FFD4C26CFF3C5AABFC14FB42" ID-DOI="http://doi.org/10.5281/zenodo.6174045" ID-GBIF-Taxon="119517085" ID-Zenodo-Dep="6174045" LSID="urn:lsid:plazi:treatment:038687A2FFD4C26CFF3C5AABFC14FB42" httpUri="http://treatment.plazi.org/id/038687A2FFD4C26CFF3C5AABFC14FB42" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
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<subSubSection id="C335653FFFD4C26FFF3C5AABFD6CF9D5" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="8B9036B4FFD4C26FFF3C5AABFE2CFA6A" blockId="37.[151,492,1490,1550]" box="[151,482,1490,1517]" pageId="37" pageNumber="38">
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<heading id="D0D881D8FFD4C26FFF3C5AABFE2CFA6A" bold="true" box="[151,482,1490,1517]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
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<emphasis id="B95BEAA6FFD4C26FFF3C5AABFE2CFA6A" bold="true" box="[151,482,1490,1517]" pageId="37" pageNumber="38">
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<taxonomicName id="4C2F4D37FFD4C26FFF3C5AABFE56FA6B" ID-CoL="7YGX2" box="[151,408,1491,1517]" class="Arachnida" family="Gonyleptidae" genus="Promitobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="trapista" status="sp. nov.">
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<emphasis id="B95BEAA6FFD4C26FFF3C5AABFE56FA6B" bold="true" box="[151,408,1491,1517]" italics="true" pageId="37" pageNumber="38">Promitobates trapista</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26857DDFFD4C26FFE0B5AAAFE2CFA6A" box="[416,482,1490,1516]" pageId="37" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9036B4FFD4C26FFF3C5A8EFE22F988" blockId="37.[151,492,1490,1550]" box="[151,492,1526,1550]" pageId="37" pageNumber="38">
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||
(
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<figureCitation id="13142A31FFD4C26FFF345A8EFF23F988" box="[159,237,1526,1550]" captionStart="FIGURE 6" captionStartId="19.[151,250,1545,1568]" captionTargetBox="[191,1401,236,1473]" captionTargetId="figure@19.[151,1436,193,1524]" captionTargetPageId="19" captionText="FIGURE 6. Dorsal view of males of Promitobates A. P. a l e sp. n. (MZSP 27404); B. P. trapista sp. n. (MZSP 27406); C. P. wiessbier sp. n. (MZSP 16171); D. lager sp. n. (MNRJ 4545). Scale bar, 1 mm." httpUri="https://zenodo.org/record/280975/files/figure.png" pageId="37" pageNumber="38">Figs. 6</figureCitation>
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B, 8D, 10D, 18G, H, I)
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</paragraph>
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<paragraph id="8B9036B4FFD4C26FFF3C5946FD6CF9D5" blockId="37.[151,674,1597,1620]" box="[151,674,1597,1620]" pageId="37" pageNumber="38">
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<treatmentCitationGroup id="AB3F119AFFD4C26FFF3C5946FD6CF9D5" box="[151,674,1597,1620]" pageId="37" pageNumber="38">
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<taxonomicName id="4C2F4D37FFD4C26FFF3C5946FED1F9D2" ID-CoL="7PVRB" box="[151,287,1598,1620]" class="Arachnida" family="Gonyleptidae" genus="Promitobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="B95BEAA6FFD4C26FFF3C5946FED1F9D2" box="[151,287,1598,1620]" italics="true" pageId="37" pageNumber="38">Promitobates</emphasis>
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</taxonomicName>
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sp.:
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<treatmentCitation id="0A8E10A5FFD4C26FFEE45945FDA4F9D5" author="Bragagnolo" box="[335,618,1597,1620]" page="393" pageId="37" pageNumber="38" year="2007">
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<bibRefCitation id="EFBE4B45FFD4C26FFEE45945FDF4F9D5" author="Bragagnolo" box="[335,570,1597,1620]" pageId="37" pageNumber="38" refString="Bragagnolo, C., Pinto-da-Rocha, R., Nogueira, A. A., Pardini, R. (2007) Harvestmen in an Atlantic forest fragmented landscape: Evaluating assemblage response to habitat quality and quantitty. Biological Conservation, 139, 389 - 400." type="journal article" year="2007">
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Bragagnolo
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<emphasis id="B95BEAA6FFD4C26FFE605947FE30F9D2" box="[459,510,1598,1620]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
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2007
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</bibRefCitation>
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: 393
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</treatmentCitation>
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(cit).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C335653FFFD4C26FFF3C59F9FF20F966" pageId="37" pageNumber="38" type="materials_examined">
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<paragraph id="8B9036B4FFD4C26FFF3C59F9FF20F966" blockId="37.[151,1436,1664,2013]" pageId="37" pageNumber="38">
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<emphasis id="B95BEAA6FFD4C26FFF3C59F9FE8AF91F" bold="true" box="[151,324,1664,1689]" pageId="37" pageNumber="38">
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<typeStatus id="54948816FFD4C26FFF3C59F9FF1CF91F" box="[151,210,1665,1689]" pageId="37" pageNumber="38">Type</typeStatus>
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material.
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</emphasis>
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<collectingCountry id="F3387624FFD4C26FFEE059F9FE7AF91F" box="[331,436,1665,1689]" name="Brazil" pageId="37" pageNumber="38">BRAZIL</collectingCountry>
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.
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<emphasis id="B95BEAA6FFD4C26FFE1559F9FDF4F91E" box="[446,570,1664,1688]" italics="true" pageId="37" pageNumber="38">São Paulo:</emphasis>
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Cotia (Reserva Morro Grande),
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<date id="FF911074FFD4C26FFC0C59F8FBDEF91E" box="[935,1040,1664,1688]" pageId="37" pageNumber="38" value="2008-07">VII.2008</date>
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, R. Pinto-da-Rocha
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<emphasis id="B95BEAA6FFD4C26FFB5959F9FAE4F91E" box="[1266,1322,1664,1688]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
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leg;
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<typeStatus id="54948816FFD4C26FFACA59F8FF09F93B" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
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3 (
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<collectionCode id="ED3EAE71FFD4C26FFF5D59DCFE8FF93A" box="[246,321,1700,1724]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="37" pageNumber="38">MZSP</collectionCode>
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31197).
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<typeStatus id="54948816FFD4C26FFE0C59DDFDD9F93B" box="[423,535,1701,1725]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
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, same locality as
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<typeStatus id="54948816FFD4C26FFD4F59DCFC87F93A" box="[740,841,1700,1724]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
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,
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<date id="FF911074FFD4C26FFCFD59DCFC2FF93A" box="[854,993,1700,1724]" pageId="37" pageNumber="38" value="2002-12-15">15.XII.2002</date>
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, A.A. Nogueira
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<emphasis id="B95BEAA6FFD4C26FFB3459DEFB14F93B" box="[1183,1242,1701,1725]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
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leg. 2 3 (
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<collectionCode id="ED3EAE71FFD4C26FFAFA59DCFA52F93A" box="[1361,1436,1700,1724]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="37" pageNumber="38">MZSP</collectionCode>
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27406).
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</paragraph>
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</subSubSection>
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<subSubSection id="C335653FFFD4C26FFF6C5995FBD7F882" box="[199,1049,1772,1797]" pageId="37" pageNumber="38" type="etymology">
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<paragraph id="8B9036B4FFD4C26FFF6C5995FBD7F882" blockId="37.[151,1436,1664,2013]" box="[199,1049,1772,1797]" pageId="37" pageNumber="38">
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<emphasis id="B95BEAA6FFD4C26FFF6C5995FE80F883" bold="true" box="[199,334,1773,1797]" pageId="37" pageNumber="38">Etymology.</emphasis>
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<taxonomicName id="4C2F4D37FFD4C26FFEFF5994FE61F882" box="[340,431,1772,1796]" class="Arachnida" family="Gonyleptidae" genus="Promitobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="46" phylum="Arthropoda" rank="species" species="trapista" status="sp. nov.">Trapista</taxonomicName>
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is a kind of
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<taxonomicName id="4C2F4D37FFD4C26FFD915994FD95F882" box="[570,603,1772,1796]" class="Arachnida" family="Gonyleptidae" genus="Promitobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ale">ale</taxonomicName>
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beer, made by Belgian Trappist monks.
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</paragraph>
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</subSubSection>
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<subSubSection id="C335653FFFD4C26FFF6C5868FF15F812" pageId="37" pageNumber="38" type="diagnosis">
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<paragraph id="8B9036B4FFD4C26FFF6C5868FF15F812" blockId="37.[151,1436,1664,2013]" pageId="37" pageNumber="38">
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<emphasis id="B95BEAA6FFD4C26FFF6C5868FE8FF8AF" bold="true" box="[199,321,1808,1833]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
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It resembles
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<emphasis id="B95BEAA6FFD4C26FFE715869FDDBF8AE" box="[474,533,1808,1832]" italics="true" pageId="37" pageNumber="38">P. a l e</emphasis>
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<emphasis id="B95BEAA6FFD4C26FFD8B5869FDAFF8AF" bold="true" box="[544,609,1809,1833]" pageId="37" pageNumber="38">sp. n.</emphasis>
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by external apophysis of coxa IV short and single-branched; also resembles
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<taxonomicName id="4C2F4D37FFD4C26FFF67584EFEA1F8CB" box="[204,367,1845,1869]" class="Arachnida" family="Gonyleptidae" genus="Promitobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hatschbachi">
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<emphasis id="B95BEAA6FFD4C26FFF67584EFEA1F8CB" box="[204,367,1845,1869]" italics="true" pageId="37" pageNumber="38">P. hatschbachi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2F4D37FFD4C26FFE0C584EFDD7F8CB" box="[423,537,1846,1869]" class="Arachnida" family="Gonyleptidae" genus="Promitobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ornatus">
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<emphasis id="B95BEAA6FFD4C26FFE0C584EFDD7F8CB" box="[423,537,1846,1869]" italics="true" pageId="37" pageNumber="38">P. ornatus</emphasis>
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</taxonomicName>
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by the pattern of tubercles on dorsal scutum. It can be distinguished form them by coloration, with dorsal scutum and legs I–III dark-yellow and apice of coxa IV, trochanter IV and femur IV black.
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</paragraph>
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</subSubSection>
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<subSubSection id="C335653FFFD4C26CFF6C58D9FC14FB42" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
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<paragraph id="8B9036B4FFD4C26FFF6C58D9FA52F85A" blockId="37.[151,1436,1664,2013]" pageId="37" pageNumber="38">
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<emphasis id="B95BEAA6FFD4C26FFF6C58D9FE56F83F" bold="true" box="[199,408,1952,1977]" pageId="37" pageNumber="38">Male description.</emphasis>
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Measurements (
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<typeStatus id="54948816FFD4C26FFDF958D8FD74F83E" box="[594,698,1952,1976]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
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). Dorsal scutum; maximum width: 4.5; total length: 4.2; prosoma length: 1.8. Leg IV; length of external apophysis of coxa: 0.95; femur IV length: 37.6. Tarsal formula: 7; 15; 10; 8.
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</paragraph>
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<paragraph id="8B9036B4FFD7C26CFF6C5FE0FBC9FE2A" blockId="38.[151,1436,151,1221]" pageId="38" pageNumber="39">
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Dorsum (
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<figureCitation id="13142A31FFD7C26CFE995FEFFE4CFF29" box="[306,386,151,175]" captionStart="FIGURE 6" captionStartId="19.[151,250,1545,1568]" captionTargetBox="[191,1401,236,1473]" captionTargetId="figure@19.[151,1436,193,1524]" captionTargetPageId="19" captionText="FIGURE 6. Dorsal view of males of Promitobates A. P. a l e sp. n. (MZSP 27404); B. P. trapista sp. n. (MZSP 27406); C. P. wiessbier sp. n. (MZSP 16171); D. lager sp. n. (MNRJ 4545). Scale bar, 1 mm." httpUri="https://zenodo.org/record/280975/files/figure.png" pageId="38" pageNumber="39">Figs. 6</figureCitation>
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B, 8D). Anterior margin of prosoma with two or three tubercles on corners. Ocularium with two high apophyses and tubercles behind apohphysis. Prosoma with tubercles behind ocularium, arranged in “U”. Area I with a row of small tubercles up to the posterior groove and two enlarged median tubercles. Area II with a row of tubercles up to the posterior groove and small scattered tubercles. Areas III and IV fused, III with two high spiniform apophyses and tubercles on median region. Lateral margin of dorsal scutum with one row of tubercles which is placed from ocularium to area II. Posterior margin with two median spiniform tubercles and one row of rounded tubercles increasing in size to the corners. Free tergites I and II with small scattered tubercles and one enlarged pair of tubercles on the corners; III with a median spiniform tubercle.
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</paragraph>
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<paragraph id="8B9036B4FFD7C26CFF625ECFFC50FE49" blockId="38.[151,1436,151,1221]" box="[201,926,439,464]" pageId="38" pageNumber="39">Chelicera. Segment I with three setiferous tubercles; II unarmed.</paragraph>
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<paragraph id="8B9036B4FFD7C26CFF6C5EA4FBF0FD91" blockId="38.[151,1436,151,1221]" pageId="38" pageNumber="39">Pedipalp. Trochanter with a small ventral spine. Femur with a small ventral basal spine and one subapical, prolateral spine. Tibia setation: mesal IiIi; ectal IiIi. Tarsus setation: mesal IIi; ectal IIi.</paragraph>
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<paragraph id="8B9036B4FFD7C26CFF625D5DFDD2FD69" blockId="38.[151,1436,151,1221]" pageId="38" pageNumber="39">Venter. Coxa I with median row of six setiferous tubercles and two apical tubercles; trochanter I with median tubercle and few granules scattered. Coxa II with median row of eight small setiferous tubercles and two apical tubercles; trochanter II with median tubercle. Coxa III with small setiferous tubercles, distributed roughly in rows, the anterior and posterior one, increasing in size distally; trochanter III with median tubercle. Coxa IV with granules scattered and three tubercles on posterior margin; genital area unarmed; free sternites with row of tubercles; anal operculum with few granules.</paragraph>
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<paragraph id="8B9036B4FFD7C26CFF625D85FBA9FC22" blockId="38.[151,1436,151,1221]" pageId="38" pageNumber="39">
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Legs. (
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<figureCitation id="13142A31FFD7C26CFEBC5D84FEA7FC92" box="[279,361,764,788]" captionStart="FIGURE 10" captionStartId="28.[151,250,1938,1961]" captionTargetBox="[169,1414,502,1868]" captionTargetId="figure@28.[151,1436,461,1918]" captionTargetPageId="28" captionText="FIGURE 10. Dorsal view of male coxa, trochanter and femur IV of Promitobates A e B. P. viridigranulatus, A. MZSP 27401; B. MHNCI 6527; C. P. ale sp. n. (MZSP 27404); D. P. trapista sp. n. (MZSP 27406); E. P. weissbier sp. n. (MZSP 16171); F. P lager sp. n. (MNRJ 04545); Scale bar, 1 mm." httpUri="https://zenodo.org/record/280979/files/figure.png" pageId="38" pageNumber="39">Fig. 10</figureCitation>
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D). Coxa I–II with one anterior dorso basal apophysis and one posterior; coxa III with anterior dorso basal apophysis; coxa IV with high tubercles, external apical apophysis directed ventrally and with one basal rounded tubercle, two small internal spiniform apophysis. Trochanters I–III with small scattered tubercles. Trochanter IV with one dorsal apical and one dorso basal rounded apophyses and one small internal apical spiniform apophysis. Femur more than 10 times longer than dorsal scutum, straight and unarmed.
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</paragraph>
|
||
<paragraph id="8B9036B4FFD7C26CFF625CD7FEB7FC6A" blockId="38.[151,1436,151,1221]" pageId="38" pageNumber="39">Coloration. Dorsal scutum, chelicerae, pedipalps and legs I–III dark-yellow. Apex of coxa IV, trochanter IV and femur IV black.</paragraph>
|
||
<paragraph id="8B9036B4FFD7C26CFF625C8FFD3BFBFA" blockId="38.[151,1436,151,1221]" pageId="38" pageNumber="39">
|
||
Penis (
|
||
<figureCitation id="13142A31FFD7C26CFEB35C8FFEA4FB89" box="[280,362,1015,1039]" captionStart="FIGURE 18" captionStartId="36.[151,250,1803,1826]" captionTargetBox="[151,1436,193,1781]" captionTargetId="figure@36.[151,1436,193,1782]" captionTargetPageId="36" captionText="FIGURE 18. Distal part of penis of Promitobates. A – C. P. viridigranulatus (MZSP- 27401); A. dorsal view; B. lateral view; C. ventral view. D – F. P. ale sp. n. (MZSP- 18649); D. dorsal view; E. lateral view; F. ventral view. G – I. P. trapista sp. n. (MZSP- 27406); G. dorsal view; H. lateral view; I. ventral view. Scale bar = 0,05 mm." httpUri="https://zenodo.org/record/280987/files/figure.png" pageId="38" pageNumber="39">Fig. 18</figureCitation>
|
||
G–I). Ventral plate convex, apex straight and distal margin corner rounded and directed ventrally; three pairs of distal setae, one pair of median setae; three pairs of basal setae, arranged in “V” and one pair of ventral basal setae. Stylus unarmed, apex straight and swollen. Ventral process almost parallel to the stylus, apex spoon-shaped, with serrate margins directed ventrally.
|
||
</paragraph>
|
||
<paragraph id="8B9036B4FFD7C26CFF6C5BF0FE6CFB26" blockId="38.[151,1436,151,1221]" box="[199,418,1160,1184]" pageId="38" pageNumber="39">
|
||
<emphasis id="B95BEAA6FFD7C26CFF6C5BF0FED0FB26" bold="true" box="[199,286,1160,1184]" pageId="38" pageNumber="39">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="8B9036B4FFD7C26CFF6C5BD3FC14FB42" blockId="38.[151,1436,151,1221]" box="[199,986,1195,1221]" pageId="38" pageNumber="39">
|
||
<emphasis id="B95BEAA6FFD7C26CFF6C5BD3FE33FB42" bold="true" box="[199,509,1195,1220]" pageId="38" pageNumber="39">Geographical distribution</emphasis>
|
||
(
|
||
<figureCitation id="13142A31FFD7C26CFDA65BD4FD81FB42" box="[525,591,1196,1220]" captionStart="FIGURE 3" captionStartId="13.[151,250,2007,2030]" captionTargetBox="[260,1316,193,1957]" captionTargetId="figure@13.[260,1317,193,1958]" captionTargetPageId="13" captionText="FIGURE 3. Geographical distribution of Promitobates spp .." httpUri="https://zenodo.org/record/280972/files/figure.png" pageId="38" pageNumber="39">Fig. 3</figureCitation>
|
||
B). Only known from
|
||
<typeStatus id="54948816FFD7C26CFCE35BD5FCB6FB43" box="[840,888,1197,1221]" pageId="38" pageNumber="39">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |