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<document id="A5B347019097902725DEE47499E18101" ID-CLB-Dataset="107842" ID-DOI="10.11646/zootaxa.5169.2.3" ID-GBIF-Dataset="12d20702-67c3-49ee-b7f4-3990c93f6951" ID-ISSN="1175-5326" ID-Zenodo-Dep="6920070" ID-ZooBank="F5E85D71-FA41-480E-BB83-05585E90D0F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658995984828" checkinUser="plazi" docAuthor="László, Gyula M." docDate="2022" docId="D82B87B5FFDDB152FF33FC36FE17FF11" docLanguage="en" docName="zootaxa.5169.2.3.pdf" docOrigin="Zootaxa 5169 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ivindonola László 2022, gen. n." docType="treatment" docVersion="7" lastPageNumber="159" masterDocId="2412FFCDFFD6B15EFFA4FFE5FF9DFFE8" masterDocTitle="Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay &amp; Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae)" masterLastPageNumber="164" masterPageNumber="147" pageNumber="158" updateTime="1699362488378" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A588D70F28F88F151D400971396E59B6">Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay &amp; Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae)</mods:title>
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<mods:namePart id="58E8F05CEFF24D6BC43B9571D1BADB84">László, Gyula M.</mods:namePart>
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<treatment id="D82B87B5FFDDB152FF33FC36FE17FF11" ID-DOI="http://doi.org/10.5281/zenodo.6920044" ID-GBIF-Taxon="197867548" ID-Zenodo-Dep="6920044" LSID="urn:lsid:plazi:treatment:D82B87B5FFDDB152FF33FC36FE17FF11" httpUri="http://treatment.plazi.org/id/D82B87B5FFDDB152FF33FC36FE17FF11" lastPageId="12" lastPageNumber="159" pageId="11" pageNumber="158">
<subSubSection id="18986528FFDDB155FF33FC36FEE8FC05" box="[151,373,979,1006]" pageId="11" pageNumber="158" type="nomenclature">
<paragraph id="503D36A3FFDDB155FF33FC36FEE8FC05" blockId="11.[151,373,979,1041]" box="[151,373,979,1006]" pageId="11" pageNumber="158">
<heading id="0B7581CFFFDDB155FF33FC36FEE8FC05" bold="true" box="[151,373,979,1006]" fontSize="11" level="1" pageId="11" pageNumber="158" reason="1">
<emphasis id="62F6EAB1FFDDB155FF33FC36FEE8FC05" bold="true" box="[151,373,979,1006]" pageId="11" pageNumber="158">
<taxonomicName id="97824D20FFDDB155FF33FC36FE86FC06" ID-CoL="BH9F3" authority="László, 2022" authorityName="László" authorityYear="2022" box="[151,283,979,1006]" class="Insecta" family="Nolidae" genus="Ivindonola" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="158" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="62F6EAB1FFDDB155FF33FC36FE86FC06" bold="true" box="[151,283,979,1006]" italics="true" pageId="11" pageNumber="158">Ivindonola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C557CAFFDDB155FE87FC36FEE8FC05" box="[291,373,979,1005]" pageId="11" pageNumber="158" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="18986528FFDDB155FF33FC12FEBAFBF9" box="[151,295,1015,1041]" pageId="11" pageNumber="158" type="description">
<paragraph id="503D36A3FFDDB155FF33FC12FEBAFBF9" blockId="11.[151,373,979,1041]" box="[151,295,1015,1041]" pageId="11" pageNumber="158">
(
<figureCitation id="C8B92A26FFDDB155FF3BFC12FF6BFBF9" box="[159,246,1015,1041]" captionStart="FIGURES 1621" captionStartId="6.[151,264,1931,1955]" captionTargetBox="[165,1429,1240,1901]" captionTargetId="figure-394@6.[155,1433,1234,1907]" captionTargetPageId="6" captionText="FIGURES 1621. adults (coll. ANHRT). 16, Brunneonola gen. n. nimba sp. n., holotype, ♂, Liberia; 17, Ibidem, paratype, ♂, Liberia; 18, Ivindonola gen. n. ipassa sp. n., holotype, ♂, Gabon; 19, Cryptonola gen. n. confundata sp. n., holotype, ♂, Ivory Coast; 20, Ibidem, paratype, ♂, Sierra Leone; 21, Ibidem, paratype, ♀, Ivory Coast." figureDoi="http://doi.org/10.5281/zenodo.6920078" httpUri="https://zenodo.org/record/6920078/files/figure.png" pageId="11" pageNumber="158">Figs 18</figureCitation>
,
<figureCitation id="C8B92A26FFDDB155FEA6FC12FE82FBF9" box="[258,287,1015,1041]" captionStart="FIGURES 2831" captionStartId="13.[151,264,1788,1812]" captionTargetBox="[157,1424,858,1757]" captionTargetId="figure-280@13.[154,1434,849,1765]" captionTargetPageId="13" captionText="FIGURES 2831. male genitalia, a: clasping apparatus, b: aedeagus (coll. ANHRT). 28, Bellanola gen. n. mikongo sp. n., holotype, Gabon, LGNA 1294; 29, Brunneonola gen. n. nimba sp. n., holotype, Liberia, LGNA 1158; 30, Ivindonola gen. n. ipassa sp. n., holotype, Gabon, LGNA 590; 31, Cryptonola gen. n. confundata sp. n., holotype, Ivory Coast, LGNA 1179." figureDoi="http://doi.org/10.5281/zenodo.6920083" httpUri="https://zenodo.org/record/6920083/files/figure.png" pageId="11" pageNumber="158">30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="18986528FFDDB155FF33FBD9FD86FBBC" box="[151,539,1084,1108]" pageId="11" pageNumber="158" type="type_taxon">
<paragraph id="503D36A3FFDDB155FF33FBD9FD86FBBC" blockId="11.[151,539,1084,1108]" box="[151,539,1084,1108]" pageId="11" pageNumber="158">
<typeStatus id="8F398801FFDDB155FF33FBD9FF54FBBC" box="[151,201,1084,1108]" pageId="11" pageNumber="158">Type</typeStatus>
species:
<taxonomicName id="97824D20FFDDB155FE82FBD9FE46FBBB" ID-CoL="BHV5X" authority="László, 2022" authorityName="László" authorityYear="2022" box="[294,475,1084,1108]" class="Insecta" family="Nolidae" genus="Ivindonola" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="158" phylum="Arthropoda" rank="species" species="ipassa" status="sp. nov.">
<emphasis id="62F6EAB1FFDDB155FE82FBD9FE46FBBB" box="[294,475,1084,1108]" italics="true" pageId="11" pageNumber="158">Ivindonola ipassa</emphasis>
</taxonomicName>
<emphasis id="62F6EAB1FFDDB155FE46FBD9FD86FBBC" bold="true" box="[482,539,1084,1108]" pageId="11" pageNumber="158">
<taxonomicNameLabel id="79C557CAFFDDB155FE46FBD9FD86FBBC" box="[482,539,1084,1108]" pageId="11" pageNumber="158" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="18986528FFDDB155FF33FB64FADCFAA6" pageId="11" pageNumber="158" type="diagnosis">
<paragraph id="503D36A3FFDDB155FF33FB64FADCFAA6" blockId="11.[151,1437,1153,2039]" pageId="11" pageNumber="158">
<emphasis id="62F6EAB1FFDDB155FF33FB64FE8FFB73" bold="true" box="[151,274,1153,1179]" pageId="11" pageNumber="158">Diagnosis.</emphasis>
The facies of the
<typeStatus id="8F398801FFDDB155FE7EFB67FD97FB74" box="[474,522,1154,1180]" pageId="11" pageNumber="158">type</typeStatus>
species of the new genus is reminiscent of
<taxonomicName id="97824D20FFDDB155FC4BFB64FB1CFB73" authorityName="László" authorityYear="2022" box="[1007,1153,1153,1179]" class="Insecta" family="Nolidae" genus="Brunneonola" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="62F6EAB1FFDDB155FC4BFB64FB1CFB73" box="[1007,1153,1153,1179]" italics="true" pageId="11" pageNumber="158">Brunneonola</emphasis>
</taxonomicName>
due to its similarly small size, brownish coloration and the red-brown terminal area of the forewing. Despite the similar habitus,
<taxonomicName id="97824D20FFDDB155FA87FB40FA01FB57" authorityName="László" authorityYear="2022" box="[1315,1436,1189,1215]" class="Insecta" family="Nolidae" genus="Ivindonola" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="62F6EAB1FFDDB155FA87FB40FA01FB57" box="[1315,1436,1189,1215]" italics="true" pageId="11" pageNumber="158">Ivindonola</emphasis>
</taxonomicName>
displays a rather unique and distinct genital morphology expressed by the long, pointed and densely setose uncus, a character which is rather prevalent in
<taxonomicName id="97824D20FFDDB155FD9DFB09FD36FAEE" box="[569,683,1260,1286]" class="Insecta" family="Nolidae" genus="Meganola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="62F6EAB1FFDDB155FD9DFB09FD36FAEE" box="[569,683,1260,1286]" italics="true" pageId="11" pageNumber="158">Meganola</emphasis>
</taxonomicName>
, the remarkably narrow tegumen, the medially strongly constricted, unilobate valva, the lack of a harpe, and the presence of a short membranous lobe of the sacculus. The combination of these characters is unknown in any other noline genera and thus a new monotypic genus is erected here.
</paragraph>
</subSubSection>
<subSubSection id="18986528FFDDB152FF63FAB2FE3FFF59" lastPageId="12" lastPageNumber="159" pageId="11" pageNumber="158" type="description">
<paragraph id="503D36A3FFDDB155FF63FAB2FEBCF93F" blockId="11.[151,1437,1153,2039]" pageId="11" pageNumber="158">
<emphasis id="62F6EAB1FFDDB155FF63FAB2FDE8FA99" bold="true" box="[199,629,1367,1394]" pageId="11" pageNumber="158">
Description. Adult. Male (
<figureCitation id="C8B92A26FFDDB155FDA8FAB2FDFBFA99" box="[524,614,1367,1393]" captionStart="FIGURES 1621" captionStartId="6.[151,264,1931,1955]" captionTargetBox="[165,1429,1240,1901]" captionTargetId="figure-394@6.[155,1433,1234,1907]" captionTargetPageId="6" captionText="FIGURES 1621. adults (coll. ANHRT). 16, Brunneonola gen. n. nimba sp. n., holotype, ♂, Liberia; 17, Ibidem, paratype, ♂, Liberia; 18, Ivindonola gen. n. ipassa sp. n., holotype, ♂, Gabon; 19, Cryptonola gen. n. confundata sp. n., holotype, ♂, Ivory Coast; 20, Ibidem, paratype, ♂, Sierra Leone; 21, Ibidem, paratype, ♀, Ivory Coast." figureDoi="http://doi.org/10.5281/zenodo.6920078" httpUri="https://zenodo.org/record/6920078/files/figure.png" pageId="11" pageNumber="158">Fig. 18</figureCitation>
).
</emphasis>
Forewing length of
<typeStatus id="8F398801FFDDB155FCCBFABDFC02FA9A" box="[879,927,1368,1394]" pageId="11" pageNumber="158">type</typeStatus>
species
<quantity id="977A9B46FFDDB155FBADFABDFBCFFA9A" box="[1033,1106,1368,1394]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="158" unit="mm" value="7.0">7 mm</quantity>
. Head small, antenna short, bipectinate with short, finely ciliate rami, distal quarter filiform; labial palp short, porrect, dark brown; frons and vertex grey. Collar and tegula dark grey, suffused by pale grey scales, mesothorax and abdomen pale greyish-brown. Forewing relatively broad, elongate-quadrangular, apically rounded, ground colour pale creamy-grey with sparse, very fine dark brown suffusion; costal margin with small, rounded patches; dorsal two-thirds of terminal area redbrown. Basal and subbasal lines deleted, antemedial line gently arched, diffuse. Medial line deleted, postmedial line very fine, greyish-brown, interrupted. Subterminal line rather broad, dark brown, interrupted, represented by dark brown patches in dorsal-, small dots in ventral area; terminal line very fine, ochreous. Cilia moderately long, red-brown in dorsal, cream in ventral half. Hindwing grey, somewhat darkened apically veins slightly darker; cilia long, proximal half dark grey, distal half pale greyish-brown. Underside of wings ochreous brown with well-defined discal spots.
</paragraph>
<paragraph id="503D36A3FFDDB155FF63F904FE08F81F" blockId="11.[151,1437,1153,2039]" pageId="11" pageNumber="158">
<emphasis id="62F6EAB1FFDDB155FF63F904FE46F913" bold="true" box="[199,475,1761,1787]" pageId="11" pageNumber="158">
Male genitalia (
<figureCitation id="C8B92A26FFDDB155FED8F904FE52F913" box="[380,463,1761,1787]" captionStart="FIGURES 2831" captionStartId="13.[151,264,1788,1812]" captionTargetBox="[157,1424,858,1757]" captionTargetId="figure-280@13.[154,1434,849,1765]" captionTargetPageId="13" captionText="FIGURES 2831. male genitalia, a: clasping apparatus, b: aedeagus (coll. ANHRT). 28, Bellanola gen. n. mikongo sp. n., holotype, Gabon, LGNA 1294; 29, Brunneonola gen. n. nimba sp. n., holotype, Liberia, LGNA 1158; 30, Ivindonola gen. n. ipassa sp. n., holotype, Gabon, LGNA 590; 31, Cryptonola gen. n. confundata sp. n., holotype, Ivory Coast, LGNA 1179." figureDoi="http://doi.org/10.5281/zenodo.6920083" httpUri="https://zenodo.org/record/6920083/files/figure.png" pageId="11" pageNumber="158">Fig. 30</figureCitation>
).
</emphasis>
Uncus relatively long, distal quarter tapered and densely setose, apex pointed.Tegumen very narrow and elongate, arms thin; subscaphium membranous with elongate, densely scobinate plate. Valva long, relatively narrow, medially strongly constricted, dorsal margin concave basally, slightly convex submedially, straight distally, weakly sclerotized. Valva apex broadly rounded, densely scobinate, ventral margin deeply concave medially, gently convex in proximal half. Harpe absent; sacculus narrow, bearing a short, longitudinal lobe distally. Transtilla weakly sclerotized, ribbon-like arms medially fused. Juxta small, U-shaped, vinculum short, V-shaped. Aedeagus long and very thin, gently S-curved, coecum long, apically bifid, Y-shaped, carina absent; vesica without cornuti or scobination.
</paragraph>
<paragraph id="503D36A3FFDAB152FF63FF72FE3FFF59" blockId="12.[151,1436,151,249]" box="[199,418,151,177]" pageId="12" pageNumber="159">
<emphasis id="62F6EAB1FFDAB152FF63FF72FEB8FF59" bold="true" box="[199,293,151,177]" pageId="12" pageNumber="159">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="18986528FFDAB152FF63FF5EFE17FF11" pageId="12" pageNumber="159" type="etymology">
<paragraph id="503D36A3FFDAB152FF63FF5EFE17FF11" blockId="12.[151,1436,151,249]" pageId="12" pageNumber="159">
<emphasis id="62F6EAB1FFDAB152FF63FF5EFED3FF3D" bold="true" box="[199,334,187,213]" pageId="12" pageNumber="159">Etymology.</emphasis>
The prefix in the generic name refers to the
<typeStatus id="8F398801FFDAB152FCE7FF5EFCEEFF3D" box="[835,883,187,213]" pageId="12" pageNumber="159">type</typeStatus>
locality of the
<typeStatus id="8F398801FFDAB152FBB9FF5EFBD0FF3D" box="[1053,1101,187,213]" pageId="12" pageNumber="159">type</typeStatus>
species of the new genus, the Ivindo National Park.
</paragraph>
</subSubSection>
</treatment>
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