379 lines
41 KiB
XML
379 lines
41 KiB
XML
<document ID-DOI="10.11646/zootaxa.5277.3.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="7890052" ID-ZooBank="749AFFB5-4A38-4409-9FB9-32263F3F6E69" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683101025327" checkinUser="plazi" docAuthor="Bezděk, Jan" docDate="2023" docId="DB6D3549CE56DF7676891BC6FD903662" docLanguage="en" docName="zootaxa.5277.3.10.pdf" docOrigin="Zootaxa 5277 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.3.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hesperopenna gilolo Bezdek 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="583" masterDocId="27544D31CE57DF75761E1B51FFB3377F" masterDocTitle="Hesperopenna gilolo sp. nov., the easternmost record of the genus from Halmahera Indonesia (Coleoptera: Chrysomelidae: Galerucinae)" masterLastPageNumber="584" masterPageNumber="581" pageNumber="582" updateTime="1683101457609" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Hesperopenna gilolo sp. nov., the easternmost record of the genus from Halmahera Indonesia (Coleoptera: Chrysomelidae: Galerucinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bezděk, Jan</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5277</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>581</mods:start>
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<mods:end>584</mods:end>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.3.10</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5277.3.10</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7890052</mods:identifier>
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<mods:identifier type="ZooBank">749AFFB5-4A38-4409-9FB9-32263F3F6E69</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7890038" ID-Zenodo-Dep="7890038" LSID="urn:lsid:plazi:treatment:DB6D3549CE56DF7676891BC6FD903662" httpUri="http://treatment.plazi.org/id/DB6D3549CE56DF7676891BC6FD903662" lastPageId="3" lastPageNumber="583" pageId="1" pageNumber="582">
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<subSubSection box="[151,610,151,178]" pageId="1" pageNumber="582" type="nomenclature">
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<paragraph blockId="1.[151,1086,151,243]" box="[151,610,151,178]" pageId="1" pageNumber="582">
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<heading bold="true" box="[151,610,151,178]" fontSize="11" level="1" pageId="1" pageNumber="582" reason="1">
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<emphasis bold="true" box="[151,610,151,178]" pageId="1" pageNumber="582">
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<taxonomicName authority="Bezdek" authorityName="Bezdek" authorityYear="2023" box="[151,503,151,178]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="gilolo" status="sp. nov.">
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<emphasis bold="true" box="[151,403,151,178]" italics="true" pageId="1" pageNumber="582">Hesperopenna gilolo</emphasis>
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Bezděk
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</taxonomicName>
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,
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<taxonomicNameLabel box="[516,610,151,177]" pageId="1" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="582" type="description">
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<paragraph blockId="1.[151,1086,151,243]" box="[151,267,187,213]" pageId="1" pageNumber="582">
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(
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<figureCitation box="[159,258,187,213]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1570,1594]" captionTargetBox="[157,1433,238,1541]" captionTargetId="figure-47@2.[151,1436,234,1544]" captionTargetPageId="2" captionText="FIGURES 1–8. Hesperopenna gilolo Bezděk, sp. nov. 1—holotype in dorsal view; 2—labels; 3—head; 4—penis in dorsal view; 5—penis in lateral view; 6—penis in ventral view; 7—pronotum; 8—apex of abdomen." figureDoi="http://doi.org/10.5281/zenodo.7890054" httpUri="https://zenodo.org/record/7890054/files/figure.png" pageId="1" pageNumber="582">Figs 1–8</figureCitation>
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)
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</paragraph>
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<paragraph blockId="1.[151,1086,151,243]" box="[151,1086,219,243]" pageId="1" pageNumber="582">
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http://zoobank.org/
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<uri box="[348,1086,219,243]" pageId="1" pageNumber="582">
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urn:lsid:zoobank.org:act:
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<uuid box="[591,1086,219,243]" pageId="1" pageNumber="582">EB0045AF-E24D-4A8D-A107-13003343CCF2</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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<subSubSection pageId="1" pageNumber="582" type="materials_examined">
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<paragraph blockId="1.[151,1437,289,2008]" box="[151,1253,289,317]" pageId="1" pageNumber="582">
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<materialsCitation box="[151,1249,289,317]" country="Indonesia" county="Kampung Pasir Putih" latitude="0.8833333" location="Halmahera Isl." longLatPrecision="1308" longitude="127.683334" municipality="Jailolo Dist." pageId="1" pageNumber="582" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[151,310,289,316]" pageId="1" pageNumber="582">
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<typeStatus box="[151,210,290,316]" pageId="1" pageNumber="582">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry box="[316,428,290,316]" name="Indonesia" pageId="1" pageNumber="582">Indonesia</collectingCountry>
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,
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<location LSID="urn:lsid:plazi:treatment:DB6D3549CE56DF7676891BC6FD903662:561BD284CE56DF7477A91A73FDED3643" box="[439,606,290,316]" country="Indonesia" county="Kampung Pasir Putih" latitude="0.8833333" longLatPrecision="1308" longitude="127.683334" municipality="Jailolo Dist." name="Halmahera Isl." pageId="1" pageNumber="582">Halmahera Isl.</location>
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,
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<collectingMunicipality box="[618,756,290,316]" pageId="1" pageNumber="582">Jailolo Dist.</collectingMunicipality>
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,
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<collectingCounty box="[767,1010,290,316]" pageId="1" pageNumber="582">Kampung Pasir Putih</collectingCounty>
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, at
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<geoCoordinate box="[1048,1130,290,317]" degrees="0" direction="north" minutes="53" orientation="latitude" pageId="1" pageNumber="582" precision="925" value="0.8833333">0°53´N</geoCoordinate>
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<geoCoordinate box="[1138,1249,290,316]" degrees="127" direction="east" minutes="41" orientation="longitude" pageId="1" pageNumber="582" precision="925" value="127.683334">127°41´E</geoCoordinate>
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
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<emphasis bold="true" box="[199,374,325,352]" pageId="1" pageNumber="582">Type material.</emphasis>
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<materialsCitation collectingDate="1981-01-01" collectingDateMax="1981-01-14" collectingDateMin="1981-01-01" collectionCode="USNM, AC, PM" collectorName="The" country="Indonesia" county="Kampung Pasir Putih" determinerName="J. Bezdek" latitude="0.8833333" location="Halmahera" longLatPrecision="1308" longitude="127.683334" municipality="Jailolo Dist." pageId="1" pageNumber="582" specimenCount="3" typeStatus="holotype">
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<typeStatus box="[383,492,326,352]" pageId="1" pageNumber="582">Holotype</typeStatus>
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: ³ (
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<collectionCode box="[540,627,326,352]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="582" type="Museum">USNM</collectionCode>
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), “
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<collectingCountry box="[657,816,326,352]" name="Indonesia" pageId="1" pageNumber="582">INDONESIA</collectingCountry>
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||
:
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||
<location LSID="urn:lsid:plazi:treatment:DB6D3549CE56DF7676891BC6FD903662:561BD284CE56DF7475221A17FC04361F" box="[828,951,326,352]" country="Indonesia" county="Kampung Pasir Putih" latitude="0.8833333" longLatPrecision="1308" longitude="127.683334" municipality="Jailolo Dist." name="Halmahera" pageId="1" pageNumber="582">Halmahera</location>
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/ Isl.,
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<collectingMunicipality box="[1026,1165,326,352]" pageId="1" pageNumber="582">Jailolo Dist.</collectingMunicipality>
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, /
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<collectingCounty box="[1194,1436,326,352]" pageId="1" pageNumber="582">Kampung Pasir Putih</collectingCounty>
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/
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<geoCoordinate box="[167,252,362,389]" degrees="0" direction="north" minutes="53" orientation="latitude" pageId="1" pageNumber="582" precision="925" value="0.8833333">0°53´N</geoCoordinate>
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,
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||
<geoCoordinate box="[265,373,362,388]" degrees="127" direction="east" minutes="41" orientation="longitude" pageId="1" pageNumber="582" precision="925" value="127.683334">127°41´E</geoCoordinate>
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[w, p] //
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<date box="[478,646,362,388]" pageId="1" pageNumber="582" value="1981-01-01" valueMax="1981-01-14" valueMin="1981-01-01">
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<collectingDate box="[478,646,362,388]" pageId="1" pageNumber="582" value="1981-01-01" valueMax="1981-01-14" valueMin="1981-01-01">1-14 Jan. 1981</collectingDate>
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</date>
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/
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<collectionCode box="[668,707,362,388]" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="1" pageNumber="582" type="University or college">AC</collectionCode>
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Messer &
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<collectionCode box="[834,874,362,388]" country="USA" name="Pratt Museum" pageId="1" pageNumber="582">PM</collectionCode>
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Taylor [w, p] //
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<typeStatus box="[1061,1230,362,388]" pageId="1" pageNumber="582">HOLOTYPUS</typeStatus>
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, /
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<taxonomicName box="[1258,1420,363,388]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[1258,1420,363,388]" italics="true" pageId="1" pageNumber="582">Hesperopenna</emphasis>
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</taxonomicName>
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||
/
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||
<taxonomicName authority="Bezděk, 2023" authorityName="Bezděk" authorityYear="2023" box="[151,216,398,424]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="gilolo" status="sp. nov.">
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<emphasis box="[151,216,398,424]" italics="true" pageId="1" pageNumber="582">gilolo</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel box="[224,313,398,424]" pageId="1" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
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||
, /
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||
<determinerName box="[338,447,398,424]" pageId="1" pageNumber="582">J. Bezděk</determinerName>
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||
det. 2023 [r, p]”.
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||
<collectorName box="[646,690,398,424]" pageId="1" pageNumber="582">The</collectorName>
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<typeStatus box="[697,795,398,424]" pageId="1" pageNumber="582">holotype</typeStatus>
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is lacking the hind right leg.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="1" pageNumber="582" type="description">
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<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
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<emphasis bold="true" box="[199,337,433,459]" pageId="1" pageNumber="582">Description</emphasis>
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||
. Body length (male,
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<typeStatus box="[583,687,434,460]" pageId="1" pageNumber="582">holotype</typeStatus>
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): 3.0 mm. Body elongate oval, moderately convex, and glabrous. Head, pronotum, ventral side, and legs orange, apices of mandibles black, maxillary palpi brownish black, antennae black with orange basal half of antennomere I and brown antennomeres II and III, scutellum brown, elytra black.
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</paragraph>
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<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
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Male (
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<typeStatus box="[270,372,542,568]" pageId="1" pageNumber="582">holotype</typeStatus>
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,
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<figureCitation box="[381,480,542,568]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1570,1594]" captionTargetBox="[157,1433,238,1541]" captionTargetId="figure-47@2.[151,1436,234,1544]" captionTargetPageId="2" captionText="FIGURES 1–8. Hesperopenna gilolo Bezděk, sp. nov. 1—holotype in dorsal view; 2—labels; 3—head; 4—penis in dorsal view; 5—penis in lateral view; 6—penis in ventral view; 7—pronotum; 8—apex of abdomen." figureDoi="http://doi.org/10.5281/zenodo.7890054" httpUri="https://zenodo.org/record/7890054/files/figure.png" pageId="1" pageNumber="582">Figs 1–2</figureCitation>
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). Head (
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<figureCitation box="[571,636,542,568]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1570,1594]" captionTargetBox="[157,1433,238,1541]" captionTargetId="figure-47@2.[151,1436,234,1544]" captionTargetPageId="2" captionText="FIGURES 1–8. Hesperopenna gilolo Bezděk, sp. nov. 1—holotype in dorsal view; 2—labels; 3—head; 4—penis in dorsal view; 5—penis in lateral view; 6—penis in ventral view; 7—pronotum; 8—apex of abdomen." figureDoi="http://doi.org/10.5281/zenodo.7890054" httpUri="https://zenodo.org/record/7890054/files/figure.png" pageId="1" pageNumber="582">Fig. 3</figureCitation>
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) with transversely oval labrum, anterior margin straight, widely rounded anterior angles, and surface with six pores in transverse row, each bearing long, pale seta. Anterior part of head slightly convex and glabrous, with several setae along anterior margin and close to eyes, anterior margin slightly concave. Interantennal space narrow, 0.62 times as wide as transverse diameter of antennal socket. Interocular space 1.36 times as wide as transverse diameter of eye. Frontal tubercles transversely subtriangular, moderately elevated, semi-opaque, and separated by thin, shallow groove. Vertex glabrous, semi-opaque, impunctate, separated from frontal tubercles by shallow bent line. Antennae filiform, as long as body, length ratios of antennomeres in sequence from first equals 100-20-30-70-70-60-60-60-60-50-50 (100 =
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<quantity box="[1018,1112,794,821]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="582" unit="mm" value="0.5">0.5 mm</quantity>
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). Antennomeres I–II almost glabrous, III–XI densely covered by short recumbent setae. Penultimate maxillary palpomere moderately enlarged, ultimate palpomere conical.
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</paragraph>
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<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
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Pronotum (
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<figureCitation box="[327,397,902,928]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1570,1594]" captionTargetBox="[157,1433,238,1541]" captionTargetId="figure-47@2.[151,1436,234,1544]" captionTargetPageId="2" captionText="FIGURES 1–8. Hesperopenna gilolo Bezděk, sp. nov. 1—holotype in dorsal view; 2—labels; 3—head; 4—penis in dorsal view; 5—penis in lateral view; 6—penis in ventral view; 7—pronotum; 8—apex of abdomen." figureDoi="http://doi.org/10.5281/zenodo.7890054" httpUri="https://zenodo.org/record/7890054/files/figure.png" pageId="1" pageNumber="582">Fig. 7</figureCitation>
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) transverse, 1.68 times as wide as long, semi-opaque, glabrous, widest in anterior quarter, and covered with microsculpture and fine punctures. Surface moderately convex, with distinct short transverse impressions from anterior angles parallel with anterior margin. Anterior margin slightly concave, lateral margins rounded, slightly posteriorly, posterior margin moderately rounded.Anterior margin unbordered, lateral and posterior margins distinctly bordered. Anterior angles rounded, slightly swollen, posterior angles obtusangulate, each angle with setigerous pore bearing long seta. Scutellum small, triangular, impunctate, and glabrous.
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</paragraph>
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<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">Elytra 1.27 times as long as wide (measured at widest, in posterior third) and 0.72 times as long as body. Surface with slight oil sheen, glabrous, densely covered with small, confused punctures. Humeral calli developed. Epipleura lustrous, glabrous, smooth, widest at anterior third, gradually narrowed towards elytral apex. Macropterous.</paragraph>
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<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
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Procoxal cavities opened behind. Abdomen sparsely covered with setae (somewhat denser towards abdominal apex), posterior margin of last abdominal ventrite widely shallowly emarginated, with fringe of long setae (
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<figureCitation box="[1354,1422,1262,1288]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1570,1594]" captionTargetBox="[157,1433,238,1541]" captionTargetId="figure-47@2.[151,1436,234,1544]" captionTargetPageId="2" captionText="FIGURES 1–8. Hesperopenna gilolo Bezděk, sp. nov. 1—holotype in dorsal view; 2—labels; 3—head; 4—penis in dorsal view; 5—penis in lateral view; 6—penis in ventral view; 7—pronotum; 8—apex of abdomen." figureDoi="http://doi.org/10.5281/zenodo.7890054" httpUri="https://zenodo.org/record/7890054/files/figure.png" pageId="1" pageNumber="582">Fig. 8</figureCitation>
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).
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</paragraph>
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<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
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All legs densely covered with short recumbent setae. Apices of meso- and metatibiae with spine. Protarsomere I narrow, subparallel, slightly wider than small and triangular protarsomere II, length ratio of protarsomeres I–III and V equals 100-50-50-86 (100 =
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||
<quantity box="[490,592,1370,1396]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="582" unit="mm" value="0.2">0.20 mm</quantity>
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||
). Mesotarsomere I elongated triangular, as wide as triangular mesotarsomere II, length ratio of mesotarsomeres I–III and V equals 100-50-50-86 (100 =
|
||
<quantity box="[978,1081,1406,1432]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="582" unit="mm" value="0.2">0.20 mm</quantity>
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). Metatarsomere I long, narrow, length ratio of metatarsomeres I–III and V equals 100-25-25-44 (100 =
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<quantity box="[952,1042,1442,1468]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="582" unit="mm" value="0.4">0.4 mm</quantity>
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). Claws appendiculate.
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</paragraph>
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<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
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Penis (
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<figureCitation box="[280,384,1478,1504]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1570,1594]" captionTargetBox="[157,1433,238,1541]" captionTargetId="figure-47@2.[151,1436,234,1544]" captionTargetPageId="2" captionText="FIGURES 1–8. Hesperopenna gilolo Bezděk, sp. nov. 1—holotype in dorsal view; 2—labels; 3—head; 4—penis in dorsal view; 5—penis in lateral view; 6—penis in ventral view; 7—pronotum; 8—apex of abdomen." figureDoi="http://doi.org/10.5281/zenodo.7890054" httpUri="https://zenodo.org/record/7890054/files/figure.png" pageId="1" pageNumber="582">Figs 4–6</figureCitation>
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) elongate, parallel, 6.50 times as long as wide, apex rounded, surface with shallow oval impression in anterior third. In lateral view, penis wide, slightly bent. Ventral side with long, wide and shallow groove, parallel in apical half, distinctly wider in middle part. Penis with one long thin internal sclerite wrapped around apex, with tip directed backward on ventral side of penis.
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</paragraph>
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<paragraph blockId="1.[151,1437,289,2008]" box="[199,401,1622,1648]" pageId="1" pageNumber="582">Female unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="582" type="diagnosis">
|
||
<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
|
||
<emphasis bold="true" box="[199,459,1657,1683]" pageId="1" pageNumber="582">Differential diagnosis.</emphasis>
|
||
<taxonomicName authority="Bezděk, 2023" authorityName="Bezděk" authorityYear="2023" box="[470,703,1658,1684]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="gilolo" status="sp. nov.">
|
||
<emphasis box="[470,703,1658,1684]" italics="true" pageId="1" pageNumber="582">Hesperopenna gilolo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[715,804,1658,1684]" pageId="1" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is characterized by completely black elytra. The black coloration of the dorsal side is known only in several species: the head in
|
||
<taxonomicName authority="(Kimoto, 2004)" baseAuthorityName="Kimoto" baseAuthorityYear="2004" box="[925,1229,1694,1720]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="nigriceps">
|
||
<emphasis box="[925,1057,1695,1720]" italics="true" pageId="1" pageNumber="582">H. nigriceps</emphasis>
|
||
(Kimoto, 2004)
|
||
</taxonomicName>
|
||
from eastern
|
||
<collectingCountry box="[1374,1433,1694,1720]" name="India" pageId="1" pageNumber="582">India</collectingCountry>
|
||
, the head and pronotum in
|
||
<taxonomicName authority="(Kimoto, 1989)" baseAuthorityName="Kimoto" baseAuthorityYear="1989" box="[430,748,1730,1756]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="nigricollis">
|
||
<emphasis box="[430,574,1730,1756]" italics="true" pageId="1" pageNumber="582">H. nigricollis</emphasis>
|
||
(Kimoto, 1989)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[811,910,1730,1756]" name="Thailand" pageId="1" pageNumber="582">Thailand</collectingCountry>
|
||
, the head and pronotum in some specimens of
|
||
<taxonomicName authority="(Medvedev, 2013)" baseAuthorityName="Medvedev" baseAuthorityYear="2013" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="bacboensis">
|
||
<emphasis italics="true" pageId="1" pageNumber="582">H. bacboensis</emphasis>
|
||
(Medvedev, 2013)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[543,638,1766,1792]" name="Vietnam" pageId="1" pageNumber="582">Vietnam</collectingCountry>
|
||
, and, finally, the head and scutellum in some specimens of
|
||
<taxonomicName authority="(Kimoto, 1989)" baseAuthorityName="Kimoto" baseAuthorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="thailandica">
|
||
<emphasis box="[1279,1436,1766,1792]" italics="true" pageId="1" pageNumber="582">H. thailandica</emphasis>
|
||
(Kimoto, 1989)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[390,489,1802,1828]" name="Thailand" pageId="1" pageNumber="582">Thailand</collectingCountry>
|
||
,
|
||
<collectingCountry box="[502,555,1802,1828]" name="Laos" pageId="1" pageNumber="582">Laos</collectingCountry>
|
||
and
|
||
<collectingCountry box="[611,676,1802,1828]" name="China" pageId="1" pageNumber="582">China</collectingCountry>
|
||
(
|
||
<collectingRegion box="[691,781,1802,1828]" country="China" name="Yunnan" pageId="1" pageNumber="582">Yunnan</collectingRegion>
|
||
). The elytra of all known species are completely pale, or at most, with a black extreme lateral margin in the basal half in
|
||
<taxonomicName authority="Bezdek, 2013" authorityName="Bezdek" authorityYear="2013" box="[817,1063,1838,1864]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="zofka">
|
||
<emphasis box="[817,907,1838,1864]" italics="true" pageId="1" pageNumber="582">H. zofka</emphasis>
|
||
<bibRefCitation author="Bezdek, J." box="[914,1063,1838,1864]" pageId="1" pageNumber="582" pagination="715 - 746" refId="ref2114" refString="Bezdek, J. (2013) Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae, 53, 715 - 746." type="journal article" year="2013">Bezděk, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[1130,1236,1838,1864]" name="Indonesia" pageId="1" pageNumber="582">Indonesia</collectingCountry>
|
||
(
|
||
<collectingRegion box="[1250,1302,1838,1864]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="582">Java</collectingRegion>
|
||
,
|
||
<collectingRegion box="[1313,1362,1838,1864]" country="Indonesia" name="Bali" pageId="1" pageNumber="582">Bali</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="582">
|
||
The penis of
|
||
<taxonomicName box="[358,613,1874,1900]" pageId="1" pageNumber="582">
|
||
<emphasis box="[358,520,1875,1900]" italics="true" pageId="1" pageNumber="582">Hesperopenna</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
is very variable in shape, from relatively simple to very complicated, always with a large, elongated basal orifice and internally with one or two large and long sclerites. The penis of
|
||
<taxonomicName authority="Bezděk, 2023" authorityName="Bezděk" authorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="gilolo" status="sp. nov.">
|
||
<emphasis italics="true" pageId="1" pageNumber="582">H. gilolo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[223,310,1946,1972]" pageId="1" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation box="[324,424,1946,1972]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1570,1594]" captionTargetBox="[157,1433,238,1541]" captionTargetId="figure-47@2.[151,1436,234,1544]" captionTargetPageId="2" captionText="FIGURES 1–8. Hesperopenna gilolo Bezděk, sp. nov. 1—holotype in dorsal view; 2—labels; 3—head; 4—penis in dorsal view; 5—penis in lateral view; 6—penis in ventral view; 7—pronotum; 8—apex of abdomen." figureDoi="http://doi.org/10.5281/zenodo.7890054" httpUri="https://zenodo.org/record/7890054/files/figure.png" pageId="1" pageNumber="582">Figs 4–6</figureCitation>
|
||
) is of relatively simple structure, sub-tubular, with one long, thin internal sclerite wrapped around the apex, with the tip directed backward on the ventral side of the penis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,624,150,176]" pageId="2" pageNumber="583" type="distribution">
|
||
<paragraph blockId="2.[199,1027,150,213]" box="[199,624,150,176]" pageId="2" pageNumber="583">
|
||
<emphasis bold="true" box="[199,353,150,176]" pageId="2" pageNumber="583">Distribution.</emphasis>
|
||
<collectingCountry box="[359,468,150,176]" name="Indonesia" pageId="2" pageNumber="583">Indonesia</collectingCountry>
|
||
(Halmahera).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1027,186,213]" pageId="2" pageNumber="583" type="etymology">
|
||
<paragraph blockId="2.[199,1027,150,213]" box="[199,1027,186,213]" pageId="2" pageNumber="583">
|
||
<emphasis bold="true" box="[199,334,186,212]" pageId="2" pageNumber="583">Etymology.</emphasis>
|
||
Gilolo is the former name of Halmahera. Noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7890054" ID-Zenodo-Dep="7890054" httpUri="https://zenodo.org/record/7890054/files/figure.png" pageId="2" pageNumber="583" startId="2.[151,264,1570,1594]" targetBox="[157,1433,238,1541]" targetPageId="2" targetType="figure">
|
||
<paragraph blockId="2.[151,1436,1570,1630]" pageId="2" pageNumber="583">
|
||
<emphasis bold="true" box="[151,317,1570,1594]" pageId="2" pageNumber="583">FIGURES 1–8.</emphasis>
|
||
<taxonomicName authority="Bezdek" authorityName="Bezdek" authorityYear="2023" box="[326,629,1570,1594]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="gilolo" status="sp. nov.">
|
||
<emphasis box="[326,541,1570,1594]" italics="true" pageId="2" pageNumber="583">Hesperopenna gilolo</emphasis>
|
||
Bezděk
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[641,722,1571,1594]" pageId="2" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
1—holotype in dorsal view; 2—labels; 3—head; 4—penis in dorsal view; 5—penis in lateral view; 6—penis in ventral view; 7—pronotum; 8—apex of abdomen.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="3" lastPageNumber="584" pageId="2" pageNumber="583" type="discussion">
|
||
<paragraph blockId="2.[151,1437,1676,2027]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="584" pageId="2" pageNumber="583">
|
||
<emphasis bold="true" box="[199,332,1676,1702]" pageId="2" pageNumber="583">Discussion.</emphasis>
|
||
For a long time, the known occurrence of the genus
|
||
<taxonomicName box="[932,1094,1677,1702]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[932,1094,1677,1702]" italics="true" pageId="2" pageNumber="583">Hesperopenna</emphasis>
|
||
</taxonomicName>
|
||
was limited to the continental part of Southeast Asia, with a few species described from Sumatra, Mentawai, Java, and Bali (
|
||
<bibRefCitation author="Bezdek, J." box="[1191,1341,1712,1738]" pageId="2" pageNumber="583" pagination="715 - 746" refId="ref2114" refString="Bezdek, J. (2013) Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group. Acta Entomologica Musei Nationalis Pragae, 53, 715 - 746." type="journal article" year="2013">Bezděk 2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Bezdek, J." box="[1351,1425,1712,1738]" pageId="2" pageNumber="583" pagination="7 - 27" refId="ref2217" refString="Bezdek, J. (2016 a) Revision of the genus Hesperopenna Medvedev et Dang, 1981 (Coleoptera: Chrysomelidae: Galerucinae). II. H. vietnamica species group and new taxonomical changes. Studies and Reports, Taxonomical Series, 12, 7 - 27." type="journal article" year="2016">2016a</bibRefCitation>
|
||
). During the extensive study of undetermined material from various institutions in recent years, the occurrence of
|
||
<taxonomicName box="[151,313,1785,1810]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[151,313,1785,1810]" italics="true" pageId="2" pageNumber="583">Hesperopenna</emphasis>
|
||
</taxonomicName>
|
||
in Borneo and the
|
||
<collectingCountry box="[518,644,1784,1810]" name="Philippines" pageId="2" pageNumber="583">Philippines</collectingCountry>
|
||
(several species awaiting descriptions) has been proven. The expanding knowledge of the distribution almost exactly copies that of the genus
|
||
<taxonomicName authorityName="Baly" authorityYear="1878" box="[938,1036,1820,1846]" class="Insecta" family="Chrysomelidae" genus="Charaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[938,1036,1820,1846]" italics="true" pageId="2" pageNumber="583">Charaea</emphasis>
|
||
</taxonomicName>
|
||
, which was also recently proven to occur on the Greater Sunda Islands (Bezděk in prep.) and the
|
||
<collectingCountry box="[831,957,1856,1882]" name="Philippines" pageId="2" pageNumber="583">Philippines</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Bezdek, J." box="[971,1124,1856,1883]" pageId="2" pageNumber="583" pagination="371 - 375" refId="ref2164" refString="Bezdek, J. (2015) Charaea luzonicum sp. nov. (Coleoptera: Chrysomelidae: Galerucinae): the first record of Charaea in the Philippines. Revue Suisse de Zoologie, 122, 371 - 375. https: // doi. org / 10.5281 / zenodo. 30005" type="journal article" year="2015">Bezděk 2015</bibRefCitation>
|
||
). Some oriental species-rich genera, such as
|
||
<taxonomicName authority="Chevrolat, 1836" authorityName="Chevrolat" authorityYear="1836" box="[334,674,1892,1918]" class="Insecta" family="Chrysomelidae" genus="Aulacophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[334,480,1892,1918]" italics="true" pageId="2" pageNumber="583">Aulacophora</emphasis>
|
||
Chevrolat, 1836
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Laboissiere, 1933" authorityName="Laboissiere" authorityYear="1933" box="[687,1069,1892,1918]" class="Insecta" family="Chrysomelidae" genus="Hoplosaenidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[687,857,1892,1918]" italics="true" pageId="2" pageNumber="583">Hoplosaenidea</emphasis>
|
||
Laboissière, 1933
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Weber, 1801" authorityName="Weber" authorityYear="1801" box="[1082,1301,1892,1918]" class="Insecta" family="Chrysomelidae" genus="Oides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1082,1147,1892,1918]" italics="true" pageId="2" pageNumber="583">Oides</emphasis>
|
||
Weber, 1801
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Chevrolat, 1836" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1315,1436,1892,1918]" italics="true" pageId="2" pageNumber="583">Monolepta</emphasis>
|
||
Chevrolat, 1836
|
||
</taxonomicName>
|
||
, occur also in the Australian Region (e.g.,
|
||
<bibRefCitation author="Reid, C. A. M." box="[812,931,1928,1954]" pageId="2" pageNumber="583" pagination="40 - 45" refId="ref2670" refString="Reid, C. A. M. (2003) Recognition of the genus Hoplosaenidea Laboissiere in Australia, with a key to the Australian genera of Galerucini (Coleoptera: Chrysomelidae: Galerucinae). Australian Journal of Entomology, 42, 40 - 45" type="journal article" year="2003">Reid 2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Bezdek, J." box="[942,1105,1928,1954]" pageId="2" pageNumber="583" pagination="504 - 524" refId="ref2268" refString="Bezdek, J. (2016 b) Revision of the genus Coeligetes from Malaysia and Indonesia, and description of Coeligetoides gen. nov. (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 4085 (4), 504 - 524. https: // doi. org / 10.11646 / zootaxa. 4085.4.3" type="journal article" year="2016">Bezděk 2016b</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Lee, Ch. - F. & Beenen, R." box="[1117,1345,1928,1955]" pageId="2" pageNumber="583" pagination="1 - 125" refId="ref2418" refString="Lee, Ch. - F. & Beenen, R. (2017) Revision of the Palaearctic and Oriental species of the genus Oides Weber, 1801 (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 4346 (1), 1 - 125. https: // doi. org / 10.11646 / zootaxa. 4346.1.1" type="journal article" year="2017">Lee & Beenen 2017</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Reid, C. A. M. & Halling, L. & Beatson, M." pageId="2" pageNumber="583" pagination="1 - 73" refId="ref2717" refString="Reid, C. A. M., Halling, L. & Beatson, M. (2021) Revision of the Australopapuan and West Pacific species of plain pumpkinbeetles, the Aulacophora indica species-complex (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 4932 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4932.1.1" type="journal article" year="2021">
|
||
Reid
|
||
<emphasis italics="true" pageId="2" pageNumber="583">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
); however, the distribution of
|
||
<taxonomicName box="[581,743,1965,1990]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[581,743,1965,1990]" italics="true" pageId="2" pageNumber="583">Hesperopenna</emphasis>
|
||
</taxonomicName>
|
||
, covering continental Asia, the Greater Sunda Islands, and the
|
||
<collectingCountry box="[151,280,2000,2026]" name="Philippines" pageId="2" pageNumber="583">Philippines</collectingCountry>
|
||
, corresponds rather to moderately numerous genera of
|
||
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[887,1025,2000,2026]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="584" phylum="Arthropoda" rank="subFamily" subFamily="Galerucinae">Galerucinae</taxonomicName>
|
||
, such as
|
||
<taxonomicName authority="Thunberg, 1814" authorityName="Thunberg" authorityYear="1814" box="[1122,1432,2000,2026]" class="Insecta" family="Chrysomelidae" genus="Taumacera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1122,1247,2001,2026]" italics="true" pageId="2" pageNumber="583">Taumacera</emphasis>
|
||
Thunberg, 1814
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Baly, 1861" authorityName="Baly" authorityYear="1861" box="[151,398,151,177]" class="Insecta" family="Chrysomelidae" genus="Palpoxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[151,271,151,177]" italics="true" pageId="3" pageNumber="584">Palpoxena</emphasis>
|
||
Baly, 1861
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Jacoby, 1884" authorityName="Jacoby" authorityYear="1884" box="[409,680,151,177]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[409,527,151,177]" italics="true" pageId="3" pageNumber="584">Coeligetes</emphasis>
|
||
Jacoby, 1884
|
||
</taxonomicName>
|
||
, or
|
||
<taxonomicName authority="Lee & Bezdek, 2020" authorityName="Lee & Bezdek" authorityYear="2020" box="[719,1126,151,177]" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[719,890,151,177]" italics="true" pageId="3" pageNumber="584">Pseudotheopea</emphasis>
|
||
<bibRefCitation author="Lee, Ch. - F. & Bezdek, J." box="[895,1126,151,177]" pageId="3" pageNumber="584" pagination="65 - 124" refId="ref2478" refString="Lee, Ch. - F. & Bezdek, J. (2020) Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species. ZooKeys, 912, 65 - 124. https: // doi. org / 10.3897 / zookeys. 912.47719" type="journal article" year="2020">Lee & Bezděk, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
, which do not cross Wallace line (
|
||
<bibRefCitation author="Wilcox, J. A." box="[209,360,187,213]" pageId="3" pageNumber="584" pagination="433 - 664" refId="ref2785" refString="Wilcox, J. A. (1973) Chrysomelidae: Galerucinae (Luperini: Luperina). In: Wilcox, J. A. (Ed.), Coleopterorum Catalogus Supplementa. Pars 78 (3). 2 nd Edition. W. Junk, ' s - Gravenhage, pp. 433 - 664." type="book chapter" year="1973">Wilcox 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Bezdek, J." box="[373,538,187,213]" pageId="3" pageNumber="584" pagination="504 - 524" refId="ref2268" refString="Bezdek, J. (2016 b) Revision of the genus Coeligetes from Malaysia and Indonesia, and description of Coeligetoides gen. nov. (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 4085 (4), 504 - 524. https: // doi. org / 10.11646 / zootaxa. 4085.4.3" type="journal article" year="2016">Bezděk 2016b</bibRefCitation>
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,
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<bibRefCitation author="Bezdek, J." box="[551,610,187,213]" pageId="3" pageNumber="584" pagination="23 - 52" refId="ref2325" refString="Bezdek, J. (2019) Taumacera revisited, with new synonyms, new combinations and a revised catalogue of the species (Coleoptera: Chrysomelidae: Galerucinae). Acta Entomologica Musei Nationalis Pragae, 59, 23 - 52. https: // doi. org / 10.2478 / aemnp- 2019 - 0003" type="journal article" year="2019">2019</bibRefCitation>
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;
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<bibRefCitation author="Lee, Ch. - F. & Bezdek, J." box="[623,857,187,213]" pageId="3" pageNumber="584" pagination="65 - 124" refId="ref2478" refString="Lee, Ch. - F. & Bezdek, J. (2020) Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species. ZooKeys, 912, 65 - 124. https: // doi. org / 10.3897 / zookeys. 912.47719" type="journal article" year="2020">Lee & Bezděk 2020</bibRefCitation>
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). The discovery of a new species of
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<taxonomicName box="[1274,1436,187,212]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="genus">
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<emphasis box="[1274,1436,187,212]" italics="true" pageId="3" pageNumber="584">Hesperopenna</emphasis>
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</taxonomicName>
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in Wallacea, even east of Weber line, is therefore extremely surprising and suggests the possible occurrence of this genus on other islands of Wallacea.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |