treatments-xml/data/8C/70/BE/8C70BE3B24521975FF676E4AAE76FC53.xml

<document id="0688D2AE30BAD8C69502D79D66E73D73" ID="10.11646/zootaxa.4273.4.3SLASH11267" ID-CLB-Dataset="32891" ID-DOI="10.11646/zootaxa.4273.4.3" ID-GBIF-Dataset="0935deaf-8f18-434e-b4a0-0468cefc9577" ID-ISSN="1175-5326" ID-Zenodo-Dep="803704" ID-ZooBank="4CD96097-1016-47CB-8DC3-AD6E9EDA330C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496820946147" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2017" docId="8C70BE3B24521975FF676E4AAE76FC53" docLanguage="en" docName="zootaxa.4273.4.3SLASH11267.pdf" docOrigin="Zootaxa 4273 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tropostreptus hamatus Demange 1977, comb. nov." docType="treatment" docVersion="7" lastPageNumber="513" masterDocId="7049C64324551979FFF06F72AF21FF9B" masterDocTitle="A new East African genus of spirostreptid millipedes (Diplopoda, Spirostreptida, Spirostreptidae), with notes on their fungal ectoparasite Rickia gigas" masterLastPageNumber="530" masterPageNumber="501" pageNumber="508" updateTime="1698440246528" updateUser="ExternalLinkService">
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<mods:title id="59D0AAE9CC3B71BF8847DB76DFADE4EF">A new East African genus of spirostreptid millipedes (Diplopoda, Spirostreptida, Spirostreptidae), with notes on their fungal ectoparasite Rickia gigas</mods:title>
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<mods:date id="D2B88129DBF3DB1C4AEF38BC9984ECAB">2017</mods:date>
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<paragraph id="04660F2D2452197EFF676E4AAC3AFEC9" blockId="7.[151,795,312,371]" box="[151,795,312,338]" pageId="7" pageNumber="508">
<heading id="5F2EB8412452197EFF676E4AAC3AFEC9" bold="true" box="[151,795,312,338]" fontSize="11" level="1" pageId="7" pageNumber="508" reason="1">
<emphasis id="36ADD33F2452197EFF676E4AAC3AFEC9" bold="true" box="[151,795,312,338]" pageId="7" pageNumber="508">
<taxonomicName id="C3D974AE2452197EFF676E4AADA6FEC9" ID-CoL="59D9W" authority="Demange, 1977" authorityName="Demange" authorityYear="1977" box="[151,647,312,338]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="hamatus" status="comb. nov.">
<emphasis id="36ADD33F2452197EFF676E4AAE8CFEC9" bold="true" box="[151,429,312,338]" italics="true" pageId="7" pageNumber="508">Tropostreptus hamatus</emphasis>
(
<bibRefCitation id="604872DC2452197EFE4C6E4AAD5EFEC9" author="Demange" box="[444,639,312,338]" pageId="7" pageNumber="508" refString="Demange, J. - M. (1977) Nouveaux Myriapodes de Tanzanie. Description de deux especes nouvelles de Diplopodes. Affinites de quelques genres d'Oxydesmidae. Bulletin du Museum national d'Histoire naturelle 3 ° serie, n ° 451, Zoologie, 314, 507 - 518." type="journal article" year="1977">Demange, 1977</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="2D9E6E442452197EFD656E4AAC3AFEC9" box="[661,795,312,338]" pageId="7" pageNumber="508">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="04660F2D2452197EFF676E28ADF5FEE8" blockId="7.[151,795,312,371]" box="[151,724,346,371]" pageId="7" pageNumber="508">
<figureCitation id="9CE213A82452197EFF676E28AFFDFEE8" box="[151,220,346,371]" captionStart="FIGURE 1" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[200,1379,237,1858]" captionTargetId="figure@4.[151,1436,193,1911]" captionTargetPageId="4" captionText="FIGURE 1. Tropostreptus hamatus, live specimens. A: pink-legged specimen (East Usambara Mts), S. Kuhlmann photo. B: a copulating pair of the pink-legged form, S. Kuhlmann photo, C: darker, dark-legged specimen (Udzungwa Mts.), N. Scharff photo." httpUri="https://zenodo.org/record/803706/files/figure.png" pageId="7" pageNumber="508">Figs 1</figureCitation>
, 
<figureCitation id="9CE213A82452197EFF1A6E29AFDBFEE8" box="[234,250,347,371]" captionStart="FIGURE 3" captionStartId="7.[151,250,1364,1386]" captionTargetBox="[198,1394,742,1302]" captionTargetId="figure@7.[151,1436,699,1387]" captionTargetPageId="7" captionText="FIGURE 3. Tropostretus hamatus, specimens from E. Usambara Mts. A: male, B: female. Scales 1 mm. Notice produced antero-ventral corner of collum in male (arrow). A. Illum photo." httpUri="https://zenodo.org/record/803712/files/figure.png" pageId="7" pageNumber="508">3</figureCitation>
, 
<figureCitation id="9CE213A82452197EFEF66E29AE34FEE8" box="[262,277,347,371]" captionStart="FIGURE 4" captionStartId="9.[151,250,1673,1695]" captionTargetBox="[212,1376,195,1600]" captionTargetId="figure@9.[151,1436,159,1715]" captionTargetPageId="9" captionText="FIGURE 4. Tropostreptus spp., mouthparts. A: T. droides sp. nov., left mandibular gnathal lobe, dorsal / anterior view, B: T. kipunji sp. nov., right mandibular gnathal lobe, mesal view. C: T. hamatus, male, gnathochilarium and right mandible, oblique ventral view. — Scales 0.1 mm. me = mentum, mf = molar furrow, msl = mandibular stipital lobe, od = odontomere, pl = pectinate lamellae, pm = prementum, ps = psectromere, se = sectile edge." httpUri="https://zenodo.org/record/803714/files/figure.png" pageId="7" pageNumber="508">4</figureCitation>
C, 5A, B, D, 6, 7 A, B, 8 A, B, 11, 12 A
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62452197EFF676ED1AD73FE4C" pageId="7" pageNumber="508" type="reference_group">
<paragraph id="04660F2D2452197EFF676ED1AD73FE4C" blockId="7.[151,606,418,471]" pageId="7" pageNumber="508">
<treatmentCitationGroup id="24C928032452197EFF676ED1AD73FE4C" pageId="7" pageNumber="508">
<taxonomicName id="C3D974AE2452197EFF676ED1AE3DFE23" ID-CoL="7Q2ML" box="[151,284,419,440]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="36ADD33F2452197EFF676ED1AE3DFE23" box="[151,284,419,440]" italics="true" pageId="7" pageNumber="508">Spirostreptus</emphasis>
</taxonomicName>
sp. 1: 
<bibRefCitation id="604872DC2452197EFEAF6ED0AD7FFE23" author="Enghoff" box="[351,606,418,440]" pageId="7" pageNumber="508" refString="Enghoff, I. B. &amp; Enghoff, H. (1976) Notes on myriapods observed and collected in Tanzania and Kenya during the summer 1974. Xeroxed report, Zoological Museum, University of Copenhagen, Copenhagen, 19 pp." type="book" year="1976">Enghoff &amp; Enghoff 1976</bibRefCitation>
<taxonomicName id="C3D974AE2452197EFF676EB0AD73FE4C" ID-CoL="4Z8T4" authority="Demange, 1977: 517" authorityName="Demange" authorityPageNumber="517" authorityYear="1977" box="[151,594,449,471]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F2452197EFF676EB0AE59FE4C" box="[151,376,449,471]" italics="true" pageId="7" pageNumber="508">Spirostreptus hamatus</emphasis>
<treatmentCitation id="8578293C2452197EFE8E6EB0AD73FE4C" author="Demange" box="[382,594,449,471]" page="517" pageId="7" pageNumber="508" year="1977">
<bibRefCitation id="604872DC2452197EFE8E6EB0AD02FE4C" author="Demange" box="[382,547,449,471]" pageId="7" pageNumber="508" refString="Demange, J. - M. (1977) Nouveaux Myriapodes de Tanzanie. Description de deux especes nouvelles de Diplopodes. Affinites de quelques genres d'Oxydesmidae. Bulletin du Museum national d'Histoire naturelle 3 ° serie, n ° 451, Zoologie, 314, 507 - 518." type="journal article" year="1977">Demange, 1977</bibRefCitation>
: 517
</treatmentCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="4CC35CA62452197EFF676D76AD43FD36" pageId="7" pageNumber="508" type="diagnosis">
<paragraph id="04660F2D2452197EFF676D76AD43FD36" blockId="7.[151,1437,516,685]" pageId="7" pageNumber="508">
<emphasis id="36ADD33F2452197EFF676D76AE30FD86" bold="true" box="[151,273,516,541]" pageId="7" pageNumber="508">Diagnosis.</emphasis>
A medium-sized species of 
<taxonomicName id="C3D974AE2452197EFD8F6D77AC39FD87" box="[639,792,517,540]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="36ADD33F2452197EFD8F6D77AC39FD87" box="[639,792,517,540]" italics="true" pageId="7" pageNumber="508">Tropostreptus</emphasis>
</taxonomicName>
(
<figureCitation id="9CE213A82452197EFCC16D76ACA2FD87" box="[817,899,516,541]" captionStart="FIGURE 2" captionStartId="5.[151,250,1182,1204]" captionTargetBox="[151,1436,193,1096]" captionTargetId="figure@5.[151,1436,193,1161]" captionTargetPageId="5" captionText="FIGURE 2. Size of Tropostreptus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males. See Figs. 11 and 13 for details on T. hamatus and T. sigmatospinus sp. nov." httpUri="https://zenodo.org/record/803710/files/figure.png" pageId="7" pageNumber="508">Figs 2</figureCitation>
, 
<figureCitation id="9CE213A82452197EFC696D76AC97FD87" box="[921,950,516,540]" captionStart="FIGURE 11" captionStartId="16.[151,250,1055,1077]" captionTargetBox="[188,1407,227,1018]" captionTargetId="figure@16.[151,1436,193,1060]" captionTargetPageId="16" captionText="FIGURE 11. Size of Tropostreptus hamatus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males." httpUri="https://zenodo.org/record/803726/files/figure.png" pageId="7" pageNumber="508">11</figureCitation>
). Differs from other species, except 
<taxonomicName id="C3D974AE2452197EFA776D77AE16FDDB" class="Diplopoda" family="Spirostreptidae" genus="Tropostretus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="sigmatospinus">
<emphasis id="36ADD33F2452197EFA776D77AE16FDDB" italics="true" pageId="7" pageNumber="508">T. sigmatospinus</emphasis>
</taxonomicName>
, by the combination of size, the presence of two (not three) apical processes (
<emphasis id="36ADD33F2452197EFB4C6D5BABCCFDDB" box="[1212,1261,553,576]" italics="true" pageId="7" pageNumber="508">map</emphasis>
and 
<emphasis id="36ADD33F2452197EFAD66D55AA63FDDB" box="[1318,1346,551,576]" italics="true" pageId="7" pageNumber="508">bh</emphasis>
) on the gonopodal metaplica, and the laterad direction and considerable size of the birdhead-shaped process 
<emphasis id="36ADD33F2452197EFAD06D3EAA1DFDFE" box="[1312,1340,588,613]" italics="true" pageId="7" pageNumber="508">bh</emphasis>
. Differs from 
<taxonomicName id="C3D974AE2452197EFF266D03AEB5FD13" box="[214,404,625,648]" class="Diplopoda" family="Spirostreptidae" genus="Tropostretus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="sigmatospinus">
<emphasis id="36ADD33F2452197EFF266D03AEB5FD13" box="[214,404,625,648]" italics="true" pageId="7" pageNumber="508">T. sigmatospinus</emphasis>
</taxonomicName>
by the larger size of the lateral proplical lobe (
<emphasis id="36ADD33F2452197EFC466D1DACF2FD13" box="[950,979,623,648]" italics="true" pageId="7" pageNumber="508">lpl</emphasis>
) and the much longer and more curved antetorsal process (
<emphasis id="36ADD33F2452197EFE9F6DE7AEB2FD37" box="[367,403,661,684]" italics="true" pageId="7" pageNumber="508">atp</emphasis>
) of the telopodite.
</paragraph>
</subSubSection>
<caption id="50A65FA52452197EFF676A26AC00FA13" httpUri="https://zenodo.org/record/803712/files/figure.png" pageId="7" pageNumber="508" targetBox="[198,1394,742,1302]" targetPageId="7">
<paragraph id="04660F2D2452197EFF676A26AC00FA13" blockId="7.[151,1435,1364,1416]" pageId="7" pageNumber="508">
<emphasis id="36ADD33F2452197EFF676A26AE37FAF2" bold="true" box="[151,278,1364,1386]" pageId="7" pageNumber="508">FIGURE 3.</emphasis>
<taxonomicName id="C3D974AE2452197EFEEF6A27AEDDFAF1" box="[287,508,1364,1386]" class="Diplopoda" family="Spirostreptidae" genus="Tropostretus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F2452197EFEEF6A27AEDDFAF1" box="[287,508,1364,1386]" italics="true" pageId="7" pageNumber="508">Tropostretus hamatus</emphasis>
</taxonomicName>
, specimens from E. Usambara Mts. A: male, B: female. Scales 1 mm. Notice produced antero-ventral corner of collum in male (arrow). A. Illum photo.
</paragraph>
</caption>
<subSubSection id="4CC35CA624521971FF376AC2AC67FC1B" lastPageId="8" lastPageNumber="509" pageId="7" pageNumber="508" type="materials_examined">
<paragraph id="04660F2D2452197EFF376AC2ADCBFA52" blockId="7.[151,1437,1456,2021]" box="[199,746,1456,1481]" pageId="7" pageNumber="508">
<emphasis id="36ADD33F2452197EFF376AC2AEACFA52" bold="true" box="[199,397,1456,1481]" pageId="7" pageNumber="508">Material studied</emphasis>
(total: 83 ♂♂, 53 ♀♀, 71 juv.)
</paragraph>
<paragraph id="04660F2D2452197EFF376AA6AC39FA76" blockId="7.[151,1437,1456,2021]" box="[199,792,1492,1517]" pageId="7" pageNumber="508">All from TANZANIA, all in ZMUC except as noted.</paragraph>
<paragraph id="04660F2D2452197EFF376A8BAB66F9C2" blockId="7.[151,1437,1456,2021]" pageId="7" pageNumber="508">
<materialsCitation id="B4B105702452197EFF376A8BAFE1F9C2" ID-GBIF-Occurrence="1563407761" collectingDate="1970" collectingDateMax="1992" collectingDateMin="1970" collectorName="M. Andersen &amp; H. Enghoff &amp; I. B. Enghoff &amp; C. Griswold &amp; R. Jacobsen &amp; O. Lomholdt &amp; O. Martin &amp; N. Scharff &amp; M. Stoltze &amp; D. Ubick" elevation="1000" location="East Usambara Mts" municipality="Amani" pageId="7" pageNumber="508" specimenCount="61" specimenCount-female="18" specimenCount-male="34">
<location id="010659F62452197EFF376A8BAE88F98B" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFF376A8BAE88F98B" box="[199,425,1528,1553]" municipality="Amani" name="East Usambara Mts" pageId="7" pageNumber="508">East Usambara Mts</location>
: 
<specimenCount id="12DFC4A42452197EFE456A8AAD24F98B" box="[437,517,1528,1552]" pageId="7" pageNumber="508" type="male">34 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A42452197EFDE26A8AAD40F98B" box="[530,609,1528,1552]" pageId="7" pageNumber="508" type="female">18 ♀♀</specimenCount>
, 
<specimenCount id="12DFC4A42452197EFD806A8AAD91F98A" box="[624,688,1528,1553]" pageId="7" pageNumber="508" type="generic">9 juv.</specimenCount>
TOPOTYPES, 
<collectingMunicipality id="E40295572452197EFC946A8AAC92F98A" box="[868,947,1528,1553]" pageId="7" pageNumber="508">Amani</collectingMunicipality>
, ca. 
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<quantity id="C321A2C82452197EFC186A8AAB1FF98B" box="[1000,1086,1528,1552]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="508" unit="m" value="1000.0">1000 m</quantity>
asl
</elevation>
, 
<collectingDate id="6023D0052452197EFB846A8AABD7F98B" box="[1140,1270,1528,1552]" pageId="7" pageNumber="508" value="1970" valueMax="1992">
<date id="706729ED2452197EFB846A8AABD7F98B" box="[1140,1270,1528,1552]" pageId="7" pageNumber="508" value="1970" valueMax="1992">1970–1992</date>
</collectingDate>
, 
<collectorName id="A92C6AFB2452197EFAF26A8BAAB9F98A" box="[1282,1432,1528,1553]" pageId="7" pageNumber="508">M. Andersen</collectorName>
, 
<collectorName id="A92C6AFB2452197EFF67696FAE39F9AE" box="[151,280,1564,1589]" pageId="7" pageNumber="508">H. Enghoff</collectorName>
, 
<collectorName id="A92C6AFB2452197EFED4696FAE9DF9AE" box="[292,444,1564,1589]" pageId="7" pageNumber="508">I. B. Enghoff</collectorName>
, 
<collectorName id="A92C6AFB2452197EFE38696EAD74F9AE" box="[456,597,1564,1589]" pageId="7" pageNumber="508">C. Griswold</collectorName>
, 
<collectorName id="A92C6AFB2452197EFD91696FADCAF9AE" box="[609,747,1564,1589]" pageId="7" pageNumber="508">R. Jacobsen</collectorName>
, 
<collectorName id="A92C6AFB2452197EFD07696EACAAF9AE" box="[759,907,1564,1589]" pageId="7" pageNumber="508">O. Lomholdt</collectorName>
, 
<collectorName id="A92C6AFB2452197EFC67696EAB28F9AE" box="[919,1033,1564,1589]" pageId="7" pageNumber="508">O. Martin</collectorName>
, 
<collectorName id="A92C6AFB2452197EFBE6696FABAFF9AE" box="[1046,1166,1564,1589]" pageId="7" pageNumber="508">N. Scharff</collectorName>
, 
<collectorName id="A92C6AFB2452197EFB6A696FAA2FF9AE" box="[1178,1294,1564,1589]" pageId="7" pageNumber="508">M. Stoltze</collectorName>
&amp; 
<collectorName id="A92C6AFB2452197EFAC4696FAABCF9AE" box="[1332,1437,1564,1589]" pageId="7" pageNumber="508">D. Ubick</collectorName>
leg.
</materialsCitation>
- 
<materialsCitation id="B4B105702452197EFF276932AB66F9C2" ID-GBIF-Occurrence="1563407801" box="[215,1095,1600,1625]" collectingDate="1977-02-07" collectorName="H. Enghoff &amp; O. Lomholdt &amp; O. Martin" elevation="500" location="Sigi River" pageId="7" pageNumber="508" specimenCount="2" specimenCount-male="2">
<specimenCount id="12DFC4A42452197EFF276932AE37F9C3" box="[215,278,1600,1624]" pageId="7" pageNumber="508" type="male">2 ♂♂</specimenCount>
at 
<location id="010659F62452197EFEC96932AE8CF9C2" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFEC96932AE8CF9C2" box="[313,429,1600,1625]" name="Sigi River" pageId="7" pageNumber="508">Sigi River</location>
, 
<elevation id="8FF4E81E2452197EFE486932AD20F9C3" box="[440,513,1600,1625]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="508" unit="m" value="500.0">
<quantity id="C321A2C82452197EFE486932AD20F9C3" box="[440,513,1600,1625]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="508" unit="m" value="500.0">500 m</quantity>
</elevation>
, 
<collectingDate id="6023D0052452197EFDFD6933AD52F9C2" box="[525,627,1600,1625]" pageId="7" pageNumber="508" value="1977-02-07">
<date id="706729ED2452197EFDFD6933AD52F9C2" box="[525,627,1600,1625]" pageId="7" pageNumber="508" value="1977-02-07">7.ii.1977</date>
</collectingDate>
, 
<collectorName id="A92C6AFB2452197EFD8E6933ADDFF9C2" box="[638,766,1600,1625]" pageId="7" pageNumber="508">H. Enghoff</collectorName>
, 
<collectorName id="A92C6AFB2452197EFCFA6932ACBCF9C2" box="[778,925,1600,1625]" pageId="7" pageNumber="508">O. Lomholdt</collectorName>
, 
<collectorName id="A92C6AFB2452197EFC586932AB37F9C2" box="[936,1046,1600,1625]" pageId="7" pageNumber="508">O. Martin</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="04660F2D2452197EFF376916ACB8F8CE" blockId="7.[151,1437,1456,2021]" pageId="7" pageNumber="508">
<materialsCitation id="B4B105702452197EFF376916ADFFF93A" ID-GBIF-Occurrence="1563407786" collectingDate="1993-10" collectorName="L. Sorensen" county="Uluguru South Forest Reserve" elevation="1500" latitude="-7.1833334" location="South Forest Reserve" longLatPrecision="1303" longitude="36.033333" municipality="Uluguru South Forest" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region">
Uluguru Mts: 
<specimenCount id="12DFC4A42452197EFE946916AEAFF9E7" box="[356,398,1636,1660]" pageId="7" pageNumber="508" type="male">1 ♂</specimenCount>
<collectingRegion id="C61DC1CF2452197EFE646917AD41F9E6" box="[404,608,1636,1661]" country="Tanzania" name="Morogoro" pageId="7" pageNumber="508">Morogoro Region</collectingRegion>
, 
<collectingCounty id="ED0777A12452197EFD9A6916AC9EF9E7" box="[618,959,1636,1661]" pageId="7" pageNumber="508">
<collectingMunicipality id="E40295572452197EFD9A6916AC7AF9E7" box="[618,859,1636,1661]" pageId="7" pageNumber="508">
Uluguru 
<location id="010659F62452197EFD3E6916AC7AF9E7" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFD3E6916AC7AF9E7" box="[718,859,1636,1661]" county="Uluguru South Forest Reserve" latitude="-7.1833334" longLatPrecision="1303" longitude="36.033333" municipality="Uluguru South Forest" name="South Forest Reserve" pageId="7" pageNumber="508" stateProvince="Morogoro Region">South Forest</location>
</collectingMunicipality>
Reserve
</collectingCounty>
, forest above 
<location id="010659F62452197EFBA96916AA32F9E6" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFBA96916AA32F9E6" box="[1113,1299,1636,1661]" county="Uluguru South Forest Reserve" latitude="-7.1833334" longLatPrecision="1303" longitude="36.033333" municipality="Uluguru South Forest" name="Ukwama village" pageId="7" pageNumber="508" stateProvince="Morogoro Region">Ukwama village</location>
, 
<elevation id="8FF4E81E2452197EFAEE6916AFF8F93A" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="7" pageNumber="508" unit="m" value="1500.0" valueMax="1550.0" valueMin="1450.0">
<quantity id="C321A2C82452197EFAEE6916AF8CF93B" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="7" pageNumber="508" unit="m" value="1500.0" valueMax="1550.0" valueMin="1450.0">1450–1550 m</quantity>
asl
</elevation>
, 
<geoCoordinate id="61ED69EA2452197EFF1869FAAE64F93B" box="[232,325,1672,1696]" direction="south" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="-7.1833334">07°11’S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA2452197EFEA569FAAE86F93A" box="[341,423,1672,1697]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="36.033333">36°2’E</geoCoordinate>
, 
<collectingDate id="6023D0052452197EFE4769FBAD30F93B" box="[439,529,1672,1696]" pageId="7" pageNumber="508" value="1993-10">
<date id="706729ED2452197EFE4769FBAD30F93B" box="[439,529,1672,1696]" pageId="7" pageNumber="508" value="1993-10">x. 1993</date>
</collectingDate>
, 
<collectorName id="A92C6AFB2452197EFDD069FBAD8BF93B" box="[544,682,1672,1696]" pageId="7" pageNumber="508">L. Sørensen</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B105702452197EFD0F69FAAE9FF95E" ID-GBIF-Occurrence="1563407800" collectingDate="1981-07-10" collectorName="M. Stoltze &amp; N. Scharff" elevation="1300" location="East" municipality="Lupanga" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A42452197EFD0F69FAAC09F93B" box="[767,808,1672,1697]" pageId="7" pageNumber="508" type="male">1 ♂</specimenCount>
<collectingMunicipality id="E40295572452197EFCC369FBACB9F93A" box="[819,920,1673,1697]" pageId="7" pageNumber="508">Lupanga</collectingMunicipality>
, 
<location id="010659F62452197EFC5769FBACFBF93B" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFC5769FBACFBF93B" box="[935,986,1673,1696]" municipality="Lupanga" name="East" pageId="7" pageNumber="508" stateProvince="Morogoro Region">East</location>
, 
<elevation id="8FF4E81E2452197EFC1969FAAB4FF93A" box="[1001,1134,1672,1697]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="508" unit="m" value="1300.0">
<quantity id="C321A2C82452197EFC1969FAAB63F93B" box="[1001,1090,1672,1696]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="508" unit="m" value="1300.0">1300 m</quantity>
asl
</elevation>
, in trunk, 
<collectingDate id="6023D0052452197EFB1B69FAAA4CF93A" box="[1259,1389,1672,1697]" pageId="7" pageNumber="508" value="1981-07-10">
<date id="706729ED2452197EFB1B69FAAA4CF93A" box="[1259,1389,1672,1697]" pageId="7" pageNumber="508" value="1981-07-10">10.vii.1981</date>
</collectingDate>
, 
<collectorName id="A92C6AFB2452197EFA8C69FBAFC5F95E" pageId="7" pageNumber="508">M. Stoltze</collectorName>
&amp; 
<collectorName id="A92C6AFB2452197EFEE169DFAEA8F95E" box="[273,393,1708,1733]" pageId="7" pageNumber="508">N. Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B105702452197EFE2E69DEAC19F896" ID-GBIF-Occurrence="1563407783" collectingDate="2000-08" collectorName="Nike Doggart &amp; Uluguru Mountains Biodiversity Conservation Project" county="Mkungwe Forest" elevation="700" latitude="-6.9" location="Mountains Biodiversity Conservation Project" longLatPrecision="1303" longitude="37.933334" municipality="Uluguru Mountains" pageId="7" pageNumber="508" specimenCount="4" specimenCount-female="1" specimenCount-male="1" stateProvince="Morogoro Region, Morogoro">
<specimenCount id="12DFC4A42452197EFE2E69DEAD2BF95F" box="[478,522,1708,1733]" pageId="7" pageNumber="508" type="male">1 ♂</specimenCount>
, 
<specimenCount id="12DFC4A42452197EFDEB69DEAD68F95F" box="[539,585,1708,1732]" pageId="7" pageNumber="508" type="female">1 ♀</specimenCount>
, 
<specimenCount id="12DFC4A42452197EFDAB69DEADBFF95E" box="[603,670,1708,1733]" pageId="7" pageNumber="508" type="generic">2 juv.</specimenCount>
<collectingRegion id="C61DC1CF2452197EFD5969DFACDBF95E" box="[681,1018,1708,1733]" isEnumeration="true" pageId="7" pageNumber="508">Morogoro Region, Morogoro</collectingRegion>
District, 
<collectingCounty id="ED0777A12452197EFB9D69DEAA0EF95F" box="[1133,1327,1708,1733]" pageId="7" pageNumber="508">Mkungwe Forest</collectingCounty>
Reserve. Lowland submontane forest. 
<geoCoordinate id="61ED69EA2452197EFE1469A2AD5AF972" box="[484,635,1744,1769]" direction="south" orientation="latitude" pageId="7" pageNumber="508" precision="15" value="-6.8569446">6°51’25”S—</geoCoordinate>
<geoCoordinate id="61ED69EA2452197EFD8B69A2ADE7F972" box="[635,710,1744,1769]" direction="south" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="-6.9">6°54’S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA2452197EFD2569A2AC73F972" box="[725,850,1744,1769]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="37.9">37°54’E—</geoCoordinate>
<geoCoordinate id="61ED69EA2452197EFCA269A2AC8FF972" box="[850,942,1744,1769]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="37.933334">37°56’E</geoCoordinate>
, 
<elevation id="8FF4E81E2452197EFC4C69A2AB52F972" box="[956,1139,1744,1769]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="7" pageNumber="508" unit="m" value="700.0" valueMax="1000.0" valueMin="400.0">
<quantity id="C321A2C82452197EFC4C69A2AB6AF973" box="[956,1099,1744,1768]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="7" pageNumber="508" unit="m" value="700.0" valueMax="1000.0" valueMin="400.0">400–1000 m</quantity>
asl
</elevation>
, 
<collectingDate id="6023D0052452197EFB7C69A2ABD2F972" box="[1164,1267,1744,1769]" pageId="7" pageNumber="508" value="2000-08">
<date id="706729ED2452197EFB7C69A2ABD2F972" box="[1164,1267,1744,1769]" pageId="7" pageNumber="508" value="2000-08">viii.2000</date>
</collectingDate>
, 
<collectorName id="A92C6AFB2452197EFAF169A2AABDF972" box="[1281,1436,1744,1769]" pageId="7" pageNumber="508">Nike Doggart</collectorName>
leg., 
<collectorName id="A92C6AFB2452197EFF3F6986AC19F896" box="[207,824,1780,1805]" pageId="7" pageNumber="508">
<collectingMunicipality id="E40295572452197EFF3F6986AE8CF896" box="[207,429,1780,1805]" pageId="7" pageNumber="508">
Uluguru 
<location id="010659F62452197EFEC56986AE8CF896" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFEC56986AE8CF896" box="[309,429,1780,1805]" county="Mkungwe Forest" latitude="-6.9" longLatPrecision="1303" longitude="37.933334" municipality="Uluguru Mountains" name="Mountains Biodiversity Conservation Project" pageId="7" pageNumber="508" stateProvince="Morogoro Region, Morogoro">Mountains</location>
</collectingMunicipality>
Biodiversity Conservation Project
</collectorName>
</materialsCitation>
.— 
<materialsCitation id="B4B105702452197EFCA86986ACB4F8CE" ID-GBIF-Occurrence="1563407805" collectingDate="2000-07-27" collectingDateMax="2000-08-02" collectingDateMin="2000-07-27" collectorName="Nike Doggart &amp; Uluguru Mountains Biodiversity Conservation Project" county="Kasanga Forest" latitude="-7.1666665" location="Mountains Biodiversity Conservation Project" longLatPrecision="1303" longitude="37.75" municipality="Uluguru Mountains" pageId="7" pageNumber="508" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Morogoro Region, Morogoro">
<specimenCount id="12DFC4A42452197EFCA86986ACB8F897" box="[856,921,1780,1805]" pageId="7" pageNumber="508" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A42452197EFC566986ACC7F897" box="[934,998,1780,1804]" pageId="7" pageNumber="508" type="female">2 ♀♀</specimenCount>
<collectingRegion id="C61DC1CF2452197EFC1E6987AA1BF896" box="[1006,1338,1780,1805]" isEnumeration="true" pageId="7" pageNumber="508">Morogoro Region, Morogoro</collectingRegion>
District, 
<collectingCounty id="ED0777A12452197EFF67686BAE6BF8AB" box="[151,330,1817,1841]" pageId="7" pageNumber="508">Kasanga Forest</collectingCounty>
Reserve. Taken from leaf litter and rotting logs. 
<geoCoordinate id="61ED69EA2452197EFC52686AACD2F8AB" box="[930,1011,1816,1840]" direction="south" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="-7.1666665">7°10’S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA2452197EFBF4686AAB45F8AA" box="[1028,1124,1816,1841]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="37.75">37°45’E</geoCoordinate>
, 
<collectingDate id="6023D0052452197EFB86686AAA75F8AA" box="[1142,1364,1816,1841]" pageId="7" pageNumber="508" value="2000-07-27" valueMax="2000-08-02" valueMin="2000-07-27">
<date id="706729ED2452197EFB86686AAA75F8AA" box="[1142,1364,1816,1841]" pageId="7" pageNumber="508" value="2000-07-27" valueMax="2000-08-02" valueMin="2000-07-27">27.vii.– 2.viii.2000</date>
</collectingDate>
, 
<collectorName id="A92C6AFB2452197EFA96686AAFD2F8CE" pageId="7" pageNumber="508">Nike Doggart</collectorName>
leg., 
<collectorName id="A92C6AFB2452197EFEC1684EACB4F8CE" box="[305,917,1852,1877]" pageId="7" pageNumber="508">
<collectingMunicipality id="E40295572452197EFEC1684EAD2CF8CE" box="[305,525,1852,1877]" pageId="7" pageNumber="508">
Uluguru 
<location id="010659F62452197EFE65684EAD2CF8CE" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFE65684EAD2CF8CE" box="[405,525,1852,1877]" county="Kasanga Forest" latitude="-7.1666665" longLatPrecision="1303" longitude="37.75" municipality="Uluguru Mountains" name="Mountains Biodiversity Conservation Project" pageId="7" pageNumber="508" stateProvince="Morogoro Region, Morogoro">Mountains</location>
</collectingMunicipality>
Biodiversity Conservation Project
</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="04660F2D2452197EFF376812AABCF87E" blockId="7.[151,1437,1456,2021]" pageId="7" pageNumber="508">
<materialsCitation id="B4B105702452197EFF376812AE82F806" ID-GBIF-Occurrence="1563407791" collectingDate="1981-08-01" collectorName="M. Stolte &amp; N. Scharff" county="Udzungwa Mts" elevation="1000" location="Sanje" municipality="Mwanihana Forest" pageId="7" pageNumber="508" specimenCount="14" specimenCount-female="3" specimenCount-male="1">
<collectingCounty id="ED0777A12452197EFF376812AE59F8E3" box="[199,376,1888,1913]" pageId="7" pageNumber="508">Udzungwa Mts</collectingCounty>
: 
<specimenCount id="12DFC4A42452197EFE736812AE91F8E3" box="[387,432,1888,1912]" pageId="7" pageNumber="508" type="male">1 ♂</specimenCount>
, 
<specimenCount id="12DFC4A42452197EFE4C6812AEDCF8E3" box="[444,509,1888,1912]" pageId="7" pageNumber="508" type="female">3 ♀♀</specimenCount>
, 
<specimenCount id="12DFC4A42452197EFDFB6812AD79F8E2" box="[523,600,1888,1913]" pageId="7" pageNumber="508" type="generic">10 juv.</specimenCount>
<collectingMunicipality id="E40295572452197EFDAF6812AC10F8E3" box="[607,817,1888,1913]" pageId="7" pageNumber="508">Mwanihana Forest</collectingMunicipality>
above 
<location id="010659F62452197EFC736812ACE5F8E2" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFC736812ACE5F8E2" box="[899,964,1888,1913]" county="Udzungwa Mts" municipality="Mwanihana Forest" name="Sanje" pageId="7" pageNumber="508">Sanje</location>
, 
<elevation id="8FF4E81E2452197EFC206812AB6EF8E2" box="[976,1103,1888,1913]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="508" unit="m" value="1000.0">
<quantity id="C321A2C82452197EFC206812AB07F8E3" box="[976,1062,1888,1912]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="508" unit="m" value="1000.0">1000 m</quantity>
asl
</elevation>
, in tree trunk, 
<collectingDate id="6023D0052452197EFB046812AA51F8E2" box="[1268,1392,1888,1913]" pageId="7" pageNumber="508" value="1981-08-01">
<date id="706729ED2452197EFB046812AA51F8E2" box="[1268,1392,1888,1913]" pageId="7" pageNumber="508" value="1981-08-01">1.viii.1981</date>
</collectingDate>
, 
<collectorName id="A92C6AFB2452197EFA8C6813AFF9F806" pageId="7" pageNumber="508">M. Stolte</collectorName>
&amp; 
<collectorName id="A92C6AFB2452197EFF0D68F7AE52F806" box="[253,371,1924,1949]" pageId="7" pageNumber="508">N. Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B105702452197EFE3268F6ACA0F806" ID-GBIF-Occurrence="1563407790" box="[450,897,1924,1949]" collectingDate="1982-08-01" collectorName="M. Stolte &amp; N. Scharff" county="Udzungwa Mts" elevation="1000" location="Sanje" municipality="Mwanihana Forest" pageId="7" pageNumber="508" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
<specimenCount id="12DFC4A42452197EFE3268F6AD23F807" box="[450,514,1924,1949]" pageId="7" pageNumber="508" type="male">3 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A42452197EFDFF68F6AD6EF807" box="[527,591,1924,1948]" pageId="7" pageNumber="508" type="female">2 ♀♀</specimenCount>
same data, but 
<collectingDate id="6023D0052452197EFCF268F6AC5CF806" box="[770,893,1924,1949]" pageId="7" pageNumber="508" value="1982-08-01">
<date id="706729ED2452197EFCF268F6AC5CF806" box="[770,893,1924,1949]" pageId="7" pageNumber="508" value="1982-08-01">1.viii.1982</date>
</collectingDate>
.
</materialsCitation>
— 
<materialsCitation id="B4B105702452197EFC5368F6AB4DF85B" ID-GBIF-Occurrence="1563407804" collectingDate="1981-01-05" collectionCode="VMNH" collectorName="K. M. Howell &amp; S. Stuart" elevation="1097" location="Sanje" municipality="Mwanihana Forest" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Iringa">
<specimenCount id="12DFC4A42452197EFC5368F6ACE8F807" box="[931,969,1924,1949]" pageId="7" pageNumber="508" type="male">1 ♂</specimenCount>
<collectingRegion id="C61DC1CF2452197EFC2168F6AB35F806" box="[977,1044,1924,1949]" country="Tanzania" name="Iringa" pageId="7" pageNumber="508">Iringa</collectingRegion>
District, 
<collectingMunicipality id="E40295572452197EFB8F68F6AA70F807" box="[1151,1361,1924,1949]" pageId="7" pageNumber="508">Mwanihana Forest</collectingMunicipality>
above 
<location id="010659F62452197EFF6768DAAFF9F85A" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFF6768DAAFF9F85A" box="[151,216,1960,1985]" municipality="Mwanihana Forest" name="Sanje" pageId="7" pageNumber="508" stateProvince="Iringa">Sanje</location>
, 
<elevation id="8FF4E81E2452197EFF1868DAAE4DF85A" box="[232,364,1960,1985]" metricMagnitude="3" metricUnit="m" metricValue="1.09728" pageId="7" pageNumber="508" unit="ft" value="3600.0">
<quantity id="C321A2C82452197EFF1868DAAE1FF85A" box="[232,318,1960,1985]" metricMagnitude="3" metricUnit="m" metricValue="1.09728" pageId="7" pageNumber="508" unit="ft" value="3600.0">3600 ft</quantity>
asl
</elevation>
, 
<collectingDate id="6023D0052452197EFE8C68DAAEFBF85A" box="[380,474,1960,1985]" pageId="7" pageNumber="508" value="1981-01-05">
<date id="706729ED2452197EFE8C68DAAEFBF85A" box="[380,474,1960,1985]" pageId="7" pageNumber="508" value="1981-01-05">5.i.1981</date>
</collectingDate>
, 
<collectorName id="A92C6AFB2452197EFE1B68DBADA7F85A" box="[491,646,1960,1985]" pageId="7" pageNumber="508">K.M. Howell</collectorName>
&amp; 
<collectorName id="A92C6AFB2452197EFD4368DAAC38F85B" box="[691,793,1960,1984]" pageId="7" pageNumber="508">S. Stuart</collectorName>
leg. (KMH 1723) (
<collectionCode id="62C897E82452197EFBF768DAAB44F85B" box="[1031,1125,1960,1984]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="7" pageNumber="508">VMNH</collectionCode>
)
</materialsCitation>
.— 
<materialsCitation id="B4B105702452197EFB7F68DAAEDFF87F" ID-GBIF-Occurrence="1563407789" collectionCode="VMNH" location="Mwanihana Forest" municipality="Mwanihana Forest Reserve" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Iringa">
<specimenCount id="12DFC4A42452197EFB7F68DAAB98F85B" box="[1167,1209,1960,1984]" pageId="7" pageNumber="508" type="male">1 ♂</specimenCount>
<collectingMunicipality id="E40295572452197EFB3668DAAFD5F87F" pageId="7" pageNumber="508">
<location id="010659F62452197EFB3668DAAABDF85B" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFB3668DAAABDF85B" box="[1222,1436,1960,1985]" municipality="Mwanihana Forest Reserve" name="Mwanihana Forest" pageId="7" pageNumber="508" stateProvince="Iringa">Mwanihana Forest</location>
Reserve
</collectingMunicipality>
, KMH 2058 (
<collectionCode id="62C897E82452197EFE6868BEAED4F87F" box="[408,501,1996,2020]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="7" pageNumber="508">VMNH</collectionCode>
)
</materialsCitation>
.— 
<materialsCitation id="B4B105702452197EFDD168BEABE1F87F" ID-GBIF-Occurrence="1563407797" box="[545,1216,1996,2021]" collectionCode="VMNH" location="Ifanre" municipality="Mwanihana forest" pageId="7" pageNumber="508" specimenCount="1" specimenCount-male="1" stateProvince="Iringa">
<specimenCount id="12DFC4A42452197EFDD168BEAD68F87F" box="[545,585,1996,2020]" pageId="7" pageNumber="508" type="male">1 ♂</specimenCount>
(incomplete) 
<collectingMunicipality id="E40295572452197EFD1D68BEAC9AF87E" box="[749,955,1996,2021]" pageId="7" pageNumber="508">Mwanihana forest</collectingMunicipality>
N of 
<location id="010659F62452197EFBF568BEAB69F87E" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F62452197EFBF568BEAB69F87E" box="[1029,1096,1996,2021]" municipality="Mwanihana forest" name="Ifanre" pageId="7" pageNumber="508" stateProvince="Iringa">Ifanre</location>
(
<collectionCode id="62C897E82452197EFBAA68BEAB99F87F" box="[1114,1208,1996,2020]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="7" pageNumber="508">VMNH</collectionCode>
)
</materialsCitation>
.—1 ♂, 1 ♀, 4 juv.
</paragraph>
<paragraph id="04660F2D245D1971FF676FEAABB3FD6B" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
<materialsCitation id="B4B10570245D1971FF676FEAAA5FFF2B" ID-GBIF-Occurrence="1563407792" box="[151,1406,151,176]" collectingDate="1984-09-07" collectingDateMax="1984-09-16" collectingDateMin="1984-09-07" collectorName="N. Scharff" elevation="625" location="Mwanihana Forest" municipality="Mwanihana Forest Reserve" pageId="8" pageNumber="509" specimenCount="1" stateProvince="Morogoro Region">
<collectingRegion id="C61DC1CF245D1971FF676FEAAE42FF2B" box="[151,355,151,176]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, 
<collectingMunicipality id="E4029557245D1971FE9D6FE5AD80FF34" box="[365,673,151,176]" pageId="8" pageNumber="509">
<location id="010659F6245D1971FE9D6FE5AD1CFF34" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FE9D6FE5AD1CFF34" box="[365,573,151,176]" municipality="Mwanihana Forest Reserve" name="Mwanihana Forest" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Mwanihana Forest</location>
Reserve
</collectingMunicipality>
, 
<elevation id="8FF4E81E245D1971FD5B6FE5AC73FF2B" box="[683,850,151,176]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="8" pageNumber="509" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">
<quantity id="C321A2C8245D1971FD5B6FE5AC0BFF34" box="[683,810,151,176]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="8" pageNumber="509" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">500–750 m</quantity>
asl
</elevation>
, 
<collectingDate id="6023D005245D1971FCAC6FE5ACD2FF2B" box="[860,1011,151,176]" pageId="8" pageNumber="509" value="1984-09-07" valueMax="1984-09-16" valueMin="1984-09-07">
<date id="706729ED245D1971FCAC6FE5ACD2FF2B" box="[860,1011,151,176]" pageId="8" pageNumber="509" value="1984-09-07" valueMax="1984-09-16" valueMin="1984-09-07">7–16.ix.1984</date>
</collectingDate>
, lowland rain forest, 
<collectorName id="A92C6AFB245D1971FB2D6FEAAA70FF2B" box="[1245,1361,151,176]" pageId="8" pageNumber="509">N. Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FF676FCEABCEFF4E" ID-GBIF-Occurrence="1563407793" box="[151,1263,188,213]" collectingDate="1981-08-01" collectorName="M. Stoltze &amp; N. Scharff" elevation="1000" location="Mwanihana Forest" municipality="Mwanihana Forest Reserve" pageId="8" pageNumber="509" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FF676FCEAFFAFF4F" box="[151,219,188,212]" pageId="8" pageNumber="509" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FF1A6FCEAE0CFF4F" box="[234,301,188,212]" pageId="8" pageNumber="509" type="female">4 ♀♀</specimenCount>
, 
<collectingMunicipality id="E4029557245D1971FECD6FCEAD58FF4F" box="[317,633,188,213]" pageId="8" pageNumber="509">
<location id="010659F6245D1971FECD6FCEAD33FF4F" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FECD6FCEAD33FF4F" box="[317,530,188,213]" municipality="Mwanihana Forest Reserve" name="Mwanihana Forest" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Mwanihana Forest</location>
Reserve
</collectingMunicipality>
, 
<elevation id="8FF4E81E245D1971FD786FCEAC2DFF4E" box="[648,780,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="509" unit="m" value="1000.0">
<quantity id="C321A2C8245D1971FD786FCEADC1FF4F" box="[648,736,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="509" unit="m" value="1000.0">1000 m</quantity>
asl
</elevation>
, 
<collectingDate id="6023D005245D1971FCEA6FCEACB7FF4E" box="[794,918,188,213]" pageId="8" pageNumber="509" value="1981-08-01">
<date id="706729ED245D1971FCEA6FCEACB7FF4E" box="[794,918,188,213]" pageId="8" pageNumber="509" value="1981-08-01">1.viii.1981</date>
</collectingDate>
, 
<collectorName id="A92C6AFB245D1971FC556FCFAB3DFF4E" box="[933,1052,188,213]" pageId="8" pageNumber="509">M. Stoltze</collectorName>
&amp; 
<collectorName id="A92C6AFB245D1971FBB66FCFAB9CFF4E" box="[1094,1213,188,213]" pageId="8" pageNumber="509">N. Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FAFD6FCEAB87FF63" ID-GBIF-Occurrence="1563407787" collectingDate="1981-04-06" collectionCode="VMNH" collectorName="J. Kielland" location="Ifakara" municipality="Mwanihana Forest" pageId="8" pageNumber="509" specimenCount="2" specimenCount-male="2" stateProvince="Iringa Region">
<specimenCount id="12DFC4A4245D1971FAFD6FCEAA6EFF4F" box="[1293,1359,188,213]" pageId="8" pageNumber="509" type="male">2 ♂♂</specimenCount>
<collectingRegion id="C61DC1CF245D1971FAA96FCEAFC9FF63" country="Tanzania" name="Iringa" pageId="8" pageNumber="509">Iringa Region</collectingRegion>
and district: 
<collectingMunicipality id="E4029557245D1971FE8B6FADAD6CFF6C" box="[379,589,223,248]" pageId="8" pageNumber="509">Mwanihana Forest</collectingMunicipality>
north of 
<location id="010659F6245D1971FD446FADAC27FF63" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FD446FADAC27FF63" box="[692,774,223,248]" municipality="Mwanihana Forest" name="Ifakara" pageId="8" pageNumber="509" stateProvince="Iringa Region">Ifakara</location>
, 
<collectingDate id="6023D005245D1971FCE16FADAC5DFF63" box="[785,892,223,248]" pageId="8" pageNumber="509" value="1981-04-06">
<date id="706729ED245D1971FCE16FADAC5DFF63" box="[785,892,223,248]" pageId="8" pageNumber="509" value="1981-04-06">6.iv.1981</date>
</collectingDate>
, 
<collectorName id="A92C6AFB245D1971FC786F92AB23FF63" box="[904,1026,223,248]" pageId="8" pageNumber="509">J. Kielland</collectorName>
leg. (
<collectionCode id="62C897E8245D1971FBB06F92ABBBFF63" box="[1088,1178,224,248]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="8" pageNumber="509">VMNH</collectionCode>
).
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FB346F92AD4FFEDB" ID-GBIF-Occurrence="1563407799" collectingDate="2013-09-01" collectorName="T. Pape &amp; N. Scharff" elevation="769" latitude="-7.4731665" location="Mizimu Camp" longLatPrecision="1" longitude="36.852024" pageId="8" pageNumber="509" specimenCount="8" specimenCount-female="2" specimenCount-male="4" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FB346F92AA24FF6C" box="[1220,1285,224,248]" pageId="8" pageNumber="509" type="male">4 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FAE16F92AA70FF6C" box="[1297,1361,224,248]" pageId="8" pageNumber="509" type="female">2 ♀♀</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FAAE6F92AABCFF63" box="[1374,1437,223,248]" pageId="8" pageNumber="509" type="generic">2 juv.</specimenCount>
<collectingRegion id="C61DC1CF245D1971FF676E77AE42FE86" box="[151,355,260,285]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, Udzungwa Mts Natl. Park, 
<location id="010659F6245D1971FD6E6E76AC62FE87" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FD6E6E76AC62FE87" box="[670,835,260,285]" latitude="-7.4731665" longLatPrecision="1" longitude="36.852024" name="Mizimu Camp" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Mizimu Camp</location>
, 
<collectingDate id="6023D005245D1971FCBE6E76AC98FE86" box="[846,953,260,285]" pageId="8" pageNumber="509" value="2013-09-01">
<date id="706729ED245D1971FCBE6E76AC98FE86" box="[846,953,260,285]" pageId="8" pageNumber="509" value="2013-09-01">1.ix.2013</date>
</collectingDate>
, 
<geoCoordinate id="61ED69EA245D1971FC356E76AB40FE87" box="[965,1121,260,284]" direction="south" orientation="latitude" pageId="8" pageNumber="509" precision="1" value="-7.4731665">7°28’23.40”S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA245D1971FB9C6E76AA39FE86" box="[1132,1304,260,285]" direction="east" orientation="longitude" pageId="8" pageNumber="509" precision="1" value="36.852024">36°51’07.29”E</geoCoordinate>
, 
<elevation id="8FF4E81E245D1971FAD26E76AAAEFE86" box="[1314,1423,260,285]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="8" pageNumber="509" unit="m" value="769.0">
<quantity id="C321A2C8245D1971FAD26E76AA48FE87" box="[1314,1385,260,285]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="8" pageNumber="509" unit="m" value="769.0">769 m</quantity>
asl
</elevation>
, hand collected, 
<collectorName id="A92C6AFB245D1971FEBB6E5AAE80FEA4" box="[331,417,296,319]" pageId="8" pageNumber="509">T. Pape</collectorName>
&amp; 
<collectorName id="A92C6AFB245D1971FE386E5AAD1FFEDB" box="[456,574,295,320]" pageId="8" pageNumber="509">N. Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FD7D6E55AE0FFE13" ID-GBIF-Occurrence="1563407798" collectingDate="1993-08" collectorName="L. L. Sorensen" county="Mahenge District" elevation="1725" latitude="-7.7833333" location="Ndundulu Mts" longLatPrecision="1302" longitude="36.483334" municipality="West Kilombero Forest Reserve" pageId="8" pageNumber="509" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Iringa Region">
<specimenCount id="12DFC4A4245D1971FD7D6E55ADECFEA4" box="[653,717,295,320]" pageId="8" pageNumber="509" type="male">2 ♂♂</specimenCount>
<specimenCount id="12DFC4A4245D1971FD266E5AAC23FEA4" box="[726,770,296,320]" pageId="8" pageNumber="509" type="female">1 ♀</specimenCount>
, 
<collectingRegion id="C61DC1CF245D1971FCE26E55AC93FEDB" box="[786,946,295,320]" country="Tanzania" name="Iringa" pageId="8" pageNumber="509">Iringa Region</collectingRegion>
, 
<collectingCounty id="ED0777A1245D1971FC4F6E55ABA6FEDB" box="[959,1159,295,320]" pageId="8" pageNumber="509">Mahenge District</collectingCounty>
, 
<collectingMunicipality id="E4029557245D1971FB646E5AAFD5FEFF" pageId="8" pageNumber="509">West Kilombero Forest Reserve</collectingMunicipality>
, 
<location id="010659F6245D1971FEF06E3EAE85FEFF" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FEF06E3EAE85FEFF" box="[256,420,332,357]" county="Mahenge District" latitude="-7.7833333" longLatPrecision="1302" longitude="36.483334" municipality="West Kilombero Forest Reserve" name="Ndundulu Mts" pageId="8" pageNumber="509" stateProvince="Iringa Region">Ndundulu Mts</location>
ESE 
<location id="010659F6245D1971FE156E3EADBFFEFE" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FE156E3EADBFFEFE" box="[485,670,332,357]" county="Mahenge District" latitude="-7.7833333" longLatPrecision="1302" longitude="36.483334" municipality="West Kilombero Forest Reserve" name="Udekwa Village" pageId="8" pageNumber="509" stateProvince="Iringa Region">Udekwa Village</location>
, 
<geoCoordinate id="61ED69EA245D1971FD5A6E3EADDBFEFF" box="[682,762,332,356]" direction="south" orientation="latitude" pageId="8" pageNumber="509" precision="925" value="-7.7833333">7°47’S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA245D1971FCF76E3EAC46FEFE" box="[775,871,332,357]" direction="east" orientation="longitude" pageId="8" pageNumber="509" precision="925" value="36.483334">36°29’E</geoCoordinate>
, 
<elevation id="8FF4E81E245D1971FC836E3EAB18FEFE" box="[883,1081,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="8" pageNumber="509" unit="m" value="1725.0" valueMax="1750.0" valueMin="1700.0">
<quantity id="C321A2C8245D1971FC836E3EAB2EFEFF" box="[883,1039,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="8" pageNumber="509" unit="m" value="1725.0" valueMax="1750.0" valueMin="1700.0">1700–1750 m</quantity>
asl
</elevation>
, in rotten wood, 
<collectingDate id="6023D005245D1971FB096E3EAA41FEFE" box="[1273,1376,332,357]" pageId="8" pageNumber="509" value="1993-08">
<date id="706729ED245D1971FB096E3EAA41FEFE" box="[1273,1376,332,357]" pageId="8" pageNumber="509" value="1993-08">viii.1993</date>
</collectingDate>
, 
<collectorName id="A92C6AFB245D1971FA9C6E3FAFDFFE13" pageId="8" pageNumber="509">L.L. Sørensen</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FEBE6E1DABC5FE13" ID-GBIF-Occurrence="1563407803" box="[334,1252,367,392]" collectingDate="1993-08" collectorName="L. L. Sorensen" county="Mahenge District" elevation="1450" latitude="-7.7833333" location="Ndundulu Mts" longLatPrecision="1302" longitude="36.483334" municipality="West Kilombero Forest Reserve" pageId="8" pageNumber="509" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Iringa Region">
<specimenCount id="12DFC4A4245D1971FEBE6E1DAE55FE1C" box="[334,372,367,392]" pageId="8" pageNumber="509" type="male">1 ♂</specimenCount>
<specimenCount id="12DFC4A4245D1971FE8B6E02AE9AFE1C" box="[379,443,368,392]" pageId="8" pageNumber="509" type="female">4 ♀♀</specimenCount>
, same data but 
<elevation id="8FF4E81E245D1971FD9A6E1DADC9FE13" box="[618,744,367,392]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="509" unit="m" value="1450.0">
<quantity id="C321A2C8245D1971FD9A6E1DAD9EFE1C" box="[618,703,367,392]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="509" unit="m" value="1450.0">1450 m</quantity>
asl
</elevation>
, in rotten wood, 
<collectingDate id="6023D005245D1971FC556E1DAB2AFE13" box="[933,1035,367,392]" pageId="8" pageNumber="509" value="1993-08">
<date id="706729ED245D1971FC556E1DAB2AFE13" box="[933,1035,367,392]" pageId="8" pageNumber="509" value="1993-08">viii.1993</date>
</collectingDate>
, 
<collectorName id="A92C6AFB245D1971FBE76E02AB97FE13" box="[1047,1206,368,392]" pageId="8" pageNumber="509">L.L. Sørensen</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FAF36E1DADF9FE4B" ID-GBIF-Occurrence="1563407807" collectingDate="2014-01-17" collectingDateMax="2014-01-19" collectingDateMin="2014-01-17" collectorName="T. Pape &amp; N. Scharff" elevation="598" latitude="-7.7735" location="Sanje Chini" longLatPrecision="1" longitude="36.896584" municipality="Udzungwa Mts Natl Park" pageId="8" pageNumber="509" specimenCount="12" specimenCount-female="2" specimenCount-male="10" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FAF36E1DAA73FE1C" box="[1283,1362,367,392]" pageId="8" pageNumber="509" type="male">10 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FAAD6E02AABDFE1C" box="[1373,1436,368,392]" pageId="8" pageNumber="509" type="female">2 ♀♀</specimenCount>
<collectingRegion id="C61DC1CF245D1971FF676EE7AE46FE36" box="[151,359,404,429]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, 
<collectingMunicipality id="E4029557245D1971FE856EE6ADBFFE36" box="[373,670,404,429]" pageId="8" pageNumber="509">Udzungwa Mts Natl Park</collectingMunicipality>
, 
<location id="010659F6245D1971FD5C6EE6AC12FE36" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FD5C6EE6AC12FE36" box="[684,819,404,429]" latitude="-7.7735" longLatPrecision="1" longitude="36.896584" municipality="Udzungwa Mts Natl Park" name="Sanje Chini" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Sanje Chini</location>
camp, 
<elevation id="8FF4E81E245D1971FC7B6EE6AB20FE36" box="[907,1025,404,429]" metricMagnitude="2" metricUnit="m" metricValue="5.9799999999999995" pageId="8" pageNumber="509" unit="m" value="598.0">
<quantity id="C321A2C8245D1971FC7B6EE6ACF4FE37" box="[907,981,404,429]" metricMagnitude="2" metricUnit="m" metricValue="5.9799999999999995" pageId="8" pageNumber="509" unit="m" value="598.0">598 m</quantity>
asl
</elevation>
, 
<geoCoordinate id="61ED69EA245D1971FBFF6EE6AB8AFE36" box="[1039,1195,404,429]" direction="south" orientation="latitude" pageId="8" pageNumber="509" precision="1" value="-7.7735">07°46’24.6”S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA245D1971FB4A6EE6AA76FE36" box="[1210,1367,404,429]" direction="east" orientation="longitude" pageId="8" pageNumber="509" precision="1" value="36.896584">36°53’47.7”E</geoCoordinate>
, hand collected, 
<collectingDate id="6023D005245D1971FEFD6EC5AE82FE4B" box="[269,419,439,464]" pageId="8" pageNumber="509" value="2014-01-17" valueMax="2014-01-19" valueMin="2014-01-17">
<date id="706729ED245D1971FEFD6EC5AE82FE4B" box="[269,419,439,464]" pageId="8" pageNumber="509" value="2014-01-17" valueMax="2014-01-19" valueMin="2014-01-17">17–19.i.2014</date>
</collectingDate>
, 
<collectorName id="A92C6AFB245D1971FE416ECAAD26FE54" box="[433,519,440,463]" pageId="8" pageNumber="509">T. Pape</collectorName>
&amp; 
<collectorName id="A92C6AFB245D1971FDC06ECAAD86FE4B" box="[560,679,439,464]" pageId="8" pageNumber="509">N. Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FD066EC5AA52FE6E" ID-GBIF-Occurrence="1563407796" collectingDate="2014-01-20" collectingDateMax="2014-01-23" collectingDateMin="2014-01-20" collectorName="Pape &amp; Scharff" elevation="850" latitude="-7.763222" location="Sanje Kati" longLatPrecision="1" longitude="36.886223" municipality="Udzungwa Mts Natl Park" pageId="8" pageNumber="509" specimenCount="5" specimenCount-male="5" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FD066EC5AC60FE54" box="[758,833,439,464]" pageId="8" pageNumber="509" type="male">4 ♂ ♂</specimenCount>
<collectingRegion id="C61DC1CF245D1971FCBB6ECAAB3BFE4B" box="[843,1050,439,464]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, 
<collectingMunicipality id="E4029557245D1971FBD86EC5AA71FE4B" box="[1064,1360,439,464]" pageId="8" pageNumber="509">Udzungwa Mts Natl Park</collectingMunicipality>
, 
<location id="010659F6245D1971FAAE6EC5AFE6FE6E" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FAAE6EC5AFE6FE6E" latitude="-7.763222" longLatPrecision="1" longitude="36.886223" municipality="Udzungwa Mts Natl Park" name="Sanje Kati" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Sanje Kati</location>
camp &amp; plot, 
<elevation id="8FF4E81E245D1971FE9C6EAEAEFFFE6E" box="[364,478,476,501]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="509" unit="m" value="850.0">
<quantity id="C321A2C8245D1971FE9C6EAEAE95FE6F" box="[364,436,476,501]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="509" unit="m" value="850.0">850 m</quantity>
asl
</elevation>
, 
<geoCoordinate id="61ED69EA245D1971FE1A6EAEADA6FE6E" box="[490,647,476,501]" direction="south" orientation="latitude" pageId="8" pageNumber="509" precision="1" value="-7.763222">07°45’47.6”S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA245D1971FD636EAEAC11FE6E" box="[659,816,476,501]" direction="east" orientation="longitude" pageId="8" pageNumber="509" precision="1" value="36.886223">36°53’10.4”E</geoCoordinate>
, hand collected, 
<collectingDate id="6023D005245D1971FC006EAEABA6FE6E" box="[1008,1159,476,501]" pageId="8" pageNumber="509" value="2014-01-20" valueMax="2014-01-23" valueMin="2014-01-20">
<date id="706729ED245D1971FC006EAEABA6FE6E" box="[1008,1159,476,501]" pageId="8" pageNumber="509" value="2014-01-20" valueMax="2014-01-23" valueMin="2014-01-20">20–23.i.2014</date>
</collectingDate>
. 
<collectorName id="A92C6AFB245D1971FB636EAFABEBFE6F" box="[1171,1226,477,500]" pageId="8" pageNumber="509">Pape</collectorName>
&amp; 
<collectorName id="A92C6AFB245D1971FB1F6EAEAA60FE6E" box="[1263,1345,476,501]" pageId="8" pageNumber="509">Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FA636EAEAD0CFDA6" ID-GBIF-Occurrence="1563407802" collectingDate="2012-09-03" collectorName="T. Pape &amp; N Scharff" elevation="250" location="Mizimu Camp" municipality="Udzungwa Mts Natl Park" pageId="8" pageNumber="509" specimenCount="3" specimenCount-female="1" specimenCount-male="1" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FA636EAEAF8FFD8C" pageId="8" pageNumber="509" type="male">1 ♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FF4F6D72AE2BFD83" box="[191,266,511,536]" pageId="8" pageNumber="509" type="generic">1 juv.</specimenCount>
<specimenCount id="12DFC4A4245D1971FEFA6E8DAE38FD83" box="[266,281,511,536]" pageId="8" pageNumber="509" type="female">♀</specimenCount>
<collectingRegion id="C61DC1CF245D1971FED56D72AED6FD83" box="[293,503,511,536]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, 
<collectingMunicipality id="E4029557245D1971FDF76E8DAC19FD83" box="[519,824,511,536]" pageId="8" pageNumber="509">Udzungwa Mts Natl Park</collectingMunicipality>
, trail to 
<location id="010659F6245D1971FC556E8DAB70FD83" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FC556E8DAB70FD83" box="[933,1105,511,536]" municipality="Udzungwa Mts Natl Park" name="Mizimu Camp" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Mizimu Camp</location>
, 
<elevation id="8FF4E81E245D1971FB916E8DABFDFD83" box="[1121,1244,511,536]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="509" unit="m" value="250.0">
<quantity id="C321A2C8245D1971FB916E8DAB8FFD8C" box="[1121,1198,511,536]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="509" unit="m" value="250.0">250 m</quantity>
asl
</elevation>
, hand collected, 
<collectingDate id="6023D005245D1971FF676D56AE25FDA6" box="[151,260,548,573]" pageId="8" pageNumber="509" value="2012-09-03">
<date id="706729ED245D1971FF676D56AE25FDA6" box="[151,260,548,573]" pageId="8" pageNumber="509" value="2012-09-03">3.ix.2012</date>
</collectingDate>
. 
<collectorName id="A92C6AFB245D1971FEE06D57AE44FDA7" box="[272,357,549,572]" pageId="8" pageNumber="509">T. Pape</collectorName>
&amp; 
<collectorName id="A92C6AFB245D1971FE7D6D57AEDDFDA6" box="[397,508,548,573]" pageId="8" pageNumber="509">N Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FDBC6D56AB04FDFB" ID-GBIF-Occurrence="1563407811" collectingDate="2012-09-03" collectorName="T. Pape &amp; N Scharff" elevation="250" location="Mizimu Camp" municipality="Udzungwa Mts Natl Park" pageId="8" pageNumber="509" specimenCount="18" specimenCount-female="2" specimenCount-male="3" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FDBC6D56ADACFDA7" box="[588,653,548,573]" pageId="8" pageNumber="509" type="male">3 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FD6B6D56ADFCFDA7" box="[667,733,548,572]" pageId="8" pageNumber="509" type="female">2 ♀♀</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FD1C6D56AC1BFDA6" box="[748,826,548,573]" pageId="8" pageNumber="509" type="generic">13 juv.</specimenCount>
<collectingRegion id="C61DC1CF245D1971FCB26D57AB30FDA6" box="[834,1041,548,573]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, 
<collectingMunicipality id="E4029557245D1971FBEE6D56AA62FDA6" box="[1054,1347,548,573]" pageId="8" pageNumber="509">Udzungwa Mts Natl Park</collectingMunicipality>
, trail to 
<location id="010659F6245D1971FF676D35AE1EFDFB" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FF676D35AE1EFDFB" box="[151,319,583,608]" municipality="Udzungwa Mts Natl Park" name="Mizimu Camp" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Mizimu Camp</location>
, 
<elevation id="8FF4E81E245D1971FEBD6D35AEE2FDFB" box="[333,451,583,608]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="509" unit="m" value="250.0">
<quantity id="C321A2C8245D1971FEBD6D35AEB6FDC4" box="[333,407,583,608]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="509" unit="m" value="250.0">250 m</quantity>
asl
</elevation>
, hand collected, 
<collectingDate id="6023D005245D1971FD776D35ADD2FDFB" box="[647,755,583,608]" pageId="8" pageNumber="509" value="2012-09-03">
<date id="706729ED245D1971FD776D35ADD2FDFB" box="[647,755,583,608]" pageId="8" pageNumber="509" value="2012-09-03">3.ix.2012</date>
</collectingDate>
. 
<collectorName id="A92C6AFB245D1971FCF26D3AAC79FDC4" box="[770,856,584,607]" pageId="8" pageNumber="509">T. Pape</collectorName>
&amp; 
<collectorName id="A92C6AFB245D1971FC726D3AACD3FDFB" box="[898,1010,583,608]" pageId="8" pageNumber="509">N Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FBB56D35AE8BFD33" ID-GBIF-Occurrence="1563407795" collectingDate="2013-03-22" collectorName="Hand &amp; T. Pape &amp; N. Scharff" elevation="800" latitude="-7.837861" location="Mito Mitatu" longLatPrecision="1" longitude="36.845997" municipality="Udzungwa Mts Natl Park" pageId="8" pageNumber="509" specimenCount="6" specimenCount-male="1" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FBB56D35AB4EFDC4" box="[1093,1135,583,608]" pageId="8" pageNumber="509" type="male">1 ♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FB8E6D35ABE1FDFB" box="[1150,1216,583,608]" pageId="8" pageNumber="509" type="generic">5 juv.</specimenCount>
<collectingRegion id="C61DC1CF245D1971FB396D3AAAB9FDFB" box="[1225,1432,583,608]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, 
<collectingMunicipality id="E4029557245D1971FF676D1EAE9EFD1E" box="[151,447,620,645]" pageId="8" pageNumber="509">Udzungwa Mts Natl Park</collectingMunicipality>
, 
<location id="010659F6245D1971FE3D6D1EAD7AFD1E" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FE3D6D1EAD7AFD1E" box="[461,603,620,645]" latitude="-7.837861" longLatPrecision="1" longitude="36.845997" municipality="Udzungwa Mts Natl Park" name="Mito Mitatu" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Mito Mitatu</location>
, 
<geoCoordinate id="61ED69EA245D1971FD986D1EAC25FD1E" box="[616,772,620,645]" direction="south" orientation="latitude" pageId="8" pageNumber="509" precision="1" value="-7.837861">07°50’16.3”S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA245D1971FCE26D1EAC91FD1E" box="[786,944,620,645]" direction="east" orientation="longitude" pageId="8" pageNumber="509" precision="1" value="36.845997">36°50’45.6”E</geoCoordinate>
, 
<elevation id="8FF4E81E245D1971FC4E6D1EAB12FD1E" box="[958,1075,620,645]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="509" unit="m" value="800.0">
<quantity id="C321A2C8245D1971FC4E6D1EAB26FD1F" box="[958,1031,620,644]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="509" unit="m" value="800.0">800 m</quantity>
asl
</elevation>
, 
<collectingDate id="6023D005245D1971FBB06D1EAB9CFD1E" box="[1088,1213,620,645]" pageId="8" pageNumber="509" value="2013-03-22">
<date id="706729ED245D1971FBB06D1EAB9CFD1E" box="[1088,1213,620,645]" pageId="8" pageNumber="509" value="2013-03-22">22.iii.2013</date>
</collectingDate>
. 
<collectorName id="A92C6AFB245D1971FB3A6D1EAA26FD1E" box="[1226,1287,620,645]" pageId="8" pageNumber="509">Hand</collectorName>
collected. 
<collectorName id="A92C6AFB245D1971FA766D1FAFECFD3C" pageId="8" pageNumber="509">T. Pape</collectorName>
&amp; 
<collectorName id="A92C6AFB245D1971FF0C6DE2AE57FD33" box="[252,374,655,680]" pageId="8" pageNumber="509">N. Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FE386DFDAB8FFD56" ID-GBIF-Occurrence="1563407788" collectingDate="2012-09-03" collectorName="T. Pape &amp; N. Scharff" elevation="769" latitude="-7.8065" location="Mizimu Camp" longLatPrecision="1" longitude="36.852024" municipality="Udzungwa Mts Natl Park" pageId="8" pageNumber="509" specimenCount="9" specimenCount-female="5" specimenCount-male="2" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FE386DFDAD2FFD3C" box="[456,526,655,680]" pageId="8" pageNumber="509" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FDD06DFDAD47FD3C" box="[544,614,655,680]" pageId="8" pageNumber="509" type="female">5 ♀♀</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FD886DE2AD9CFD33" box="[632,701,655,680]" pageId="8" pageNumber="509" type="generic">2 juv.</specimenCount>
<collectingRegion id="C61DC1CF245D1971FD386DE2ACBBFD33" box="[712,922,655,680]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, 
<collectingMunicipality id="E4029557245D1971FC5B6DFDABFDFD33" box="[939,1244,655,680]" pageId="8" pageNumber="509">Udzungwa Mts Natl Park</collectingMunicipality>
, 
<location id="010659F6245D1971FB1D6DFDAAB9FD33" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FB1D6DFDAAB9FD33" box="[1261,1432,655,680]" latitude="-7.8065" longLatPrecision="1" longitude="36.852024" municipality="Udzungwa Mts Natl Park" name="Mizimu Camp" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Mizimu Camp</location>
, 
<geoCoordinate id="61ED69EA245D1971FF676DC6AE60FD57" box="[151,321,692,716]" direction="south" orientation="latitude" pageId="8" pageNumber="509" precision="1" value="-7.8065">07°48’23.40”S</geoCoordinate>
, 
<geoCoordinate id="61ED69EA245D1971FEBD6DC6AECBFD56" box="[333,490,692,717]" direction="east" orientation="longitude" pageId="8" pageNumber="509" precision="1" value="36.852024">36°51’7.29”E</geoCoordinate>
, 
<elevation id="8FF4E81E245D1971FE056DC6AD46FD56" box="[501,615,692,717]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="8" pageNumber="509" unit="m" value="769.0">
<quantity id="C321A2C8245D1971FE056DC6AD1DFD57" box="[501,572,692,717]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="8" pageNumber="509" unit="m" value="769.0">769 m</quantity>
asl
</elevation>
., 
<collectingDate id="6023D005245D1971FD876DC6ADC5FD56" box="[631,740,692,717]" pageId="8" pageNumber="509" value="2012-09-03">
<date id="706729ED245D1971FD876DC6ADC5FD56" box="[631,740,692,717]" pageId="8" pageNumber="509" value="2012-09-03">3.ix.2012</date>
</collectingDate>
, and collected. 
<collectorName id="A92C6AFB245D1971FC626DC7ACC4FD57" box="[914,997,693,716]" pageId="8" pageNumber="509">T. Pape</collectorName>
&amp; 
<collectorName id="A92C6AFB245D1971FBFA6DC7AB5FFD56" box="[1034,1150,692,717]" pageId="8" pageNumber="509">N. Scharff</collectorName>
leg.
</materialsCitation>
— 
<materialsCitation id="B4B10570245D1971FB3F6DC6ABB3FD6B" ID-GBIF-Occurrence="1563407814" collectingDate="1998-07-15" collectorName="L. A. Hansen" county="Luhega Forest Reserve" elevation="454" latitude="-8.383333" location="Udzungwa Scarp" longLatPrecision="1301" longitude="35.966667" municipality="Luhega Forest" pageId="8" pageNumber="509" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region">
<specimenCount id="12DFC4A4245D1971FB3F6DC6ABD5FD57" box="[1231,1268,692,717]" pageId="8" pageNumber="509" type="male">1 ♂</specimenCount>
<collectingCounty id="ED0777A1245D1971FB0B6DC6AFD0FD74" pageId="8" pageNumber="509">
<collectingMunicipality id="E4029557245D1971FB0B6DC6AABDFD57" box="[1275,1436,692,717]" pageId="8" pageNumber="509">Luhega Forest</collectingMunicipality>
Reserve
</collectingCounty>
(
<location id="010659F6245D1971FEF06DA5AEE7FD6B" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FEF06DA5AEE7FD6B" box="[256,454,727,752]" county="Luhega Forest Reserve" latitude="-8.383333" longLatPrecision="1301" longitude="35.966667" municipality="Luhega Forest" name="Udzungwa Scarp" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Udzungwa Scarp</location>
), 
<geoCoordinate id="61ED69EA245D1971FE296DA5AD18FD6B" box="[473,569,727,752]" direction="east" orientation="longitude" pageId="8" pageNumber="509" precision="925" value="35.966667">35°58’E</geoCoordinate>
, 
<geoCoordinate id="61ED69EA245D1971FDB46DAAAD82FD6B" box="[580,675,728,752]" direction="south" orientation="latitude" pageId="8" pageNumber="509" precision="925" value="-8.383333">08°23’S</geoCoordinate>
, 
<elevation id="8FF4E81E245D1971FD5E6DAAAC1BFD6B" box="[686,826,727,752]" metricMagnitude="2" metricUnit="m" metricValue="4.54" metricValueMax="9.0" metricValueMin="0.08" pageId="8" pageNumber="509" unit="m" value="454.0" valueMax="900.0" valueMin="8.0">
<quantity id="C321A2C8245D1971FD5E6DAAAC30FD74" box="[686,785,728,752]" metricMagnitude="2" metricUnit="m" metricValue="4.54" metricValueMax="9.0" metricValueMin="0.08" pageId="8" pageNumber="509" unit="m" value="454.0" valueMax="900.0" valueMin="8.0">8–900 m</quantity>
asl
</elevation>
, 
<collectingDate id="6023D005245D1971FCCF6DA5ACE1FD6B" box="[831,960,727,752]" pageId="8" pageNumber="509" value="1998-07-15">
<date id="706729ED245D1971FCCF6DA5ACE1FD6B" box="[831,960,727,752]" pageId="8" pageNumber="509" value="1998-07-15">15.vii.1998</date>
</collectingDate>
, 
<collectorName id="A92C6AFB245D1971FC3C6DAAAB40FD74" box="[972,1121,728,752]" pageId="8" pageNumber="509">L. A. Hansen</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="04660F2D245D1971FF376D8FAEA8FCA3" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
<materialsCitation id="B4B10570245D1971FF376D8FAEA8FCA3" ID-GBIF-Occurrence="1563407816" collectingDate="1984-11-22" collectingDateMax="1984-11-25" collectingDateMin="1984-11-22" collectorName="N. Scharff" county="Kanga Mt." elevation="450" location="Kanga Forest" municipality="Kanga Forest Reserve" pageId="8" pageNumber="509" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Morogoro Region">
<collectingCounty id="ED0777A1245D1971FF376D8FAE65FC8F" box="[199,324,765,789]" pageId="8" pageNumber="509">Kanga Mt.</collectingCounty>
: 
<specimenCount id="12DFC4A4245D1971FEA16D8EAE5FFC8F" box="[337,382,764,788]" pageId="8" pageNumber="509" type="male">1 ♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FE7C6D8EAE96FC8F" box="[396,439,764,788]" pageId="8" pageNumber="509" type="female">1 ♀</specimenCount>
<collectingRegion id="C61DC1CF245D1971FE306D8FADAFFC8E" box="[448,654,764,789]" country="Tanzania" name="Morogoro" pageId="8" pageNumber="509">Morogoro Region</collectingRegion>
, 
<collectingMunicipality id="E4029557245D1971FD6B6D8FACB9FC8F" box="[667,920,765,789]" pageId="8" pageNumber="509">
<location id="010659F6245D1971FD6B6D8FAC13FC8F" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FD6B6D8FAC13FC8F" box="[667,818,765,789]" county="Kanga Mt." municipality="Kanga Forest Reserve" name="Kanga Forest" pageId="8" pageNumber="509" stateProvince="Morogoro Region">Kanga Forest</location>
Reserve
</collectingMunicipality>
, 
<elevation id="8FF4E81E245D1971FC556D8EAB71FC8E" box="[933,1104,764,789]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="509" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<quantity id="C321A2C8245D1971FC556D8EAB04FC8F" box="[933,1061,764,789]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="509" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400–500 m</quantity>
asl
</elevation>
, 
<collectingDate id="6023D005245D1971FBAC6D8EAA21FC8E" box="[1116,1280,764,789]" pageId="8" pageNumber="509" value="1984-11-22" valueMax="1984-11-25" valueMin="1984-11-22">
<date id="706729ED245D1971FBAC6D8EAA21FC8E" box="[1116,1280,764,789]" pageId="8" pageNumber="509" value="1984-11-22" valueMax="1984-11-25" valueMin="1984-11-22">22–25.xi.1984</date>
</collectingDate>
, lowland rain forest, 
<collectorName id="A92C6AFB245D1971FF146C52AE78FCA3" box="[228,345,799,824]" pageId="8" pageNumber="509">N. Scharff</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="04660F2D245D1971FF376C36ABD0FCC6" blockId="8.[151,1437,151,2013]" box="[199,1265,836,861]" pageId="8" pageNumber="509">
<materialsCitation id="B4B10570245D1971FF376C36ABD0FCC6" ID-GBIF-Occurrence="1563407806" box="[199,1265,836,861]" collectingDate="1993-09-10" collectingDateMax="1993-09-24" collectingDateMin="1993-09-10" collectorName="M. Andersen" location="Rubeho Mts." pageId="8" pageNumber="509" specimenCount="35" specimenCount-female="5" specimenCount-male="2">
<location id="010659F6245D1971FF376C36AE7DFCC7" LSID="urn:lsid:plazi:treatment:8C70BE3B24521975FF676E4AAE76FC53:010659F6245D1971FF376C36AE7DFCC7" box="[199,348,836,861]" name="Rubeho Mts." pageId="8" pageNumber="509">Rubeho Mts.</location>
: 
<specimenCount id="12DFC4A4245D1971FE966C36AE86FCC7" box="[358,423,836,860]" pageId="8" pageNumber="509" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FE436C36AED2FCC7" box="[435,499,836,861]" pageId="8" pageNumber="509" type="female">5 ♀♀</specimenCount>
, 
<specimenCount id="12DFC4A4245D1971FDF06C36AD6CFCC6" box="[512,589,836,861]" pageId="8" pageNumber="509" type="generic">28 juv.</specimenCount>
forest 
<locationDeviation id="15D4160F245D1971FD6A6C36AC52FCC6" box="[666,883,836,861]" pageId="8" pageNumber="509">
<quantity id="C321A2C8245D1971FD6A6C36ADF2FCC6" box="[666,723,836,861]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="509" unit="km" value="5.0">5 km</quantity>
SW Madizini
</locationDeviation>
, 
<collectingDate id="6023D005245D1971FC8E6C36AB03FCC6" box="[894,1058,836,861]" pageId="8" pageNumber="509" value="1993-09-10" valueMax="1993-09-24" valueMin="1993-09-10">
<date id="706729ED245D1971FC8E6C36AB03FCC6" box="[894,1058,836,861]" pageId="8" pageNumber="509" value="1993-09-10" valueMax="1993-09-24" valueMin="1993-09-10">10–24.ix.1993</date>
</collectingDate>
, 
<collectorName id="A92C6AFB245D1971FBDE6C37ABE1FCC6" box="[1070,1216,836,861]" pageId="8" pageNumber="509">M. Andersen</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="04660F2D245D1971FF376C15AC67FC1B" blockId="8.[151,1437,151,2013]" box="[199,838,871,896]" pageId="8" pageNumber="509">
<emphasis id="36ADD33F245D1971FF376C15AE44FC1B" bold="true" box="[199,357,871,896]" pageId="8" pageNumber="509">Type locality.</emphasis>
TANZANIA: East Usambara Mts, Amani.
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA6245D1972FF376CFEAD72FA5E" lastPageId="11" lastPageNumber="512" pageId="8" pageNumber="509" type="description">
<paragraph id="04660F2D245D1971FF376CFEAB2FFBAE" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
<emphasis id="36ADD33F245D1971FF376CFEAE78FC3E" bold="true" box="[199,345,908,933]" pageId="8" pageNumber="509">Description.</emphasis>
Although no consistent morphological differences could be found between specimens from the East Usambara, Kanga, Rubeho Uluguru and Udzungwas Mts there seem to be some colour differences (
<figureCitation id="9CE213A8245D1971FABE6CDDAAB0FC53" box="[1358,1425,943,968]" captionStart="FIGURE 1" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[200,1379,237,1858]" captionTargetId="figure@4.[151,1436,193,1911]" captionTargetPageId="4" captionText="FIGURE 1. Tropostreptus hamatus, live specimens. A: pink-legged specimen (East Usambara Mts), S. Kuhlmann photo. B: a copulating pair of the pink-legged form, S. Kuhlmann photo, C: darker, dark-legged specimen (Udzungwa Mts.), N. Scharff photo." httpUri="https://zenodo.org/record/803706/files/figure.png" pageId="8" pageNumber="509">Fig. 1</figureCitation>
) and in part also differences in body dimensions. Therefor a separate description is given for specimens from the 
<typeStatus id="DB62B18F245D1971FF676C8AAFE6FB8B" box="[151,199,1016,1040]" pageId="8" pageNumber="509">type</typeStatus>
locality, Amani (East Usambara Mts) and its immediate surroundings, followed by notes on specimens from the other mountains, focusing on differences 
<emphasis id="36ADD33F245D1971FD606B6FAD86FBAF" box="[656,679,1053,1076]" italics="true" pageId="8" pageNumber="509">vs</emphasis>
. the East Usambara population.
</paragraph>
<paragraph id="04660F2D245D1971FF376B4DACCFFB3B" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
<emphasis id="36ADD33F245D1971FF376B4DAC2EFBC3" bold="true" box="[199,783,1087,1112]" pageId="8" pageNumber="509">Specimens from East Usambara Mts (topotypes).</emphasis>
SIZE (
<figureCitation id="9CE213A8245D1971FCA86B4DAC88FBC3" box="[856,937,1087,1112]" captionStart="FIGURE 11" captionStartId="16.[151,250,1055,1077]" captionTargetBox="[188,1407,227,1018]" captionTargetId="figure@16.[151,1436,193,1060]" captionTargetPageId="16" captionText="FIGURE 11. Size of Tropostreptus hamatus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males." httpUri="https://zenodo.org/record/803726/files/figure.png" pageId="8" pageNumber="509">Fig. 11</figureCitation>
). Length 
<quantity id="C321A2C8245D1971FBE56B32AB55FBCC" box="[1045,1140,1088,1112]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="8" pageNumber="509" unit="cm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 cm</quantity>
(up to 
<quantity id="C321A2C8245D1971FB356B32AA2BFBCC" box="[1221,1290,1088,1112]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="509" unit="cm" value="14.0">14 cm</quantity>
according to 
<bibRefCitation id="604872DC245D1971FF676B16AE1BFBE7" author="Sigling" box="[151,314,1124,1149]" pageId="8" pageNumber="509" refString="Sigling, S. (2010) PraxisRatgeber Tausendfusser. Edition Chimaira, Frankfurt am Main, 206 pp." type="book" year="2010">Sigling [2010]</bibRefCitation>
but this may refer to a different species); vertical body diameter 
<quantity id="C321A2C8245D1971FBE56B16ABBDFBE7" box="[1045,1180,1124,1149]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="8.1" metricValueMin="6.6" pageId="8" pageNumber="509" unit="mm" value="7.35" valueMax="8.1" valueMin="6.6">6.6–8.1 mm</quantity>
(males), up to 
<quantity id="C321A2C8245D1971FAB66B16AABCFBE7" box="[1350,1437,1124,1148]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="8" pageNumber="509" unit="mm" value="9.1">9.1 mm</quantity>
(females); 53–56 podous rings (one exceptional male with 59 podous rings).
</paragraph>
<paragraph id="04660F2D245D1971FF376BDEAC8EFA96" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
COLOUR (
<figureCitation id="9CE213A8245D1971FEC76BDEAE5BFB5F" box="[311,378,1196,1221]" captionStart="FIGURE 1" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[200,1379,237,1858]" captionTargetId="figure@4.[151,1436,193,1911]" captionTargetPageId="4" captionText="FIGURE 1. Tropostreptus hamatus, live specimens. A: pink-legged specimen (East Usambara Mts), S. Kuhlmann photo. B: a copulating pair of the pink-legged form, S. Kuhlmann photo, C: darker, dark-legged specimen (Udzungwa Mts.), N. Scharff photo." httpUri="https://zenodo.org/record/803706/files/figure.png" pageId="8" pageNumber="509">Fig. 1</figureCitation>
A, B). Live specimens ringed in light and dark brown; head and antennae reddish, legs pink. After 24 years in alcohol head and antennae yellow, collum yellow with dark margins, body rings yellowish back to ozopore, then dark brown, then amber, legs light brown, telson yellow.
</paragraph>
<paragraph id="04660F2D245D1971FF376A6AADABF9EE" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
GONOPODS (
<figureCitation id="9CE213A8245D1971FEA56A65AEBCFAB4" box="[341,413,1303,1328]" captionStart="FIGURE 7" captionStartId="13.[151,250,1869,1891]" captionTargetBox="[207,1387,199,1819]" captionTargetId="figure@13.[151,1436,155,1848]" captionTargetPageId="13" captionText="FIGURE 7. Tropostretus spp., gonopods, anterior view. A: T. hamatus, East Usambara Mts, B: T. hamatus, Udzungwa Mts., left telopodite missing, C: T. sigmatospinus sp. nov., D: T. microcephalus sp. nov., E: T. droides sp. nov., F: T. kipunji sp. nov. Scales 0.5 mm. atp = antetorsal process, bh = birdhead-shaped metaplical process, lpl = apicolateral proplical lobe, map = mesapical metaplical process, mp = metaplica, mpl = mesapical proplical lobe, pp = proplica, px = paracoxite, st = sternum, tlp = telopodite." httpUri="https://zenodo.org/record/803720/files/figure.png" pageId="8" pageNumber="509">Figs 7</figureCitation>
A, 8 A, 12 A). Sternum triangular, not reaching as far distad as paracoxites, lateral edges slightly angled at mid-length, giving the sternum a somewhat “houselike” outline. Proplica ending in blunttriangular mesapical lobe (
<emphasis id="36ADD33F245D1971FE3F6A2DAED8FAE3" box="[463,505,1375,1400]" italics="true" pageId="8" pageNumber="509">mpl</emphasis>
) separated by conspicuous sinus from large apico-lateral, sub-semicircular lobe (
<emphasis id="36ADD33F245D1971FF506AF6AF9FFA06" box="[160,190,1412,1437]" italics="true" pageId="8" pageNumber="509">lpl</emphasis>
). Mesapical and apico-lateral lobes reaching equally far distad. Metaplica in anterior view basally with straight ridge flanking sternum and continuing ca. to mid-length of coxite, apically with rounded mesal process (
<emphasis id="36ADD33F245D1971FA926ADBAAB3FA5B" box="[1378,1426,1449,1472]" italics="true" pageId="8" pageNumber="509">map</emphasis>
) separated by deep sinus from lateral birdhead-shaped process (
<emphasis id="36ADD33F245D1971FC9D6ABEACA8FA7E" box="[877,905,1484,1509]" italics="true" pageId="8" pageNumber="509">bh</emphasis>
), lateral “beak” of the latter moderately long, pointed, directed laterad but curving slightly anteriad. Mesal process not reaching as far distad as birdhead-shaped process. Telopodite shortly after emergence from coxa with very long, slender, pointed, antetorsal process (
<emphasis id="36ADD33F245D1971FA9F6967AAB2F9B7" box="[1391,1427,1557,1580]" italics="true" pageId="8" pageNumber="509">atp</emphasis>
) making a broad 180° loop; direction of process first basad, then laterad, then apicad. Telopodite distal to antetorsal process slender, simple, without outgrowths.
</paragraph>
<paragraph id="04660F2D245D1971FF37690DAED5F926" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
<emphasis id="36ADD33F245D1971FF37690DAD6FF903" bold="true" box="[199,590,1663,1688]" pageId="8" pageNumber="509">Specimens from Udzungwa Mts.</emphasis>
SIZE (
<figureCitation id="9CE213A8245D1971FD6A690DADCCF903" box="[666,749,1663,1688]" captionStart="FIGURE 11" captionStartId="16.[151,250,1055,1077]" captionTargetBox="[188,1407,227,1018]" captionTargetId="figure@16.[151,1436,193,1060]" captionTargetPageId="16" captionText="FIGURE 11. Size of Tropostreptus hamatus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males." httpUri="https://zenodo.org/record/803726/files/figure.png" pageId="8" pageNumber="509">Fig. 11</figureCitation>
). Vertical body diameter 
<quantity id="C321A2C8245D1971FBFE690DABB6F90C" box="[1038,1175,1663,1688]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="7.1" metricValueMin="5.4" pageId="8" pageNumber="509" unit="mm" value="6.25" valueMax="7.1" valueMin="5.4">5.4–7.1 mm</quantity>
(males), up to 
<quantity id="C321A2C8245D1971FAB569F2AABCF90C" box="[1349,1437,1664,1687]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="8" pageNumber="509" unit="mm" value="7.7">7.7 mm</quantity>
(females); 53–57 podous rings.
</paragraph>
<paragraph id="04660F2D245D1971FF3769B5ACE1F97B" blockId="8.[151,1437,151,2013]" box="[199,960,1735,1760]" pageId="8" pageNumber="509">Structural characters as in topotypes, with the following exceptions:</paragraph>
<paragraph id="04660F2D245D1971FF37699EAB89F8EB" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
COLOUR (
<figureCitation id="9CE213A8245D1971FEC5699EAE56F89F" box="[309,375,1772,1797]" captionStart="FIGURE 1" captionStartId="4.[151,250,1933,1955]" captionTargetBox="[200,1379,237,1858]" captionTargetId="figure@4.[151,1436,193,1911]" captionTargetPageId="4" captionText="FIGURE 1. Tropostreptus hamatus, live specimens. A: pink-legged specimen (East Usambara Mts), S. Kuhlmann photo. B: a copulating pair of the pink-legged form, S. Kuhlmann photo, C: darker, dark-legged specimen (Udzungwa Mts.), N. Scharff photo." httpUri="https://zenodo.org/record/803706/files/figure.png" pageId="8" pageNumber="509">Fig. 1</figureCitation>
C). Live specimens dark brown to blackish, telson slightly lighter, legs dark brown to blackish. After 2 years in alcohol head yellow, collum blackish brown, rings yellowish brown back to ozopores, then blackish, then amber. After 3 years in alcohol: overall colour dark brown, anterior half of prozona brownish orange, telson brownish orange, but posterior part of anal valves dark brown to blackish; legs brown.
</paragraph>
<paragraph id="04660F2D245D1971FF37680EAC3AF846" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
GONOPODS (
<figureCitation id="9CE213A8245D1971FEA3680EAEBBF80F" box="[339,410,1916,1941]" captionStart="FIGURE 7" captionStartId="13.[151,250,1869,1891]" captionTargetBox="[207,1387,199,1819]" captionTargetId="figure@13.[151,1436,155,1848]" captionTargetPageId="13" captionText="FIGURE 7. Tropostretus spp., gonopods, anterior view. A: T. hamatus, East Usambara Mts, B: T. hamatus, Udzungwa Mts., left telopodite missing, C: T. sigmatospinus sp. nov., D: T. microcephalus sp. nov., E: T. droides sp. nov., F: T. kipunji sp. nov. Scales 0.5 mm. atp = antetorsal process, bh = birdhead-shaped metaplical process, lpl = apicolateral proplical lobe, map = mesapical metaplical process, mp = metaplica, mpl = mesapical proplical lobe, pp = proplica, px = paracoxite, st = sternum, tlp = telopodite." httpUri="https://zenodo.org/record/803720/files/figure.png" pageId="8" pageNumber="509">Figs 7</figureCitation>
B, 8 B). Sternum triangular, lateral edges tending to be more or less regularly arched, rather than angled. Mesal process of metaplica (
<emphasis id="36ADD33F245D1971FD8068D3AD81F823" box="[624,672,1953,1976]" italics="true" pageId="8" pageNumber="509">map</emphasis>
) often separated from lateral birdhead-shaped process (
<emphasis id="36ADD33F245D1971FAE868D2AA15F822" box="[1304,1332,1952,1977]" italics="true" pageId="8" pageNumber="509">bh</emphasis>
) only by narrow sinus and often reaching as far distad as the latter.
</paragraph>
<caption id="50A65FA5245C1970FF6769FBAC5DF960" httpUri="https://zenodo.org/record/803714/files/figure.png" pageId="9" pageNumber="510" targetBox="[212,1376,195,1600]" targetPageId="9">
<paragraph id="04660F2D245C1970FF6769FBAC5DF960" blockId="9.[151,1437,1673,1787]" pageId="9" pageNumber="510">
<emphasis id="36ADD33F245C1970FF6769FBAE34F905" bold="true" box="[151,277,1673,1695]" pageId="9" pageNumber="510">FIGURE 4.</emphasis>
<taxonomicName id="C3D974AE245C1970FEED69F8AE89F904" box="[285,424,1674,1695]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="528" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="36ADD33F245C1970FEED69F8AE89F904" box="[285,424,1674,1695]" italics="true" pageId="9" pageNumber="510">Tropostreptus</emphasis>
</taxonomicName>
spp., mouthparts. A: 
<taxonomicName id="C3D974AE245C1970FD7B69F8ADD0F904" box="[651,753,1673,1695]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="droides" status="sp. nov.">
<emphasis id="36ADD33F245C1970FD7B69F8ADD0F904" box="[651,753,1673,1695]" italics="true" pageId="9" pageNumber="510">T. droides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2D9E6E44245C1970FD0969FBAC6AF905" box="[761,843,1673,1694]" pageId="9" pageNumber="510" rank="species">
<emphasis id="36ADD33F245C1970FD0969FBAC6AF905" bold="true" box="[761,843,1673,1694]" pageId="9" pageNumber="510">sp. nov.</emphasis>
</taxonomicNameLabel>
, left mandibular gnathal lobe, dorsal/anterior view, B: 
<taxonomicName id="C3D974AE245C1970FA7969F8AFFCF926" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="kipunji" status="sp. nov.">
<emphasis id="36ADD33F245C1970FA7969F8AFFCF926" italics="true" pageId="9" pageNumber="510">T. kipunji</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2D9E6E44245C1970FF1469DAAE14F926" box="[228,309,1704,1725]" pageId="9" pageNumber="510" rank="species">
<emphasis id="36ADD33F245C1970FF1469DAAE14F926" bold="true" box="[228,309,1704,1725]" pageId="9" pageNumber="510">sp. nov.</emphasis>
</taxonomicNameLabel>
, right mandibular gnathal lobe, mesal view. C: 
<taxonomicName id="C3D974AE245C1970FCEB69DBACA8F926" box="[795,905,1703,1726]" class="Diplopoda" family="Spirostreptidae" genus="Tropostretus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F245C1970FCEB69DBACA8F926" box="[795,905,1703,1726]" italics="true" pageId="9" pageNumber="510">T. hamatus</emphasis>
</taxonomicName>
, male, gnathochilarium and right mandible, oblique ventral view.—Scales 0.1 mm. 
<emphasis id="36ADD33F245C1970FE1369B5AD21F947" box="[483,512,1735,1756]" italics="true" pageId="9" pageNumber="510">me</emphasis>
= mentum, 
<emphasis id="36ADD33F245C1970FD7669B4AD9BF947" box="[646,698,1734,1756]" italics="true" pageId="9" pageNumber="510">mf =</emphasis>
molar furrow, 
<emphasis id="36ADD33F245C1970FCAD69B4ACA1F947" box="[861,896,1734,1756]" italics="true" pageId="9" pageNumber="510">msl</emphasis>
= mandibular stipital lobe, 
<emphasis id="36ADD33F245C1970FB5B69B4ABE5F947" box="[1195,1220,1734,1756]" italics="true" pageId="9" pageNumber="510">od</emphasis>
= odontomere, 
<emphasis id="36ADD33F245C1970FA8069B4AAA5F947" box="[1392,1412,1734,1756]" italics="true" pageId="9" pageNumber="510">pl</emphasis>
= pectinate lamellae, 
<emphasis id="36ADD33F245C1970FEAB6994AE5BF960" box="[347,378,1766,1787]" italics="true" pageId="9" pageNumber="510">pm</emphasis>
= prementum, 
<emphasis id="36ADD33F245C1970FDE56994AD0AF960" box="[533,555,1766,1787]" italics="true" pageId="9" pageNumber="510">ps</emphasis>
= psectromere, 
<emphasis id="36ADD33F245C1970FD3E6994ADC2F960" box="[718,739,1766,1787]" italics="true" pageId="9" pageNumber="510">se</emphasis>
= sectile edge.
</paragraph>
</caption>
<paragraph id="04660F2D245C1970FF376855AED2F8FF" blockId="9.[151,1437,1831,2036]" pageId="9" pageNumber="510">
<emphasis id="36ADD33F245C1970FF376855AD1DF8DB" bold="true" box="[199,572,1831,1856]" pageId="9" pageNumber="510">Specimens from Uluguru Mts.</emphasis>
SIZE (
<figureCitation id="9CE213A8245C1970FD7E6855ADC5F8DB" box="[654,740,1831,1856]" captionStart="FIGURE 11" captionStartId="16.[151,250,1055,1077]" captionTargetBox="[188,1407,227,1018]" captionTargetId="figure@16.[151,1436,193,1060]" captionTargetPageId="16" captionText="FIGURE 11. Size of Tropostreptus hamatus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males." httpUri="https://zenodo.org/record/803726/files/figure.png" pageId="9" pageNumber="510">Fig. 11</figureCitation>
). Vertical body diameter 
<quantity id="C321A2C8245C1970FBE16855ABA9F8A4" box="[1041,1160,1831,1856]" metricMagnitude="-3" metricUnit="m" metricValue="6.95" metricValueMax="7.9" metricValueMin="6.0" pageId="9" pageNumber="510" unit="mm" value="6.95" valueMax="7.9" valueMin="6.0">6–7.9 mm</quantity>
(males), up to 
<quantity id="C321A2C8245C1970FAB26855AABCF8A4" box="[1346,1437,1831,1856]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="9" pageNumber="510" unit="mm" value="8.6">8.6 mm</quantity>
(females), 53–60 podous rings.
</paragraph>
<paragraph id="04660F2D245C1970FF37681DAD5EF837" blockId="9.[151,1437,1831,2036]" pageId="9" pageNumber="510">Similar to topotypes in all respects, including detailed gonopod shape (except that the lateral margins of sternum are slightly arched in some males).</paragraph>
<paragraph id="04660F2D245C1970FF3768C5AE0FF86F" blockId="9.[151,1437,1831,2036]" pageId="9" pageNumber="510">
<emphasis id="36ADD33F245C1970FF3768C5AD08F84B" bold="true" box="[199,553,1975,2000]" pageId="9" pageNumber="510">Specimens from Rubeho Mts.</emphasis>
SIZE (
<figureCitation id="9CE213A8245C1970FD8368C5ADE4F84B" box="[627,709,1975,2000]" captionStart="FIGURE 11" captionStartId="16.[151,250,1055,1077]" captionTargetBox="[188,1407,227,1018]" captionTargetId="figure@16.[151,1436,193,1060]" captionTargetPageId="16" captionText="FIGURE 11. Size of Tropostreptus hamatus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males." httpUri="https://zenodo.org/record/803726/files/figure.png" pageId="9" pageNumber="510">Fig. 11</figureCitation>
). Body diameter 
<quantity id="C321A2C8245C1970FC7968C5ACC1F854" box="[905,992,1975,2000]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="510" unit="mm" value="7.6">7.6 mm</quantity>
(male), up to 
<quantity id="C321A2C8245C1970FB7068CAABF7F854" box="[1152,1238,1976,2000]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="510" unit="mm" value="8.7">8.7 mm</quantity>
(females), 56–57 podous rings.
</paragraph>
<caption id="50A65FA5245F1973FF67684FAE6BF877" httpUri="https://zenodo.org/record/803716/files/figure.png" pageId="10" pageNumber="511" targetBox="[196,1396,193,1802]" targetPageId="10">
<paragraph id="04660F2D245F1973FF67684FAE6BF877" blockId="10.[151,1436,1853,2028]" pageId="10" pageNumber="511">
<emphasis id="36ADD33F245F1973FF67684FAE33F8C8" bold="true" box="[151,274,1853,1875]" pageId="10" pageNumber="511">FIGURE 5.</emphasis>
Details of non-gonopodal morphology. A–C: ventral party of body rings, A–B: 
<taxonomicName id="C3D974AE245F1973FBC8684CAA3EF8C8" box="[1080,1311,1853,1875]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F245F1973FBC8684CAA3EF8C8" box="[1080,1311,1853,1875]" italics="true" pageId="10" pageNumber="511">Tropostreptus hamatus</emphasis>
</taxonomicName>
, A: external view, showing touching posterior pleurotergal lobes, B: internal view, showing overlapping posterior pleurotergal lobes, C: 
<taxonomicName id="C3D974AE245F1973FF676808AD4CF80B" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[151,621,1914,1936]" class="Diplopoda" family="Spirostreptidae" genus="Dendrostreptus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="macracanthus">
<emphasis id="36ADD33F245F1973FF676808AEEBF80B" box="[151,458,1914,1936]" italics="true" pageId="10" pageNumber="511">Dendrostreptus macracanthus</emphasis>
(Attems, 1914)
</taxonomicName>
for comparison, external view, showing separated posterior pleurotergal lobes. D–F: 
<emphasis id="36ADD33F245F1973FF2168E8AE9EF834" box="[209,447,1945,1967]" italics="true" pageId="10" pageNumber="511">
<taxonomicName id="C3D974AE245F1973FF2168E8AE9AF834" box="[209,443,1945,1967]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="hamatus">Tropostreptus hamatus</taxonomicName>
.
</emphasis>
D: detail of prozonal sculpture, showing fine furrows with enlarged cuticular scutes. E: limbus. F: male, midbody legs.—Scales 0.2 mm (A–C, F), 0.02 mm (D), 0.005 mm (E). 
<emphasis id="36ADD33F245F1973FC5268CAACFEF855" box="[930,991,1976,1998]" italics="true" pageId="10" pageNumber="511">pad =</emphasis>
pads on postfemur and tibia, 
<emphasis id="36ADD33F245F1973FAF868CAAA08F855" box="[1288,1321,1976,1998]" italics="true" pageId="10" pageNumber="511">ppl</emphasis>
= posterior pleurotergal lobe.
</paragraph>
</caption>
<caption id="50A65FA5245E1972FF676B97ADC6FA81" httpUri="https://zenodo.org/record/803718/files/figure.png" pageId="11" pageNumber="512" targetBox="[183,1413,189,1203]" targetPageId="11">
<paragraph id="04660F2D245E1972FF676B97ADC6FA81" blockId="11.[151,1437,1253,1306]" pageId="11" pageNumber="512">
<emphasis id="36ADD33F245E1972FF676B97AE32FB61" bold="true" box="[151,275,1253,1275]" pageId="11" pageNumber="512">FIGURE 6.</emphasis>
<taxonomicName id="C3D974AE245E1972FEEA6B94AD23FB60" box="[282,514,1253,1275]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F245E1972FEEA6B94AD23FB60" box="[282,514,1253,1275]" italics="true" pageId="11" pageNumber="512">Tropostreptus hamatus</emphasis>
</taxonomicName>
, first pair of male legs. A: oral aspect. B: latero-basal field of setae on right coxosternum. Scales 0.5 mm (A), 0.2 mm (B). 
<emphasis id="36ADD33F245E1972FE136A76AD25FA81" box="[483,516,1284,1306]" italics="true" pageId="11" pageNumber="512">pfp</emphasis>
= prefemoral process.
</paragraph>
</caption>
<paragraph id="04660F2D245E1972FF376A30ADAEFAC1" blockId="11.[151,1437,1345,2018]" box="[199,655,1345,1370]" pageId="11" pageNumber="512">Legs yellowish after 23 years in alcohol.</paragraph>
<paragraph id="04660F2D245E1972FF376A17AFF7FA39" blockId="11.[151,1437,1345,2018]" pageId="11" pageNumber="512">
<emphasis id="36ADD33F245E1972FF376A17AD32FAE5" bold="true" box="[199,531,1381,1406]" pageId="11" pageNumber="512">Specimens from Kanga Mt.</emphasis>
SIZE (
<figureCitation id="9CE213A8245E1972FDAE6A17AD93FAE5" box="[606,690,1381,1406]" captionStart="FIGURE 11" captionStartId="16.[151,250,1055,1077]" captionTargetBox="[188,1407,227,1018]" captionTargetId="figure@16.[151,1436,193,1060]" captionTargetPageId="16" captionText="FIGURE 11. Size of Tropostreptus hamatus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males." httpUri="https://zenodo.org/record/803726/files/figure.png" pageId="11" pageNumber="512">Fig. 11</figureCitation>
). Body diameter 
<quantity id="C321A2C8245E1972FC876A14ACEEFAE6" box="[887,975,1382,1406]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="11" pageNumber="512" unit="mm" value="8.1">8.1 mm</quantity>
(male), 
<quantity id="C321A2C8245E1972FBDF6A17ABA7FAE6" box="[1071,1158,1381,1406]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="11" pageNumber="512" unit="mm" value="8.6">8.6 mm</quantity>
(female), 58–59 podous rings.
</paragraph>
<paragraph id="04660F2D245E1972FF376ADFAD72FA5E" blockId="11.[151,1437,1345,2018]" box="[199,595,1453,1478]" pageId="11" pageNumber="512">Similar to topotypes in all respects.</paragraph>
</subSubSection>
<subSubSection id="4CC35CA6245E1972FF376AA3AB05F97D" pageId="11" pageNumber="512" type="distribution">
<paragraph id="04660F2D245E1972FF376AA3AECCF9CD" blockId="11.[151,1437,1345,2018]" pageId="11" pageNumber="512">
<emphasis id="36ADD33F245E1972FF376AA3AED6FA71" bold="true" box="[199,503,1489,1514]" pageId="11" pageNumber="512">Distribution and habitat.</emphasis>
<taxonomicName id="C3D974AE245E1972FDF26AA0AC25FA71" box="[514,772,1489,1514]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F245E1972FDF26AA0AC25FA71" box="[514,772,1489,1514]" italics="true" pageId="11" pageNumber="512">Tropostreptus hamatus</emphasis>
</taxonomicName>
is known from the East Usambara, Uluguru, Udzungwa, Rubeho and Kanga Mts., recorded at altitudes 250–1700 (1750?) m asl. 
<bibRefCitation id="604872DC245E1972FC4A6A87ABC4F995" author="Enghoff" box="[954,1253,1525,1550]" pageId="11" pageNumber="512" refString="Enghoff, I. B. &amp; Enghoff, H. (1976) Notes on myriapods observed and collected in Tanzania and Kenya during the summer 1974. Xeroxed report, Zoological Museum, University of Copenhagen, Copenhagen, 19 pp." type="book" year="1976">Enghoff &amp; Enghoff (1976)</bibRefCitation>
most frequently found 
<taxonomicName id="C3D974AE245E1972FF2F6968AE7BF9A9" box="[223,346,1561,1586]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F245E1972FF2F6968AE7BF9A9" box="[223,346,1561,1586]" italics="true" pageId="11" pageNumber="512">T. hamatus</emphasis>
</taxonomicName>
(recorded as “ 
<taxonomicName id="C3D974AE245E1972FE0F6968ADB3F9AA" box="[511,658,1562,1585]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="36ADD33F245E1972FE0F6968ADB3F9AA" box="[511,658,1562,1585]" italics="true" pageId="11" pageNumber="512">Spirostreptus</emphasis>
</taxonomicName>
sp. 1”) in decaying wood; several of the newly studied samples were taken in similar microhabitats.
</paragraph>
<paragraph id="04660F2D245E1972FF376913AB05F97D" blockId="11.[151,1437,1345,2018]" pageId="11" pageNumber="512">
<emphasis id="36ADD33F245E1972FF376913AEECF9E1" bold="true" box="[199,461,1633,1658]" pageId="11" pageNumber="512">Coexisting congeners.</emphasis>
Coexisting with 
<taxonomicName id="C3D974AE245E1972FD656910AC31F9E2" box="[661,784,1634,1657]" class="Diplopoda" family="Spirostreptidae" genus="Epistreptus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="austerus">
<emphasis id="36ADD33F245E1972FD656910AC31F9E2" box="[661,784,1634,1657]" italics="true" pageId="11" pageNumber="512">T. austerus</emphasis>
</taxonomicName>
in the East Usambara Mts, with 
<taxonomicName id="C3D974AE245E1972FB636910AA70F9E2" box="[1171,1361,1634,1657]" class="Diplopoda" family="Spirostreptidae" genus="Tropostretus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="sigmatospinus">
<emphasis id="36ADD33F245E1972FB636910AA70F9E2" box="[1171,1361,1634,1657]" italics="true" pageId="11" pageNumber="512">T. sigmatospinus</emphasis>
</taxonomicName>
in the Uluguru Mts (Lupanga East and Mkungwe Forest Reserve) and the Udzungwa Mts (Mwanihana Forest Reserve / Udzungwa Mts National Park), with 
<taxonomicName id="C3D974AE245E1972FDC169D8ADCDF95A" box="[561,748,1706,1729]" class="Diplopoda" family="Spirostreptidae" genus="Tropostretus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="sigmatospinus">
<emphasis id="36ADD33F245E1972FDC169D8ADCDF95A" box="[561,748,1706,1729]" italics="true" pageId="11" pageNumber="512">T. sigmatospinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C3D974AE245E1972FCD269D8AA10F95A" authority="in Kanga Forest Reserve, Kanga Mt." authorityName="in Kanga Forest Reserve, Kanga Mt." box="[802,1329,1705,1730]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="severus">
<emphasis id="36ADD33F245E1972FCD269D8ACB0F95A" box="[802,913,1706,1729]" italics="true" pageId="11" pageNumber="512">T. severus</emphasis>
in Kanga Forest Reserve, Kanga Mt.
</taxonomicName>
, and with 
<taxonomicName id="C3D974AE245E1972FF6769BCAE77F97D" box="[151,342,1741,1766]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="microcephalus">
<emphasis id="36ADD33F245E1972FF6769BCAE77F97D" box="[151,342,1741,1766]" italics="true" pageId="11" pageNumber="512">T. microcephalus</emphasis>
</taxonomicName>
in the Udzungwa Mts. (
<collectingRegion id="C61DC1CF245E1972FD9469BCADBCF97D" box="[612,669,1742,1766]" country="Tanzania" name="Mjini Magharibi" pageId="11" pageNumber="512">West</collectingRegion>
Kilombero Scarp Forest Reserve).
</paragraph>
</subSubSection>
<subSubSection id="4CC35CA6245E1975FF376983AE76FC53" lastPageId="12" lastPageNumber="513" pageId="11" pageNumber="512" type="description">
<paragraph id="04660F2D245E1975FF376983AE34FDA6" blockId="11.[151,1437,1345,2018]" lastBlockId="12.[151,1437,151,968]" lastPageId="12" lastPageNumber="513" pageId="11" pageNumber="512">
<emphasis id="36ADD33F245E1972FF376983AD2DF891" bold="true" box="[199,524,1777,1802]" pageId="11" pageNumber="512">
Infection with 
<taxonomicName id="C3D974AE245E1972FE866983AD24F891" box="[374,517,1777,1802]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" kingdom="Fungi" order="Laboulbeniales" pageId="11" pageNumber="512" phylum="Ascomycota" rank="species" species="gigas">
<emphasis id="36ADD33F245E1972FE866983AD24F891" bold="true" box="[374,517,1777,1802]" italics="true" pageId="11" pageNumber="512">Rickia gigas</emphasis>
</taxonomicName>
.
</emphasis>
Several specimens were found to be infected with 
<taxonomicName id="C3D974AE245E1972FBBC6983ABF6F892" box="[1100,1239,1777,1802]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" kingdom="Fungi" order="Laboulbeniales" pageId="11" pageNumber="512" phylum="Ascomycota" rank="species" species="gigas">
<emphasis id="36ADD33F245E1972FBBC6983ABF6F892" box="[1100,1239,1777,1802]" italics="true" pageId="11" pageNumber="512">Rickia gigas</emphasis>
</taxonomicName>
. In a sample of 
<specimenCount id="12DFC4A4245E1972FA7E6983AFE0F8B6" pageId="11" pageNumber="512" type="male">6 ♂♂</specimenCount>
and 
<specimenCount id="12DFC4A4245E1972FF096864AE18F8B6" box="[249,313,1814,1838]" pageId="11" pageNumber="512" type="female">4 ♀♀</specimenCount>
from Amani, all specimens were found to be infected: Two males were quite heavily infected with the long-celled 
<typeStatus id="DB62B18F245E1972FEB46848AE55F8C9" box="[324,372,1850,1874]" pageId="11" pageNumber="512">type</typeStatus>
on the head, antennae, collum, pre-gonopodal and post-gonopodal legs (in one case all the way back to the last pair) and adjacent parts of the body rings. In addition these specimens had many thalli of the shortcelled morphotype on the hind margin of several body rings. The other four males had lighter infections with longcelled 
<taxonomicName id="C3D974AE245E1972FF1168D4AE1CF826" box="[225,317,1958,1981]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" kingdom="Fungi" order="Laboulbeniales" pageId="11" pageNumber="512" phylum="Ascomycota" rank="species" species="gigas">
<emphasis id="36ADD33F245E1972FF1168D4AE1CF826" box="[225,317,1958,1981]" italics="true" pageId="11" pageNumber="512">R. gigas</emphasis>
</taxonomicName>
, confined to pre-gonopodal and anterior postgonopodal legs; two of these also had the short-celled form on posterior body ring margins. The four females all had long-celled 
<taxonomicName id="C3D974AE245E1972FC1A68B8AB66F87A" box="[1002,1095,1994,2017]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" kingdom="Fungi" order="Laboulbeniales" pageId="11" pageNumber="512" phylum="Ascomycota" rank="species" species="gigas">
<emphasis id="36ADD33F245E1972FC1A68B8AB66F87A" box="[1002,1095,1994,2017]" italics="true" pageId="11" pageNumber="512">R. gigas</emphasis>
</taxonomicName>
on some anterior legs, two of them also on adjacent parts of the body rings, one of them also on the head; three of them had the short-celled 
<typeStatus id="DB62B18F24591975FA9C6FEAAABDFF2B" box="[1388,1436,152,176]" pageId="12" pageNumber="513">type</typeStatus>
on the margin of several body rings. Four further males from Amani were more or less heavily infected with the long-celled 
<typeStatus id="DB62B18F24591975FEEE6F92AE6FFF63" box="[286,334,224,248]" pageId="12" pageNumber="513">type</typeStatus>
on pre-gonopodal and anterior post-gonopods legs and adjacent parts of the body rings. A male from Udzungwa Mts National Park, Sanje Chini camp, was infected with the long-celled 
<typeStatus id="DB62B18F24591975FB236E77AA22FE86" box="[1235,1283,261,285]" pageId="12" pageNumber="513">type</typeStatus>
on pre- and postgonopodal leg back to midbody, on adjacent parts of the body rings and even (just a few thalli) on the gonopods. A female from Udzungwa National Park, trail to Mizimo Camp, had a few long-celled thalli on several anterior legs and adjacent parts of the body rings. A male from Mwanihana Forest Reserve was infected with the long-celled 
<typeStatus id="DB62B18F24591975FEEC6EE7AE6DFE36" box="[284,332,405,429]" pageId="12" pageNumber="513">type</typeStatus>
on pre- and anterior post-gonopodal legs and also had several thalli of the short-celled 
<typeStatus id="DB62B18F24591975FAD06EE7AA71FE36" box="[1312,1360,405,429]" pageId="12" pageNumber="513">type</typeStatus>
on the posterior margin of body rings; a female from the same sample had a few long-celled thalli on anterior legs and some short-celled thalli on posterior margin of body rings. A male from Luhega Forest Reserve (Udzungwa Scarp) had many long-celled thalli on the pregonopodal and anterior post-gonopodal legs, and on adjacent parts of the body rings.
</paragraph>
<paragraph id="04660F2D24591975FF376D3AACC2FD1E" blockId="12.[151,1437,151,968]" pageId="12" pageNumber="513">
Two males and four females from Rubeho Mts. all had long-celled 
<taxonomicName id="C3D974AE24591975FC4F6D3BAB3DFDFB" box="[959,1052,585,608]" class="Laboulbeniomycetes" family="Laboulbeniaceae" genus="Rickia" kingdom="Fungi" order="Laboulbeniales" pageId="12" pageNumber="513" phylum="Ascomycota" rank="species" species="gigas">
<emphasis id="36ADD33F24591975FC4F6D3BAB3DFDFB" box="[959,1052,585,608]" italics="true" pageId="12" pageNumber="513">R. gigas</emphasis>
</taxonomicName>
on anterior legs, one of the males also had the short-celled 
<typeStatus id="DB62B18F24591975FE5F6D1FAEFEFD1E" box="[431,479,621,645]" pageId="12" pageNumber="513">type</typeStatus>
on the posterior margin of certain body rings.
</paragraph>
<paragraph id="04660F2D24591975FF376DFDAE76FC53" blockId="12.[151,1437,151,968]" pageId="12" pageNumber="513">
<emphasis id="36ADD33F24591975FF376DFDAE30FD33" bold="true" box="[199,273,655,680]" pageId="12" pageNumber="513">Notes.</emphasis>
The wide distribution and the considerable morphological variability, in combination with the close resemblance between 
<taxonomicName id="C3D974AE24591975FE686DC7AD37FD56" box="[408,534,692,717]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F24591975FE686DC7AD37FD56" box="[408,534,692,717]" italics="true" pageId="12" pageNumber="513">T. hamatus</emphasis>
</taxonomicName>
and the even more widespread 
<emphasis id="36ADD33F24591975FC676DC7AB7EFD57" box="[919,1119,693,716]" italics="true" pageId="12" pageNumber="513">
<taxonomicName id="C3D974AE24591975FC676DC7AB7AFD57" box="[919,1115,693,716]" class="Diplopoda" family="Spirostreptidae" genus="Tropostretus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="sigmatospinus">T. sigmatospinus</taxonomicName>
,
</emphasis>
suggest that either several species may hide under 
<taxonomicName id="C3D974AE24591975FE5B6DABAD06FD6B" box="[427,551,727,752]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F24591975FE5B6DABAD06FD6B" box="[427,551,727,752]" italics="true" pageId="12" pageNumber="513">T. hamatus</emphasis>
</taxonomicName>
as here understood, or that 
<taxonomicName id="C3D974AE24591975FC976DABACC2FD6B" box="[871,995,727,752]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F24591975FC976DABACC2FD6B" box="[871,995,727,752]" italics="true" pageId="12" pageNumber="513">T. hamatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C3D974AE24591975FBED6DABABFBFD6B" box="[1053,1242,729,752]" class="Diplopoda" family="Spirostreptidae" genus="Tropostretus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="sigmatospinus">
<emphasis id="36ADD33F24591975FBED6DABABFBFD6B" box="[1053,1242,729,752]" italics="true" pageId="12" pageNumber="513">T. sigmatospinus</emphasis>
</taxonomicName>
may belong to a single widespread, very variable species. The coexistence of both species at several sites (even in the same samples) however, speaks against conspecificity. Apart from the characters which have been given as diagnostic of the two species recognized here, the generally higher number of body rings in 
<taxonomicName id="C3D974AE24591975FC0E6C37AB59FCC6" box="[1022,1144,836,861]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F24591975FC0E6C37AB59FCC6" box="[1022,1144,836,861]" italics="true" pageId="12" pageNumber="513">T. hamatus</emphasis>
</taxonomicName>
(
<figureCitation id="9CE213A824591975FB766C36ABEBFCC7" box="[1158,1226,836,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1182,1204]" captionTargetBox="[151,1436,193,1096]" captionTargetId="figure@5.[151,1436,193,1161]" captionTargetPageId="5" captionText="FIGURE 2. Size of Tropostreptus males, expressed as number of podous rings and midbody vertical diameter. Females reach higher diameters than males. See Figs. 11 and 13 for details on T. hamatus and T. sigmatospinus sp. nov." httpUri="https://zenodo.org/record/803710/files/figure.png" pageId="12" pageNumber="513">Fig. 2</figureCitation>
), and the pink legs which seem to be unique for the East Usambaran population of 
<taxonomicName id="C3D974AE24591975FC976C1BACC3FC1B" box="[871,994,871,896]" class="Diplopoda" family="Spirostreptidae" genus="Tropostreptus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="36ADD33F24591975FC976C1BACC3FC1B" box="[871,994,871,896]" italics="true" pageId="12" pageNumber="513">T. hamatus</emphasis>
</taxonomicName>
, no consistent pattern in the variability has been perceived. This complex of large, easily collected millipedes is an obvious candidate for a molecular taxonomic study.
</paragraph>
</subSubSection>
</treatment>
</document>