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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1136.72744" ID-Pensoft-Pub="1313-2970-1136-1" ID-Pensoft-UUID="5D0E6613B1D05E4293ABEE4BFD1CA18A" ID-ZooBank="11C6BBFB339A4672AE662CE2B1E6321E" ModsDocID="1313-2970-1136-1" checkinTime="1671287389350" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2022" docId="5AE030400D3A529CA54FEBA78AC635D7" docLanguage="en" docName="ZooKeys 1136: 1-56" docOrigin="ZooKeys 1136" docPubDate="2022-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1136.72744" docTitle="Desmopachria bifurcita Miller 2022, sp. nov." docType="treatment" docUuid="B3351A38-F6E8-41FB-BCAA-DC78CB00A8F8" docUuidSource="ZooBank" docVersion="1" id="5D0E6613B1D05E4293ABEE4BFD1CA18A" lastPageNumber="1" masterDocId="5D0E6613B1D05E4293ABEE4BFD1CA18A" masterDocTitle="Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="1" updateTime="1671287389350" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Kelly B.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
<mods:nameIdentifier type="email">kbmiller@unm.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1136</mods:number>
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<mods:start>1</mods:start>
<mods:end>56</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1136.72744</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1136.72744</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1136-1</mods:identifier>
<mods:identifier type="ZooBank">11C6BBFB339A4672AE662CE2B1E6321E</mods:identifier>
<mods:identifier type="Pensoft-UUID">5D0E6613B1D05E4293ABEE4BFD1CA18A</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:B3351A38-F6E8-41FB-BCAA-DC78CB00A8F8" httpUri="http://treatment.plazi.org/id/5AE030400D3A529CA54FEBA78AC635D7" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/B3351A38-F6E8-41FB-BCAA-DC78CB00A8F8" authority="Miller, 2022" authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita" status="sp. nov.">Desmopachria bifurcita</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">Figures 28-30</figureCitation>
<figureCitation captionStart="Figure 78" captionStartId="F8" captionText="Figure 78. Desmopachria species distributions. Desmopachria bifurcita species group and Desmopachria convexa species group." figureDoi="10.3897/zookeys.1136.72744.figure78" httpUri="https://binary.pensoft.net/fig/782460" pageId="0" pageNumber="1">, 78</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Peru,
<normalizedToken originalValue="Junín">Junin</normalizedToken>
, Sani Beni. The type locality is ambiguous. According to
<bibRefCitation DOI="https://doi.org/10.15381/rpb.v24i1.13107" author="Otto, RL" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="1" refId="B17" refString="Otto, RL, 2017. Beetles of Peru: a survey of the Families. Eucnemidae Eschscholtz, 1829. Revista Peruana de Biologia 24(1): e11. https://doi.org/10.15381/rpb.v24i1.13107" title="Beetles of Peru: a survey of the Families. Eucnemidae Eschscholtz, 1829. Revista Peruana de Biologia 24 (1): e 11." url="https://doi.org/10.15381/rpb.v24i1.13107" year="2017">Otto (2017)</bibRefCitation>
this collection locality by P. Woytkowski may refer to a misspelling of &quot;Sani Benu&quot; which is at approximately
<geoCoordinate degrees="11.253917" direction="south" orientation="latitude" precision="1" value="-11.253917">11.253917°S</geoCoordinate>
,
<geoCoordinate degrees="74.565565" direction="west" orientation="longitude" precision="1" value="-74.56557">74.565565°W</geoCoordinate>
(see below).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Males of this species have the male genitalia very distinctive with the median lobe very short and strongly bifurcate (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">29</figureCitation>
). The lateral lobes in dorsal aspect and broad, parallel-sided, and apically subtruncate (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">29</figureCitation>
). In lateral aspect, they are slender and abruptly bent medially (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">30</figureCitation>
). Specimens are very broad in dorsal aspect and dark reddish-brown dorsally. A number of species of
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria</emphasis>
</taxonomicName>
have the median lobe short with long, medially bent lateral lobes like
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifurcita</emphasis>
</taxonomicName>
, including
<taxonomicName authorityName="Zimmermann" authorityYear="1921" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifasciata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifasciata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifasciata</emphasis>
</taxonomicName>
Zimmermann,
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bolivari" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bolivari</emphasis>
</taxonomicName>
Miller,
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria chei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chei">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria chei</emphasis>
</taxonomicName>
Miller,
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lata</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName authorityName="Wehncke" authorityYear="1877" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria varians" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="varians">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria varians</emphasis>
</taxonomicName>
Wehncke, and
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria ovalis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria ovalis</emphasis>
</taxonomicName>
Sharp. Among these,
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifurcita</emphasis>
</taxonomicName>
is the only one with a bifid median lobe (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">29</figureCitation>
). These species may together form another species group within
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria</emphasis>
</taxonomicName>
, but more thorough examination of specimens will be needed to determine this.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Measurements.</emphasis>
TL = 1.8-1.9 mm, GW = 1.3 mm, PW = 1.0 mm, HW = 0.6 mm, EW = 0.3 mm, TL/GW = 1.4, HW/EW = 2.1-2.4. Body very broad, rounded, laterally broadly curved, lateral margins continuous between pronotum and elytron, body broadest across elytra anterior midlength of body (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">28</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
(Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">28</figureCitation>
). Dorsal surface of head dark red, paler red along anterior margin. Pronotum evenly red. Elytron red, vaguely darker along anterior and sutural margins. Head appendages, legs, and ventral surfaces red to dark red.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sculpture and structure.</emphasis>
Head broad, anteriorly produced in rounded lobe; anterior margin of clypeus curved, flattened, margined with conspicuous, continuous flattened narrow bead; surface of head shiny, finely and sparsely punctate; eyes large (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">28</figureCitation>
, HW/EW = 2.1-2.4); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, lobed at anterodorsal angle, antennomere XI elongate, apically pointed. Pronotum very short, lateral margins short, slightly curved with continuous narrow bead, slightly wider medially; surface shiny, impunctate medially, punctate around margins, punctation variable, fine to course. Elytron moderately broad, laterally broadly curved; surface shiny, more coarsely and evenly punctate than pronotum, punctation distinctive and prominent. Prosternum extremely short, longitudinally compressed, medially slightly carinate; prosternal process slender anteriorly, with distinctive, small medial tubercle, apically short and broad, medially slightly carinate, concave, apically acutely pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, moderately and irregularly punctate; metaventrite wings extremely slender. Metacoxa with medial portion short, &lt;1/3 length of metaventrite medially, metacoxal lines slightly divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, evenly punctate, punctures evenly distributed. Metatrochanter large, subequal to length of ventral margin of metafemur; legs otherwise not noticeably modified. Abdomen with surfaces shiny and smooth, very finely and sparsely punctate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia.</emphasis>
Male median lobe in lateral aspect extremely short, apically distinctly bifid, each branch apicolaterally pointed, (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">29</figureCitation>
). Lateral lobe in ventral aspect evenly broad throughout length, lateral margins parallel, apically broadly rounded (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">30</figureCitation>
) in lateral aspect slender throughout length, abruptly curved medially (Fig.
<figureCitation captionStart="Figures 28–45" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">31</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sexual dimorphism.</emphasis>
No obvious sexual dimorphic features were discovered.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Variation.</emphasis>
No characteristic variation was examined among the specimens examined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is named
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifurcita</emphasis>
</taxonomicName>
, Latin for the short, bifurcate male median lobe.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
The exact locality of collection of this species is somewhat ambiguous. The locality Sani Beni is probably &quot;Sani Benu&quot; at coordinates
<geoCoordinate degrees="11.253917" direction="south" orientation="latitude" precision="1" value="-11.253917">11.253917°S</geoCoordinate>
,
<geoCoordinate degrees="74.565565" direction="west" orientation="longitude" precision="1" value="-74.56557">74.565565°W</geoCoordinate>
, as with
<taxonomicName authorityName="Eschscholtz" authorityYear="1829" class="Insecta" family="Eucnemidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Eucnemidae</taxonomicName>
Eschscholtz specimens collected by the same collector as documented by
<bibRefCitation DOI="https://doi.org/10.15381/rpb.v24i1.13107" author="Otto, RL" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="1" refId="B17" refString="Otto, RL, 2017. Beetles of Peru: a survey of the Families. Eucnemidae Eschscholtz, 1829. Revista Peruana de Biologia 24(1): e11. https://doi.org/10.15381/rpb.v24i1.13107" title="Beetles of Peru: a survey of the Families. Eucnemidae Eschscholtz, 1829. Revista Peruana de Biologia 24 (1): e 11." url="https://doi.org/10.15381/rpb.v24i1.13107" year="2017">Otto (2017)</bibRefCitation>
(Fig.
<figureCitation captionStart="Figure 78" captionStartId="F8" captionText="Figure 78. Desmopachria species distributions. Desmopachria bifurcita species group and Desmopachria convexa species group." figureDoi="10.3897/zookeys.1136.72744.figure78" httpUri="https://binary.pensoft.net/fig/782460" pageId="0" pageNumber="1">78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype in SEMC, male labeled, &quot;PERU Dept. Juni Sani Beni 1 Aug. 1935 [&quot;1 Aug.&quot; handwritten] P. Wyotkowski co/ HOLOTYPE
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifurcita</emphasis>
</taxonomicName>
Miller, 2021 [red label with black line border].&quot; Paratypes, 5 in SEMC and MSBA labeled same as holotype except with dates, 10 Aug 1935, 11 Aug 1935, 19 Aug 1935, 20 Aug 1935, and 6 Nov 1935 and
<normalizedToken originalValue="“/…">&quot;/...</normalizedToken>
PARATYPE
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifurcita</emphasis>
</taxonomicName>
Miller, 2021 [blue label with black line border].&quot;
</paragraph>
<caption doi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1" start="Figures 28–45" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 28-45.</emphasis>
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria</emphasis>
</taxonomicName>
species. (
<emphasis bold="true" pageId="0" pageNumber="1">28-30</emphasis>
)
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifurcita</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">28</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="1">29</emphasis>
male median and lateral lobes, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="1">30</emphasis>
male lateral lobe, right lateral aspect (
<emphasis bold="true" pageId="0" pageNumber="1">31-33</emphasis>
)
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">31</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="1">32</emphasis>
male genitalia, ventral aspect
<emphasis bold="true" pageId="0" pageNumber="1">33</emphasis>
right lateral lobe, right lateral aspect. (
<emphasis bold="true" pageId="0" pageNumber="1">34, 35</emphasis>
)
<taxonomicName genus="Desompachria" lsidName="Desompachria bifasciata" pageId="0" pageNumber="1" rank="species" species="bifasciata">
<emphasis italics="true" pageId="0" pageNumber="1">Desompachria bifasciata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">34</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="1">35</emphasis>
male genitalia, ventral aspect (
<emphasis bold="true" pageId="0" pageNumber="1">36-38</emphasis>
)
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bolivari" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bolivari</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">36</emphasis>
male genitalia, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="1">37</emphasis>
right lateral lobe, right lateral aspect
<emphasis bold="true" pageId="0" pageNumber="1">38</emphasis>
median lobe, right lateral aspect
<emphasis bold="true" pageId="0" pageNumber="1">39</emphasis>
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria ovalis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria ovalis</emphasis>
</taxonomicName>
, male genitalia, dorsal aspect (
<emphasis bold="true" pageId="0" pageNumber="1">40-42</emphasis>
)
<taxonomicName authorityName="Wehncke" authorityYear="1877" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria varians" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="varians">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria varians</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">40</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="1">41</emphasis>
male genitalia, ventral aspect
<emphasis bold="true" pageId="0" pageNumber="1">42</emphasis>
right lateral lobe, right lateral aspect (
<emphasis bold="true" pageId="0" pageNumber="1">43-45</emphasis>
)
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria pseudocavia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pseudocavia">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria pseudocavia</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">43</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="1">44</emphasis>
abdominal sternite VI, ventral aspect
<emphasis bold="true" pageId="0" pageNumber="1">45</emphasis>
male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (
<emphasis bold="true" pageId="0" pageNumber="1">44</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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