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<document id="B7270B87D9C738524D762449C40159EA" ID="10.11646/zootaxa.4178.3.3" ID-DOI="10.11646/zootaxa.4178.3.3" ID-GBIF-Dataset="e86d5f4d-f965-4329-91c5-cfcd7c8ed33b" ID-ISSN="1175-5326" ID-Zenodo-Dep="163367" ID-ZooBank="70509AD2-640A-497C-83EA-85B3EBEA35A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477466785637" checkinUser="plazi" docAuthor="Gilligan, Todd, Huemer, Peter &amp; Wiesmair, Benjamin" docDate="2016" docId="7A0E87C6E97AFFBC16B416B8F28F3939" docLanguage="en" docName="zootaxa.4178.3.3.pdf" docOrigin="Zootaxa 4178 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ancylis pacificana Walsingham 1879" docType="treatment" docVersion="8" lastPageNumber="356" masterDocId="8637FFBEE972FFB51623102FF7133D29" masterDocTitle="Different continents, same species? Resolving the taxonomy of some Holarctic Ancylis Hübner (Lepidoptera: Tortricidae)" masterLastPageNumber="370" masterPageNumber="347" pageNumber="355" updateTime="1698390823962" updateUser="plazi">
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<mods:title id="E6AD76CF19B6D7CE91C2B54661F9FE2D">Different continents, same species? Resolving the taxonomy of some Holarctic Ancylis Hübner (Lepidoptera: Tortricidae)</mods:title>
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<treatment id="7A0E87C6E97AFFBC16B416B8F28F3939" ID-DOI="http://doi.org/10.5281/zenodo.6087636" ID-GBIF-Taxon="124558273" ID-Zenodo-Dep="6087636" LSID="urn:lsid:plazi:treatment:7A0E87C6E97AFFBC16B416B8F28F3939" httpUri="http://treatment.plazi.org/id/7A0E87C6E97AFFBC16B416B8F28F3939" lastPageId="9" lastPageNumber="356" pageId="8" pageNumber="355">
<subSubSection id="BABD655BE97AFFBD16B416B8F6983BFA" pageId="8" pageNumber="355" type="nomenclature">
<paragraph id="F21836D0E97AFFBD16B416B8F5653B9B" blockId="8.[151,630,1687,1747]" box="[151,630,1687,1714]" pageId="8" pageNumber="355">
<heading id="A95081BCE97AFFBD16B416B8F5653B9B" bold="true" box="[151,630,1687,1714]" fontSize="11" level="1" pageId="8" pageNumber="355" reason="1">
<taxonomicName id="35A74D53E97AFFBD16B416B8F5653B9B" ID-CoL="DNJG" authority="Walsingham, 1879" authorityName="Walsingham" authorityYear="1879" box="[151,630,1687,1714]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97AFFBD16B416B8F5653B9B" bold="true" box="[151,630,1687,1714]" pageId="8" pageNumber="355">
<emphasis id="C0D3EAC2E97AFFBD16B416B8F6643B98" bold="true" box="[151,375,1687,1713]" italics="true" pageId="8" pageNumber="355">Ancylis pacificana</emphasis>
(Walsingham, 1879)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F21836D0E97AFFBD16B41695F6983BFA" blockId="8.[151,630,1687,1747]" box="[151,395,1722,1747]" pageId="8" pageNumber="355">Figs. 710, 6162, 76</paragraph>
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<subSubSection id="BABD655BE97AFFBD16B4172DF3FB3A31" box="[151,1256,1794,1816]" pageId="8" pageNumber="355" type="description">
<paragraph id="F21836D0E97AFFBD16B4172DF3FB3A31" blockId="8.[151,1256,1794,1816]" box="[151,1256,1794,1816]" pageId="8" pageNumber="355">
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<taxonomicName id="35A74D53E97AFFBD16B4172DF5543A31" authority="Walsingham 1879" authorityName="Walsingham" authorityYear="1879" box="[151,583,1794,1816]" class="Insecta" family="Tortricidae" genus="Phoxopteryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97AFFBD16B4172DF69B3A31" box="[151,392,1794,1816]" italics="true" pageId="8" pageNumber="355">Phoxopteryx pacificana</emphasis>
Walsingham 1879
</taxonomicName>
,
<emphasis id="C0D3EAC2E97AFFBD1471172DF37F3A31" box="[594,1132,1794,1816]" italics="true" pageId="8" pageNumber="355">
Illust. typical Specimens Lepid. Heterocera Colln.
<collectingRegion id="3063F832E97AFFBD1271172CF37F3A31" box="[1106,1132,1795,1816]" country="Turkey" name="Bolu" pageId="8" pageNumber="355">Br</collectingRegion>
</emphasis>
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.
<materialsCitation id="42CF3C8DE97AFFBD1255172CF3FB3A31" ID-GBIF-Occurrence="1324202964" box="[1142,1256,1794,1816]" location="Mus" pageId="8" pageNumber="355" specimenCount="1" stateProvince="Mus">
<collectingRegion id="3063F832E97AFFBD1255172CF3B63A31" box="[1142,1189,1795,1816]" country="Turkey" name="Mus" pageId="8" pageNumber="355">Mus</collectingRegion>
. 4: 73.
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</paragraph>
</subSubSection>
<subSubSection id="BABD655BE97AFFBD16B4176AF3B93AE0" pageId="8" pageNumber="355" type="diagnosis">
<paragraph id="F21836D0E97AFFBD16B4176AF3B93AE0" blockId="8.[151,1436,1861,2030]" pageId="8" pageNumber="355">
<emphasis id="C0D3EAC2E97AFFBD16B4176AF6193A77" bold="true" box="[151,266,1861,1886]" pageId="8" pageNumber="355">Diagnosis</emphasis>
.
<taxonomicName id="35A74D53E97AFFBD173B176AF6F43A77" box="[280,487,1861,1886]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97AFFBD173B176AF6F43A77" box="[280,487,1861,1886]" italics="true" pageId="8" pageNumber="355">Ancylis pacificana</emphasis>
</taxonomicName>
is identical in wing pattern to
<taxonomicName id="35A74D53E97AFFBD151F1769F4C23A77" box="[828,977,1861,1886]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="unguicella">
<emphasis id="C0D3EAC2E97AFFBD151F1769F4C23A77" box="[828,977,1861,1886]" italics="true" pageId="8" pageNumber="355">A. unguicella</emphasis>
</taxonomicName>
, and the two species cannot be separated without dissection. In the male, the cucullus of
<taxonomicName id="35A74D53E97AFFBD14E01745F44F3AA8" box="[707,860,1897,1921]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97AFFBD14E01745F44F3AA8" box="[707,860,1897,1921]" italics="true" pageId="8" pageNumber="355">A. pacificana</emphasis>
</taxonomicName>
is more elongate with a rounded apex versus the shorter, blunt cucullus in
<taxonomicName id="35A74D53E97AFFBD179B17A1F55C3A8F" box="[440,591,1933,1958]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="unguicella">
<emphasis id="C0D3EAC2E97AFFBD179B17A1F55C3A8F" box="[440,591,1933,1958]" italics="true" pageId="8" pageNumber="355">A. unguicella</emphasis>
</taxonomicName>
, and the phallus (and caulis) is shorter in
<taxonomicName id="35A74D53E97AFFBD120D17A1F3D73A8F" box="[1070,1220,1934,1958]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97AFFBD120D17A1F3D73A8F" box="[1070,1220,1934,1958]" italics="true" pageId="8" pageNumber="355">A. pacificana</emphasis>
</taxonomicName>
. In the female, the antrum of
<taxonomicName id="35A74D53E97AFFBD172F179DF6B23AE0" box="[268,417,1969,1993]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97AFFBD172F179DF6B23AE0" box="[268,417,1969,1993]" italics="true" pageId="8" pageNumber="355">A. pacificana</emphasis>
</taxonomicName>
is narrower posteriorly and the signa are larger than in
<taxonomicName id="35A74D53E97AFFBD122E179DF3B13AE3" box="[1037,1186,1969,1994]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="unguicella">
<emphasis id="C0D3EAC2E97AFFBD122E179DF3B13AE3" box="[1037,1186,1969,1994]" italics="true" pageId="8" pageNumber="355">A. unguicella</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BABD655BE97AFFBC16E417FAF6B13CDC" lastPageId="9" lastPageNumber="356" pageId="8" pageNumber="355" type="description">
<paragraph id="F21836D0E97AFFBD16E417FAF28F3AC7" blockId="8.[151,1436,1861,2030]" box="[199,1436,2005,2030]" pageId="8" pageNumber="355">
<emphasis id="C0D3EAC2E97AFFBD16E417FAF67E3AC7" bold="true" box="[199,365,2005,2030]" pageId="8" pageNumber="355">Redescription</emphasis>
.
<emphasis id="C0D3EAC2E97AFFBD17A217F9F6FE3AC4" box="[385,493,2006,2029]" italics="true" pageId="8" pageNumber="355">Forewing</emphasis>
. FWL Ƌ
<quantity id="355F9B35E97AFFBD144917FAF5E33AC4" box="[618,752,2005,2030]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="10.0" metricValueMin="7.5" pageId="8" pageNumber="355" unit="mm" value="8.75" valueMax="10.0" valueMin="7.5">7.510 mm</quantity>
(n=18), ♀
<quantity id="355F9B35E97AFFBD155D17F9F4C93AC4" box="[894,986,2006,2030]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="8" pageNumber="355" unit="mm" value="8.3">8.3 mm</quantity>
(n=1). Forewing description as in A.
</paragraph>
<paragraph id="F21836D0E97BFFBC16B410B8F6B13CDC" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="356">
<taxonomicName id="35A74D53E97BFFBC16B410B8F61E3D99" box="[151,269,151,176]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="unguicella">
<emphasis id="C0D3EAC2E97BFFBC16B410B8F61E3D99" box="[151,269,151,176]" italics="true" pageId="9" pageNumber="356">unguicella</emphasis>
</taxonomicName>
.
<emphasis id="C0D3EAC2E97BFFBC173C10B8F6DA3D99" box="[287,457,151,176]" italics="true" pageId="9" pageNumber="356">Male genitalia</emphasis>
. Uncus bifid to approximately half its length. Valva with shallow basal excavation extending to just beyond sacculus; saccular angle weakly to moderately developed with variable triangular terminal projection; neck of uniform width or slightly narrowed from sacculus to cucullus; cucullus elongate, densely setose, outer margin convex with several rows of short setae near the anal angle, apex rounded; caulis 0.5 as long as phallus; phallus 0.50.6 as long as valva, with small triangular tooth just proximal to apex; vesica with ca. 4060 deciduous lanceolate cornuti. The length of the cucullus, the width of the valval neck, and the saccular terminal projections all vary between individuals; we have attempted to illustrate the range of variation in Figs. 5859.
<emphasis id="C0D3EAC2E97BFFBC16B411BBF6443C84" box="[151,343,404,429]" italics="true" pageId="9" pageNumber="356">Female genitalia</emphasis>
. Antrum sclerotized to 0.4 length of ductus bursae, widened at ostium to 0.5 distance between apophyses anteriores. Corpus bursae oval, expanding gradually from ductus bursae; signa large, blade- or hornshaped, unequal in size.
</paragraph>
</subSubSection>
<subSubSection id="BABD655BE97BFFBC16E411D0F5113E11" pageId="9" pageNumber="356" type="distribution">
<paragraph id="F21836D0E97BFFBC16E411D0F28E3E3C" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="356">
<emphasis id="C0D3EAC2E97BFFBC16E411D0F64A3F31" bold="true" box="[199,345,511,536]" pageId="9" pageNumber="356">Distribution</emphasis>
. Because of its similarity with
<taxonomicName id="35A74D53E97BFFBC14F1122FF47F3F31" box="[722,876,511,536]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="unguicella">
<emphasis id="C0D3EAC2E97BFFBC14F1122FF47F3F31" box="[722,876,511,536]" italics="true" pageId="9" pageNumber="356">A. unguicella</emphasis>
</taxonomicName>
, the distribution of
<taxonomicName id="35A74D53E97BFFBC127F122FF3E53F31" box="[1116,1270,512,536]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97BFFBC127F122FF3E53F31" box="[1116,1270,512,536]" italics="true" pageId="9" pageNumber="356">A. pacificana</emphasis>
</taxonomicName>
is difficult to determine from undissected museum specimens.
<bibRefCitation id="96364B21E97BFFBC14FB120BF4823F15" author="Heinrich" box="[728,913,548,573]" pageId="9" pageNumber="356" refString="Heinrich, C. (1923) Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum, 123, 1 - 128." type="journal article" year="1923">Heinrich (1923)</bibRefCitation>
reported specimens from
<collectingRegion id="3063F832E97BFFBC12E8120BF22C3F14" box="[1227,1343,548,573]" country="United States of America" name="California" pageId="9" pageNumber="356">California</collectingRegion>
,
<collectingRegion id="3063F832E97BFFBC1373120BF61B3F49" country="Canada" name="British Columbia" pageId="9" pageNumber="356">British Columbia</collectingRegion>
, and
<collectingRegion id="3063F832E97BFFBC17651268F6A23F49" box="[326,433,583,608]" country="United States of America" name="Colorado" pageId="9" pageNumber="356">Colorado</collectingRegion>
, whereas
<bibRefCitation id="96364B21E97BFFBC14021268F42F3F49" author="Powell" box="[545,828,583,608]" pageId="9" pageNumber="356" refString="Powell, J. A. &amp; Opler, P. A. (2009) Moths of Western North America. University of California Press. Berkeley, Los Angeles, London. 369 pp. http: // dx. doi. org / 10.1525 / california / 9780520251977.001.0001" type="book" year="2009">Powell and Opler (2009)</bibRefCitation>
reported a distribution limited to montane habitats in
<collectingRegion id="3063F832E97BFFBC16B41243F7F83FAD" box="[151,235,620,644]" country="United States of America" name="Oregon" pageId="9" pageNumber="356">Oregon</collectingRegion>
and northern
<collectingRegion id="3063F832E97BFFBC17A51243F6E83FAC" box="[390,507,620,645]" country="United States of America" name="California" pageId="9" pageNumber="356">California</collectingRegion>
. The specimens that we determined as
<taxonomicName id="35A74D53E97BFFBC158F1243F3523FAD" box="[940,1089,620,644]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97BFFBC158F1243F3523FAD" box="[940,1089,620,644]" italics="true" pageId="9" pageNumber="356">A. pacificana</emphasis>
</taxonomicName>
were from
<collectingRegion id="3063F832E97BFFBC12E11243F2253FAC" box="[1218,1334,620,645]" country="United States of America" name="California" pageId="9" pageNumber="356">California</collectingRegion>
,
<collectingRegion id="3063F832E97BFFBC13621243F28B3FAD" box="[1345,1432,620,644]" country="United States of America" name="Oregon" pageId="9" pageNumber="356">Oregon</collectingRegion>
, and
<collectingRegion id="3063F832E97BFFBC16EA12A0F6833F81" box="[201,400,655,680]" country="Canada" name="British Columbia" pageId="9" pageNumber="356">British Columbia</collectingRegion>
, although we discovered one male in the EME from San Bernardo,
<collectingRegion id="3063F832E97BFFBC128712BFF3E53F81" box="[1188,1270,656,680]" country="Mexico" name="Sonora" pageId="9" pageNumber="356">Sonora</collectingRegion>
,
<collectingRegion id="3063F832E97BFFBC132712A0F24E3F81" box="[1284,1373,655,680]" country="Mexico" name="Mexico" pageId="9" pageNumber="356">
<collectingCountry id="8AB07640E97BFFBC132712A0F24E3F81" box="[1284,1373,655,680]" name="Mexico" pageId="9" pageNumber="356">Mexico</collectingCountry>
</collectingRegion>
. This suggests that the distribution of
<taxonomicName id="35A74D53E97BFFBC142A129BF5B33FE5" box="[521,672,692,716]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97BFFBC142A129BF5B33FE5" box="[521,672,692,716]" italics="true" pageId="9" pageNumber="356">A. pacificana</emphasis>
</taxonomicName>
is more extensive than assumed by previous authors, although we have not confirmed
<taxonomicName id="35A74D53E97BFFBC17A812F7F5353FD9" box="[395,550,728,752]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97BFFBC17A812F7F5353FD9" box="[395,550,728,752]" italics="true" pageId="9" pageNumber="356">A. pacificana</emphasis>
</taxonomicName>
from any other locations (including
<collectingRegion id="3063F832E97BFFBC15CB12F8F3463FD9" box="[1000,1109,727,752]" country="United States of America" name="Colorado" pageId="9" pageNumber="356">Colorado</collectingRegion>
). In any event, undissected museum specimens identified as
<taxonomicName id="35A74D53E97BFFBC142B12D3F58E3E3D" box="[520,669,764,788]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97BFFBC142B12D3F58E3E3D" box="[520,669,764,788]" italics="true" pageId="9" pageNumber="356">A. pacificana</emphasis>
</taxonomicName>
should be treated as suspect until verified using genitalic characters.
</paragraph>
<paragraph id="F21836D0E97BFFBC16E41330F5113E11" blockId="9.[151,1437,151,1040]" box="[199,514,799,824]" pageId="9" pageNumber="356">
<emphasis id="C0D3EAC2E97BFFBC16E41330F6963E11" bold="true" box="[199,389,799,824]" pageId="9" pageNumber="356">Molecular data.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="BABD655BE97BFFBC16E4136BF5013E56" pageId="9" pageNumber="356" type="biology_ecology">
<paragraph id="F21836D0E97BFFBC16E4136BF5013E56" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="356">
<emphasis id="C0D3EAC2E97BFFBC16E4136BF60C3E74" bold="true" box="[199,287,836,861]" pageId="9" pageNumber="356">Biology</emphasis>
. Adults are present May to July. The larval host is unknown, although
<bibRefCitation id="96364B21E97BFFBC121F136BF2413E75" author="Powell" box="[1084,1362,836,861]" pageId="9" pageNumber="356" refString="Powell, J. A. &amp; Opler, P. A. (2009) Moths of Western North America. University of California Press. Berkeley, Los Angeles, London. 369 pp. http: // dx. doi. org / 10.1525 / california / 9780520251977.001.0001" type="book" year="2009">Powell and Opler (2009)</bibRefCitation>
report adults associated with
<taxonomicName id="35A74D53E97BFFBC17B71348F51F3EA9" box="[404,524,871,896]" class="Magnoliopsida" family="Rhamnaceae" genus="Ceanothus" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="356" phylum="Tracheophyta" rank="genus">
<emphasis id="C0D3EAC2E97BFFBC17B71348F51F3EA9" box="[404,524,871,896]" italics="true" pageId="9" pageNumber="356">Ceanothus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BABD655BE97BFFBC16E413A3F28F3939" pageId="9" pageNumber="356" type="discussion">
<paragraph id="F21836D0E97BFFBC16E413A3F28F3939" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="356">
<emphasis id="C0D3EAC2E97BFFBC16E413A3F6273E8C" bold="true" box="[199,308,908,933]" pageId="9" pageNumber="356">Remarks</emphasis>
.
<bibRefCitation id="96364B21E97BFFBC176113A3F6E53E8D" author="Heinrich" box="[322,502,908,933]" pageId="9" pageNumber="356" refString="Heinrich, C. (1923) Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum, 123, 1 - 128." type="journal article" year="1923">Heinrich (1923)</bibRefCitation>
stated that
<taxonomicName id="35A74D53E97BFFBC145B13A3F41E3E8D" box="[632,781,908,932]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97BFFBC145B13A3F41E3E8D" box="[632,781,908,932]" italics="true" pageId="9" pageNumber="356">A. pacificana</emphasis>
</taxonomicName>
was larger and less darkly marked than
<taxonomicName id="35A74D53E97BFFBC12F013A3F27A3E8C" box="[1235,1385,908,933]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="unguicella">
<emphasis id="C0D3EAC2E97BFFBC12F013A3F27A3E8C" box="[1235,1385,908,933]" italics="true" pageId="9" pageNumber="356">A. unguicella</emphasis>
</taxonomicName>
. We found that, whereas some specimens of
<taxonomicName id="35A74D53E97BFFBC1471139FF5F43EE1" box="[594,743,944,968]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97BFFBC1471139FF5F43EE1" box="[594,743,944,968]" italics="true" pageId="9" pageNumber="356">A. pacificana</emphasis>
</taxonomicName>
are slightly larger, both species overlap in size. We also found smaller specimens of
<taxonomicName id="35A74D53E97BFFBC17BD13FBF5293EC5" box="[414,570,980,1004]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="pacificana">
<emphasis id="C0D3EAC2E97BFFBC17BD13FBF5293EC5" box="[414,570,980,1004]" italics="true" pageId="9" pageNumber="356">A. pacificana</emphasis>
</taxonomicName>
with dark markings (e.g., Fig. 9) that were incorrectly identified as
<taxonomicName id="35A74D53E97BFFBC13A713FBF61E3939" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="unguicella">
<emphasis id="C0D3EAC2E97BFFBC13A713FBF61E3939" italics="true" pageId="9" pageNumber="356">A. unguicella</emphasis>
</taxonomicName>
. Species-level determinations of either species from the Pacific Northwest should rely on male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>