treatments-xml/data/C6/56/0C/C6560C5A6B2FFF82E7FEFF3AFF62FC56.xml

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<mods:title id="BDE2584ADD1AB5914D44D822C6AE13C9">On the identity of Sundathelphusa philippina (von Martens, 1868) (Decapoda: Brachyura: Gecarcinucidae) from the Philippines, with descriptions of two new species</mods:title>
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<mods:namePart id="61DC44E7063E7E7B5D996DE088F6AD9F">Husana, Daniel Edison M.</mods:namePart>
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<taxonomicName id="89FFC6CF6B2FFF8EE7FEFF3AFE12FF2F" ID-CoL="84GBM" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[151,412,151,177]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B2FFF8EE7FEFF3AFE12FF2F" bold="true" box="[151,412,151,177]" italics="true" pageId="7" pageNumber="322">Sundathelphusa cebu</emphasis>
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<taxonomicNameLabel id="67B8DC256B2FFF8EE6CDFF35FD8FFF2C" box="[420,513,152,178]" pageId="7" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="4E40BD4C6B2FFF8EE7FEFF17FDBAFF4D" blockId="7.[151,564,151,211]" box="[151,564,186,211]" pageId="7" pageNumber="322">
(
<figureCitation id="D6C4A1C96B2FFF8EE7F6FF17FE85FF4D" box="[159,267,186,211]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="8.[151,250,1944,1966]" captionStartId-1="9.[151,250,1401,1423]" captionTargetBox-0="[150,1436,448,1922]" captionTargetBox-1="[153,1430,213,1363]" captionTargetId-0="figure@8.[150,1436,448,1922]" captionTargetId-1="figure@9.[150,1436,193,1380]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." captionText-1="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri-0="https://zenodo.org/record/2637374/files/figure.png" httpUri-1="https://zenodo.org/record/2637376/files/figure.png" pageId="7" pageNumber="322">Figs. 4–5</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE67EFF17FE52FF4C" box="[279,476,186,211]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="7" pageNumber="322">6B, D, F, H, K, L</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE681FF16FDA5FF4D" box="[488,555,187,211]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: A–F = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="7" pageNumber="322">9C, D</figureCitation>
)
</paragraph>
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<taxonomicName id="89FFC6CF6B2FFF8EE7FEFEAFFE27FE86" ID-CoL="53GC2" authorityName="Husana &amp; Ng" authorityYear="2019" box="[151,425,258,280]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2FFF8EE7FEFEAFFE27FE86" box="[151,425,258,280]" italics="true" pageId="7" pageNumber="322">Sundathelphusa philippina</emphasis>
</taxonomicName>
— 
<treatmentCitation id="CF5E9B5D6B2FFF8EE6ABFEAFFDF2FE86" author="Takeda, M." box="[450,636,258,280]" page="101" pageId="7" pageNumber="322" year="1987">
<bibRefCitation id="2A6EC0BD6B2FFF8EE6ABFEAFFDC7FE86" author="Takeda, M." box="[450,585,258,280]" pageId="7" pageNumber="322" pagination="89 - 106" refId="ref9849" refString="Takeda, M. (1987) Freshwater crabs from Peninsu 1 ar Malaysia, with notes on some species from the Philippines. In: Kawashima, K. (Ed.), Paragonimus in Asia: Biology, Genetic Variation and Speciation. Paragonimus Research Report, Kyushu University, 1, pp. 89 - 106." type="journal article" year="1987">Takeda 1987</bibRefCitation>
: 101
</treatmentCitation>
(part), 102; 
<treatmentCitation id="CF5E9B5D6B2FFF8EE468FEAEFC6CFE86" author="Ng, P. K. L. &amp; Sket, B." box="[769,994,258,280]" page="697" pageId="7" pageNumber="322" year="1996">
<bibRefCitation id="2A6EC0BD6B2FFF8EE468FEAEFC3EFE86" author="Ng, P. K. L. &amp; Sket, B." box="[769,944,258,280]" pageId="7" pageNumber="322" pagination="695 - 706" refId="ref9587" refString="Ng, P. K. L. &amp; Sket, B. (1996) The freshwater crab fauna (Crustacea: Decapoda: Brachyura) of the Philippines. IV. On the collection of Parathelphusidae from Bohol. Proceeding of the Biological Society of Washington, 109, 695 - 706." type="journal article" year="1996">Ng &amp; Sket 1996</bibRefCitation>
: 697
</treatmentCitation>
(discussion) (in part); 
<treatmentCitation id="CF5E9B5D6B2FFF8EE3B8FEAFFEAEFEA9" author="Mendoza, J. C. E. &amp; Naruse, T." page="63" pageId="7" pageNumber="322" year="2010">
<bibRefCitation id="2A6EC0BD6B2FFF8EE3B8FEAFFF70FEA9" author="Mendoza, J. C. E. &amp; Naruse, T." pageId="7" pageNumber="322" pagination="61 - 70" refId="ref9108" refString="Mendoza, J. C. E. &amp; Naruse, T. (2010) A new species of riverine crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from northeastern Luzon, Philippines. Philippine Journal of Science, 139 (1), 61 - 70." type="journal article" year="2010">Mendoza &amp; Naruse 2010</bibRefCitation>
: 63
</treatmentCitation>
(table 1), 67 (comparative material), (in part). Not 
<taxonomicName id="89FFC6CF6B2FFF8EE44EFE8FFB0FFEA9" ID-CoL="53GC2" authority="(von Martens, 1868)" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[807,1153,289,311]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2FFF8EE44EFE8FFC26FEA9" box="[807,936,289,311]" italics="true" pageId="7" pageNumber="322">S. philippina</emphasis>
(von 
<bibRefCitation id="2A6EC0BD6B2FFF8EE48AFE8FFBF7FEA9" author="Martens, E. von" box="[995,1145,289,311]" pageId="7" pageNumber="322" pagination="608 - 615" refId="ref9072" refString="Martens, E. von (1868) Ueber einige neue crustaceen. Monatsberichte der Koniglichen Preussische Akademie der Akademie der Wissenschaften zu Berlin, Sitzung der Physikalisch-mathematische Klasse, 1868, 608 - 615." type="journal article" year="1868">Martens, 1868</bibRefCitation>
)
</taxonomicName>
.
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<emphasis id="7C8B615E6B2FFF8EE7FEFEC9FEF4FEE3" bold="true" box="[151,378,356,381]" pageId="7" pageNumber="322">Material examined</emphasis>
. 
<materialsCitation id="FE97B7116B2FFF8EE6E0FEC9FD25FE3E" ID-GBIF-Occurrence="2235592118" collectingDate="1977-06-22" collectionCode="ZRC" collectorName="S. I. Ueno" elevation="450" location="Cebu Island" pageId="7" pageNumber="322" specimenCode="ZRC 2017.1254" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="914403EE6B2FFF8EE6E0FEC9FE78FEE3" box="[393,502,356,381]" pageId="7" pageNumber="322" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="58F976C56B2FFF8EE568FEC8FD98FEE2" box="[513,534,357,380]" pageId="7" pageNumber="322" type="male">♂</specimenCount>
(38.1 × 
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), 
<specimenCode id="1E5915376B2FFF8EE59CFEC9FC23FEE3" box="[757,941,356,381]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2017.1254</specimenCode>
, Campo Siete (Camp 7, Minglanilla, 
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<quantity id="890710A96B2FFF8EE23DFEC9FA12FEE2" box="[1364,1436,356,381]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="322" unit="m" value="450.0">450 m</quantity>
asl
</elevation>
, 
<location id="4B20EB976B2FFF8EE7ACFE2AFEDEFE3E" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE7ACFE2AFEDEFE3E" box="[197,336,391,416]" name="Cebu Island" pageId="7" pageNumber="322">Cebu Island</location>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE6FFFE25FD8BFE01" box="[406,517,392,416]" pageId="7" pageNumber="322">S.I. Ueno</collectorName>
, 
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<collectingDate id="2A0562646B2FFF8EE578FE25FD25FE3E" box="[529,683,392,416]" pageId="7" pageNumber="322" value="1977-06-22">22 June 1977</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FE97B7116B2FFF8EE5D1FE25FB73FE3E" ID-GBIF-Occurrence="2235592130" box="[696,1277,391,416]" collectionCode="ZRC" pageId="7" pageNumber="322" specimenCode="ZRC 2009.0102" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="914403EE6B2FFF8EE5D1FE25FCA7FE3E" box="[696,809,392,416]" pageId="7" pageNumber="322" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="58F976C56B2FFF8EE45CFE25FCEEFE01" box="[821,864,392,416]" pageId="7" pageNumber="322" type="male">1 ♂</specimenCount>
(32.9 × 
<quantity id="890710A96B2FFF8EE4AAFE2AFBA2FE3E" box="[963,1068,391,416]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="7" pageNumber="322" unit="mm" value="26.2">26.2 mm</quantity>
), 
<specimenCode id="1E5915376B2FFF8EE329FE25FB77FE3E" box="[1088,1273,392,416]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2009.0102</specimenCode>
,
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<materialsCitation id="FE97B7116B2FFF8EE26CFE25FDD2FE5B" ID-GBIF-Occurrence="2235592121" collectingDate="1977-06-22" collectionCode="ZRC" collectorName="S. I. Ueno" elevation="450" location="Cebu Island" pageId="7" pageNumber="322" specimenCount="9" specimenCount-female="5" specimenCount-male="4" typeStatus="paratype">
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, 
<specimenCount id="58F976C56B2FFF8EE23CFE2AFA18FE01" box="[1365,1430,391,416]" pageId="7" pageNumber="322" type="female">5 ♀♀</specimenCount>
, NSMT-Cr 25887, same data as holotype
</materialsCitation>
; 
<specimenCount id="58F976C56B2FFF8EE501FE01FD28FE5A" box="[616,678,428,452]" pageId="7" pageNumber="322" type="male">2 ♂♂</specimenCount>
(larger 29.3 × 24.0 mm), 
<materialsCitation id="FE97B7116B2FFF8EE4A1FE01FA09FE5B" ID-GBIF-Occurrence="2235592112" box="[968,1415,428,453]" collectionCode="ZRC" pageId="7" pageNumber="322" specimenCode="ZRC 2017.1061" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE4A1FE01FC7CFE5A" box="[968,1010,428,452]" pageId="7" pageNumber="322" type="female">1 ♀</specimenCount>
(29.5 × 
<quantity id="890710A96B2FFF8EE338FE01FB37FE5A" box="[1105,1209,428,452]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" pageId="7" pageNumber="322" unit="mm" value="24.2">24.2 mm</quantity>
), 
<specimenCode id="1E5915376B2FFF8EE3A5FE01FA0DFE5B" box="[1228,1411,428,453]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2017.1061</specimenCode>
,
</materialsCitation>
<materialsCitation id="FE97B7116B2FFF8EE2E7FE01FAECFE76" ID-GBIF-Occurrence="2235592115" collectingDate="1985-07-29" collectorName="M. Takeda" location="Mananga River" municipality="Cebu Island" pageId="7" pageNumber="322" specimenCount="14" specimenCount-female="6" specimenCount-male="8" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE2E7FE01FF4DFE79" pageId="7" pageNumber="322" type="male">8 ♂♂</specimenCount>
, 
<specimenCount id="58F976C56B2FFF8EE7B8FE62FE9DFE79" box="[209,275,463,488]" pageId="7" pageNumber="322" type="female">6 ♀♀</specimenCount>
, NSMT-Cr 25888, upper stream of 
<location id="4B20EB976B2FFF8EE5C6FE7DFCEEFE76" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE5C6FE7DFCEEFE76" box="[687,864,463,488]" municipality="Cebu Island" name="Mananga River" pageId="7" pageNumber="322">Mananga River</location>
, 
<collectingMunicipality id="AE2427366B2FFF8EE407FE62FC77FE76" box="[878,1017,463,488]" pageId="7" pageNumber="322">Cebu Island</collectingMunicipality>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE328FE7DFB30FE76" box="[1089,1214,463,488]" pageId="7" pageNumber="322">M. Takeda</collectorName>
, 
<date id="3A419B8C6B2FFF8EE3A3FE7DFAECFE76" box="[1226,1378,463,488]" pageId="7" pageNumber="322" value="1985-07-29">
<collectingDate id="2A0562646B2FFF8EE3A3FE7DFAECFE76" box="[1226,1378,463,488]" pageId="7" pageNumber="322" value="1985-07-29">29 July 1985</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="58F976C56B2FFF8EE219FE7DFA12FE79" box="[1392,1436,464,488]" pageId="7" pageNumber="322" type="male">1 ♂</specimenCount>
(39.3 × 
<quantity id="890710A96B2FFF8EE799FE59FED6FD92" box="[240,344,500,524]" metricMagnitude="-2" metricUnit="m" metricValue="3.1399999999999997" pageId="7" pageNumber="322" unit="mm" value="31.4">31.4 mm</quantity>
), 3 young 
<materialsCitation id="FE97B7116B2FFF8EE6A7FE58FDC1FDAE" ID-GBIF-Occurrence="2235592127" collectingDate="2000-12-14" collectionCode="ZRC" collectorName="Y. Cai" county="Cebu Island" location="Cantipla" municipality="Cebu City" pageId="7" pageNumber="322" specimenCode="ZRC 2006.0055" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE6A7FE58FE74FD92" box="[462,506,501,524]" pageId="7" pageNumber="322" type="male">♂♂</specimenCount>
, 
<specimenCode id="1E5915376B2FFF8EE56EFE59FD31FD93" box="[519,703,500,525]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2006.0055</specimenCode>
, 
<location id="4B20EB976B2FFF8EE5A3FE59FCA7FD93" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE5A3FE59FCA7FD93" box="[714,809,500,525]" county="Cebu Island" municipality="Cebu City" name="Cantipla" pageId="7" pageNumber="322">Cantipla</location>
(mislabeled as 
<location id="4B20EB976B2FFF8EE4B0FE59FBB5FD93" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE4B0FE59FBB5FD93" box="[985,1083,500,525]" county="Cebu Island" municipality="Cebu City" name="Caufipla" pageId="7" pageNumber="322">Caufipla</location>
) 
<location id="4B20EB976B2FFF8EE322FE59FB11FD93" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE322FE59FB11FD93" box="[1099,1183,500,525]" county="Cebu Island" municipality="Cebu City" name="Village" pageId="7" pageNumber="322">Village</location>
, near 
<collectingMunicipality id="AE2427366B2FFF8EE38BFE59FADBFD93" box="[1250,1365,500,525]" pageId="7" pageNumber="322">Cebu City</collectingMunicipality>
, 
<collectingCounty id="A721C5C06B2FFF8EE208FE59FF50FDAE" pageId="7" pageNumber="322">Cebu Island</collectingCounty>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE64BFDB5FEE4FDAE" box="[290,362,535,560]" pageId="7" pageNumber="322">Y. Cai</collectorName>
, 
<date id="3A419B8C6B2FFF8EE61FFDB5FDC1FDAE" box="[374,591,535,560]" pageId="7" pageNumber="322" value="2000-12-14">
<collectingDate id="2A0562646B2FFF8EE61FFDB5FDC1FDAE" box="[374,591,535,560]" pageId="7" pageNumber="322" value="2000-12-14">14 December 2000</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="58F976C56B2FFF8EE532FDB5FD0BFDB1" box="[603,645,536,560]" pageId="7" pageNumber="322" type="male">1 ♂</specimenCount>
(37.5 × 30.0 mm), 
<materialsCitation id="FE97B7116B2FFF8EE409FDB5FC2FFDCA" ID-GBIF-Occurrence="2235592123" collectingDate="2000-12-14" collectionCode="ZRC" collectorName="Y. Cai" county="Cebu Island" location="Malbubog Vilage" municipality="Cebu City" pageId="7" pageNumber="322" specimenCode="ZRC 2017.1255" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE409FDB5FC05FDB1" box="[864,907,536,560]" pageId="7" pageNumber="322" type="female">1 ♀</specimenCount>
(52.2 × 
<quantity id="890710A96B2FFF8EE483FDB5FBDDFDAE" box="[1002,1107,536,560]" metricMagnitude="-2" metricUnit="m" metricValue="4.029999999999999" pageId="7" pageNumber="322" unit="mm" value="40.3">40.3 mm</quantity>
), 
<specimenCode id="1E5915376B2FFF8EE30FFDB5FA90FDAE" box="[1126,1310,535,560]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2017.1255</specimenCode>
, 
<location id="4B20EB976B2FFF8EE240FDBAFF6DFDCB" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE240FDBAFF6DFDCB" county="Cebu Island" municipality="Cebu City" name="Malbubog Vilage" pageId="7" pageNumber="322">Malbubog Vilage</location>
, near 
<collectingMunicipality id="AE2427366B2FFF8EE64FFD91FE14FDCB" box="[294,410,572,597]" pageId="7" pageNumber="322">Cebu City</collectingMunicipality>
, 
<collectingCounty id="A721C5C06B2FFF8EE6CFFD91FDA1FDCB" box="[422,559,572,597]" pageId="7" pageNumber="322">Cebu Island</collectingCounty>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE51AFD90FD32FDCB" box="[627,700,572,597]" pageId="7" pageNumber="322">Y. Cai</collectorName>
, 
<date id="3A419B8C6B2FFF8EE5A1FD91FC2FFDCA" box="[712,929,572,597]" pageId="7" pageNumber="322" value="2000-12-14">
<collectingDate id="2A0562646B2FFF8EE5A1FD91FC2FFDCA" box="[712,929,572,597]" pageId="7" pageNumber="322" value="2000-12-14">14 December 2000</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="58F976C56B2FFF8EE4C4FD90FC63FDCA" box="[941,1005,573,596]" pageId="7" pageNumber="322" type="male">7 ♂♂</specimenCount>
(largest 37.5 × 
<quantity id="890710A96B2FFF8EE3F6FD91FA86FDCA" box="[1183,1288,572,596]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="7" pageNumber="322" unit="mm" value="31.2">31.2 mm</quantity>
), 
<materialsCitation id="FE97B7116B2FFF8EE272FD91FA0EFDE6" ID-GBIF-Occurrence="2235592116" collectingDate="2000-12-14" collectionCode="ZRC" collectorName="Y. Cai" county="Cebu Island" location="Tabunan Village" municipality="Cebu City" pageId="7" pageNumber="322" specimenCode="ZRC 2017.1256" specimenCount="9" specimenCount-female="9" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE272FD91FAD4FDCA" box="[1307,1370,572,596]" pageId="7" pageNumber="322" type="female">9 ♀♀</specimenCount>
(32.6 × 
<quantity id="890710A96B2FFF8EE7D9FDF2FE94FDE6" box="[176,282,607,632]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="7" pageNumber="322" unit="mm" value="27.5">27.5 mm</quantity>
), 
<specimenCode id="1E5915376B2FFF8EE647FDCDFE66FDE6" box="[302,488,607,632]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2017.1256</specimenCode>
, 
<location id="4B20EB976B2FFF8EE69CFDF2FD3CFDE6" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE69CFDF2FD3CFDE6" box="[501,690,607,632]" county="Cebu Island" municipality="Cebu City" name="Tabunan Village" pageId="7" pageNumber="322">Tabunan Village</location>
, near 
<collectingMunicipality id="AE2427366B2FFF8EE591FDF2FCE3FDE6" box="[760,877,607,632]" pageId="7" pageNumber="322">Cebu City</collectingMunicipality>
, 
<collectingCounty id="A721C5C06B2FFF8EE413FDF2FB88FDE6" box="[890,1030,607,632]" pageId="7" pageNumber="322">Cebu Island</collectingCounty>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE324FDCDFB19FDE6" box="[1101,1175,607,632]" pageId="7" pageNumber="322">Y. Cai</collectorName>
, 
<date id="3A419B8C6B2FFF8EE3CDFDCDFA0EFDE6" box="[1188,1408,607,632]" pageId="7" pageNumber="322" value="2000-12-14">
<collectingDate id="2A0562646B2FFF8EE3CDFDCDFA0EFDE6" box="[1188,1408,607,632]" pageId="7" pageNumber="322" value="2000-12-14">14 December 2000</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="FE97B7116B2FFF8EE2E7FDCDFEA4FD5E" ID-GBIF-Occurrence="2235592120" collectingDate="2003-07-30" collectionCode="ZRC" collectorName="P. K. L. Ng" county="Cebu Island" location="Kawasan Falls" municipality="Badian town" pageId="7" pageNumber="322" specimenCode="ZRC 2009.0098" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE2E7FDCDFF4DFD02" pageId="7" pageNumber="322" type="male">3 ♂♂</specimenCount>
, 
<specimenCount id="58F976C56B2FFF8EE7A6FD29FF77FD02" box="[207,249,644,668]" pageId="7" pageNumber="322" type="female">1 ♀</specimenCount>
, 
<specimenCode id="1E5915376B2FFF8EE66EFD29FE31FD02" box="[263,447,644,668]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2009.0098</specimenCode>
, 
<location id="4B20EB976B2FFF8EE6A0FD28FDFEFD03" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE6A0FD28FDFEFD03" box="[457,624,644,669]" county="Cebu Island" municipality="Badian town" name="Kawasan Falls" pageId="7" pageNumber="322">Kawasan Falls</location>
, along 
<location id="4B20EB976B2FFF8EE5A8FD29FCFCFD03" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE5A8FD29FCFCFD03" box="[705,882,644,669]" county="Cebu Island" municipality="Badian town" name="Matutinao river" pageId="7" pageNumber="322">Matutinao river</location>
, 
<collectingMunicipality id="AE2427366B2FFF8EE414FD29FB80FD02" box="[893,1038,644,669]" pageId="7" pageNumber="322">Badian town</collectingMunicipality>
, 
<collectingCounty id="A721C5C06B2FFF8EE373FD29FB2DFD03" box="[1050,1187,644,669]" pageId="7" pageNumber="322">Cebu Island</collectingCounty>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE38FFD28FAD8FD03" box="[1254,1366,645,669]" pageId="7" pageNumber="322">P.K.L. Ng</collectorName>
<emphasis id="7C8B615E6B2FFF8EE234FD28FA1BFD03" box="[1373,1429,644,669]" italics="true" pageId="7" pageNumber="322">et al.</emphasis>
, 
<date id="3A419B8C6B2FFF8EE7FEFD05FEA4FD5E" box="[151,298,679,704]" pageId="7" pageNumber="322" value="2003-07-30">
<collectingDate id="2A0562646B2FFF8EE7FEFD05FEA4FD5E" box="[151,298,679,704]" pageId="7" pageNumber="322" value="2003-07-30">30 July 2003</collectingDate>
</date>
</materialsCitation>
; 1 young 
<specimenCount id="58F976C56B2FFF8EE6FEFD05FE20FD21" box="[407,430,680,703]" pageId="7" pageNumber="322" type="male">♂</specimenCount>
, 6 young 
<materialsCitation id="FE97B7116B2FFF8EE575FD05FEC3FD7A" ID-GBIF-Occurrence="2235592125" collectingDate="2001-12-02" collectionCode="ZRC" collectorName="C. Liu" county="Cebu Island" location="Kawasan Falls" municipality="Matutinao" pageId="7" pageNumber="322" specimenCode="ZRC 2009.0099" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE575FD05FDC9FD21" box="[540,583,680,703]" pageId="7" pageNumber="322" type="female">♀♀</specimenCount>
, 
<specimenCode id="1E5915376B2FFF8EE53DFD05FC85FD5E" box="[596,779,680,704]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2009.0099</specimenCode>
, 
<location id="4B20EB976B2FFF8EE47FFD05FC32FD5E" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE47FFD05FC32FD5E" box="[790,956,679,704]" county="Cebu Island" municipality="Matutinao" name="Kawasan Falls" pageId="7" pageNumber="322">Kawasan Falls</location>
, 
<collectingMunicipality id="AE2427366B2FFF8EE4A1FD0AFBB0FD5E" box="[968,1086,679,704]" pageId="7" pageNumber="322">Matutinao</collectingMunicipality>
, 
<collectingCounty id="A721C5C06B2FFF8EE320FD0AFB5CFD5E" box="[1097,1234,679,704]" pageId="7" pageNumber="322">Cebu Island</collectingCounty>
, coll. H.- 
<collectorName id="E30AD89A6B2FFF8EE250FD05FA0DFD5E" box="[1337,1411,679,704]" pageId="7" pageNumber="322">C. Liu</collectorName>
, 
<date id="3A419B8C6B2FFF8EE2E7FD05FEC3FD7A" pageId="7" pageNumber="322" value="2001-12-02">
<collectingDate id="2A0562646B2FFF8EE2E7FD05FEC3FD7A" pageId="7" pageNumber="322" value="2001-12-02">2 December 2001</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="FE97B7116B2FFF8EE633FD61FEFCFC96" ID-GBIF-Occurrence="2235592117" collectingDate="2006-12-20" collectionCode="ZRC" collectorName="Y. Cai" county="Cebu Island" location="Kawasan Falls" municipality="Matutinao" pageId="7" pageNumber="322" specimenCode="ZRC 2006.0056" specimenCount="10" specimenCount-female="5" specimenCount-juvenile="3" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE633FD61FE12FD7A" box="[346,412,716,740]" pageId="7" pageNumber="322" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="58F976C56B2FFF8EE6C0FD61FE64FD7A" box="[425,490,716,741]" pageId="7" pageNumber="322" type="female">5 ♀♀</specimenCount>
, 
<specimenCount id="58F976C56B2FFF8EE690FD61FDF9FD7B" box="[505,631,716,741]" pageId="7" pageNumber="322" type="juvenile">3 juveniles</specimenCount>
, 
<specimenCode id="1E5915376B2FFF8EE5EAFD61FCB2FD7B" box="[643,828,716,741]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2006.0056</specimenCode>
, 
<location id="4B20EB976B2FFF8EE421FD60FC7EFD7B" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE421FD60FC7EFD7B" box="[840,1008,716,741]" county="Cebu Island" municipality="Matutinao" name="Kawasan Falls" pageId="7" pageNumber="322">Kawasan Falls</location>
, 
<collectingMunicipality id="AE2427366B2FFF8EE495FD61FBFCFD7B" box="[1020,1138,716,741]" pageId="7" pageNumber="322">Matutinao</collectingMunicipality>
, 
<collectingCounty id="A721C5C06B2FFF8EE316FD61FA87FD7B" box="[1151,1289,716,741]" pageId="7" pageNumber="322">Cebu Island</collectingCounty>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE226FD60FA16FD7B" box="[1359,1432,716,741]" pageId="7" pageNumber="322">Y. Cai</collectorName>
, 
<date id="3A419B8C6B2FFF8EE7FEFD5DFEFCFC96" box="[151,370,751,776]" pageId="7" pageNumber="322" value="2006-12-20">
<collectingDate id="2A0562646B2FFF8EE7FEFD5DFEFCFC96" box="[151,370,751,776]" pageId="7" pageNumber="322" value="2006-12-20">20 December 2006</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="FE97B7116B2FFF8EE616FD5DFDFDFCB2" ID-GBIF-Occurrence="2235592128" collectingDate="2012-12" collectionCode="ZRC" collectorName="J. C. Y. Lai" county="Cebu Island" location="Kawasan Falls" municipality="Matutinao" pageId="7" pageNumber="322" specimenCode="ZRC 2011.0002" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE616FD5DFE31FC99" box="[383,447,752,776]" pageId="7" pageNumber="322" type="male">2 ♂♂</specimenCount>
(36.6 × 
<quantity id="890710A96B2FFF8EE54BFD5DFD04FC99" box="[546,650,752,776]" metricMagnitude="-2" metricUnit="m" metricValue="3.03" pageId="7" pageNumber="322" unit="mm" value="30.3">30.3 mm</quantity>
, 40.5 × 32.0 mm), 
<specimenCode id="1E5915376B2FFF8EE40FFD5DFB93FC96" box="[870,1053,752,776]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2011.0002</specimenCode>
, 
<location id="4B20EB976B2FFF8EE343FD5DFB5CFC96" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE343FD5DFB5CFC96" box="[1066,1234,751,776]" county="Cebu Island" municipality="Matutinao" name="Kawasan Falls" pageId="7" pageNumber="322">Kawasan Falls</location>
, 
<collectingMunicipality id="AE2427366B2FFF8EE3B6FD42FADBFC96" box="[1247,1365,751,776]" pageId="7" pageNumber="322">Matutinao</collectingMunicipality>
, 
<collectingCounty id="A721C5C06B2FFF8EE208FD42FF50FCB3" pageId="7" pageNumber="322">Cebu Island</collectingCounty>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE644FCB9FE28FCB3" box="[301,422,788,813]" pageId="7" pageNumber="322">J.C.Y. Lai</collectorName>
, 
<date id="3A419B8C6B2FFF8EE6DEFCB9FDFDFCB2" box="[439,627,788,813]" pageId="7" pageNumber="322" value="2012-12">
<collectingDate id="2A0562646B2FFF8EE6DEFCB9FDFDFCB2" box="[439,627,788,813]" pageId="7" pageNumber="322" value="2012-12">December 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="FE97B7116B2FFF8EE5EDFCB9FBE1FCB3" ID-GBIF-Occurrence="2235592106" box="[644,1135,788,813]" collectionCode="ZRC" pageId="7" pageNumber="322" specimenCode="ZRC 2017.1253" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE5EDFCB9FD3BFCB2" box="[644,693,788,812]" pageId="7" pageNumber="322" type="male">1 ♂</specimenCount>
(36.5 × 
<quantity id="890710A96B2FFF8EE44FFCB9FC1AFCB2" box="[806,916,788,812]" metricMagnitude="-2" metricUnit="m" metricValue="2.8899999999999997" pageId="7" pageNumber="322" unit="mm" value="28.9">28.9 mm</quantity>
), 
<specimenCode id="1E5915376B2FFF8EE4C4FCB9FBE5FCB3" box="[941,1131,788,813]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="322">ZRC 2017.1253</specimenCode>
,
</materialsCitation>
<materialsCitation id="FE97B7116B2FFF8EE315FCB9FCCDFCCE" ID-GBIF-Occurrence="2235592111" collectingDate="1985-07-28" collectionCode="NSMT" collectorName="M. Takeda" location="Kantabako" municipality="Cebu Island" pageId="7" pageNumber="322" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="58F976C56B2FFF8EE315FCB9FB20FCB2" box="[1148,1198,788,812]" pageId="7" pageNumber="322" type="male">1 ♂</specimenCount>
, 
<specimenCount id="58F976C56B2FFF8EE3A9FCB9FAB6FCB3" box="[1216,1336,788,813]" pageId="7" pageNumber="322" type="juvenile">1 juvenile</specimenCount>
, 
<collectionCode id="28EE25896B2FFF8EE223FCB9FA17FCB2" box="[1354,1433,788,812]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="7" pageNumber="322">NSMT</collectionCode>
, 
<location id="4B20EB976B2FFF8EE7FEFC9AFE9DFCCE" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE7FEFC9AFE9DFCCE" box="[151,275,823,848]" municipality="Cebu Island" name="Kantabako" pageId="7" pageNumber="322">Kantabako</location>
, 
<collectingMunicipality id="AE2427366B2FFF8EE677FC9AFE29FCCE" box="[286,423,823,848]" pageId="7" pageNumber="322">Cebu Island</collectingMunicipality>
, coll. 
<collectorName id="E30AD89A6B2FFF8EE682FC95FDECFCCE" box="[491,610,823,848]" pageId="7" pageNumber="322">M. Takeda</collectorName>
<emphasis id="7C8B615E6B2FFF8EE500FC94FD2FFCCE" box="[617,673,823,848]" italics="true" pageId="7" pageNumber="322">et al.</emphasis>
, 
<date id="3A419B8C6B2FFF8EE5C6FC95FCCDFCCE" box="[687,835,823,848]" pageId="7" pageNumber="322" value="1985-07-28">
<collectingDate id="2A0562646B2FFF8EE5C6FC95FCCDFCCE" box="[687,835,823,848]" pageId="7" pageNumber="322" value="1985-07-28">28 July 1985</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="FE97B7116B2FFF8EE424FC9AFB22FCCE" ID-GBIF-Occurrence="2235592104" box="[845,1196,823,848]" country="Philippines" location="All" pageId="7" pageNumber="322" specimenCount="1" typeStatus="paratype">
<location id="4B20EB976B2FFF8EE424FC9AFCFFFCCE" LSID="urn:lsid:plazi:treatment:C6560C5A6B2FFF82E7FEFF3AFF62FC56:4B20EB976B2FFF8EE424FC9AFCFFFCCE" box="[845,881,823,848]" country="Philippines" name="All" pageId="7" pageNumber="322">All</location>
locations in the 
<collectingCountry id="36E8FDDC6B2FFF8EE343FC9AFB22FCCE" box="[1066,1196,823,848]" name="Philippines" pageId="7" pageNumber="322">Philippines</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="06E5EEC76B2FFF81E7AEFCF1FB83FEDE" lastPageId="8" lastPageNumber="323" pageId="7" pageNumber="322" type="description">
<paragraph id="4E40BD4C6B2FFF8EE7AEFCF1FBA3FA0E" blockId="7.[151,1437,356,2037]" pageId="7" pageNumber="322">
<emphasis id="7C8B615E6B2FFF8EE7AEFCF1FEDFFCEB" bold="true" box="[199,337,860,885]" pageId="7" pageNumber="322">Description</emphasis>
. Carapace trapezoidal, widest at anterior quarter, dorsal surface convex longitudinally, dorsoventrally depressed, regions distinct (
<figureCitation id="D6C4A1C96B2FFF8EE51CFCD2FD77FC09" box="[629,761,895,920]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Figs. 4A, B</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE46CFCD2FCC8FC06" box="[773,838,895,920]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="7" pageNumber="322">6B, F</figureCitation>
). Frontal region sloping anteroventrally; anterolateral regions inflated dorsolaterally, rugose; cervical grooves deep; H-shaped gastric groove deep; epigastric cristae distinct, edges rough, separated by distinct median furrow; postorbital cristae, sharp but low; epigastric and postorbital cristae, confluent; epibranchial teeth and postorbital cristae not confluent, separated by gaps (
<figureCitation id="D6C4A1C96B2FFF8EE254FC41FF22FBB6" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Fig. 4A, B</figureCitation>
). Frontal margin broadly protruded, two lobes clearly separated with broad median concavity; external orbital tooth produced anteriorly, outer margin longer than inner margin; epibranchial tooth distinct, triangular, well separated from external orbital tooth, margin gently curving dorsally, tapering anteriorly; anterolateral margin convex, crest low but distinct when viewed laterally, not clearly demarcated from posterolateral margin; posterolateral margin gently concave, converging gradually towards posterior margin of carapace (
<figureCitation id="D6C4A1C96B2FFF8EE38DFB32FAE6FB26" box="[1252,1384,1183,1208]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Figs. 4A, B</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE21DFB32FF63FB42" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="7" pageNumber="322">6B, D, F, H</figureCitation>
). Frontal median triangle almost complete; dorsal and lateral margins distinct, smooth; dorsal margin more produced anteriorly than lateral margins (
<figureCitation id="D6C4A1C96B2FFF8EE500FB4AFD31FA9E" box="[617,703,1255,1280]" captionStart="FIGURE 5" captionStartId="9.[151,250,1401,1423]" captionTargetBox="[153,1430,213,1363]" captionTargetId="figure@9.[150,1436,193,1380]" captionTargetPageId="9" captionText="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="7" pageNumber="322">Fig. 5F</figureCitation>
); orbit well demarcated; supraorbital margin smooth; infraorbital margin protruded anteriorly, granulated; outer edge reaching and fused with anterolateral margin; suborbital and subbranchial regions covered with scattered oblique long, short striae; pterygostomial region smooth with oblique ridges on upper outer part (
<figureCitation id="D6C4A1C96B2FFF8EE689FAF9FDC6FAF2" box="[480,584,1364,1389]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Figs. 4C</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE530FAF9FDF4FAF3" box="[601,634,1364,1389]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="7" pageNumber="322">6F</figureCitation>
). Posterior margin of epistome with three lobes, median lobe large, subtriangular, with notch; lateral lobes wider and protruded (
<figureCitation id="D6C4A1C96B2FFF8EE456FADAFC2FFA0E" box="[831,929,1399,1424]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Figs. 4C</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE4C4FADAFC5CFA0E" box="[941,978,1399,1424]" captionStart="FIGURE 5" captionStartId="9.[151,250,1401,1423]" captionTargetBox="[153,1430,213,1363]" captionTargetId="figure@9.[150,1436,193,1380]" captionTargetPageId="9" captionText="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="7" pageNumber="322">5H</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE4B6FADAFBAEFA0E" box="[991,1056,1399,1424]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="7" pageNumber="322">6F, H</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B2FFF8EE7AEFA30FCD6F9BE" blockId="7.[151,1437,356,2037]" pageId="7" pageNumber="322">
Eyes well developed, occupying entire orbit (
<figureCitation id="D6C4A1C96B2FFF8EE58DFA31FC17FA2A" box="[740,921,1436,1461]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Figs. 4A, B, C</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE4C2FA31FBCEFA2A" box="[939,1088,1436,1461]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="7" pageNumber="322">6B, D, F, H</figureCitation>
). Ischium of third maxilliped rectangular, bearing distinct submedian sulcus close to mesial margin; merus quadrate, anteroexternal angle convex, anterior margin slightly concave; tip of exopod reaches midpoint of outer margin of merus, with long flagellum reaching slightly beyond mesial margin (
<figureCitation id="D6C4A1C96B2FFF8EE5BAF9AAFCC5F9BE" box="[723,843,1543,1568]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Fig. 4C, E</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B2FFF8EE7AEF981FC76F92E" blockId="7.[151,1437,356,2037]" pageId="7" pageNumber="322">
Adult male chelipeds stout, subequal; dorsal margin of merus serrated, dorsal margin with distinct subdistal tooth; carpus with strong distal inner angle, flattened dorsoventrally, laterally fringed with proximal teeth; palm surface smooth, equal in length with finger; ventral margin granulated; fingers robust, cutting edges with teeth of various sizes, largest medially, smaller on distal and proximal parts (
<figureCitation id="D6C4A1C96B2FFF8EE4FCF93AFC65F92E" box="[917,1003,1687,1712]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Fig. 4F</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B2FFF8EE7AEF911FD9EF816" blockId="7.[151,1437,356,2037]" pageId="7" pageNumber="322">
Ambulatory legs not elongate (
<figureCitation id="D6C4A1C96B2FFF8EE549F911FD0BF94A" box="[544,645,1724,1749]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Figs. 4A</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE5F9F911FD5BF94A" box="[656,725,1724,1748]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: A–F = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="7" pageNumber="322">9C, D</figureCitation>
), second leg longest; anterior margin of merus serrated, without subdistal tooth or spine, posterior margins smooth; carpus short, with longitudinal submedian ridge on dorsal and ventral surfaces of all legs except on fourth leg that lacks ventral ridge, dorsal ridge barely visible, widened distally, outer margins indistinctly serrated; propodus with rows of spines on inner and outer margins, shorter on outer margin; dactylus with rows of spines on all margins, spines of both outer and inner margins of dactylus almost equal in length (
<figureCitation id="D6C4A1C96B2FFF8EE624F8C2FE3FF816" box="[333,433,1903,1928]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Figs. 4A</figureCitation>
; 
<figureCitation id="D6C4A1C96B2FFF8EE6D4F8DDFD8AF816" box="[445,516,1904,1928]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: A–F = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="7" pageNumber="322">9C, D</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B2FFF81E7AEF839FBE8FF4A" blockId="7.[151,1437,356,2037]" lastBlockId="8.[151,1436,151,396]" lastPageId="8" lastPageNumber="323" pageId="7" pageNumber="322">
Male sternopleonal cavity reaching to level of proximal quarter of coxae of chelipeds (
<figureCitation id="D6C4A1C96B2FFF8EE3C2F839FA86F832" box="[1195,1288,1940,1965]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="7" pageNumber="322">Fig. 4D</figureCitation>
). Adult male pleon narrow, T-shaped; somite 1 very short, proximal and distal margins sinuous; somite 2 transversely subrectangular; somites 3–5 narrow gradually; lateral margins of somite 3 convex, lateral margins of somites 4 and 5 slightly concave; somite 6 rectangular, longer than broad, lateral margins slightly concave; telson subtriangular, longer than broad, lateral margin concave medially, rounded distally (
<figureCitation id="D6C4A1C96B20FF81E4CAFF11FB89FF4A" box="[931,1031,188,213]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="8" pageNumber="323">Figs. 4D</figureCitation>
; 
<figureCitation id="D6C4A1C96B20FF81E37AFF11FBD7FF4A" box="[1043,1113,188,213]" captionStart="FIGURE 5" captionStartId="9.[151,250,1401,1423]" captionTargetBox="[153,1430,213,1363]" captionTargetId="figure@9.[150,1436,193,1380]" captionTargetPageId="9" captionText="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="8" pageNumber="323">5F, M</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B20FF81E7AEFF4DFB83FEDE" blockId="8.[151,1436,151,396]" pageId="8" pageNumber="323">
G1 relatively slender; subterminal segment, outer margin concave, almost straight, tapering; terminal segment gently bent outwards, outer margin concave, tapering, cylindrical, slightly setose (
<figureCitation id="D6C4A1C96B20FF81E344FEA9FBE0FE83" box="[1069,1134,260,285]" captionStart="FIGURE 5" captionStartId="9.[151,250,1401,1423]" captionTargetBox="[153,1430,213,1363]" captionTargetId="figure@9.[150,1436,193,1380]" captionTargetPageId="9" captionText="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="8" pageNumber="323">Fig. 5</figureCitation>
A–D, I–L). G2 subequal in length to G1, distal segment long, about half length of basal segment (
<figureCitation id="D6C4A1C96B20FF81E4C0FE8AFB8EFEDE" box="[937,1024,295,320]" captionStart="FIGURE 5" captionStartId="9.[151,250,1401,1423]" captionTargetBox="[153,1430,213,1363]" captionTargetId="figure@9.[150,1436,193,1380]" captionTargetPageId="9" captionText="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="8" pageNumber="323">Fig. 5E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="06E5EEC76B20FF81E7AEFEE2FE0CFE12" pageId="8" pageNumber="323" type="etymology">
<paragraph id="4E40BD4C6B20FF81E7AEFEE2FE0CFE12" blockId="8.[151,1436,151,396]" pageId="8" pageNumber="323">
<emphasis id="7C8B615E6B20FF81E7AEFEE2FEC8FEF6" bold="true" box="[199,326,335,360]" pageId="8" pageNumber="323">Etymology</emphasis>
. The species is named after the island of 
<collectingRegion id="8C3B73AE6B20FF81E47DFEE2FCC1FEF6" box="[788,847,335,360]" country="Philippines" name="Cebu" pageId="8" pageNumber="323">Cebu</collectingRegion>
where it is apparently endemic. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<caption id="1A80EDC46B20FF81E7FEF835FBFAF875" httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="8" pageNumber="323" startId="8.[151,250,1944,1966]" targetBox="[150,1436,448,1922]" targetPageId="8">
<paragraph id="4E40BD4C6B20FF81E7FEF835FBFAF875" blockId="8.[151,1436,1944,2027]" pageId="8" pageNumber="323">
<emphasis id="7C8B615E6B20FF81E7FEF835FE98F833" bold="true" box="[151,278,1944,1966]" pageId="8" pageNumber="323">FIGURE 4.</emphasis>
<taxonomicName id="89FFC6CF6B20FF81E676F835FE75F830" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[287,507,1944,1966]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="323" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B20FF81E676F835FE75F830" box="[287,507,1944,1966]" italics="true" pageId="8" pageNumber="323">Sundathelphusa cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B20FF81E56DF835FDD4F833" bold="true" box="[516,602,1944,1965]" pageId="8" pageNumber="323">
<taxonomicNameLabel id="67B8DC256B20FF81E56DF835FDD4F833" box="[516,602,1944,1965]" pageId="8" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela.
</paragraph>
</caption>
<caption id="1A80EDC46B21FF80E7FEFAD4FE6EF994" httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="9" pageNumber="324" startId="9.[151,250,1401,1423]" targetBox="[153,1430,213,1363]" targetPageId="9">
<paragraph id="4E40BD4C6B21FF80E7FEFAD4FE6EF994" blockId="9.[151,1436,1401,1546]" pageId="9" pageNumber="324">
<emphasis id="7C8B615E6B21FF80E7FEFAD4FE83FA11" bold="true" box="[151,269,1401,1423]" pageId="9" pageNumber="324">FIGURE 5</emphasis>
. 
<taxonomicName id="89FFC6CF6B21FF80E670FAD4FE7FFA11" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[281,497,1401,1423]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B21FF80E670FAD4FE7FFA11" box="[281,497,1401,1423]" italics="true" pageId="9" pageNumber="324">Sundathelphusa cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B21FF80E69EFAD7FDC4FA11" bold="true" box="[503,586,1402,1423]" pageId="9" pageNumber="324">
<taxonomicNameLabel id="67B8DC256B21FF80E69EFAD7FDC4FA11" box="[503,586,1402,1423]" pageId="9" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm.
</paragraph>
</caption>
<subSubSection id="06E5EEC76B21FF82E7AEF997FF62FC56" lastPageId="11" lastPageNumber="326" pageId="9" pageNumber="324" type="discussion">
<paragraph id="4E40BD4C6B21FF80E7AEF997FB84F899" blockId="9.[151,1436,1594,2015]" pageId="9" pageNumber="324">
<emphasis id="7C8B615E6B21FF80E7AEF997FEBAF9CD" bold="true" box="[199,308,1594,1619]" pageId="9" pageNumber="324">Remarks</emphasis>
. 
<taxonomicName id="89FFC6CF6B21FF80E62DF997FDBAF9CD" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[324,564,1594,1619]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B21FF80E62DF997FDBAF9CD" box="[324,564,1594,1619]" italics="true" pageId="9" pageNumber="324">Sundathelphusa cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B21FF80E554F997FD17F9CD" bold="true" box="[573,665,1594,1619]" pageId="9" pageNumber="324">
<taxonomicNameLabel id="67B8DC256B21FF80E554F997FD17F9CD" box="[573,665,1594,1619]" pageId="9" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has long been confused with 
<taxonomicName id="89FFC6CF6B21FF80E49BF996FB0DF9CD" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1010,1155,1594,1619]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B21FF80E49BF996FB0DF9CD" box="[1010,1155,1594,1619]" italics="true" pageId="9" pageNumber="324">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B21FF80E3E5F996FA94F9CD" box="[1164,1306,1594,1619]" pageId="9" pageNumber="324" sensu="stricto">sensu stricto</taxonomicNameLabel>
because of their close morphological resemblance (e.g. 
<bibRefCitation id="2A6EC0BD6B21FF80E5E2F9F3FC90F9E9" author="Takeda, M." box="[651,798,1630,1655]" pageId="9" pageNumber="324" pagination="89 - 106" refId="ref9849" refString="Takeda, M. (1987) Freshwater crabs from Peninsu 1 ar Malaysia, with notes on some species from the Philippines. In: Kawashima, K. (Ed.), Paragonimus in Asia: Biology, Genetic Variation and Speciation. Paragonimus Research Report, Kyushu University, 1, pp. 89 - 106." type="journal article" year="1987">Takeda 1987</bibRefCitation>
; 
<bibRefCitation id="2A6EC0BD6B21FF80E443F9F2FC68F9E9" author="Ng, P. K. L. &amp; Sket, B." box="[810,998,1630,1655]" pageId="9" pageNumber="324" pagination="695 - 706" refId="ref9587" refString="Ng, P. K. L. &amp; Sket, B. (1996) The freshwater crab fauna (Crustacea: Decapoda: Brachyura) of the Philippines. IV. On the collection of Parathelphusidae from Bohol. Proceeding of the Biological Society of Washington, 109, 695 - 706." type="journal article" year="1996">Ng &amp; Sket 1996</bibRefCitation>
). An initial unpublished DNA analysis of the large subunit rRNA (16S) of this group of species done by the first author suggests that the species populations from the islands of 
<collectingRegion id="8C3B73AE6B21FF80E697F90AFDCBF921" box="[510,581,1703,1727]" country="Philippines" name="Samar" pageId="9" pageNumber="324">Samar</collectingRegion>
and 
<collectingRegion id="8C3B73AE6B21FF80E514F90BFD36F921" box="[637,696,1702,1727]" country="Philippines" name="Cebu" pageId="9" pageNumber="324">Cebu</collectingRegion>
represent distinct genetic groups. The detailed study of the many specimens of both species on hand shows that there is a suite of small but distinct morphological characters that can be used to separate 
<taxonomicName id="89FFC6CF6B21FF80E61BF942FD8FF899" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[370,513,1774,1799]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B21FF80E61BF942FD8FF899" box="[370,513,1774,1799]" italics="true" pageId="9" pageNumber="324">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B21FF80E561F942FD1DF899" box="[520,659,1774,1799]" pageId="9" pageNumber="324" sensu="stricto">sensu stricto</taxonomicNameLabel>
and 
<taxonomicName id="89FFC6CF6B21FF80E5A3F942FC95F899" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[714,795,1774,1799]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B21FF80E5A3F942FC95F899" box="[714,795,1774,1799]" italics="true" pageId="9" pageNumber="324">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B21FF80E44BF943FCF5F899" bold="true" box="[802,891,1774,1799]" pageId="9" pageNumber="324">
<taxonomicNameLabel id="67B8DC256B21FF80E44BF943FCF5F899" box="[802,891,1774,1799]" pageId="9" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D6C4A1C96B21FF80E4E3F943FC42F899" box="[906,972,1774,1799]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="9" pageNumber="324">Fig. 6</figureCitation>
A–J).
</paragraph>
<paragraph id="4E40BD4C6B21FF83E7AEF8BFFCC0FF4B" blockId="9.[151,1436,1594,2015]" lastBlockId="10.[151,1436,151,213]" lastPageId="10" lastPageNumber="325" pageId="9" pageNumber="324">
When viewed laterally, the side of the external orbital tooth gently curves dorsally in 
<taxonomicName id="89FFC6CF6B21FF80E314F8BEFB40F8B5" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1149,1230,1810,1835]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B21FF80E314F8BEFB40F8B5" box="[1149,1230,1810,1835]" italics="true" pageId="9" pageNumber="324">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B21FF80E3BDF8BFFAA0F8B5" bold="true" box="[1236,1326,1810,1835]" pageId="9" pageNumber="324">
<taxonomicNameLabel id="67B8DC256B21FF80E3BDF8BFFAA0F8B5" box="[1236,1326,1810,1835]" pageId="9" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D6C4A1C96B21FF80E255F8BFFA1DF8B5" box="[1340,1427,1810,1835]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="9" pageNumber="324">Fig. 6D</figureCitation>
) while in 
<taxonomicName id="89FFC6CF6B21FF80E790F89AFE07F8D1" authorityName="Husana &amp; Ng" authorityYear="2019" box="[249,393,1846,1871]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B21FF80E790F89AFE07F8D1" box="[249,393,1846,1871]" italics="true" pageId="9" pageNumber="324">S. philippina</emphasis>
</taxonomicName>
, the entire tooth and margin is almost flat (
<figureCitation id="D6C4A1C96B21FF80E406F89BFC46F8D1" box="[879,968,1846,1871]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="9" pageNumber="324">Fig. 6C</figureCitation>
). The crest along the anterolateral margin is relatively sharper and more clearly defined in 
<taxonomicName id="89FFC6CF6B21FF80E5D0F8F6FC84F8ED" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[697,778,1882,1907]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B21FF80E5D0F8F6FC84F8ED" box="[697,778,1882,1907]" italics="true" pageId="9" pageNumber="324">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B21FF80E47BF8F7FCE2F8ED" bold="true" box="[786,876,1882,1907]" pageId="9" pageNumber="324">
<taxonomicNameLabel id="67B8DC256B21FF80E47BF8F7FCE2F8ED" box="[786,876,1882,1907]" pageId="9" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D6C4A1C96B21FF80E412F8F7FB8FF8EC" box="[891,1025,1882,1907]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="9" pageNumber="324">Figs. 4A, B</figureCitation>
; 
<figureCitation id="D6C4A1C96B21FF80E364F8F7FBA1F8ED" box="[1037,1071,1882,1907]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="9" pageNumber="324">6B</figureCitation>
) than in 
<taxonomicName id="89FFC6CF6B21FF80E3FDF8F6FAADF8ED" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1172,1315,1882,1907]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B21FF80E3FDF8F6FAADF8ED" box="[1172,1315,1882,1907]" italics="true" pageId="9" pageNumber="324">S. philippina</emphasis>
</taxonomicName>
(
<figureCitation id="D6C4A1C96B21FF80E25AF8F7FF22F808" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, C–F, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="9" pageNumber="324">Figs. 1A, B</figureCitation>
; 
<figureCitation id="D6C4A1C96B21FF80E7D0F8D3FF51F809" box="[185,223,1918,1943]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="9" pageNumber="324">6A</figureCitation>
). When viewed frontally and laterally, the branchial and gastric regions of the carapace of 
<taxonomicName id="89FFC6CF6B21FF80E38EF8D2FAB7F809" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1255,1337,1918,1943]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B21FF80E38EF8D2FAB7F809" box="[1255,1337,1918,1943]" italics="true" pageId="9" pageNumber="324">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B21FF80E228F8D3FA12F809" bold="true" box="[1345,1436,1918,1943]" pageId="9" pageNumber="324">
<taxonomicNameLabel id="67B8DC256B21FF80E228F8D3FA12F809" box="[1345,1436,1918,1943]" pageId="9" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are also proportionately higher than those of 
<taxonomicName id="89FFC6CF6B21FF80E5C9F80EFCBCF825" authorityName="Husana &amp; Ng" authorityYear="2019" box="[672,818,1954,1979]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B21FF80E5C9F80EFCBCF825" box="[672,818,1954,1979]" italics="true" pageId="9" pageNumber="324">S. philippina</emphasis>
</taxonomicName>
(
<figureCitation id="D6C4A1C96B21FF80E42DF80FFC33F824" box="[836,957,1954,1979]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="9" pageNumber="324">Fig. 6D, F</figureCitation>
vs. 
<figureCitation id="D6C4A1C96B21FF80E499F80FFBE4F825" box="[1008,1130,1954,1979]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="9" pageNumber="324">Fig. 6C, E</figureCitation>
) of comparable sizes. The oblique striations on the anterolateral regions are generally more pronounced in 
<taxonomicName id="89FFC6CF6B21FF80E347F86AFB0FF841" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1070,1153,1990,2015]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B21FF80E347F86AFB0FF841" box="[1070,1153,1990,2015]" italics="true" pageId="9" pageNumber="324">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B21FF80E3E3F86BFB68F841" bold="true" box="[1162,1254,1990,2015]" pageId="9" pageNumber="324">
<taxonomicNameLabel id="67B8DC256B21FF80E3E3F86BFB68F841" box="[1162,1254,1990,2015]" pageId="9" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D6C4A1C96B21FF80E39EF86BFADFF841" box="[1271,1361,1990,2015]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="9" pageNumber="324">Fig. 6B</figureCitation>
); in 
<taxonomicName id="89FFC6CF6B21FF83E2EEF86AFE84FF2E" authorityName="Husana &amp; Ng" authorityYear="2019" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="10" lastPageNumber="325" order="Decapoda" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B21FF83E2EEF86AFE84FF2E" italics="true" lastPageId="10" lastPageNumber="325" pageId="9" pageNumber="324">S. philippina</emphasis>
</taxonomicName>
, these are relatively lower and the surface appears much smoother (
<figureCitation id="D6C4A1C96B22FF83E351FF3AFB1BFF2E" box="[1080,1173,151,176]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="10" pageNumber="325">Fig. 6B</figureCitation>
). These characters are consistent across all size-groups and for both sexes examined.
</paragraph>
<caption id="1A80EDC46B22FF83E7FEF892FEC9F870" httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="10" pageNumber="325" startId="10.[151,250,1855,1877]" targetBox="[150,1436,265,1833]" targetPageId="10">
<paragraph id="4E40BD4C6B22FF83E7FEF892FEC9F870" blockId="10.[151,1436,1855,2030]" pageId="10" pageNumber="325">
<emphasis id="7C8B615E6B22FF83E7FEF892FE9DF8CA" bold="true" box="[151,275,1855,1877]" pageId="10" pageNumber="325">FIGURE 6.</emphasis>
A, C, E, G, I, J, 
<taxonomicName id="89FFC6CF6B22FF83E6ABF892FC21F8CB" authority="(von Martens, 1868)" authorityName="von Martens" authorityYear="1868" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[450,943,1855,1877]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B22FF83E6ABF892FD5DF8CB" box="[450,723,1855,1877]" italics="true" pageId="10" pageNumber="325">Sundathelphusa philippina</emphasis>
(von Martens, 1868)
</taxonomicName>
, ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, 
<taxonomicName id="89FFC6CF6B22FF83E6FBF8F3FE55F8ED" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[402,475,1885,1907]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B22FF83E6FBF8F3FE55F8ED" box="[402,475,1885,1907]" italics="true" pageId="10" pageNumber="325">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B22FF83E688F8F3FDBFF8ED" bold="true" box="[481,561,1886,1907]" pageId="10" pageNumber="325">
<taxonomicNameLabel id="67B8DC256B22FF83E688F8F3FDBFF8ED" box="[481,561,1886,1907]" pageId="10" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm.
</paragraph>
</caption>
<paragraph id="4E40BD4C6B23FF82E7AEFF3AFE8CFE4E" blockId="11.[151,1436,151,968]" pageId="11" pageNumber="326">
Another useful character that works in most cases is the structure of the median lobe of the posterior margin of the epistome. In 
<taxonomicName id="89FFC6CF6B23FF82E63AFF10FE2BFF4B" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[339,421,188,213]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B23FF82E63AFF10FE2BFF4B" box="[339,421,188,213]" italics="true" pageId="11" pageNumber="326">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B23FF82E6C5FF11FD88FF4B" bold="true" box="[428,518,188,213]" pageId="11" pageNumber="326">
<taxonomicNameLabel id="67B8DC256B23FF82E6C5FF11FD88FF4B" box="[428,518,188,213]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the lateral margins are prominently cristate and where they meet at the tip, there is almost always a deep notch, giving it a bilobed appearance (
<figureCitation id="D6C4A1C96B23FF82E403FF72FC41FF66" box="[874,975,223,248]" captionStart="FIGURE 4" captionStartId="8.[151,250,1944,1966]" captionTargetBox="[150,1436,448,1922]" captionTargetId="figure@8.[150,1436,448,1922]" captionTargetPageId="8" captionText="FIGURE 4. Sundathelphusa cebu sp. nov. A, C–F, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; B, paratype ♂ (largest 39.3 × 31.4 mm), ZRC 2017.1062, Cebu. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637374/files/figure.png" pageId="11" pageNumber="326">Figs. 4C</figureCitation>
; 
<figureCitation id="D6C4A1C96B23FF82E4B4FF72FB8CFF66" box="[989,1026,223,248]" captionStart="FIGURE 5" captionStartId="9.[151,250,1401,1423]" captionTargetBox="[153,1430,213,1363]" captionTargetId="figure@9.[150,1436,193,1380]" captionTargetPageId="9" captionText="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="11" pageNumber="326">5G</figureCitation>
; 
<figureCitation id="D6C4A1C96B23FF82E379FF72FBDDFF66" box="[1040,1107,223,248]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="11" pageNumber="326">6F, H</figureCitation>
). In 
<taxonomicName id="89FFC6CF6B23FF82E3E0FF4DFA94FF66" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[1161,1306,223,248]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B23FF82E3E0FF4DFA94FF66" box="[1161,1306,223,248]" italics="true" pageId="11" pageNumber="326">S. philippina</emphasis>
</taxonomicName>
, the lateral margins of the median lobe are not as prominent and where they meet at the tip, they appear to be more confluent, hardly appearing bilobed (
<figureCitation id="D6C4A1C96B23FF82E6AFFE8AFDA4FEDE" box="[454,554,295,320]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, C–F, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="11" pageNumber="326">Figs. 1C</figureCitation>
, 
<figureCitation id="D6C4A1C96B23FF82E550FE85FDD5FEDE" box="[569,603,296,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). A–G, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H– K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; L–O, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; P–T, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 3–6 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; U–W, male pleonal somites 4–6 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="11" pageNumber="326">2G</figureCitation>
, 
<figureCitation id="D6C4A1C96B23FF82E500FE8AFD01FEDE" box="[617,655,295,320]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="11" pageNumber="326">6G</figureCitation>
). The G1s differ slightly in the curvature and shape, with that of 
<taxonomicName id="89FFC6CF6B23FF82E2EEFE85FE84FEFB" baseAuthorityName="von Martens" baseAuthorityYear="1868" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B23FF82E2EEFE85FE84FEFB" italics="true" pageId="11" pageNumber="326">S. philippina</emphasis>
</taxonomicName>
being usually relatively straighter (
<figureCitation id="D6C4A1C96B23FF82E5F5FEE1FCD0FEFA" box="[668,862,332,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). A–G, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H– K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; L–O, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; P–T, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 3–6 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; U–W, male pleonal somites 4–6 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="11" pageNumber="326">Figs. 2A, B, H, I</figureCitation>
; 
<figureCitation id="D6C4A1C96B23FF82E402FEE1FC05FEFA" box="[875,907,332,356]" captionStart="FIGURE 3" captionStartId="5.[151,250,1509,1531]" captionTargetBox="[150,1436,171,1487]" captionTargetId="figure@5.[150,1436,171,1488]" captionTargetPageId="5" captionText="FIGURE 3. Sundathelphusa philippina (von Martens, 1868), variation in male carapace and G1 form. A, A’, 14.1 × 12.3 mm; B, 16.9 × 13.9 mm; C, 18.0 × 15.0 mm; D, D’, 20.3 × 16.7 mm; E, E’, 30.8 × 24.8 mm [all ZRC 2016.0107, Samar]. A–E, overall habitus; A’, D’, E’, left G1. A–E same scale; A’, D’, E’ same scale." httpUri="https://zenodo.org/record/2637372/files/figure.png" pageId="11" pageNumber="326">3F</figureCitation>
; 
<figureCitation id="D6C4A1C96B23FF82E4F1FEE1FC44FEFA" box="[920,970,332,357]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="11" pageNumber="326">
<date id="3A419B8C6B23FF82E4F1FEE1FC3FFEFB" box="[920,945,332,357]" pageId="11" pageNumber="326">6I</date>
, J
</figureCitation>
) while in 
<taxonomicName id="89FFC6CF6B23FF82E354FEE0FB1EFEFB" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1085,1168,332,357]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B23FF82E354FEE0FB1EFEFB" box="[1085,1168,332,357]" italics="true" pageId="11" pageNumber="326">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B23FF82E3FEFEE1FB7FFEFB" bold="true" box="[1175,1265,332,357]" pageId="11" pageNumber="326">
<taxonomicNameLabel id="67B8DC256B23FF82E3FEFEE1FB7FFEFB" box="[1175,1265,332,357]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, it is relatively more curved (
<figureCitation id="D6C4A1C96B23FF82E65DFEC2FE64FE19" box="[308,490,367,392]" captionStart="FIGURE 5" captionStartId="9.[151,250,1401,1423]" captionTargetBox="[153,1430,213,1363]" captionTargetId="figure@9.[150,1436,193,1380]" captionTargetPageId="9" captionText="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="11" pageNumber="326">Figs. 5A, B, I, J</figureCitation>
; 
<figureCitation id="D6C4A1C96B23FF82E691FEC2FDB2FE16" box="[504,572,367,392]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="11" pageNumber="326">6K, L</figureCitation>
). The differences in the G1 structures, however, are not always reliable as in some specimens of 
<taxonomicName id="89FFC6CF6B23FF82E618FE38FE71FE33" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[369,511,404,429]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B23FF82E618FE38FE71FE33" box="[369,511,404,429]" italics="true" pageId="11" pageNumber="326">S. philippina</emphasis>
</taxonomicName>
, the G1 also appears to be relatively more curved (
<figureCitation id="D6C4A1C96B23FF82E35DFE39FB66FE32" box="[1076,1256,404,429]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). A–G, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H– K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; L–O, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; P–T, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 3–6 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; U–W, male pleonal somites 4–6 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="11" pageNumber="326">Fig. 2L, M, P, Q</figureCitation>
vs. 
<figureCitation id="D6C4A1C96B23FF82E27CFE39FF43FE51" captionStart="FIGURE 5" captionStartId="9.[151,250,1401,1423]" captionTargetBox="[153,1430,213,1363]" captionTargetId="figure@9.[150,1436,193,1380]" captionTargetPageId="9" captionText="FIGURE 5. Sundathelphusa cebu sp. nov. A–H, holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; I–M, paratype ♂ (29.3 × 24.0 mm), ZRC 2017.1061, Cebu. A, I, ventral view of left G1; B, J, dorsal view of left G1; C, K, ventral view of terminal segment of left G1; D, L, dorsal view of terminal segment of left G1; E, left G2; F, frontal median triangle; G, posterior margin of epistome; H, male pleonal somites 3–6 and telson; M, pleonal somites 4–6 and telson. Scales: A, B, E, F, G = 1.0 mm; C, D, K, L = 0.5 mm; H, M = 3.0 mm." httpUri="https://zenodo.org/record/2637376/files/figure.png" pageId="11" pageNumber="326">Figs. 5A, C, G, H</figureCitation>
; 
<figureCitation id="D6C4A1C96B23FF82E7B0FE1AFF7BFE4E" box="[217,245,439,464]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="11" pageNumber="326">6J</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B23FF82E7AEFE71FD67FD36" blockId="11.[151,1436,151,968]" pageId="11" pageNumber="326">
The maximum adult sizes of 
<taxonomicName id="89FFC6CF6B23FF82E573FE70FDE3FE6B" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[538,621,476,501]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B23FF82E573FE70FDE3FE6B" box="[538,621,476,501]" italics="true" pageId="11" pageNumber="326">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B23FF82E51FFE71FD5CFE6B" bold="true" box="[630,722,476,501]" pageId="11" pageNumber="326">
<taxonomicNameLabel id="67B8DC256B23FF82E51FFE71FD5CFE6B" box="[630,722,476,501]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="89FFC6CF6B23FF82E467FE70FC2EFE6B" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[782,928,476,501]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B23FF82E467FE70FC2EFE6B" box="[782,928,476,501]" italics="true" pageId="11" pageNumber="326">S. philippina</emphasis>
</taxonomicName>
appear to differ. The largest specimen of 
<taxonomicName id="89FFC6CF6B23FF82E2EEFE70FE84FD86" baseAuthorityName="von Martens" baseAuthorityYear="1868" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B23FF82E2EEFE70FE84FD86" italics="true" pageId="11" pageNumber="326">S. philippina</emphasis>
</taxonomicName>
examined is a male 53.4 × 
<quantity id="890710A96B23FF82E52BFE52FD28FD89" box="[578,678,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" pageId="11" pageNumber="326" unit="mm" value="42.5">42.5 mm</quantity>
(ZRC 2009.0409) from Binangkaan river in Camarines Sur, Luzon Island and adult females can also exceed 
<quantity id="890710A96B23FF82E503FD89FD37FDA2" box="[618,697,548,573]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="326" unit="mm" value="50.0">50 mm</quantity>
in carapace width. The largest male specimen of 
<taxonomicName id="89FFC6CF6B23FF82E380FD88FAB4FDA3" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1257,1338,548,573]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B23FF82E380FD88FAB4FDA3" box="[1257,1338,548,573]" italics="true" pageId="11" pageNumber="326">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B23FF82E22BFD89FA12FDA3" bold="true" box="[1346,1436,548,573]" pageId="11" pageNumber="326">
<taxonomicNameLabel id="67B8DC256B23FF82E22BFD89FA12FDA3" box="[1346,1436,548,573]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
measured only 40.5 × 32.0 mm (ZRC 2011.0002), from Kawasan Falls, Matutinao, Cebu Island, but specimens up to 
<quantity id="890710A96B23FF82E7DDFDC1FE8CFD1A" box="[180,258,620,644]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="326" unit="mm" value="30.0">30 mm</quantity>
in carapace width are already mature. The largest female examined is 52.2 × 
<quantity id="890710A96B23FF82E30DFDC1FB46FD1A" box="[1124,1224,620,644]" metricMagnitude="-2" metricUnit="m" metricValue="4.029999999999999" pageId="11" pageNumber="326" unit="mm" value="40.3">40.3 mm</quantity>
(ZRC 2017.1255), from Malbubog Vilage, near Cebu City, Cebu Island.
</paragraph>
<paragraph id="4E40BD4C6B23FF82E7AEFD19FF62FC56" blockId="11.[151,1436,151,968]" pageId="11" pageNumber="326">
The distributions of the three species treated in this paper generally correspond to biogeographic zones recognized in 
<bibRefCitation id="2A6EC0BD6B23FF82E65DFD75FE63FD6E" author="Ong, P. S. &amp; Afuang, L. E. &amp; Roseii-Ambal, R. G." box="[308,493,727,752]" pageId="11" pageNumber="326" refId="ref9642" refString="Ong, P. S., Afuang, L. E. &amp; Roseii-Ambal, R. G. (Eds.) (2002) Philippine biodiversity conservation priorities: a second iteration of the national biodiversity strategy and action plan. Department of Environment and Natural Resources-Protected Areas and Wildlife Bureau, Conservation International Philippines, Biodiversity Conservation Program, University of the Philippines Center for Integrative and Development Studies, and Foundation for the Philippine Environment, Quezon City, 113 pp." type="book" year="2002">Ong et al. (2002</bibRefCitation>
: 2, fig. 4). 
<taxonomicName id="89FFC6CF6B23FF82E50AFD7AFC1EFD6E" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[611,912,727,752]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B23FF82E50AFD7AFC1EFD6E" box="[611,912,727,752]" italics="true" pageId="11" pageNumber="326">Sundathelphusa philippina</emphasis>
</taxonomicName>
is known from the islands of 
<collectingRegion id="8C3B73AE6B23FF82E3B0FD75FAACFD6E" box="[1241,1314,728,752]" country="Philippines" name="Samar" pageId="11" pageNumber="326">Samar</collectingRegion>
, 
<collectingRegion id="8C3B73AE6B23FF82E244FD75FAE3FD6E" box="[1325,1389,728,752]" country="Philippines" name="Southern Leyte" pageId="11" pageNumber="326">Leyte</collectingRegion>
and southernmost Luzon. 
<collectingRegion id="8C3B73AE6B23FF82E6E2FD51FE5CFC8A" box="[395,466,764,788]" country="Philippines" name="Samar" pageId="11" pageNumber="326">Samar</collectingRegion>
and 
<collectingRegion id="8C3B73AE6B23FF82E560FD50FDC7FC8B" box="[521,585,765,789]" country="Philippines" name="Southern Leyte" pageId="11" pageNumber="326">Leyte</collectingRegion>
are part of Greater Mindanao Region, and while southern Luzon belongs to the Greater Luzon Region, its proximity to 
<collectingRegion id="8C3B73AE6B23FF82E5EAFC8DFD45FCA6" box="[643,715,800,824]" country="Philippines" name="Samar" pageId="11" pageNumber="326">Samar</collectingRegion>
means that these areas were probably connected during the last ice age 15,000–20,000 years ago. 
<taxonomicName id="89FFC6CF6B23FF82E57AFCE9FC8CFCC3" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[531,770,836,861]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B23FF82E57AFCE9FC8CFCC3" box="[531,770,836,861]" italics="true" pageId="11" pageNumber="326">Sundathelphusa cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B23FF82E460FCE9FCECFCC3" bold="true" box="[777,866,836,861]" pageId="11" pageNumber="326">
<taxonomicNameLabel id="67B8DC256B23FF82E460FCE9FCECFCC3" box="[777,866,836,861]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occurs in the Greater Negros-Panay Region, while 
<taxonomicName id="89FFC6CF6B23FF82E7FEFCC5FE89FC1E" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[151,263,872,896]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
<emphasis id="7C8B615E6B23FF82E7FEFCC5FE89FC1E" box="[151,263,872,896]" italics="true" pageId="11" pageNumber="326">S. quirino</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B23FF82E666FCCAFEE7FC1E" bold="true" box="[271,361,871,896]" pageId="11" pageNumber="326">
<taxonomicNameLabel id="67B8DC256B23FF82E666FCCAFEE7FC1E" box="[271,361,871,896]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is in the Greater Luzon Region. In any case, a number of 
<taxonomicName id="89FFC6CF6B23FF82E36DFCCAFB39FC1E" authorityName="Bott" authorityYear="1969" box="[1028,1207,871,896]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="326" phylum="Arthropoda" rank="genus">
<emphasis id="7C8B615E6B23FF82E36DFCCAFB39FC1E" box="[1028,1207,871,896]" italics="true" pageId="11" pageNumber="326">Sundathelphusa</emphasis>
</taxonomicName>
species from 
<collectingRegion id="8C3B73AE6B23FF82E23EFCCAFA15FC1E" box="[1367,1435,871,896]" country="Philippines" name="Bohol" pageId="11" pageNumber="326">Bohol</collectingRegion>
are known to have rapid speciation rates (see 
<bibRefCitation id="2A6EC0BD6B23FF82E5FDFC21FCEEFC3A" author="Klaus, S. &amp; Mendoza, J. C. E. &amp; Liew, J. H. &amp; Plath, M. &amp; Meier, R &amp; Yeo, D. C. J." box="[660,864,908,933]" pageId="11" pageNumber="326" pagination="20121098" refId="ref8874" refString="Klaus, S., Mendoza, J. C. E., Liew, J. H., Plath, M., Meier, R &amp; Yeo, D. C. J. (2013 a) Rapid evolution of troglomorphic characters suggests selection rather than neutral mutation as a driver of eye reduction in cave crabs. Biology Letters, 9, 20121098. https: // doi. org / 10.1098 / rsbl. 2012.1098" type="journal article" year="2013">
Klaus 
<emphasis id="7C8B615E6B23FF82E5B5FC20FC9AFC3B" box="[732,788,908,933]" italics="true" pageId="11" pageNumber="326">et al.</emphasis>
2013a
</bibRefCitation>
) and this is likely to be true for species in the other islands.
</paragraph>
</subSubSection>
</treatment>
</document>