575 lines
51 KiB
XML
575 lines
51 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.993.53805" ID-GBIF-Dataset="153d057d-c190-4db9-aa8b-4da3cb04ae64" ID-PMC="PMC7683496" ID-Pensoft-Pub="1313-2970-993-47" ID-Pensoft-UUID="2D7B2EB8CB285CBCA90A27064C6B1F17" ID-PubMed="33262675" ID-ZooBank="C40989DB80634C9FA481E7AA82CA924B" ModsDocID="1313-2970-993-47" checkinTime="1605571755153" checkinUser="pensoft" docAuthor="Ruan, Yongying, Douglas, Hume B., Qiu, Lu, Chen, Xiaoqin & Jiang, Shihong" docDate="2020" docId="2996AC3928D75A2C9557DD0AD04C3F9D" docLanguage="en" docName="ZooKeys 993: 47-120" docOrigin="ZooKeys 993" docSource="http://dx.doi.org/10.3897/zookeys.993.53805" docTitle="Phorocardius zhiweii Ruan, Douglas & Qiu 2020, sp. nov." docType="treatment" docUuid="6DFDF68D-2147-4055-8034-243A6FEBEA72" docUuidSource="ZooBank" docVersion="4" id="2D7B2EB8CB285CBCA90A27064C6B1F17" lastPageNumber="47" masterDocId="2D7B2EB8CB285CBCA90A27064C6B1F17" masterDocTitle="Revision of Chinese Phorocardius species (Coleoptera, Elateridae, Cardiophorinae)" masterLastPageNumber="120" masterPageNumber="47" pageNumber="47" updateTime="1668169908327" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of Chinese Phorocardius species (Coleoptera, Elateridae, Cardiophorinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ruan, Yongying</mods:namePart>
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<mods:affiliation>School of Applied Chemistry and Biological Technology, Shenzhen Polytechnic, Shenzhen, Guangdong 518055, China</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5025-5592</mods:nameIdentifier>
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<mods:nameIdentifier type="email">yongyingruan@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Douglas, Hume B.</mods:namePart>
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<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Ave., Ottawa, Ontario, K 1 A 0 C 6, Canada</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-1722-7554</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Qiu, Lu</mods:namePart>
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<mods:affiliation>Institute of Entomology, College of Plant Protection, Southwest University, Beibei, Chongqing 400716, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chen, Xiaoqin</mods:namePart>
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<mods:affiliation>School of Applied Chemistry and Biological Technology, Shenzhen Polytechnic, Shenzhen, Guangdong 518055, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jiang, Shihong</mods:namePart>
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<mods:affiliation>School of Applied Chemistry and Biological Technology, Shenzhen Polytechnic, Shenzhen, Guangdong 518055, China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>993</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>47</mods:start>
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<mods:end>120</mods:end>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.993.53805</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.993.53805</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-993-47</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2D7B2EB8CB285CBCA90A27064C6B1F17</mods:identifier>
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<treatment ID-GBIF-Taxon="169091766" LSID="urn:lsid:zoobank.org:act:6DFDF68D-2147-4055-8034-243A6FEBEA72" httpUri="http://treatment.plazi.org/id/2996AC3928D75A2C9557DD0AD04C3F9D" lastPageNumber="47" pageId="0" pageNumber="47">
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<subSubSection pageId="0" pageNumber="47" type="nomenclature">
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<paragraph pageId="0" pageNumber="47">
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11.
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<taxonomicName LSID="2996AC39-28D7-5A2C-9557-DD0AD04C3F9D" authority="Ruan, Douglas & Qiu" authorityName="Ruan, Douglas & Qiu" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius zhiweii" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="zhiweii" status="sp. nov.">Phorocardius zhiweii Ruan, Douglas & Qiu</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="47">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov., holotype, male. A habitus, dorsal view. B habitus, ventral view C habitus, lateral view D aedeagus ventral view, arrow indicating apex and apical mesal callus of paramere E aedeagus, dorsal view F aedeagus, lateral view, arrow indicating apices of paramere and median lobe G tergite 7, dorsal view, hand drawing, arrow indicating posterior concavity." figureDoi="10.3897/zookeys.993.53805.figure21" httpUri="https://binary.pensoft.net/fig/475454" pageId="0" pageNumber="47">Figs 21</figureCitation>
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov. A pronotum and scutellar shield, dorsal view B left posterior edge of pronotum, showing sublateral incision (arrow) C head, frontal view D metacoxal plate, ventral view E tarsal claws F male abdominal tergites IX-X, dorsal view G male abdominal sternite IX, dorsal view." figureDoi="10.3897/zookeys.993.53805.figure22" httpUri="https://binary.pensoft.net/fig/475455" pageId="0" pageNumber="47">, 22</figureCitation>
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Shape of supraantennal carina of Chinese Phorocardius species. Arrows indicating supraantennal carina forked near eye. A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan & Douglas, sp. nov. J P. zhiweii Ruan, Douglas & Qiu, sp. nov. K head, frontal-lateral view, arrows indicating forked supraantennal carina." figureDoi="10.3897/zookeys.993.53805.figure23" httpUri="https://binary.pensoft.net/fig/475457" pageId="0" pageNumber="47">, 23J</figureCitation>
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Lateral view of prosternal process and mesosternum of Chinese Phorocardius species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i. e., on anteroventral angle of mesosternal fossa according to Douglas 2003). A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan & Douglas, sp. nov. J P. zhiweii Ruan, Douglas & Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47">, 24J</figureCitation>
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Ventral view of pro- and mesothorax of Chinese Phorocardius species. Showing shape of ventral surface of prosternal process (upper, yellow dashed line), mesepisternum (middle, green arrow and dashed line) and projections on posterior edge of mesosternum (lower, red arrow and solid line) middle, green arrows indicating anterior-mesal angle of mesepisternum. A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. zhiweii Ruan, Douglas & Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure25" httpUri="https://binary.pensoft.net/fig/475459" pageId="0" pageNumber="47">, 25I</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="47" type="type locality">
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<paragraph pageId="0" pageNumber="47">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="47">Yunnan: Long-chuan county, Hu-sa township, Gun-bang-jian-shan [i.e., Bang-gun-jian-shan邦棍尖山].</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="47" type="etymology">
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<paragraph pageId="0" pageNumber="47">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="47">This species is named after its collector, Mr Zhiwei Dong, who generously provided specimens for this study.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="47" type="distribution">
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<paragraph pageId="0" pageNumber="47">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="47">China (Yunnan).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="47" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="47">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="47">Body length greater than 7.0 mm; pronotum with integument red, elytra metallic green. Prothorax: procoxal cavities open; prosternal process not strongly narrowed posterad to ventral apex in ventral view, with ventral apex rounded. Pterothorax: scutellar shield with posterior edge pointed. Tarsal claw with ventral apex not smaller than dorsal apex. Male genitalia: paramere without preapical lateral expansion, but with apical mesal callus. Female unknown.</paragraph>
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<paragraph pageId="0" pageNumber="47">
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This species is distinct among
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<taxonomicName authorityName="Fleutiaux" authorityYear="1931" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius</emphasis>
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</taxonomicName>
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in its entirely metallic green elytra and red pronotum. It is the second species having metallic color on elytra, as the type species of the genus (i.e.,
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<taxonomicName lsidName="P. florentini" pageId="0" pageNumber="47" rank="species" species="florentini">
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<emphasis italics="true" pageId="0" pageNumber="47">P. florentini</emphasis>
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</taxonomicName>
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(Fleutiaux, 1895)) also has slight blue metallic luster on elytra.
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</paragraph>
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<paragraph pageId="0" pageNumber="47">
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<taxonomicName authorityName="Ruan, Douglas & Qiu" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius zhiweii" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="zhiweii">
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<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius zhiweii</emphasis>
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</taxonomicName>
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Ruan, Douglas & Qiu, sp. nov. resembles
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<taxonomicName lsidName="P. manuleatus" pageId="0" pageNumber="47" rank="species" species="manuleatus">
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<emphasis italics="true" pageId="0" pageNumber="47">P. manuleatus</emphasis>
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</taxonomicName>
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most in body shape and color of leg and prothorax. They can be easily separated by elytral color and aedeagus shape. In
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<taxonomicName lsidName="P. zhiweii" pageId="0" pageNumber="47" rank="species" species="zhiweii">
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<emphasis italics="true" pageId="0" pageNumber="47">P. zhiweii</emphasis>
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</taxonomicName>
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Ruan, Douglas & Qiu, sp. nov., elytra are metallic green with slight purple luster; in ventral view, the aedeagus has apical fourth slightly narrowed, robust in both ventral and lateral views; and the paramere has preapical lateral expansion absent, apical mesal callus present; while in
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<taxonomicName lsidName="P. manuleatus" pageId="0" pageNumber="47" rank="species" species="manuleatus">
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<emphasis italics="true" pageId="0" pageNumber="47">P. manuleatus</emphasis>
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</taxonomicName>
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, elytra are black to yellow, without metallic color; and the aedeagus in ventral view has the apical fourth greatly narrowed. The aedeagus is slender in both ventral and lateral views with parameres lacking acute preapical lateral expansions or apical mesal calli.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="47" type="description">
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<paragraph pageId="0" pageNumber="47">Description.</paragraph>
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<paragraph pageId="0" pageNumber="47">
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(based on holotype) Dorsum shiny. Head black, with mouthparts red-brown to brown. Antennae brown. Pronotum orange, with posterior edge dark brown (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov. A pronotum and scutellar shield, dorsal view B left posterior edge of pronotum, showing sublateral incision (arrow) C head, frontal view D metacoxal plate, ventral view E tarsal claws F male abdominal tergites IX-X, dorsal view G male abdominal sternite IX, dorsal view." figureDoi="10.3897/zookeys.993.53805.figure22" httpUri="https://binary.pensoft.net/fig/475455" pageId="0" pageNumber="47">22A</figureCitation>
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). Scutellar shield black. Elytra entirely metallic green, with purple luster. Hypomera orange; rest of ventral surface black (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov., holotype, male. A habitus, dorsal view. B habitus, ventral view C habitus, lateral view D aedeagus ventral view, arrow indicating apex and apical mesal callus of paramere E aedeagus, dorsal view F aedeagus, lateral view, arrow indicating apices of paramere and median lobe G tergite 7, dorsal view, hand drawing, arrow indicating posterior concavity." figureDoi="10.3897/zookeys.993.53805.figure21" httpUri="https://binary.pensoft.net/fig/475454" pageId="0" pageNumber="47">21B</figureCitation>
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). Epipleura metallic purple. Legs black on coxa, pale brown on femur and basal half of tibia, dark brown from mid-length of tibia to apex. Body with short, yellow-grey pubescence, brown setae also present on disc of pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="47" type="description">
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<paragraph pageId="0" pageNumber="47">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="47">(based on holotype) Body length 8.0 mm, width 2.8 mm. Body length to width ratio 2.9. Pronotal width to length ratio 1.1. Pronotal width to body width ratio 0.80. Elytral length to pronotal length ratio 2.5; elytron length to width ratio 3.8.</paragraph>
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<paragraph pageId="0" pageNumber="47">
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<emphasis bold="true" pageId="0" pageNumber="47">
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<emphasis italics="true" pageId="0" pageNumber="47">Head</emphasis>
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.
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</emphasis>
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Frons and vertex with interspaces between punctures 1-3
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<normalizedToken originalValue="×">x</normalizedToken>
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average puncture diameter (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov. A pronotum and scutellar shield, dorsal view B left posterior edge of pronotum, showing sublateral incision (arrow) C head, frontal view D metacoxal plate, ventral view E tarsal claws F male abdominal tergites IX-X, dorsal view G male abdominal sternite IX, dorsal view." figureDoi="10.3897/zookeys.993.53805.figure22" httpUri="https://binary.pensoft.net/fig/475455" pageId="0" pageNumber="47">22C</figureCitation>
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). Frontal carina in frontal view convex. Antenna with apex extending to basal edge of elytron. Distance between eyes to width of eye ratio 2.7. Antenna length to body length ratio 0.42. Proportions of antennomere lengths (male): 100 (scape); 71; 76; 100; 100; 105; 103; 90; 90; 88; 116.
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</paragraph>
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<paragraph pageId="0" pageNumber="47">
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<emphasis bold="true" pageId="0" pageNumber="47">
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<emphasis italics="true" pageId="0" pageNumber="47">Prothorax</emphasis>
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.
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</emphasis>
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Pronotum in dorsal view (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov. A pronotum and scutellar shield, dorsal view B left posterior edge of pronotum, showing sublateral incision (arrow) C head, frontal view D metacoxal plate, ventral view E tarsal claws F male abdominal tergites IX-X, dorsal view G male abdominal sternite IX, dorsal view." figureDoi="10.3897/zookeys.993.53805.figure22" httpUri="https://binary.pensoft.net/fig/475455" pageId="0" pageNumber="47">22A</figureCitation>
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): sides slightly convex from anterior edge to concavity near posterior fourth, widest near posterior third; posterior angles with lateral sides almost straight, not bulged; surface with interspaces between punctures 2-2.5
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<normalizedToken originalValue="×">x</normalizedToken>
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average puncture diameter. In ventral view, ventral surface of prosternal process with sides carinate and slightly and gradually narrow from anterior to mid-length, parallel from mid-length to near posterior end, apex broadly rounded. In lateral view, prosternal process with ventral surface curved slightly dorsad, posterior end concave (Fig.
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Lateral view of prosternal process and mesosternum of Chinese Phorocardius species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i. e., on anteroventral angle of mesosternal fossa according to Douglas 2003). A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan & Douglas, sp. nov. J P. zhiweii Ruan, Douglas & Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47">24J</figureCitation>
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). Procoxal cavities open.
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</paragraph>
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<paragraph pageId="0" pageNumber="47">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Pterothorax</emphasis>
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(Figs
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Lateral view of prosternal process and mesosternum of Chinese Phorocardius species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i. e., on anteroventral angle of mesosternal fossa according to Douglas 2003). A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan & Douglas, sp. nov. J P. zhiweii Ruan, Douglas & Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47">24J</figureCitation>
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,
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Ventral view of pro- and mesothorax of Chinese Phorocardius species. Showing shape of ventral surface of prosternal process (upper, yellow dashed line), mesepisternum (middle, green arrow and dashed line) and projections on posterior edge of mesosternum (lower, red arrow and solid line) middle, green arrows indicating anterior-mesal angle of mesepisternum. A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. zhiweii Ruan, Douglas & Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure25" httpUri="https://binary.pensoft.net/fig/475459" pageId="0" pageNumber="47">25I</figureCitation>
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). Mesepisternum in ventral view with antero-mesal corner right-angled (Fig.
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Ventral view of pro- and mesothorax of Chinese Phorocardius species. Showing shape of ventral surface of prosternal process (upper, yellow dashed line), mesepisternum (middle, green arrow and dashed line) and projections on posterior edge of mesosternum (lower, red arrow and solid line) middle, green arrows indicating anterior-mesal angle of mesepisternum. A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. zhiweii Ruan, Douglas & Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure25" httpUri="https://binary.pensoft.net/fig/475459" pageId="0" pageNumber="47">25I</figureCitation>
|
||
, upper, green arrow). Projections on posterior edge of mesosternum: in ventral view present (Fig.
|
||
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Ventral view of pro- and mesothorax of Chinese Phorocardius species. Showing shape of ventral surface of prosternal process (upper, yellow dashed line), mesepisternum (middle, green arrow and dashed line) and projections on posterior edge of mesosternum (lower, red arrow and solid line) middle, green arrows indicating anterior-mesal angle of mesepisternum. A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. zhiweii Ruan, Douglas & Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure25" httpUri="https://binary.pensoft.net/fig/475459" pageId="0" pageNumber="47">25I</figureCitation>
|
||
, red arrow); in lateral view present, strongly developed, produced anteriorly (Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Lateral view of prosternal process and mesosternum of Chinese Phorocardius species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i. e., on anteroventral angle of mesosternal fossa according to Douglas 2003). A P. alterlineatus Ruan & Douglas, sp. nov. B P. flavistriolatus Ruan & Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan & Douglas, sp. nov. G P. rufiposterus Ruan & Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan & Douglas, sp. nov. J P. zhiweii Ruan, Douglas & Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47">24J</figureCitation>
|
||
, red arrow). Scutellar shield: width to length ratio 1.0; anterolateral edges slightly sinuate; posterior apex pointed. Elytra: upper edge of epipleura with minute serrations.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Length ratio of metatarsomeres I-V (excluding claws): 100; 71; 53; 50; 120. Claw with ventral apex almost as large as dorsal apex.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Abdomen</emphasis>
|
||
.
|
||
</emphasis>
|
||
Lateral edges of visible abdominal ventrites I-V with minute serrations.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Robust in ventral and lateral views. Median lobe in ventral view with sides nearly parallel, slightly narrowed near apex; apex broadly rounded (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov., holotype, male. A habitus, dorsal view. B habitus, ventral view C habitus, lateral view D aedeagus ventral view, arrow indicating apex and apical mesal callus of paramere E aedeagus, dorsal view F aedeagus, lateral view, arrow indicating apices of paramere and median lobe G tergite 7, dorsal view, hand drawing, arrow indicating posterior concavity." figureDoi="10.3897/zookeys.993.53805.figure21" httpUri="https://binary.pensoft.net/fig/475454" pageId="0" pageNumber="47">21D</figureCitation>
|
||
). Median lobe in lateral view gently bent ventrad, apex dilated and broadly rounded (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov., holotype, male. A habitus, dorsal view. B habitus, ventral view C habitus, lateral view D aedeagus ventral view, arrow indicating apex and apical mesal callus of paramere E aedeagus, dorsal view F aedeagus, lateral view, arrow indicating apices of paramere and median lobe G tergite 7, dorsal view, hand drawing, arrow indicating posterior concavity." figureDoi="10.3897/zookeys.993.53805.figure21" httpUri="https://binary.pensoft.net/fig/475454" pageId="0" pageNumber="47">21F</figureCitation>
|
||
). Paramere in ventral view: wide, widest near mid-length, sides convex to near apex; preapical ventral expansion absent, mesal side of apex with ovoid disc-shaped callus (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov., holotype, male. A habitus, dorsal view. B habitus, ventral view C habitus, lateral view D aedeagus ventral view, arrow indicating apex and apical mesal callus of paramere E aedeagus, dorsal view F aedeagus, lateral view, arrow indicating apices of paramere and median lobe G tergite 7, dorsal view, hand drawing, arrow indicating posterior concavity." figureDoi="10.3897/zookeys.993.53805.figure21" httpUri="https://binary.pensoft.net/fig/475454" pageId="0" pageNumber="47">21D</figureCitation>
|
||
, indicated by blue arrow); paramere 2-3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
wider than median lobe (measured at mid-length of paramere and median lobe respectively). Paramere in lateral view: robust, almost straight at basal half, bent ventrad at apical half; preapical ventral expansion hook-like, with acute tip facing ventrad (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Phorocardius zhiweii Ruan, Douglas & Qiu, sp. nov., holotype, male. A habitus, dorsal view. B habitus, ventral view C habitus, lateral view D aedeagus ventral view, arrow indicating apex and apical mesal callus of paramere E aedeagus, dorsal view F aedeagus, lateral view, arrow indicating apices of paramere and median lobe G tergite 7, dorsal view, hand drawing, arrow indicating posterior concavity." figureDoi="10.3897/zookeys.993.53805.figure21" httpUri="https://binary.pensoft.net/fig/475454" pageId="0" pageNumber="47">21F</figureCitation>
|
||
, indicated by blue arrow).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.993.53805.figure21" httpUri="https://binary.pensoft.net/fig/475454" pageId="0" pageNumber="47" start="Figure 21" startId="F21">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 21.</emphasis>
|
||
<taxonomicName authorityName="Ruan, Douglas & Qiu" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius zhiweii" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="zhiweii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius zhiweii</emphasis>
|
||
</taxonomicName>
|
||
Ruan, Douglas & Qiu, sp. nov., holotype, male.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
|
||
habitus, dorsal view.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||
habitus, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||
habitus, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
|
||
aedeagus ventral view, arrow indicating apex and apical mesal callus of paramere
|
||
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
|
||
aedeagus, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="47">F</emphasis>
|
||
aedeagus, lateral view, arrow indicating apices of paramere and median lobe
|
||
<emphasis bold="true" pageId="0" pageNumber="47">G</emphasis>
|
||
tergite 7, dorsal view, hand drawing, arrow indicating posterior concavity.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="47" type="type material">
|
||
<paragraph pageId="0" pageNumber="47">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Holotype</emphasis>
|
||
. male (IZCAS, ex. LQCC), labels: 1)
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
|
||
, Long-chuan county, Hu-sa township (户撒), Mang-dong road, Gun-bang-jian-shan Mt., leg. Zhiwei Dong, 2018-VI-5 [label in Chinese]; 2)
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Holotype</emphasis>
|
||
,
|
||
<taxonomicName authorityName="Ruan, Douglas & Qiu" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius zhiweii" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="zhiweii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius zhiweii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., Des. Ruan et al., 2019.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.993.53805.figure22" httpUri="https://binary.pensoft.net/fig/475455" pageId="0" pageNumber="47" start="Figure 22" startId="F22">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 22.</emphasis>
|
||
<taxonomicName authorityName="Ruan, Douglas & Qiu" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius zhiweii" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="zhiweii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius zhiweii</emphasis>
|
||
</taxonomicName>
|
||
Ruan, Douglas & Qiu, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
|
||
pronotum and scutellar shield, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||
left posterior edge of pronotum, showing sublateral incision (arrow)
|
||
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||
head, frontal view
|
||
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
|
||
metacoxal plate, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
|
||
tarsal claws
|
||
<emphasis bold="true" pageId="0" pageNumber="47">F</emphasis>
|
||
male abdominal tergites IX-X, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="47">G</emphasis>
|
||
male abdominal sternite IX, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.993.53805.figure23" httpUri="https://binary.pensoft.net/fig/475457" pageId="0" pageNumber="47" start="Figure 23" startId="F23">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 23.</emphasis>
|
||
Shape of supraantennal carina of Chinese
|
||
<emphasis italics="true" pageId="0" pageNumber="47">
|
||
<taxonomicName authorityName="Fleutiaux" authorityYear="1931" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">Phorocardius</taxonomicName>
|
||
species
|
||
</emphasis>
|
||
. Arrows indicating supraantennal carina forked near eye.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
|
||
<taxonomicName lsidName="P. alterlineatus" pageId="0" pageNumber="47" rank="species" species="alterlineatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. alterlineatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||
<taxonomicName lsidName="P. flavistriolatus" pageId="0" pageNumber="47" rank="species" species="flavistriolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. flavistriolatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||
<taxonomicName lsidName="P. florentini" pageId="0" pageNumber="47" rank="species" species="florentini">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. florentini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
|
||
<taxonomicName lsidName="P. magnus" pageId="0" pageNumber="47" rank="species" species="magnus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. magnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
|
||
<taxonomicName lsidName="P. manuleatus" pageId="0" pageNumber="47" rank="species" species="manuleatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. manuleatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">F</emphasis>
|
||
<taxonomicName lsidName="P. minutus" pageId="0" pageNumber="47" rank="species" species="minutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. minutus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">G</emphasis>
|
||
<taxonomicName lsidName="P. rufiposterus" pageId="0" pageNumber="47" rank="species" species="rufiposterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. rufiposterus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">H</emphasis>
|
||
<taxonomicName lsidName="P. unguicularis" pageId="0" pageNumber="47" rank="species" species="unguicularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. unguicularis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">I</emphasis>
|
||
<taxonomicName lsidName="P. yunnanensis" pageId="0" pageNumber="47" rank="species" species="yunnanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">J</emphasis>
|
||
<taxonomicName lsidName="P. zhiweii" pageId="0" pageNumber="47" rank="species" species="zhiweii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. zhiweii</emphasis>
|
||
</taxonomicName>
|
||
Ruan, Douglas & Qiu, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">K</emphasis>
|
||
head, frontal-lateral view, arrows indicating forked supraantennal carina.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="47" type="remarks">
|
||
<paragraph pageId="0" pageNumber="47">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="47">This species inhabits middle elevations (around 1500 m) in Yunnan Prov., south China. This area is rainy, subtropical to tropical, with subtropical evergreen broad-leaf forest or tropical rain forest. This species is known only from the Oriental Region.</paragraph>
|
||
<caption doi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47" start="Figure 24" startId="F24">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 24.</emphasis>
|
||
Lateral view of prosternal process and mesosternum of Chinese
|
||
<taxonomicName authorityName="Fleutiaux" authorityYear="1931" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius</emphasis>
|
||
</taxonomicName>
|
||
species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i.e., on anteroventral angle of mesosternal fossa according to
|
||
<bibRefCitation DOI="https://doi.org/10.4039/n02-003" author="Douglas, HB" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="47" pagination="493 - 548" refId="B11" refString="Douglas, HB, 2003. Revision of the Cardiophorus (Coleoptera: Elateridae) species of eastern Canada and United States of America. The Canadian Entomologist 135: 493 - 548, DOI: https://doi.org/10.4039/n02-003" title="Revision of the Cardiophorus (Coleoptera: Elateridae) species of eastern Canada and United States of America." url="https://doi.org/10.4039/n02-003" volume="135" year="2003">Douglas 2003</bibRefCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
|
||
<taxonomicName lsidName="P. alterlineatus" pageId="0" pageNumber="47" rank="species" species="alterlineatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. alterlineatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||
<taxonomicName lsidName="P. flavistriolatus" pageId="0" pageNumber="47" rank="species" species="flavistriolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. flavistriolatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||
<taxonomicName lsidName="P. florentini" pageId="0" pageNumber="47" rank="species" species="florentini">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. florentini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
|
||
<taxonomicName lsidName="P. magnus" pageId="0" pageNumber="47" rank="species" species="magnus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. magnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
|
||
<taxonomicName lsidName="P. manuleatus" pageId="0" pageNumber="47" rank="species" species="manuleatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. manuleatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">F</emphasis>
|
||
<taxonomicName lsidName="P. minutus" pageId="0" pageNumber="47" rank="species" species="minutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. minutus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">G</emphasis>
|
||
<taxonomicName lsidName="P. rufiposterus" pageId="0" pageNumber="47" rank="species" species="rufiposterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. rufiposterus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">H</emphasis>
|
||
<taxonomicName lsidName="P. unguicularis" pageId="0" pageNumber="47" rank="species" species="unguicularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. unguicularis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">I</emphasis>
|
||
<taxonomicName lsidName="P. yunnanensis" pageId="0" pageNumber="47" rank="species" species="yunnanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">J</emphasis>
|
||
<taxonomicName lsidName="P. zhiweii" pageId="0" pageNumber="47" rank="species" species="zhiweii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. zhiweii</emphasis>
|
||
</taxonomicName>
|
||
Ruan, Douglas & Qiu, sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.993.53805.figure25" httpUri="https://binary.pensoft.net/fig/475459" pageId="0" pageNumber="47" start="Figure 25" startId="F25">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 25.</emphasis>
|
||
Ventral view of pro- and mesothorax of Chinese
|
||
<taxonomicName authorityName="Fleutiaux" authorityYear="1931" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius</emphasis>
|
||
</taxonomicName>
|
||
species. Showing shape of ventral surface of prosternal process (upper, yellow dashed line), mesepisternum (middle, green arrow and dashed line) and projections on posterior edge of mesosternum (lower, red arrow and solid line) middle, green arrows indicating anterior-mesal angle of mesepisternum.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
|
||
<taxonomicName lsidName="P. alterlineatus" pageId="0" pageNumber="47" rank="species" species="alterlineatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. alterlineatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||
<taxonomicName lsidName="P. flavistriolatus" pageId="0" pageNumber="47" rank="species" species="flavistriolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. flavistriolatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||
<taxonomicName lsidName="P. florentini" pageId="0" pageNumber="47" rank="species" species="florentini">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. florentini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
|
||
<taxonomicName lsidName="P. magnus" pageId="0" pageNumber="47" rank="species" species="magnus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. magnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
|
||
<taxonomicName lsidName="P. manuleatus" pageId="0" pageNumber="47" rank="species" species="manuleatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. manuleatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">F</emphasis>
|
||
<taxonomicName lsidName="P. minutus" pageId="0" pageNumber="47" rank="species" species="minutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. minutus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">G</emphasis>
|
||
<taxonomicName lsidName="P. rufiposterus" pageId="0" pageNumber="47" rank="species" species="rufiposterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. rufiposterus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">H</emphasis>
|
||
<taxonomicName lsidName="P. unguicularis" pageId="0" pageNumber="47" rank="species" species="unguicularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. unguicularis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">I</emphasis>
|
||
<taxonomicName lsidName="P. zhiweii" pageId="0" pageNumber="47" rank="species" species="zhiweii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. zhiweii</emphasis>
|
||
</taxonomicName>
|
||
Ruan, Douglas & Qiu, sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="47">
|
||
There are two undetermined female specimens (SZPT) from Xi-shuang-ban-na (西双版纳), Yunnan that have metallic blue elytra and a black pronotum. One male specimen (SZPT) from Ying-jiang (盈江), Yunnan also has metallic blue elytra and a black pronotum. Its aedeagus is extremely close to that of
|
||
<taxonomicName authorityName="Ruan, Douglas & Qiu" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius zhiweii" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="zhiweii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius zhiweii</emphasis>
|
||
</taxonomicName>
|
||
Ruan, Douglas & Qiu, sp. nov. However, it is still not entirely clear whether these are conspecific. More specimens should be studied before a reliable determination can be made.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.993.53805.figure26" httpUri="https://binary.pensoft.net/fig/475460" pageId="0" pageNumber="47" start="Figure 26" startId="F26">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 26.</emphasis>
|
||
Female abdominal ventrite V in Chinese
|
||
<taxonomicName authorityName="Fleutiaux" authorityYear="1931" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius</emphasis>
|
||
</taxonomicName>
|
||
species (ventral view).
|
||
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
|
||
<taxonomicName lsidName="P. alterlineatus" pageId="0" pageNumber="47" rank="species" species="alterlineatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. alterlineatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
|
||
<taxonomicName lsidName="P. flavistriolatus" pageId="0" pageNumber="47" rank="species" species="flavistriolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. flavistriolatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
|
||
<taxonomicName lsidName="P. florentini" pageId="0" pageNumber="47" rank="species" species="florentini">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. florentini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
|
||
<taxonomicName lsidName="P. magnus" pageId="0" pageNumber="47" rank="species" species="magnus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. magnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
|
||
<taxonomicName lsidName="P. manuleatus" pageId="0" pageNumber="47" rank="species" species="manuleatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. manuleatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="47">F</emphasis>
|
||
<taxonomicName lsidName="P. rufiposterus" pageId="0" pageNumber="47" rank="species" species="rufiposterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. rufiposterus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">G</emphasis>
|
||
<taxonomicName lsidName="P. unguicularis" pageId="0" pageNumber="47" rank="species" species="unguicularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. unguicularis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="47">H</emphasis>
|
||
<taxonomicName lsidName="P. yunnanensis" pageId="0" pageNumber="47" rank="species" species="yunnanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov, arrows showing incisions near apex.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.993.53805.figure27" httpUri="https://binary.pensoft.net/fig/475461" pageId="0" pageNumber="47" start="Figure 27" startId="F27">
|
||
<paragraph pageId="0" pageNumber="47">
|
||
<emphasis bold="true" pageId="0" pageNumber="47">Figure 27.</emphasis>
|
||
A distribution map of Chinese
|
||
<taxonomicName authorityName="Fleutiaux" authorityYear="1931" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">1</emphasis>
|
||
<taxonomicName lsidName="P. alterlineatus" pageId="0" pageNumber="47" rank="species" species="alterlineatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. alterlineatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">2</emphasis>
|
||
<taxonomicName lsidName="P. flavistriolatus" pageId="0" pageNumber="47" rank="species" species="flavistriolatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. flavistriolatus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">3</emphasis>
|
||
<taxonomicName lsidName="P. florentini" pageId="0" pageNumber="47" rank="species" species="florentini">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. florentini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="47">4</emphasis>
|
||
<taxonomicName lsidName="P. magnus" pageId="0" pageNumber="47" rank="species" species="magnus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. magnus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="47">5</emphasis>
|
||
<taxonomicName lsidName="P. manuleatus" pageId="0" pageNumber="47" rank="species" species="manuleatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. manuleatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="47">6</emphasis>
|
||
<taxonomicName lsidName="P. minutus" pageId="0" pageNumber="47" rank="species" species="minutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. minutus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">7</emphasis>
|
||
<taxonomicName lsidName="P. rufiposterus" pageId="0" pageNumber="47" rank="species" species="rufiposterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. rufiposterus</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">8</emphasis>
|
||
<taxonomicName lsidName="P. unguicularis" pageId="0" pageNumber="47" rank="species" species="unguicularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. unguicularis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="47">9</emphasis>
|
||
<taxonomicName lsidName="P. yanagiharae" pageId="0" pageNumber="47" rank="species" species="yanagiharae">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. yanagiharae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="47">10</emphasis>
|
||
<taxonomicName lsidName="P. yunnanensis" pageId="0" pageNumber="47" rank="species" species="yunnanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
Ruan & Douglas, sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="47">11</emphasis>
|
||
<taxonomicName lsidName="P. zhiweii" pageId="0" pageNumber="47" rank="species" species="zhiweii">
|
||
<emphasis italics="true" pageId="0" pageNumber="47">P. zhiweii</emphasis>
|
||
</taxonomicName>
|
||
Ruan, Douglas & Qiu, sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |