478 lines
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478 lines
55 KiB
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<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="59C3770EFAE55DADBBC2115A71AABB2B" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina siamensis Moellendorff 1902" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:title id="068D2C61A6259A73F97C52FB20E09BC6">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:namePart id="7CA21AC13388F8C4D42053E4B3217476">Jirapatrasilp, Parin</mods:namePart>
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<mods:affiliation id="8D39D1CE46EA3C232103E3B5CEEE45D9">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="A7E7254374F762E4BFF215970AD0CE6A">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="6AD876EE8A14CD113DE643C759FEC7BC">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="2AF749146B1C731574A4D6FA562C1FA8" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="F073A1BA1FCB89BA681FEE68D13AF2FE">Panha, Somsak</mods:namePart>
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<mods:affiliation id="3CF2E5E6A130B336DEAE091C2748BC26">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="628C9687F47193E83901587C6FDBA0DA" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:date id="092120FCAAE11ABBCB084F545D27889E">2022</mods:date>
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<treatment id="59C3770EFAE55DADBBC2115A71AABB2B" ID-GBIF-Taxon="199838977" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B" httpUri="http://treatment.plazi.org/id/59C3770EFAE55DADBBC2115A71AABB2B" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupininae" scope_superFamily="Cyclophoroidea">
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<subSubSection id="E45ABFA482A892A7F89206303DD0DBF9" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="EB14C53935638531DED5F8B3D7759C24" pageId="0" pageNumber="1">
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<taxonomicName id="519DF5601812F0ED186F49CD3AA6380D" ID-CoL="7SWWN" LSID="59C3770E-FAE5-5DAD-BBC2-115A71AABB2B" authority="Moellendorff, 1902" authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
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Pupina siamensis
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<normalizedToken id="3BF3569BC2F75D77B334C643EBBAEF4A" originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1902
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="29FA652059B11B9710D7DE04B20A2996" pageId="0" pageNumber="1" type="description">
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<paragraph id="C91A45D8E9C9AFD1D7298C2319F25DB4" pageId="0" pageNumber="1">
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<figureCitation id="1D83ED344217C35C8B999D4075322B2E" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. External shell morphology of three mainland Southeast Asian Pupina species groups A Pupina artata from the Pupina artata species group, CUMZ 12003 from Ban Ping Khong, Chiang Mai B Pupina peguensis from the Pupina arula species group, CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai C Pupina siamensis from the Pupina arula species group, CUMZ 12052 from Sri Thong Cave, Sra Keo, and D Pupina tchehelensis from the Pupina aureola species group, CUMZ 12136 from limestone mountain, Phang Nga. Red frames focus on the parietal tooth and posterior canal; blue frames focus on the curvature of the apertural lip when observed from lateral view." figureDoi="10.3897/zookeys.1119.85400.figure18" httpUri="https://binary.pensoft.net/fig/735379" pageId="0" pageNumber="1">Figs 18C</figureCitation>
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<figureCitation id="66C7E9DA4FC56155FF999168C622F6EB" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Opercula of three mainland Southeast Asian Pupina species groups A Pupina bensoni sp. nov. from the Pupina artata species group, specimen CUMZ 12047 B Pupina siamensis from the Pupina arula species group, specimen CUMZ 12067, and C Pupina aureola from the Pupina aureola species group, specimen CUMZ 12116. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure19" httpUri="https://binary.pensoft.net/fig/735380" pageId="0" pageNumber="1">, 19B</figureCitation>
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<figureCitation id="D8D3B0EDFB9354E93090A08BDC52320A" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">, 25H</figureCitation>
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<figureCitation id="51F04E0DE227AAE3A62FDABAC0994696" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Shells of Pupina arula species group from mainland Southeast Asia A-G Pupina peguensis A syntype of Pupina blanfordi NHMUK 1888.12.4.100 and specimens B CUMZ 12105 C CUMZ 12050 D CUMZ 12103 E NHMUK ex. Cuming coll. F CUMZ 12094, and G CUMZ 12108 H-J Pupina crosseana H lectotype MNHN-IM- 2000 - 35834 I paralectotype RBINS MT 966 / 10591, and J specimen CUMZ 12049 K, L Pupina siamensis K lectotype SMF 109948 and L specimen CUMZ 12052 M-P Pupina bilabiata sp. nov. M holotype CUMZ 12073 / 1 and specimens N CUMZ 12079 O CUMZ 12081, and P CUMZ 12086 Q, R Pupina godwinausteni sp. nov. Q holotype CUMZ 12090 / 1 and R paratype CUMZ 12090 / 2. Photo: P. Maestrati, MNHN (H), F. Trus, RBINS (I)." figureDoi="10.3897/zookeys.1119.85400.figure28" httpUri="https://binary.pensoft.net/fig/735389" pageId="0" pageNumber="1">, 28K, L</figureCitation>
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<figureCitation id="73278E67822665A05B204B244F6C0A68" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. A Pupina crosseana, specimen CUMZ 12049 from Khao Jedee Temple, Nakhon Sawan B, C Pupina siamensis: B lectotype SMF 109948 and C specimen CUMZ 12052 from Sri Thong Cave, Sra Keo D-F Pupina bilabiata sp. nov. D holotype CUMZ 12073 / 1, and specimens E CUMZ 12079 from Tham Pha Ya Temple, Loei and F CUMZ 12081 from Pha Jor Cave, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure31" httpUri="https://binary.pensoft.net/fig/735392" pageId="0" pageNumber="1">, 31B, C</figureCitation>
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<figureCitation id="DB6BDFE1A25861EE1DFFB4CA35F2D8F1" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">, 32A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B368E615365A2D5C06EA925D5F18BEA8" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="3C33C3E11F9138EC5D8C511001178F3D" pageId="0" pageNumber="1">
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<taxonomicName id="604C7C66578571842CB126725F0801FD" authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis" subGenus="Tylotoechus">Pupina (Tylotoechus) siamensis</taxonomicName>
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<normalizedToken id="95D6167CB2812AFB4781EDBE2650E8B9" originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1902b: 160. Type locality:
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<normalizedToken id="7C14A3B4652BF1EA81140562F31DEBAB" originalValue="“Bangkok”">"Bangkok"</normalizedToken>
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[see Remarks].
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<bibRefCitation id="E7B69CE5DA1E72EE10C1B57CAF91360B" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="41 - 56" refId="B198" refString="Zilch, A, 1957. Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae. Archiv fuer Molluskenkunde 86: 41 - 56" title="Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae." volume="86" year="1957">Zilch 1957</bibRefCitation>
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: 47, pl. 2, fig. 15.
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<bibRefCitation id="09BC251F6673A62F638E38A666F3CE4A" author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
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: 39.
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</paragraph>
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<paragraph id="B28CD8DC0CC095BE8C491138B049813D" pageId="0" pageNumber="1">
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<taxonomicName id="6BE8237CF972AAC8030005A90B9A454A" authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">Pupina siamensis</taxonomicName>
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-
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<bibRefCitation id="179337A95BDB9330E5F2FBDE53811457" author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 432, Muok-Lek, Siam [Muak Lek District, Saraburi Province, Thailand].
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<bibRefCitation id="200A5A0A1C8D1D278ACC55FA21812DEF" author="Boonngam, P" journalOrPublisher="Witthayasan Kasetsat Witthayasat" pageId="0" pageNumber="1" pagination="256 - 263" refId="B14" refString="Boonngam, P, Dumrongrojwattana, P, Matchacheep, S, 2008. The diversity of land snail fauna in Chonburi Province, Eastern Thailand. Witthayasan Kasetsat Witthayasat 42: 256 - 263" title="The diversity of land snail fauna in Chonburi Province, Eastern Thailand." volume="42" year="2008">Boonngam et al. 2008</bibRefCitation>
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: 258, Chonburi Province, Thailand.
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<bibRefCitation id="36CC269B56A476435021AAD75CBA8AA7" author="Chanyapate, C" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" refId="B20" refString="Chanyapate, C, Promkerd, P, Klakhaeng, S, Rintarak, D, Nookarn, P, 2008. Biodiversity of land snail and slug in Biosphere Reserve Sakaerat. Research Report 2008, Plant Protection Research and Development Office, Department of Agriculture, Ministry of Agriculture and Cooperatives, Bangkok, 60-72. [in Thai]" title="Biodiversity of land snail and slug in Biosphere Reserve Sakaerat. Research Report 2008, Plant Protection Research and Development Office, Department of Agriculture, Ministry of Agriculture and Cooperatives, Bangkok, 60 - 72. [in Thai]" year="2008">Chanyapate et al. 2008</bibRefCitation>
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: 2116, with text fig., Sakaerat Biosphere Reserves, Nakhon Ratchasima Province.
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<bibRefCitation id="C779056AF810E475D109FEBF4F2DA9EA" author="Chidchua, W" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" refId="B22" refString="Chidchua, W, Dumrongrojwattana, P, 2010. Taxonomy of land snails in Klaeng District Rayong Province and Kaenghangmaew District Chanthaburi Province, Eastern Thailand (Gastropoda: Prosobranchia, Pulmonata). In: The Proceedings of 48th Kasetsart University Annual Conference: Science. Kasetsart University, Bangkok, 161-170. [in Thai]" title="Taxonomy of land snails in Klaeng District Rayong Province and Kaenghangmaew District Chanthaburi Province, Eastern Thailand (Gastropoda: Prosobranchia, Pulmonata). In: The Proceedings of 48 th Kasetsart University Annual Conference: Science. Kasetsart University, Bangkok, 161 - 170. [in Thai]" year="2010">Chidchua and Dumrongrojwattana 2010</bibRefCitation>
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: 164, fig. 2, Klaeng District, Rayong Province and Kaenghangmaew District, Chanthaburi Province.
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<bibRefCitation id="958E626619C6702434A682743DE09456" author="Dumrongrojwattana, P" journalOrPublisher="Department of Biology, Burapha University, Bangkok" pageId="0" pageNumber="1" refId="B37" refString="Dumrongrojwattana, P, 2016. Malacofauna in Pracheenburi Province (Under the Plant Genetic Conservation Project Under the Royal Initiative of Her Highness Princess Maha Chakri Sirindhorn). Department of Biology, Burapha University, Bangkok" title="Malacofauna in Pracheenburi Province (Under the Plant Genetic Conservation Project Under the Royal Initiative of Her Highness Princess Maha Chakri Sirindhorn)." year="2016">Dumrongrojwattana 2016</bibRefCitation>
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: 17, 18, fig. 4-4, Kaeng Hin Poeng, Thap Lan National Park, Prachin Buri Province.
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<bibRefCitation id="83CAB9F714F7C2DD033B1127B16FC47D" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
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: 94.
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<bibRefCitation id="DF8AAE2F6C4191D8A45D2C95E74E7DF0" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
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: fig. 5-13j.
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</paragraph>
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</subSubSection>
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<subSubSection id="78D337D0A12567CE60B94BA67EB1DC2E" pageId="0" pageNumber="1" type="type material">
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<paragraph id="6D107A850D43CFE0F2F2C8182739B566" pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph id="8FB080127E37999BCC80F80DF2D74729" pageId="0" pageNumber="1">
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<emphasis id="C9272265A20C2C4E34A0088BEF8C856B" bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
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SMF 109948 (Figs
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<figureCitation id="2D38B0432583D27DF6CFDA1904B2B8C7" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Shells of Pupina arula species group from mainland Southeast Asia A-G Pupina peguensis A syntype of Pupina blanfordi NHMUK 1888.12.4.100 and specimens B CUMZ 12105 C CUMZ 12050 D CUMZ 12103 E NHMUK ex. Cuming coll. F CUMZ 12094, and G CUMZ 12108 H-J Pupina crosseana H lectotype MNHN-IM- 2000 - 35834 I paralectotype RBINS MT 966 / 10591, and J specimen CUMZ 12049 K, L Pupina siamensis K lectotype SMF 109948 and L specimen CUMZ 12052 M-P Pupina bilabiata sp. nov. M holotype CUMZ 12073 / 1 and specimens N CUMZ 12079 O CUMZ 12081, and P CUMZ 12086 Q, R Pupina godwinausteni sp. nov. Q holotype CUMZ 12090 / 1 and R paratype CUMZ 12090 / 2. Photo: P. Maestrati, MNHN (H), F. Trus, RBINS (I)." figureDoi="10.3897/zookeys.1119.85400.figure28" httpUri="https://binary.pensoft.net/fig/735389" pageId="0" pageNumber="1">28K</figureCitation>
|
|
,
|
|
<figureCitation id="0964150EE900D64CE3B63F982862A9AA" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. A Pupina crosseana, specimen CUMZ 12049 from Khao Jedee Temple, Nakhon Sawan B, C Pupina siamensis: B lectotype SMF 109948 and C specimen CUMZ 12052 from Sri Thong Cave, Sra Keo D-F Pupina bilabiata sp. nov. D holotype CUMZ 12073 / 1, and specimens E CUMZ 12079 from Tham Pha Ya Temple, Loei and F CUMZ 12081 from Pha Jor Cave, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure31" httpUri="https://binary.pensoft.net/fig/735392" pageId="0" pageNumber="1">31B</figureCitation>
|
|
) from
|
|
<normalizedToken id="7AC48B9FB3B25EB4128D5CC1F6E51907" originalValue="“Bangkok”">"Bangkok"</normalizedToken>
|
|
, Thailand.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7F4F3412AE87D575907C22C13A59E680" pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph id="6793EB910F9EB2F86AFEF4327A7DDBBF" pageId="0" pageNumber="1">Other material examined.</paragraph>
|
|
<paragraph id="9FEBF358A1EC81E6A77D79CDA32867F1" pageId="0" pageNumber="1">
|
|
<materialsCitation id="A1BF18AF01B44625FE53FCE17865F458" collectingDate="2006-11-25" collectorName="Figs" county="Sra Keo Province" location="Sri Thong Cave" municipality="Klong Had District" specimenCount="1">
|
|
CUMZ 12052 (15 shells;
|
|
<collectorName id="24CA270CC5BE554BE05F6FE13037A521">Figs</collectorName>
|
|
<figureCitation id="FD2C626D7232305739FF02D45D9312A2" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. External shell morphology of three mainland Southeast Asian Pupina species groups A Pupina artata from the Pupina artata species group, CUMZ 12003 from Ban Ping Khong, Chiang Mai B Pupina peguensis from the Pupina arula species group, CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai C Pupina siamensis from the Pupina arula species group, CUMZ 12052 from Sri Thong Cave, Sra Keo, and D Pupina tchehelensis from the Pupina aureola species group, CUMZ 12136 from limestone mountain, Phang Nga. Red frames focus on the parietal tooth and posterior canal; blue frames focus on the curvature of the apertural lip when observed from lateral view." figureDoi="10.3897/zookeys.1119.85400.figure18" httpUri="https://binary.pensoft.net/fig/735379" pageId="0" pageNumber="1">18C</figureCitation>
|
|
,
|
|
<figureCitation id="146DFC3EA53174D5C794EE8E586839BE" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Shells of Pupina arula species group from mainland Southeast Asia A-G Pupina peguensis A syntype of Pupina blanfordi NHMUK 1888.12.4.100 and specimens B CUMZ 12105 C CUMZ 12050 D CUMZ 12103 E NHMUK ex. Cuming coll. F CUMZ 12094, and G CUMZ 12108 H-J Pupina crosseana H lectotype MNHN-IM- 2000 - 35834 I paralectotype RBINS MT 966 / 10591, and J specimen CUMZ 12049 K, L Pupina siamensis K lectotype SMF 109948 and L specimen CUMZ 12052 M-P Pupina bilabiata sp. nov. M holotype CUMZ 12073 / 1 and specimens N CUMZ 12079 O CUMZ 12081, and P CUMZ 12086 Q, R Pupina godwinausteni sp. nov. Q holotype CUMZ 12090 / 1 and R paratype CUMZ 12090 / 2. Photo: P. Maestrati, MNHN (H), F. Trus, RBINS (I)." figureDoi="10.3897/zookeys.1119.85400.figure28" httpUri="https://binary.pensoft.net/fig/735389" pageId="0" pageNumber="1">28L</figureCitation>
|
|
,
|
|
<figureCitation id="53205B6C9DCA0CAA88DB8CB7CA7769E6" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. A Pupina crosseana, specimen CUMZ 12049 from Khao Jedee Temple, Nakhon Sawan B, C Pupina siamensis: B lectotype SMF 109948 and C specimen CUMZ 12052 from Sri Thong Cave, Sra Keo D-F Pupina bilabiata sp. nov. D holotype CUMZ 12073 / 1, and specimens E CUMZ 12079 from Tham Pha Ya Temple, Loei and F CUMZ 12081 from Pha Jor Cave, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure31" httpUri="https://binary.pensoft.net/fig/735392" pageId="0" pageNumber="1">31C</figureCitation>
|
|
) from
|
|
<location id="2F1816670DEDA01214392AA0B1110B23" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:2F1816670DEDA01214392AA0B1110B23" county="Sra Keo Province" municipality="Klong Had District" name="Sri Thong Cave">Sri Thong Cave</location>
|
|
,
|
|
<collectingMunicipality id="39D84FF672C22FAFC496D3268614EE06">Klong Had District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="0E8C569991F96E2374D6E901AEFB32A5">Sra Keo Province</collectingCounty>
|
|
,
|
|
<collectingDate id="36D50F23FCEB94D0CAB3AD80E7307153" value="2006-11-25">25 Nov. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="F00A375E957ABC7891DB352DB177BF06" collectingDate="2006-11-25" county="Sra Keo Province" location="Liam Cave" municipality="Klong Had District" specimenCount="1">
|
|
CUMZ 12053 (3 shells) from
|
|
<location id="6378E27689E12DD67B4526B75C42C7B3" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:6378E27689E12DD67B4526B75C42C7B3" county="Sra Keo Province" municipality="Klong Had District" name="Liam Cave">Liam Cave</location>
|
|
,
|
|
<collectingMunicipality id="F20583E2AEDC9D2F200874102A925E1E">Klong Had District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="DC55F2654B7A2EEBB360C0ED7889979A">Sra Keo Province</collectingCounty>
|
|
,
|
|
<collectingDate id="B26717860E1222938B0410E2CFF5FAF5" value="2006-11-25">25 Nov. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="D13290D0919C9F60F6EEA4ACE32F51CC" collectingDate="2012-05-21" county="Sra Keo Province" location="Khao Pha Pheung Temple" municipality="Klong Had District" specimenCount="9">
|
|
CUMZ 12054 (7 shells and
|
|
<specimenCount id="B3844899CEE59DC247197B3637D887F9" type="generic">9 specimens</specimenCount>
|
|
in ethanol) from
|
|
<location id="E0B618EDA99678B10BAA7B1D6FDF0317" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:E0B618EDA99678B10BAA7B1D6FDF0317" county="Sra Keo Province" municipality="Klong Had District" name="Khao Pha Pheung Temple">Khao Pha Pheung Temple</location>
|
|
,
|
|
<collectingMunicipality id="36F024B361E0F7B5CCAD89A5CDFBD0E9">Klong Had District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="B8AD588356BF2159B315B8591370C2D6">Sra Keo Province</collectingCounty>
|
|
,
|
|
<collectingDate id="EC778A3C8CF0A819E19BD6EC1CB0EFEC" value="2012-05-21">21 May 2012</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="9EC5338224EC5ACD0FE7EB9636B74439" collectingDate="2008-11-02" county="Sa Kaeo Province" location="Tham Khao Maka Temple" municipality="Mueang Sa Kaeo District" specimenCount="1">
|
|
CUMZ 12055 (12 shells and
|
|
<specimenCount id="45041902F2A52C3A9DDC8CA7B9637F29" type="generic">1 specimen</specimenCount>
|
|
in ethanol) from
|
|
<location id="236A2A54CE70241BE03746E185057E61" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:236A2A54CE70241BE03746E185057E61" county="Sa Kaeo Province" municipality="Mueang Sa Kaeo District" name="Tham Khao Maka Temple">Tham Khao Maka Temple</location>
|
|
,
|
|
<collectingMunicipality id="D7C03D61D7D4434BCB67DE6B543A1200">Mueang Sa Kaeo District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="76DF1F3F8454C64F0F9989A85868300F">Sa Kaeo Province</collectingCounty>
|
|
,
|
|
<collectingDate id="6F3F2553312A5879D6AEEE30CD649C92" value="2008-11-02">2 Nov. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="BE0E23032D0A62B06057F9F22880DA0F" collectingDate="2000-04-07" collectorName="Tham Khao Chakan Temple" location="Khao Chakan District" municipality="Sa Kaeo Province" specimenCount="1">
|
|
CUMZ 12056 (9 shells) from
|
|
<collectorName id="24984A9188BA0A78C50B6ADCFCF15CAE">Tham Khao Chakan Temple</collectorName>
|
|
,
|
|
<location id="0155D6638605649E781CD4276C280DEC" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:0155D6638605649E781CD4276C280DEC" municipality="Sa Kaeo Province" name="Khao Chakan District">Khao Chakan District</location>
|
|
,
|
|
<collectingMunicipality id="A4C865469276261E17AC163889D287D7">Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="11EE97F3B5F8A4D828D5B6F6B6B4CB59" value="2000-04-07">7 Apr. 2000</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="87D3CD887A02D72F0CEFD551240F0B2E" collectingDate="2018-07-25" collectorName="Tham Khao Chakan Temple" location="Khao Chakan District" municipality="Sa Kaeo Province" specimenCount="9">
|
|
CUMZ 12057 (
|
|
<specimenCount id="50D2B14B672B6297023F7269084083D0" type="generic">9 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="FF425BAD9A7746DD66A2D21F03761061">Tham Khao Chakan Temple</collectorName>
|
|
,
|
|
<location id="0CF9213FD812EC6E9565DAF897E3F12B" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:0CF9213FD812EC6E9565DAF897E3F12B" municipality="Sa Kaeo Province" name="Khao Chakan District">Khao Chakan District</location>
|
|
,
|
|
<collectingMunicipality id="33A23506EF024485A971BC8DBA889FAD">Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="CB435740849F65E10787D05B598B82D2" value="2018-07-25">25 July 2018</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="74D3E8732162D6CD98D2C88BC384256C" collectingDate="2012-05-22" collectorName="Khao Chakan" location="Khao Chakan District" municipality="Sa Kaeo Province" specimenCount="3">
|
|
CUMZ 12058 (
|
|
<specimenCount id="421247376E6AB28D0FD2D992A76DFA90" type="generic">3 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="F83028AF32C1EF524257CC36E2A74E48">Khao Chakan</collectorName>
|
|
,
|
|
<location id="F7A22B5C3AF584691CC9419D29651B32" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:F7A22B5C3AF584691CC9419D29651B32" municipality="Sa Kaeo Province" name="Khao Chakan District">Khao Chakan District</location>
|
|
,
|
|
<collectingMunicipality id="B321A8A1322F93596D89351A55460C58">Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="C18943B98D07E3CF991312C301782B30" value="2012-05-22">22 May 2012</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="36A039FBF0047CF089A374CE2CB62CE8" collectingDate="2014-08-10" collectorName="Makok Waterfall" location="Khlung District" municipality="Chanthaburi Province" specimenCount="1">
|
|
CUMZ 12059 (1 shell) from
|
|
<collectorName id="CCE4A4B239E5DF15CF5289DD14A6099F">Makok Waterfall</collectorName>
|
|
,
|
|
<location id="C899EAEBD60779C9081EBA13F4251115" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:C899EAEBD60779C9081EBA13F4251115" municipality="Chanthaburi Province" name="Khlung District">Khlung District</location>
|
|
,
|
|
<collectingMunicipality id="3F4DD82891AED4D05B5A33B8912DF30A">Chanthaburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="606E16F34A6DE357E231FCC8BE6F2391" value="2014-08-10">10 Aug. 2014</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="E1EDAA6AE922199D2CEF1397990B51C3" collectingDate="2010-10-20" collectorName="Phlio Waterfall" location="Mueang Chanthaburi District" municipality="Chanthaburi Province" specimenCount="2">
|
|
CUMZ 12060 (
|
|
<specimenCount id="C853C39A523335D8DE6E35882D9D5A7E" type="generic">2 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="BBFF869BF5AFCED277E274D1A3F9FE28">Phlio Waterfall</collectorName>
|
|
,
|
|
<location id="237265147925484551ED5E2FBFCE25E2" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:237265147925484551ED5E2FBFCE25E2" municipality="Chanthaburi Province" name="Mueang Chanthaburi District">Mueang Chanthaburi District</location>
|
|
,
|
|
<collectingMunicipality id="9EDFCFF04B32CCE562284404134757F8">Chanthaburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="1281DDE7E983A3E28E7AA7EBF9CDB9C0" value="2010-10-20">20 Oct. 2010</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="346C13A24878455DB2CED969D0850C6C" collectingDate="2011-08-09" collectorName="Khao Sukim Temple" location="Tha Mai District" municipality="Chanthaburi Province" specimenCount="3">
|
|
CUMZ 12061 (
|
|
<specimenCount id="69DA3B9F984BE3DD2675FA480708749F" type="generic">3 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="7E901619B4846EC9D4BC1BDE05EBE7CC">Khao Sukim Temple</collectorName>
|
|
,
|
|
<location id="6E9870D4566906322BFBC7DF4B6130D4" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:6E9870D4566906322BFBC7DF4B6130D4" municipality="Chanthaburi Province" name="Tha Mai District">Tha Mai District</location>
|
|
,
|
|
<collectingMunicipality id="A520D941BC82A748BBDDE73DE808D361">Chanthaburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="7939584C248709ECEA77F29EB9C96950" value="2011-08-09">9 Aug. 2011</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="4DCBA9E3AC7B1298B2C6D142439CB3C5" collectingDate="2018-07-24" county="Chanthaburi Province" location="Tham Krong Thip Bureau of Monks" municipality="Tha Mai District" specimenCount="2">
|
|
CUMZ 12062 (
|
|
<specimenCount id="386889229EB9257A47AB9CA73A16FC7B" type="generic">2 specimens</specimenCount>
|
|
in ethanol) from
|
|
<location id="670E52D48D4C45472CA9C45EADC83985" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:670E52D48D4C45472CA9C45EADC83985" county="Chanthaburi Province" municipality="Tha Mai District" name="Tham Krong Thip Bureau of Monks">Tham Krong Thip Bureau of Monks</location>
|
|
,
|
|
<collectingMunicipality id="5CF3361C252628DFE457F128EB4C63D3">Tha Mai District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="B77F43A88E7E15D3CF98E24E63145843">Chanthaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="A3CDF1A5B10D067F1902ED6F21CA4127" value="2018-07-24">24 July 2018</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="E88684F4E4FBC3DDF963B9397ADEAADD" collectingDate="2016-08-04" collectorName="Tham Khao Wong Temple" location="Kaeng Hang Maeo District" municipality="Chanthaburi Province" specimenCount="2">
|
|
CUMZ 12063 (
|
|
<specimenCount id="F50E2E08C252414C8CC811AD277FEE20" type="generic">2 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="0FF6CAE6994ABA08848962AF2300EF73">Tham Khao Wong Temple</collectorName>
|
|
,
|
|
<location id="4660FD374B0A4691227A08208045270D" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:4660FD374B0A4691227A08208045270D" municipality="Chanthaburi Province" name="Kaeng Hang Maeo District">Kaeng Hang Maeo District</location>
|
|
,
|
|
<collectingMunicipality id="5C7099ADFFD7E17AA363EB264EF69219">Chanthaburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="D25D2CF60AA5AB0D2F38C17C613F50E7" value="2016-08-04">4 Aug. 2016</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3C7EC53261A51CE32E0760C8C6789618" collectingDate="2008-01-02" collectorName="Tham Khao Charoensuk Temple" location="Phanom Sarakham District" municipality="Chachoengsao Province" specimenCount="1">
|
|
CUMZ 12064 (8 shells) from
|
|
<collectorName id="71547A1D5C5AF5518CCFC37B6786FA68">Tham Khao Charoensuk Temple</collectorName>
|
|
,
|
|
<location id="8FEE2E7C9FA941A90F733376AFEFC87A" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:8FEE2E7C9FA941A90F733376AFEFC87A" municipality="Chachoengsao Province" name="Phanom Sarakham District">Phanom Sarakham District</location>
|
|
,
|
|
<collectingMunicipality id="F635BFBCFB56F62AADB77C8F296E517E">Chachoengsao Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="C80CC85CFC803F306B3036690DE209D6" value="2008-01-02">2 Jan. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3A9B0C24634B9A3B79D8C994730E61E4" collectingDate="2006-08-17" county="Chonburi Province" location="Tham Khao Cha Ang On Temple" municipality="Bo Thong District" specimenCount="3">
|
|
CUMZ 12065 (18 shells and
|
|
<specimenCount id="44B27ACBA8D160F96F12D655E9E0D83D" type="generic">3 specimens</specimenCount>
|
|
in ethanol) from
|
|
<location id="D9AEE78E573A0FE072C70FB8F6F0BD54" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:D9AEE78E573A0FE072C70FB8F6F0BD54" county="Chonburi Province" municipality="Bo Thong District" name="Tham Khao Cha Ang On Temple">Tham Khao Cha Ang On Temple</location>
|
|
,
|
|
<collectingMunicipality id="723786CE4DC6A6CD9D236C32EBACD0A6">Bo Thong District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="41ACE87C3CECD80F609F8814F487898B">Chonburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="8C2891EF8BC31C0A2B2DB676D99F911A" value="2006-08-17">17 Aug. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="BDF084A1E1DF9A02D9E3D3CA4C964761" collectingDate="2008-05-09" location="Bo Thong District" municipality="Chonburi Province" specimenCount="1">
|
|
CUMZ 12066 (5 shells) from
|
|
<location id="24106B78FEDA7EFF0D518DBE9A2C2748" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:24106B78FEDA7EFF0D518DBE9A2C2748" municipality="Chonburi Province" name="Bo Thong District">Bo Thong District</location>
|
|
,
|
|
<collectingMunicipality id="B13C81793D1BFE3179F611DCED0EE81D">Chonburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="BF80E3A4CB310750A41C2F8CAF741B7B" value="2008-05-09">9 May 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="A414829B9085C6082FD18F61E31C6596" collectingDate="2022-02-06" collectorName="Khao Ha Yot Temple" location="Bo Thong District" municipality="Chonburi Province" specimenCount="2">
|
|
CUMZ 12194 (
|
|
<specimenCount id="544D04B06703BE30EEABA140CBD48327" type="generic">2 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="DE488DE557EFEF3109A005B667AD8230">Khao Ha Yot Temple</collectorName>
|
|
,
|
|
<location id="9A38138C8AD1AE5C05E191F6B6D91F0A" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:9A38138C8AD1AE5C05E191F6B6D91F0A" municipality="Chonburi Province" name="Bo Thong District">Bo Thong District</location>
|
|
,
|
|
<collectingMunicipality id="568DCC644E865E444FB3B627E2CD6B3B">Chonburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="58650022B055463A9774784986078D09" value="2022-02-06">6 Feb. 2022</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="7BB5D71C882EDE3475BF63D6D890F82B" collectingDate="2020-09-19" collectorName="Fig." county="Chonburi Province" location="Phromawat Temple" municipality="Si Racha District" specimenCount="20">
|
|
CUMZ 12067 (2 shells and
|
|
<specimenCount id="7111D44F02D4DC664D57C0334710E800" type="generic">20 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="09C9BACFF83A2C719AC10181075C1E0F">Fig.</collectorName>
|
|
<figureCitation id="5B0CE4ACE6E7E7DF285B375FBEA048B2" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Opercula of three mainland Southeast Asian Pupina species groups A Pupina bensoni sp. nov. from the Pupina artata species group, specimen CUMZ 12047 B Pupina siamensis from the Pupina arula species group, specimen CUMZ 12067, and C Pupina aureola from the Pupina aureola species group, specimen CUMZ 12116. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure19" httpUri="https://binary.pensoft.net/fig/735380" pageId="0" pageNumber="1">19B</figureCitation>
|
|
) from
|
|
<location id="3F2000083C6E576C7FF1F9D9A488376D" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:3F2000083C6E576C7FF1F9D9A488376D" county="Chonburi Province" municipality="Si Racha District" name="Phromawat Temple">Phromawat Temple</location>
|
|
,
|
|
<collectingMunicipality id="33B947B58BCBB474B541CDC394441451">Si Racha District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="D75CE4CF37213938A251BF3ADD2D646C">Chonburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="AD2416BB395AA6E171D5F061A1CFC140" value="2020-09-19">19 Sept. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="9D91510DE899582AB32420297A17AEA7" collectingDate="2020-09-19" collectorName="Pa Lilaiyawan Temple" location="Si Racha District" municipality="Chonburi Province" specimenCount="10">
|
|
CUMZ 12068 (
|
|
<specimenCount id="A602596C404492989AE8CB752223ED58" type="generic">10 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="C363504DE7155C845A384C5AED85FDE8">Pa Lilaiyawan Temple</collectorName>
|
|
,
|
|
<location id="E047964B3B56F6BCBC7E7C0E44C18659" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:E047964B3B56F6BCBC7E7C0E44C18659" municipality="Chonburi Province" name="Si Racha District">Si Racha District</location>
|
|
,
|
|
<collectingMunicipality id="7F2E62945454A97D204FDCAE355BEA4D">Chonburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="31D488FB12304CC64D03EBE694011294" value="2020-09-19">19 Sept. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="116587DA2EC1276D71C6AD410A8129B9" collectingDate="2020-09-19" collectorName="Fig." county="Chonburi Province" location="Khao Chi Chan Buddha Image" municipality="Sattahip District" specimenCount="9">
|
|
CUMZ 12069 (1 shell and
|
|
<specimenCount id="65A71D48BF8B0E908B9DC94C8DCBE8DF" type="generic">9 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="223EC9D0A1CCBBE5764BB2ED33DFA716">Fig.</collectorName>
|
|
<figureCitation id="7C608873A6EEF5E3A09C26034175B1F8" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">25H</figureCitation>
|
|
) from
|
|
<location id="0BA8A0BD8FB0C7363E838FAD1B40FCAF" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:0BA8A0BD8FB0C7363E838FAD1B40FCAF" county="Chonburi Province" municipality="Sattahip District" name="Khao Chi Chan Buddha Image">Khao Chi Chan Buddha Image</location>
|
|
,
|
|
<collectingMunicipality id="F0B22F25923C85E909820378EA0637E7">Sattahip District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="DB28508BA13095667C7C6EC6BED46726">Chonburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="33D0287E6EBD238A4912194AFEA8F283" value="2020-09-19">19 Sept. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="956EF8FE16EE678A7B338CF06ECB2E11" collectingDate="2020-09-19" county="Chonburi Province" location="Ban Klong Wan Pen" municipality="Sattahip District" specimenCount="8">
|
|
CUMZ 12070 (1 shell and
|
|
<specimenCount id="A3DA60DE3201E96A0213144C95F87823" type="generic">8 specimens</specimenCount>
|
|
in ethanol) from
|
|
<location id="58FCA0B9698F00E3F9FFFE56465AF67D" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:58FCA0B9698F00E3F9FFFE56465AF67D" county="Chonburi Province" municipality="Sattahip District" name="Ban Klong Wan Pen">Ban Klong Wan Pen</location>
|
|
,
|
|
<collectingMunicipality id="970B26A65C61FA3421B7BB295FB9E1D3">Sattahip District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="E93D0C5F483E3B8B532835FC77DE6662">Chonburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="2A3E8803FF26791791DF7A2D0995A593" value="2020-09-19">19 Sept. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="7C94B60EA0188FFE9846883F60DFAEB9" collectingDate="2008-09-05" collectorName="Fig." county="Rayong Province" location="Tham Khao Loi Temple" municipality="Khao Chamao District" specimenCount="10">
|
|
CUMZ 12071 (
|
|
<specimenCount id="88F650C5D285DB608FA35728D5FEDC5E" type="generic">10 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="8F8456F55AB82B61407CBE4CFF6A5EDA">Fig.</collectorName>
|
|
<figureCitation id="756ABE5E91CFE42DFEFCE1E86C28EB7D" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">32A</figureCitation>
|
|
) from
|
|
<location id="3C613D893DE6021A67AC45326A20BFCD" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:3C613D893DE6021A67AC45326A20BFCD" county="Rayong Province" municipality="Khao Chamao District" name="Tham Khao Loi Temple">Tham Khao Loi Temple</location>
|
|
,
|
|
<collectingMunicipality id="FF03B301859BDF18F9B604B559FAFC4C">Khao Chamao District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="B823F209C714F14A50A5C459F35474D3">Rayong Province</collectingCounty>
|
|
,
|
|
<collectingDate id="E87226E8BA7C66B62F36B7457F0980DD" value="2008-09-05">5 Sept. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="DC635CD82D3D4FEBE594EFCBE7F56B15" collectingDate="2019-06-09" county="Rayong Province" location="Khao Hin Tang Bureau of Monks" municipality="Klaeng District" specimenCount="1">
|
|
CUMZ 12072 (
|
|
<specimenCount id="14C93CA25D7E9B28EF41B92D2294A7D5" type="generic">1 specimen</specimenCount>
|
|
in ethanol) from
|
|
<location id="2E275F558529956B96951C03DAD074E6" LSID="urn:lsid:plazi:treatment:59C3770EFAE55DADBBC2115A71AABB2B:2E275F558529956B96951C03DAD074E6" county="Rayong Province" municipality="Klaeng District" name="Khao Hin Tang Bureau of Monks">Khao Hin Tang Bureau of Monks</location>
|
|
,
|
|
<collectingMunicipality id="2A36E25A92449284A903667F08EAAE01">Klaeng District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="C54EE8EAA8B36B6595062D43725590C1">Rayong Province</collectingCounty>
|
|
,
|
|
<collectingDate id="1C173283BB565EE56F2F8FFABA5058FE" value="2019-06-09">9 June 2019</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5495D17D25F64642B9BD741D53BE1CB0" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="76B789087FA84D02F8C55DA616AD8A55" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="20ED751C35966919CA052AAA9E467042" pageId="0" pageNumber="1">Shell globose; last whorl ca. 80% of shell height. Apertural lip thickened, but not expanded; apertural lip when observed from lateral view almost straight. Columellar tooth fin-shaped.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2267AB7B80B5F2F5BB8DF9B72CAAB582" pageId="0" pageNumber="1" type="differential diagnosis">
|
|
<paragraph id="40008C83689C38B2BEB5A0647B18219B" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
|
<paragraph id="E87E3CC9CCF720C688EB2EF0FFD82BC1" pageId="0" pageNumber="1">
|
|
<taxonomicName id="CC72266F741552E06143CF28A58C919E" authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
|
|
<emphasis id="D5C5DC0AEBE0E661E00974A8EE1DF71F" italics="true" pageId="0" pageNumber="1">Pupina siamensis</emphasis>
|
|
</taxonomicName>
|
|
is most similar to
|
|
<taxonomicName id="5845090FC643ED606794BDFD33E7A553" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mouhoti">
|
|
<emphasis id="64600F7881DA53AD2EDF80428C9692A1" italics="true" pageId="0" pageNumber="1">P. mouhoti</emphasis>
|
|
</taxonomicName>
|
|
in having an almost straight apertural lip when observed from lateral view, but differs in having a more globose shell shape and a thicker, more distinct, parietal tooth.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="47353A2DB1F658B2654A893D057F6BAA" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="C5FE6A5F79012225C2CE9BE14032A674" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="46CBDA8A7A95097F2917C3B27731EBAA" pageId="0" pageNumber="1">
|
|
Eastern and northeastern Thailand (
|
|
<bibRefCitation id="5B988B267E75596037EE9576C3C7CAB5" author="Boonngam, P" journalOrPublisher="Witthayasan Kasetsat Witthayasat" pageId="0" pageNumber="1" pagination="256 - 263" refId="B14" refString="Boonngam, P, Dumrongrojwattana, P, Matchacheep, S, 2008. The diversity of land snail fauna in Chonburi Province, Eastern Thailand. Witthayasan Kasetsat Witthayasat 42: 256 - 263" title="The diversity of land snail fauna in Chonburi Province, Eastern Thailand." volume="42" year="2008">Boonngam et al. 2008</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="41307A6B1E14E02FE9AA22E7007D9B4D" author="Chanyapate, C" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" refId="B20" refString="Chanyapate, C, Promkerd, P, Klakhaeng, S, Rintarak, D, Nookarn, P, 2008. Biodiversity of land snail and slug in Biosphere Reserve Sakaerat. Research Report 2008, Plant Protection Research and Development Office, Department of Agriculture, Ministry of Agriculture and Cooperatives, Bangkok, 60-72. [in Thai]" title="Biodiversity of land snail and slug in Biosphere Reserve Sakaerat. Research Report 2008, Plant Protection Research and Development Office, Department of Agriculture, Ministry of Agriculture and Cooperatives, Bangkok, 60 - 72. [in Thai]" year="2008">Chanyapate et al. 2008</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7DDB65C6F1CA6EA1026C80B7353238C8" author="Chidchua, W" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" refId="B22" refString="Chidchua, W, Dumrongrojwattana, P, 2010. Taxonomy of land snails in Klaeng District Rayong Province and Kaenghangmaew District Chanthaburi Province, Eastern Thailand (Gastropoda: Prosobranchia, Pulmonata). In: The Proceedings of 48th Kasetsart University Annual Conference: Science. Kasetsart University, Bangkok, 161-170. [in Thai]" title="Taxonomy of land snails in Klaeng District Rayong Province and Kaenghangmaew District Chanthaburi Province, Eastern Thailand (Gastropoda: Prosobranchia, Pulmonata). In: The Proceedings of 48 th Kasetsart University Annual Conference: Science. Kasetsart University, Bangkok, 161 - 170. [in Thai]" year="2010">Chidchua and Dumrongrojwattana 2010</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="2F5F189B312F9661680AA1134C3F9AA6" author="Dumrongrojwattana, P" journalOrPublisher="Department of Biology, Burapha University, Bangkok" pageId="0" pageNumber="1" refId="B37" refString="Dumrongrojwattana, P, 2016. Malacofauna in Pracheenburi Province (Under the Plant Genetic Conservation Project Under the Royal Initiative of Her Highness Princess Maha Chakri Sirindhorn). Department of Biology, Burapha University, Bangkok" title="Malacofauna in Pracheenburi Province (Under the Plant Genetic Conservation Project Under the Royal Initiative of Her Highness Princess Maha Chakri Sirindhorn)." year="2016">Dumrongrojwattana 2016</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9B5C78D5D32BFB92EB4408431BCAC72B" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="3A325DE7818D622EC41E413249B89735" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="81F06340A31DFE88A4751FF133E1C4AC" pageId="0" pageNumber="1">
|
|
As the original description did not explicitly state that the description of this species was based on a single specimen (nor could this be inferred), the designation of a holotype by
|
|
<bibRefCitation id="2C08534F568D7672CD2AD083D172C729" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="41 - 56" refId="B198" refString="Zilch, A, 1957. Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae. Archiv fuer Molluskenkunde 86: 41 - 56" title="Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae." volume="86" year="1957">Zilch (1957)</bibRefCitation>
|
|
in fact constitutes a lectotype designation (
|
|
<bibRefCitation id="8DBC7CA177C06EC30384688F75B414BF" author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
|
|
: Art. 74.6).
|
|
</paragraph>
|
|
<paragraph id="E0853EB5B03B77479D9632BE77877703" pageId="0" pageNumber="1">
|
|
The type locality of this species in Bangkok, the capital city of Thailand, as designated by
|
|
<bibRefCitation id="04E3608C22C4A5561DD132DC045846B3" author="von Moellendorff, OF" journalOrPublisher="Nachrichtsbatt der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="153 - 160" refId="B195" refString="von Moellendorff, OF, 1902b. Binnenmollusken aus Hinterindien. 2. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsbatt der Deutschen Malakozoologischen Gesellschaft 34: 153 - 160" title="Binnenmollusken aus Hinterindien. 2. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt." volume="34" year="1902 b">
|
|
von
|
|
<normalizedToken id="42048C9C8B34962D392C201D78BD6CF7" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
|
(1902b)
|
|
</bibRefCitation>
|
|
is dubious. This species was described based on a collection made by the butterfly collector, H. Fruhstorfer, who made an expedition in Thailand (
|
|
<bibRefCitation id="5F644F15797F3397FF0E7214D0688AA3" author="Lamas, G" journalOrPublisher="Entomofauna" pageId="0" pageNumber="1" pagination="57 - 100" refId="B83" refString="Lamas, G, 2005. A bibliography of the zoological publications of Hans Fruhstorfer (1866-1922). Entomofauna 26: 57 - 100" title="A bibliography of the zoological publications of Hans Fruhstorfer (1866 - 1922)." volume="26" year="2005">Lamas 2005</bibRefCitation>
|
|
). The type locality
|
|
<normalizedToken id="1C89608ED47FEBC4D9ED855D1A432861" originalValue="“Bangkok”">"Bangkok"</normalizedToken>
|
|
is probably not the location where the type specimen was collected. The probable type locality would be
|
|
<normalizedToken id="4634EF5C3EAB553C5C63650774444CD3" originalValue="“Muok-Lek”">"Muok-Lek"</normalizedToken>
|
|
[Muak Lek District, Saraburi Province, Thailand] as indicated in
|
|
<bibRefCitation id="ECBE35EFFD0F0BB063BC7AC8E99628CE" author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg (1904)</bibRefCitation>
|
|
, and several butterfly specimens were also collected from this site by H. Fruhstorfer. This locality is in the same vicinity as recent records and our collecting localities of
|
|
<taxonomicName id="E5AABB642D1EC0CC17A773F8C3BB984B" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
|
|
<emphasis id="8F5A013CE28334DF53F40BF0AD1692B1" italics="true" pageId="0" pageNumber="1">P. siamensis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |