226 lines
20 KiB
XML
226 lines
20 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4469962" ID-GBIF-Dataset="6784a642-4e5b-4726-b560-b7b6ed83e0bc" ID-GBIF-Taxon="177128893" ID-Zenodo-Dep="4469962" approvedBy="poa" checkinTime="1611690174320" checkinUser="tatiana" docAuthor="Fedosov, Alexander E, Malcolm, Gavin, Terryn, Yves, Gorson, Juliette, Modica, Maria Vittoria, Holford, Mandë & Puillandre, Nicolas" docDate="2019" docId="D42087ADFF9088662B80E3C7C7D3FEC4" docLanguage="en" docName="jMolluscanStudies.85.359-388.pdf.imf" docOrigin="Journal of Molluscan Studies The Malacological Society of London 85 (4)" docTitle="Bathyterebra zhongshaensis Malcolm, Terryn & Fedosov 2020, newspecies" docType="treatment" docVersion="7" lastPageNumber="23" masterDocId="2819FFD5FF8688702B2FE32FC42CFF8A" masterDocTitle="Phylogenetic classification of the family Terebridae (Neogastropoda: Conoidea)" masterLastPageNumber="388" masterPageNumber="359" pageId="22" pageNumber="23" updateTime="1634699503310" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Phylogenetic classification of the family Terebridae (Neogastropoda: Conoidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fedosov, Alexander E</mods:namePart>
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<mods:affiliation>A. N. Severtsov Institute of Ecology and Evolution of Russian Academy of Sciences, Leninskiy Prospect, 33, Moscow 119071, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Malcolm, Gavin</mods:namePart>
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<mods:affiliation>Bird Hill, Barnes Lane, Milford on Sea, Hampshire, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Terryn, Yves</mods:namePart>
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<mods:affiliation>Kapiteinstraat 27, 9000 Gent, Belgium</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gorson, Juliette</mods:namePart>
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<mods:affiliation>Department of Chemistry, Hunter College Belfer Research Center, New York, NY 10021, USA; 5 Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024, USA; 6 Program in Biology, Graduate Center, City University of New York, New York, NY 10016, USA;</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Modica, Maria Vittoria</mods:namePart>
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<mods:affiliation>Stazione Zoologica Anton Dohrn, Naples, Italy</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Holford, Mandë</mods:namePart>
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<mods:affiliation>Department of Chemistry, Hunter College Belfer Research Center, New York, NY 10021, USA; 5 Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024, USA; 6 Program in Biology, Graduate Center, City University of New York, New York, NY 10016, USA; Programs in Biology, Chemistry & Biochemistry, Graduate Center, City University of New York, New York, NY 10016, USA; 9 Department of Biochemistry, Weill Cornell Medicine, Cornell University, New York, NY 10021, USA.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Puillandre, Nicolas</mods:namePart>
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<mods:affiliation>Institut Systématique Evolution Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, 57 rue Cuvier, CP 26, 75005 Paris, France</mods:affiliation>
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<mods:nameIdentifier type="email">nicolaspuillandre@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Molluscan Studies The Malacological Society of London</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-01-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>85</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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||
</mods:detail>
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<mods:extent unit="page">
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<mods:start>359</mods:start>
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<mods:end>388</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1093/mollus/eyz004</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">6784a642-4e5b-4726-b560-b7b6ed83e0bc</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4469844</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4469962" ID-GBIF-Taxon="177128893" ID-Zenodo-Dep="4469962" LSID="urn:lsid:plazi:treatment:D42087ADFF9088662B80E3C7C7D3FEC4" httpUri="http://treatment.plazi.org/id/D42087ADFF9088662B80E3C7C7D3FEC4" lastPageNumber="23" pageId="22" pageNumber="23">
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<subSubSection pageId="22" pageNumber="23" type="nomenclature">
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||
<paragraph blockId="22.[175,799,232,308]" pageId="22" pageNumber="23">
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||
<heading level="4" pageId="22" pageNumber="23">
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||
<emphasis bold="true" pageId="22" pageNumber="23">
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||
<taxonomicName authority="Malcolm, Terryn & Fedosov" authorityName="Malcolm, Terryn & Fedosov" authorityYear="2020" box="[175,799,232,254]" class="Gastropoda" family="Terebridae" genus="Bathyterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="zhongshaensis" status="new species">
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||
<emphasis bold="true" box="[175,487,232,254]" italics="true" pageId="22" pageNumber="23">Bathyterebra zhongshaensis</emphasis>
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Malcolm, Terryn & Fedosov
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</taxonomicName>
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<taxonomicNameLabel box="[428,563,258,280]" pageId="22" pageNumber="23" rank="species">newspecies</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[465,549,285,308]" pageId="22" pageNumber="23" type="description">
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||
<paragraph blockId="22.[175,799,232,308]" box="[465,549,285,308]" pageId="22" pageNumber="23">
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||
(
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<figureCitation box="[471,541,285,308]" captionStart="Figure 2" captionStartId="5.[158,223,1208,1227]" captionTargetBox="[161,1475,148,1179]" captionTargetId="figure-6@5.[161,1476,148,1179]" captionTargetPageId="5" captionText="Figure 2. Subfamily Pellifroniinae. A–C. Pellifronia jungi. A. MNHN-IM-2013-52275, NANHAI 2014 Stn DW4102, 15◦03'N, 116◦31'E, 339–533 m, 25.5 mm. B. MNHN-IM-2013-52249, EXBODI Stn CP3831, 22◦02'S, 167◦09'E, 523–560 m, 24 mm. C. MNHN-IM-2007-30591, SALOMON 2 Stn CP2195, 08◦26'S, 159◦26'E, 543–593 m, 29 mm (broken), D. Pellifronia brianhayesi holotype, MNHN-IM-2000-20800, S Mozambique, 22.6 mm. E–G. Bathyterebra benthalis. E. MNHN-IM-2013-60185, KARUBENTHOS 2015 Stn CP4524, 16◦29'N, 61◦42'W, 500–550 m, 35.5 mm. F, G. MNHN-IM- 2013-61124, KARUBENTHOS 2015 Stn DW4608, 9.3 mm. H. Bathyterebra zhongshaensis n. sp. holotype, MNHN-IM-2013-61800, ZhongSha 2015 Stn DW4138, 19◦13'N, 113◦56'E, 470–494 m, 17 mm. I. Bathyterebra coriolisi, MNHN-IM-2013-52331, CONCALIS Stn DW3001 18◦32'S, 163◦09'E, 390–400 m, 12.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4469848" httpUri="https://zenodo.org/record/4469848/files/figure.png" pageId="22" pageNumber="23">Fig. 2H</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="materials_examined">
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<paragraph blockId="22.[151,789,339,414]" pageId="22" pageNumber="23">
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<emphasis box="[151,265,339,360]" italics="true" pageId="22" pageNumber="23">
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<typeStatus box="[151,191,339,360]" pageId="22" pageNumber="23">Type</typeStatus>
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material
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</emphasis>
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:
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<materialsCitation ID-GBIF-Occurrence="3026465310" collectingDate="2015" collectionCode="MNHN" country="South China Sea" elevation="482" latitude="19.216667" location="Helen Shoal" longLatPrecision="1272" longitude="113.916664" pageId="22" pageNumber="23" specimenCode="IM-2013-61800" specimenCount="1" stateProvince="Helen Shoal" typeStatus="holotype">
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<typeStatus box="[279,372,339,361]" pageId="22" pageNumber="23">Holotype</typeStatus>
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:
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<collectionCode box="[382,459,339,360]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="22" pageNumber="23">MNHN</collectionCode>
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<specimenCode box="[466,622,339,361]" pageId="22" pageNumber="23">IM-2013-61800</specimenCode>
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,
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<quantity box="[632,722,339,360]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="22" pageNumber="23" unit="mm" value="17.4">17.4 mm</quantity>
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;
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<collectingCountry pageId="22" pageNumber="23">South China Sea</collectingCountry>
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,
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<collectingRegion box="[289,423,365,387]" pageId="22" pageNumber="23">Helen Shoal</collectingRegion>
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,
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<geoCoordinate box="[445,529,360,387]" degrees="19" direction="north" minutes="13" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="19.216667">19°13'N</geoCoordinate>
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,
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<geoCoordinate box="[551,645,360,387]" degrees="113" direction="east" minutes="55" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="113.916664">113°55'E</geoCoordinate>
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,
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<quantity box="[667,789,365,387]" metricMagnitude="2" metricUnit="m" metricValue="4.82" metricValueMax="4.9399999999999995" metricValueMin="4.7" pageId="22" pageNumber="23" unit="m" value="482.0" valueMax="494.0" valueMin="470.0">
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<elevation box="[667,789,365,387]" metricMagnitude="2" metricUnit="m" metricValue="4.82" metricValueMax="4.9399999999999995" metricValueMin="4.7" pageId="22" pageNumber="23" unit="m" value="482.0" valueMax="494.0" valueMin="470.0">470–494 m</elevation>
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</quantity>
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(ZhongSha
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<collectingDate box="[264,312,392,414]" pageId="22" pageNumber="23">2015</collectingDate>
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Stn DW4138).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="22" pageNumber="23" type="description">
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||
<paragraph blockId="22.[151,789,445,493]" pageId="22" pageNumber="23">
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||
<emphasis box="[151,345,445,467]" italics="true" pageId="22" pageNumber="23">Zoobank registration:</emphasis>
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||
<uri pageId="22" pageNumber="23">
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||
urn:lsid:zoobank.org:act:
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||
<uuid pageId="22" pageNumber="23">C8461C7A-BF53- 48D4-BE1B-EF23F17413F8</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[151,501,520,542]" pageId="22" pageNumber="23" type="diagnosis">
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||
<paragraph blockId="22.[151,501,520,542]" box="[151,501,520,542]" pageId="22" pageNumber="23">
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||
<emphasis box="[151,404,520,541]" italics="true" pageId="22" pageNumber="23">Diagnostic nucleotide positions:</emphasis>
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<tableCitation box="[410,495,520,542]" captionStart="Table 11" captionStartId="22.[153,209,1247,1266]" captionTargetBox="[153,787,1328,1931]" captionText="Table 11. Diagnostic combinations of nucleotides in COI alignment for new species of Terebridae described herein." httpUri="http://table.plazi.org/id/08F66633FF9088662BB6E7F0C63DFA80" pageId="22" pageNumber="23" tableUuid="08F66633FF9088662BB6E7F0C63DFA80">Table 11</tableCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="22" pageNumber="23" type="description">
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<paragraph blockId="22.[151,790,568,856]" pageId="22" pageNumber="23">
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<emphasis box="[151,249,568,589]" italics="true" pageId="22" pageNumber="23">Description:</emphasis>
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Shell small, with orthocline spire, wide apical angle. Protoconch wide, paucispiral, 2.5 whorls, with small nucleus. Teleoconch 9 whorls. Early whorls convex in outline with strong, sharply convex ribs; subsequent whorls with narrow constriction formed by indentation, with straight ribs below suture; suture distinct, ragged; mature whorls with less convex outline, weaker and more numerous ribs, 30 on penultimate whorl. Spiral sculpture of very weak threads. Last whorl bulbous at base, but elongate with wide aperture. Columella straight, with weak fold. Overall colour white, with extensive patches of light-tan axial flammules on mature whorls.
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,448,882,904]" pageId="22" pageNumber="23" type="biology_ecology">
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<paragraph blockId="22.[151,448,882,904]" box="[151,448,882,904]" pageId="22" pageNumber="23">
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<emphasis box="[151,223,882,903]" italics="true" pageId="22" pageNumber="23">Habitat:</emphasis>
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Depths of
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<quantity box="[332,444,882,904]" metricMagnitude="2" metricUnit="m" metricValue="4.82" metricValueMax="4.9399999999999995" metricValueMin="4.7" pageId="22" pageNumber="23" unit="m" value="482.0" valueMax="494.0" valueMin="470.0">
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470–
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<specimenCount box="[380,444,882,904]" pageId="22" pageNumber="23" type="generic">494 m</specimenCount>
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</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,560,930,952]" pageId="22" pageNumber="23" type="distribution">
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<paragraph blockId="22.[151,560,930,952]" box="[151,560,930,952]" pageId="22" pageNumber="23">
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<emphasis box="[151,256,930,951]" italics="true" pageId="22" pageNumber="23">Distribution:</emphasis>
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Only known from
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<typeStatus box="[440,481,931,952]" pageId="22" pageNumber="23">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="etymology">
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<paragraph blockId="22.[151,790,978,1027]" pageId="22" pageNumber="23">
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<emphasis box="[151,243,978,999]" italics="true" pageId="22" pageNumber="23">Etymology:</emphasis>
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Named for the Zhongsha sand bars, which form a subsurface atoll on top of the Macclesfield Bank.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="discussion">
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<paragraph blockId="22.[151,789,1053,1181]" lastBlockId="22.[837,1476,152,334]" pageId="22" pageNumber="23">
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<emphasis box="[151,459,1053,1074]" italics="true" pageId="22" pageNumber="23">
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Remarks:
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<taxonomicName authority="Alexander E Fedosov & Gavin Malcolm & Yves Terryn & Juliette Gorson & Maria Vittoria Modica & Mandë Holford & Nicolas Puillandre, 2019" authorityName="Fedosov & Malcolm & Terryn & Gorson & Modica & Holford & Puillandre" authorityYear="2019" box="[235,459,1053,1074]" class="Gastropoda" family="Terebridae" genus="Bathyterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="zhongshaensis" status="n. sp.">Bathyterebra zhongshaensis</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[466,519,1053,1074]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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is similar to the deep-water species,
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<taxonomicName baseAuthorityName="Aubry" baseAuthorityYear="1999" box="[232,324,1079,1101]" class="Gastropoda" family="Terebridae" genus="Bathyterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="coriolisi">
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<emphasis box="[232,324,1079,1101]" italics="true" pageId="22" pageNumber="23">B. coriolisi</emphasis>
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</taxonomicName>
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and
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<taxonomicName baseAuthorityName="Dall" baseAuthorityYear="1889" box="[380,482,1079,1101]" class="Gastropoda" family="Terebridae" genus="Bathyterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="benthalis">
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<emphasis box="[380,482,1079,1101]" italics="true" pageId="22" pageNumber="23">B. benthalis</emphasis>
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</taxonomicName>
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.
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<taxonomicName baseAuthorityName="Aubry" baseAuthorityYear="1999" box="[498,673,1079,1100]" class="Gastropoda" family="Terebridae" genus="Bathyterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="coriolisi">
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<emphasis box="[498,673,1079,1100]" italics="true" pageId="22" pageNumber="23">Bathyterebra coriolisi</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[689,759,1079,1102]" captionStart="Figure 2" captionStartId="5.[158,223,1208,1227]" captionTargetBox="[161,1475,148,1179]" captionTargetId="figure-6@5.[161,1476,148,1179]" captionTargetPageId="5" captionText="Figure 2. Subfamily Pellifroniinae. A–C. Pellifronia jungi. A. MNHN-IM-2013-52275, NANHAI 2014 Stn DW4102, 15◦03'N, 116◦31'E, 339–533 m, 25.5 mm. B. MNHN-IM-2013-52249, EXBODI Stn CP3831, 22◦02'S, 167◦09'E, 523–560 m, 24 mm. C. MNHN-IM-2007-30591, SALOMON 2 Stn CP2195, 08◦26'S, 159◦26'E, 543–593 m, 29 mm (broken), D. Pellifronia brianhayesi holotype, MNHN-IM-2000-20800, S Mozambique, 22.6 mm. E–G. Bathyterebra benthalis. E. MNHN-IM-2013-60185, KARUBENTHOS 2015 Stn CP4524, 16◦29'N, 61◦42'W, 500–550 m, 35.5 mm. F, G. MNHN-IM- 2013-61124, KARUBENTHOS 2015 Stn DW4608, 9.3 mm. H. Bathyterebra zhongshaensis n. sp. holotype, MNHN-IM-2013-61800, ZhongSha 2015 Stn DW4138, 19◦13'N, 113◦56'E, 470–494 m, 17 mm. I. Bathyterebra coriolisi, MNHN-IM-2013-52331, CONCALIS Stn DW3001 18◦32'S, 163◦09'E, 390–400 m, 12.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4469848" httpUri="https://zenodo.org/record/4469848/files/figure.png" pageId="22" pageNumber="23">Fig. 2E</figureCitation>
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) is similar in colour, in the sculpture of the early whorls and in the shape of its aperture. However,
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<taxonomicName baseAuthorityName="Aubry" baseAuthorityYear="1999" box="[459,549,1133,1154]" class="Gastropoda" family="Terebridae" genus="Bathyterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="coriolisi">
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<emphasis box="[459,549,1133,1154]" italics="true" pageId="22" pageNumber="23">B. coriolisi</emphasis>
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</taxonomicName>
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differs in having a much narrower shape, with a strong subsutural groove and fewer ribs on the early whorls, creating a band with nodes. Its subsutural groove becomes weaker, with just a slight indentation on mature whorls.
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<taxonomicName baseAuthorityName="Dall" baseAuthorityYear="1889" box="[837,1023,206,227]" class="Gastropoda" family="Terebridae" genus="Bathyterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="benthalis">
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<emphasis box="[837,1023,206,227]" italics="true" pageId="22" pageNumber="23">Bathyterebra benthalis</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[1042,1108,206,228]" captionStart="Figure 2" captionStartId="5.[158,223,1208,1227]" captionTargetBox="[161,1475,148,1179]" captionTargetId="figure-6@5.[161,1476,148,1179]" captionTargetPageId="5" captionText="Figure 2. Subfamily Pellifroniinae. A–C. Pellifronia jungi. A. MNHN-IM-2013-52275, NANHAI 2014 Stn DW4102, 15◦03'N, 116◦31'E, 339–533 m, 25.5 mm. B. MNHN-IM-2013-52249, EXBODI Stn CP3831, 22◦02'S, 167◦09'E, 523–560 m, 24 mm. C. MNHN-IM-2007-30591, SALOMON 2 Stn CP2195, 08◦26'S, 159◦26'E, 543–593 m, 29 mm (broken), D. Pellifronia brianhayesi holotype, MNHN-IM-2000-20800, S Mozambique, 22.6 mm. E–G. Bathyterebra benthalis. E. MNHN-IM-2013-60185, KARUBENTHOS 2015 Stn CP4524, 16◦29'N, 61◦42'W, 500–550 m, 35.5 mm. F, G. MNHN-IM- 2013-61124, KARUBENTHOS 2015 Stn DW4608, 9.3 mm. H. Bathyterebra zhongshaensis n. sp. holotype, MNHN-IM-2013-61800, ZhongSha 2015 Stn DW4138, 19◦13'N, 113◦56'E, 470–494 m, 17 mm. I. Bathyterebra coriolisi, MNHN-IM-2013-52331, CONCALIS Stn DW3001 18◦32'S, 163◦09'E, 390–400 m, 12.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4469848" httpUri="https://zenodo.org/record/4469848/files/figure.png" pageId="22" pageNumber="23">
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Fig.
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<date box="[1088,1108,206,228]" pageId="22" pageNumber="23">2I</date>
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</figureCitation>
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) is similar in colour and in overall sculpture of the shell, but differs from
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<taxonomicName authority="Alexander E Fedosov & Gavin Malcolm & Yves Terryn & Juliette Gorson & Maria Vittoria Modica & Mandë Holford & Nicolas Puillandre, 2019" authorityName="Fedosov & Malcolm & Terryn & Gorson & Modica & Holford & Puillandre" authorityYear="2019" box="[1212,1353,232,254]" class="Gastropoda" family="Terebridae" genus="Bathyterebra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="zhongshaensis" status="n. sp.">
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<emphasis box="[1212,1353,232,254]" italics="true" pageId="22" pageNumber="23">B. zhongshaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1360,1412,233,254]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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in the outline of its mature whorls being flat and the ribs on the early whorls extremely convex, creating nodes that dominate the profile of the early whorls.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |