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<document id="0033C322034BBDD2517A2E75713EF2E0" ID-CLB-Dataset="8243" ID-DOI="10.11646/zootaxa.4923.1.1" ID-GBIF-Dataset="0a027439-c601-4533-bd03-f0e5d4442801" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496757" ID-ZooBank="41D057A7-F855-47C2-A598-8000CFD83A5B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612332986149" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2021" docId="1A3236786206223609D9FAC9FA836ED0" docLanguage="en" docName="zootaxa.4923.1.1.pdf" docOrigin="Zootaxa 4923 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lepanus williamsi Gunter &amp; Weir 2021, new species" docType="treatment" docVersion="10" lastPageNumber="43" masterDocId="E60B4E00622E221C094EFFBDFFC26960" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species" masterLastPageNumber="66" masterPageNumber="1" pageNumber="41" updateTime="1698926692322" updateUser="ExternalLinkService">
<mods:mods id="46D181A85C79EC3C925E91A1B8DE60FA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E20AEFEB0931676A8AC5B5682A86E4E8">
<mods:title id="0A2D7DD51CAB18012BC0BC4D14C4D043">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species</mods:title>
</mods:titleInfo>
<mods:name id="F17DA11130CE1193B21B39DCE0524B1A" type="personal">
<mods:role id="6F6BB6A6F9D9AC2B29D7DEAEC2CE7814">
<mods:roleTerm id="D10FBBDB41A004D7817C6E88B56A9493">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="90D7787E4489BE52C5E3BCE3F1602D05">Gunter, Nicole L.</mods:namePart>
<mods:nameIdentifier id="2ECDFB39D38D2B01CA6FD68BF11491E3" type="ORCID">0000-0002-8355-0862</mods:nameIdentifier>
<mods:nameIdentifier id="81E7169AEFEE21F654D0F1341A281605" type="email">ngunter@cmnh.org</mods:nameIdentifier>
</mods:name>
<mods:name id="A23FE3D4CE67A0B1DEC93426C1F6433A" type="personal">
<mods:role id="FB2C2E9150FC03C92397EF62332E1E69">
<mods:roleTerm id="2B9113380FC510B1F9E960E3B88DC892">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19EC0779FA4443506E94B9DC75F0467C">Weir, Thomas A.</mods:namePart>
<mods:nameIdentifier id="08E8FB2C742CD214AF67B0160631FCE9" type="ORCID">0000-0002-8355-0862</mods:nameIdentifier>
<mods:nameIdentifier id="887D3DA791ABA3BC18D089D269947B97" type="email">ngunter@cmnh.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="D6F65A2CC08F020CA4373CC8CDB28574">text</mods:typeOfResource>
<mods:relatedItem id="A44A2C1CBCF02D0B083CEB341D2CE0B7" type="host">
<mods:titleInfo id="31B801FF292012981740C40212895DDC">
<mods:title id="768DC495820E566D4BE50D8DBF12B09D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4A71BD7E727786E0FE6AEC7F13972A7E">
<mods:date id="94C5C739B869D2EC71174ACAFE562270">2021</mods:date>
<mods:detail id="27CF6673EA10D664BD181CC31E86D9AE" type="pubDate">
<mods:number id="3DD48AF367D11957AC7824DEE33EF9DF">2021-02-03</mods:number>
</mods:detail>
<mods:detail id="854D7CA2B316AFEEB921409128AB5D00" type="volume">
<mods:number id="48DEC76932620076BC143B510BDCABCF">4923</mods:number>
</mods:detail>
<mods:detail id="86C3CEF72489153F53351ECABA8D8391" type="issue">
<mods:number id="BACE119C2628087FA761F9AA79BB4086">1</mods:number>
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<mods:start id="179EFC2993A62AC7C22FFFC45D975C5E">1</mods:start>
<mods:end id="F9AC2D8835EAC49EBD0BAB7DE692D77C">66</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="C4440C31C62948CA979885088E3617F9">journal article</mods:classification>
<mods:identifier id="8BA367E24CC8B90B84626D890143E5A4" type="CLB-Dataset">8243</mods:identifier>
<mods:identifier id="0E5FF3DEBAEF9360A1D73785974790FC" type="DOI">10.11646/zootaxa.4923.1.1</mods:identifier>
<mods:identifier id="072E9BF27F82C2BD51E53F9C0DD7045C" type="GBIF-Dataset">0a027439-c601-4533-bd03-f0e5d4442801</mods:identifier>
<mods:identifier id="AE93C3993429834CF41F57EE8D5F5C4A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="20A8AEE738BF5A93716B7BC999789A6E" type="Zenodo-Dep">4496757</mods:identifier>
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</mods:mods>
<treatment id="1A3236786206223609D9FAC9FA836ED0" ID-DOI="http://doi.org/10.5281/zenodo.4537968" ID-GBIF-Taxon="177324116" ID-Zenodo-Dep="4537968" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0" httpUri="http://treatment.plazi.org/id/1A3236786206223609D9FAC9FA836ED0" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="DA81D4E56206223409D9FAC9FD146CED" box="[151,726,1395,1422]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="9224876E6206223409D9FAC9FD146CED" blockId="40.[151,726,1395,1458]" box="[151,726,1395,1422]" pageId="40" pageNumber="41">
<heading id="C96C30026206223409D9FAC9FD146CED" bold="true" box="[151,726,1395,1422]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="4">
<emphasis id="A0EF5B7C6206223409D9FAC9FD146CED" bold="true" box="[151,726,1395,1422]" pageId="40" pageNumber="41">
<taxonomicName id="559BFCED6206223409D9FAC9FDF86CED" ID-CoL="7WWVW" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[151,570,1395,1422]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
<emphasis id="A0EF5B7C6206223409D9FAC9FEB66CEE" bold="true" box="[151,372,1395,1422]" italics="true" pageId="40" pageNumber="41">Lepanus williamsi</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="BBDCE607620622340B09FAC9FD146CED" box="[583,726,1395,1422]" pageId="40" pageNumber="41" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E56206223409D9FA25FE026CD2" box="[151,448,1432,1458]" pageId="40" pageNumber="41" type="description">
<paragraph id="9224876E6206223409D9FA25FE026CD2" blockId="40.[151,726,1395,1458]" box="[151,448,1432,1458]" pageId="40" pageNumber="41">
(
<figureCitation id="0AA09BEB6206223409D1FA25FEC16CD2" box="[159,259,1432,1458]" captionStart="FIGURE 9" captionStartId="40.[151,250,1253,1277]" captionTargetBox="[159,1430,568,1216]" captionTargetId="figure-219@40.[151,1436,561,1230]" captionTargetPageId="40" captionText="FIGURE 9. Dorsal habitus of species within the Lepanus storeyi species group. A, Lepanus meierae; B, Lepanus storeyi; C, Lepanus williamsi. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4496777" httpUri="https://zenodo.org/record/4496777/files/figure.png" pageId="40" pageNumber="41">Figs. 9C</figureCitation>
;
<figureCitation id="0AA09BEB620622340840FA25FEFC6CD2" box="[270,318,1432,1458]" captionStart="FIGURE 10" captionStartId="41.[151,250,1239,1263]" captionTargetBox="[158,1430,191,1207]" captionTargetId="figure-19@41.[151,1436,181,1215]" captionTargetPageId="41" captionText="FIGURE 10. Pygidial configuration of species within the Lepanus storeyi species group. A, Lepanus meierae male; B, Lepanus meierae female; C, Lepanus storeyi male; D, Lepanus storeyi female; E, Lepanus williamsi male. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496779" httpUri="https://zenodo.org/record/4496779/files/figure.png" pageId="40" pageNumber="41">10E</figureCitation>
;
<figureCitation id="0AA09BEB620622340804FA25FEB86CD2" box="[330,378,1432,1458]" captionStart="FIGURE 11" captionStartId="42.[151,250,1439,1463]" captionTargetBox="[159,1423,189,1407]" captionTargetId="figure-26@42.[151,1436,181,1415]" captionTargetPageId="42" captionText="FIGURE 11. Left, dorsal and right view of aedeagi of species within the Lepanus storeyi species group. A, Lepanus meierae; B, Lepanus storeyi; C, Lepanus williamsi. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496781" httpUri="https://zenodo.org/record/4496781/files/figure.png" pageId="40" pageNumber="41">11C</figureCitation>
;
<figureCitation id="0AA09BEB6206223408C8FA25FE7A6CD2" box="[390,440,1432,1458]" captionStart="FIGURE 12" captionStartId="43.[151,250,1221,1245]" captionTargetBox="[155,1427,186,1191]" captionTargetId="figure-19@43.[151,1436,181,1197]" captionTargetPageId="43" captionText="FIGURE 12. A, Map of known records of Lepanus meierae, Lepanus storeyi, and Lepanus williamsi, B, predicted distribution of Lepanus meierae." figureDoi="http://doi.org/10.5281/zenodo.4496783" httpUri="https://zenodo.org/record/4496783/files/figure.png" pageId="40" pageNumber="41">12A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E56206223409D9FA5DFC276ECA" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="9224876E6206223409D9FA5DFC276ECA" blockId="40.[151,1437,1503,1963]" pageId="40" pageNumber="41">
<emphasis id="A0EF5B7C6206223409D9FA5DFEE16C99" bold="true" box="[151,291,1503,1530]" pageId="40" pageNumber="41">Type series.</emphasis>
<materialsCitation id="22F38D33620622340865FA5DFE7A6F22" ID-GBIF-Occurrence="3034608701" collectingDate="1970-09-03" collectingDateMax="1970-09-04" collectingDateMin="1970-09-03" collectionCode="ANIC" collectorName="K. R. Pullen &amp; Det T. A. Weir" latitude="-31.466667" location="Sea Acres Reserve" longLatPrecision="20" longitude="152.93333" municipality="Port Macquarie" pageId="40" pageNumber="41" specimenCount="1" typeStatus="holotype">
<emphasis id="A0EF5B7C620622340865FA5DFE7C6C9A" bold="true" box="[299,446,1504,1530]" pageId="40" pageNumber="41">
<typeStatus id="4D2039CC620622340865FA5DFE5E6C9A" box="[299,412,1504,1530]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
: ♂
</emphasis>
“
<location id="9744D1B5620622340881FA5DFD646C9A" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0:9744D1B5620622340881FA5DFD646C9A" box="[463,678,1504,1530]" latitude="-31.466667" longLatPrecision="20" longitude="152.93333" municipality="Port Macquarie" name="Sea Acres Reserve" pageId="40" pageNumber="41">Sea Acres Reserve</location>
,
<collectingMunicipality id="72401D14620622340BFCFA5DFCA26C9A" box="[690,864,1504,1530]" pageId="40" pageNumber="41">Port Macquarie</collectingMunicipality>
NSW,
<date id="E625A1AE620622340AFFFA5DFB956C9A" box="[945,1111,1504,1530]" pageId="40" pageNumber="41" value="1970-09-03" valueMax="1970-09-04" valueMin="1970-09-03">
<collectingDate id="F6615846620622340AFFFA5DFB956C9A" box="[945,1111,1504,1530]" pageId="40" pageNumber="41" value="1970-09-03" valueMax="1970-09-04" valueMin="1970-09-03">3-4 Sept. 1970</collectingDate>
</date>
<collectorName id="3F6EE2B8620622340D10FA5DFB276C9A" box="[1118,1253,1504,1530]" pageId="40" pageNumber="41">K.R. Pullen</collectorName>
” / “
<collectionCode id="F48A1FAB620622340C56FA5DFA9E6C9A" box="[1304,1372,1504,1530]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="40" pageNumber="41">ANIC</collectionCode>
database 25 054070” / “
<taxonomicName id="559BFCED62062234083CF9B9FDDC6F7E" authority="NSW" authorityName="NSW" box="[370,542,1540,1566]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Lepanus NSW</taxonomicName>
3,
<collectorName id="3F6EE2B8620622340B7BF9B9FD136F7E" box="[565,721,1540,1566]" pageId="40" pageNumber="41">Det T.A. Weir</collectorName>
2012” / “
<collectionCode id="F48A1FAB620622340A72F9B9FC416F7E" box="[828,899,1540,1566]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="40" pageNumber="41">ANIC</collectionCode>
Image” (in
<collectionCode id="F48A1FAB620622340D46F9B9FB8C6F7E" box="[1032,1102,1540,1566]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="40" pageNumber="41">ANIC</collectionCode>
) [
<geoCoordinate id="F7AFE1A9620622340D2DF9B9FB2E6F7E" box="[1123,1260,1540,1566]" degrees="31" direction="south" minutes="28" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="-31.466667">31°28’00”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A9620622340DB6F9BEFA526F7E" box="[1272,1424,1539,1566]" degrees="152" direction="east" minutes="56" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="152.93333">152°56’00”E</geoCoordinate>
], (missing one metatarsus)
</materialsCitation>
;
<materialsCitation id="22F38D3362062234088BF995FAA46FEA" ID-GBIF-Occurrence="3034608729" collectingDate="1999-01-01" collectionCode="ANIC, GWC, QM" collectorName="G. Williams &amp; T. Williams" latitude="-31.466667" location="Sea Acres Nature Reserve" longLatPrecision="20" longitude="152.93333" municipality="Port Macquarie" pageId="40" pageNumber="41" specimenCount="15" specimenCount-female="13" typeStatus="paratype">
<emphasis id="A0EF5B7C62062234088BF995FC6A6F21" bold="true" box="[453,936,1575,1604]" pageId="40" pageNumber="41">
<typeStatus id="4D2039CC62062234088BF995FDFF6F22" box="[453,573,1576,1602]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
(14 ♂,
<specimenCount id="849D4CE7620622340BD4F995FD046F24" box="[666,710,1576,1604]" pageId="40" pageNumber="41" type="female">8 ♀</specimenCount>
, 1 undetermined):
</emphasis>
<location id="9744D1B5620622340AFCF995FB1D6F22" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0:9744D1B5620622340AFCF995FB1D6F22" box="[946,1247,1576,1602]" latitude="-31.466667" longLatPrecision="20" longitude="152.93333" municipality="Port Macquarie" name="Sea Acres Nature Reserve" pageId="40" pageNumber="41">Sea Acres Nature Reserve</location>
,
<collectingMunicipality id="72401D14620622340DA3F995FA5E6F22" box="[1261,1436,1576,1602]" pageId="40" pageNumber="41">Port Macquarie</collectingMunicipality>
[
<geoCoordinate id="F7AFE1A96206223409D1F9F1FEE56F06" box="[159,295,1612,1638]" degrees="31" direction="south" minutes="28" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="-31.466667">31°28’00”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A962062234087DF9F6FE0E6F06" box="[307,460,1611,1638]" degrees="152" direction="east" minutes="56" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="152.93333">152°56’00”E</geoCoordinate>
],
<date id="E625A1AE620622340891F9F1FDFF6F06" box="[479,573,1612,1638]" pageId="40" pageNumber="41" value="1999-01-01">
<collectingDate id="F6615846620622340891F9F1FDFF6F06" box="[479,573,1612,1638]" pageId="40" pageNumber="41" value="1999-01-01">1.i.1999</collectingDate>
</date>
,
<collectorName id="3F6EE2B8620622340B07F9F1FD136F06" box="[585,721,1612,1638]" pageId="40" pageNumber="41">G. Williams</collectorName>
&amp;
<collectorName id="3F6EE2B8620622340BBAF9F1FCB56F06" box="[756,887,1612,1638]" pageId="40" pageNumber="41">T. Williams</collectorName>
(25-054073
<specimenCount id="849D4CE7620622340D49F9F1FB6C6F06" box="[1031,1198,1612,1638]" pageId="40" pageNumber="41" type="female">2 specimens ♀</specimenCount>
in
<collectionCode id="F48A1FAB620622340D9EF9F1FAD56F06" box="[1232,1303,1612,1638]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="40" pageNumber="41">ANIC</collectionCode>
, 25-058084 6 ♂,
<specimenCount id="849D4CE762062234099DF9CDFEBE6FEA" box="[211,380,1648,1674]" pageId="40" pageNumber="41" type="female">2 specimens ♀</specimenCount>
,
<specimenCount id="849D4CE76206223408C5F9CDFDCE6FEA" box="[395,524,1648,1674]" pageId="40" pageNumber="41" type="generic">1 specimen</specimenCount>
with sex undetermined in
<collectionCode id="F48A1FAB620622340A70F9CDFCBD6FEA" box="[830,895,1648,1674]" pageId="40" pageNumber="41">GWC</collectionCode>
and
<specimenCount id="849D4CE7620622340AF7F9CDFBF86FEA" box="[953,1082,1648,1674]" pageId="40" pageNumber="41" type="generic">1 specimen</specimenCount>
♂,
<specimenCount id="849D4CE7620622340D29F9CDFAC46FEA" box="[1127,1286,1648,1674]" pageId="40" pageNumber="41" type="female">1 specimen ♀</specimenCount>
in
<collectionCode id="F48A1FAB620622340C63F9CDFA9D6FEA" box="[1325,1375,1648,1674]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="40" pageNumber="41" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22F38D33620622340C3AF9CDFF276FB2" ID-GBIF-Occurrence="3034608709" collectingDate="1999-11-25" collectingDateMax="1999-11-27" collectingDateMin="1999-11-25" collectionCode="GWC" latitude="-31.466667" location="Sea Acres Nature Reserve" longLatPrecision="20" longitude="152.93333" municipality="Port Macquarie" pageId="40" pageNumber="41" specimenCount="3" typeStatus="paratype">
<location id="9744D1B5620622340C3AF9CDFE506FCE" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0:9744D1B5620622340C3AF9CDFE506FCE" latitude="-31.466667" longLatPrecision="20" longitude="152.93333" municipality="Port Macquarie" name="Sea Acres Nature Reserve" pageId="40" pageNumber="41">Sea Acres Nature Reserve</location>
,
<collectingMunicipality id="72401D146206223408D1F929FD8C6FCE" box="[415,590,1684,1710]" pageId="40" pageNumber="41">Port Macquarie</collectingMunicipality>
[
<geoCoordinate id="F7AFE1A9620622340B10F929FD256FCE" box="[606,743,1684,1710]" degrees="31" direction="south" minutes="28" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="-31.466667">31°28’00”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A9620622340BBAF92EFC4E6FCE" box="[756,908,1683,1710]" degrees="152" direction="east" minutes="56" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="152.93333">152°56’00”E</geoCoordinate>
],
<date id="E625A1AE620622340AEEF929FB806FCF" box="[928,1090,1684,1711]" pageId="40" pageNumber="41" value="1999-11-25" valueMax="1999-11-27" valueMin="1999-11-25">
<collectingDate id="F6615846620622340AEEF929FB806FCF" box="[928,1090,1684,1711]" pageId="40" pageNumber="41" value="1999-11-25" valueMax="1999-11-27" valueMin="1999-11-25">25–27.xi.1999</collectingDate>
</date>
(25-058083
<specimenCount id="849D4CE7620622340D98F929FAA36FCE" box="[1238,1377,1684,1710]" pageId="40" pageNumber="41" type="generic">3 specimens</specimenCount>
♂ in
<collectionCode id="F48A1FAB6206223409D9F905FF1F6FB2" box="[151,221,1720,1746]" pageId="40" pageNumber="41">GWC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22F38D336206223409BEF905FC946F96" ID-GBIF-Occurrence="3034608377" collectingDate="1998-12-30" collectionCode="ANIC" collectorName="G. Williams" county="Laurieton" elevation="400" latitude="-31.65" location="Dooragan National Park" longLatPrecision="20" longitude="152.76666" municipality="Brother Mountain" pageId="40" pageNumber="41" specimenCount="1" typeStatus="paratype">
<location id="9744D1B56206223409BEF905FDC26FB2" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0:9744D1B56206223409BEF905FDC26FB2" box="[240,512,1720,1746]" county="Laurieton" latitude="-31.65" longLatPrecision="20" longitude="152.76666" municipality="Brother Mountain" name="Dooragan National Park" pageId="40" pageNumber="41">Dooragan National Park</location>
, eastern slope, N
<collectingMunicipality id="72401D14620622340B8CF905FC4C6FB2" box="[706,910,1720,1746]" pageId="40" pageNumber="41">Brother Mountain</collectingMunicipality>
,
<collectingCounty id="7B45FFE2620622340AD6F905FBC76FB2" box="[920,1029,1720,1746]" pageId="40" pageNumber="41">Laurieton</collectingCounty>
[
<geoCoordinate id="F7AFE1A9620622340D5CF905FB596FB2" box="[1042,1179,1720,1746]" degrees="31" direction="south" minutes="39" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="-31.65">31°39’00”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A9620622340DEBF90AFAFF6FB2" box="[1189,1341,1719,1746]" degrees="152" direction="east" minutes="46" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="152.76666">152°46’00”E</geoCoordinate>
],
<quantity id="55632A8B620622340C01F905FA556FB2" box="[1359,1431,1720,1746]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="40" pageNumber="41" unit="m" value="400.0">
<elevation id="19B6605D620622340C01F905FA556FB2" box="[1359,1431,1720,1746]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="40" pageNumber="41" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="E625A1AE6206223409D9F961FEDB6F96" box="[151,281,1756,1782]" pageId="40" pageNumber="41" value="1998-12-30">
<collectingDate id="F66158466206223409D9F961FEDB6F96" box="[151,281,1756,1782]" pageId="40" pageNumber="41" value="1998-12-30">30.xii.1998</collectingDate>
</date>
,
<collectorName id="3F6EE2B8620622340868F961FE6D6F96" box="[294,431,1756,1782]" pageId="40" pageNumber="41">G. Williams</collectorName>
(25-054072
<specimenCount id="849D4CE7620622340B0CF961FD016F96" box="[578,707,1756,1782]" pageId="40" pageNumber="41" type="generic">1 specimen</specimenCount>
♂ in
<collectionCode id="F48A1FAB620622340A4BF961FC8C6F96" box="[773,846,1756,1782]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="40" pageNumber="41">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22F38D33620622340A2DF961FC366E7A" ID-GBIF-Occurrence="3034609228" collectingDate="1998-01-18" collectingDateMax="1998-01-20" collectingDateMin="1998-01-18" collectionCode="GWC" collectorName="G. Williams" country="Gibraltar" latitude="-31.65" location="Mount" longLatPrecision="20" longitude="152.43333" municipality="Comboyne Plateau" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<location id="9744D1B5620622340A2DF961FC6C6F96" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0:9744D1B5620622340A2DF961FC6C6F96" box="[867,942,1756,1782]" country="Gibraltar" latitude="-31.65" longLatPrecision="20" longitude="152.43333" municipality="Comboyne Plateau" name="Mount" pageId="40" pageNumber="41">Mount</location>
<collectingCountry id="EA8CC7FE620622340AF8F961FBDF6F96" box="[950,1053,1756,1782]" name="Gibraltar" pageId="40" pageNumber="41">Gibraltar</collectingCountry>
,
<collectingMunicipality id="72401D14620622340D67F961FB3C6F96" box="[1065,1278,1756,1782]" pageId="40" pageNumber="41">Comboyne Plateau</collectingMunicipality>
[
<geoCoordinate id="F7AFE1A9620622340C41F961FA5A6F96" box="[1295,1432,1756,1782]" degrees="31" direction="south" minutes="39" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="-31.65">31°39’00”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A96206223409D9F942FEED6E7A" box="[151,303,1791,1818]" degrees="152" direction="east" minutes="26" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="152.43333">152°26’00”E</geoCoordinate>
],
<date id="E625A1AE62062234080FF8BDFE156E7A" box="[321,471,1792,1818]" pageId="40" pageNumber="41" value="1998-01-18" valueMax="1998-01-20" valueMin="1998-01-18">
<collectingDate id="F661584662062234080FF8BDFE156E7A" box="[321,471,1792,1818]" pageId="40" pageNumber="41" value="1998-01-18" valueMax="1998-01-20" valueMin="1998-01-18">18–20.i.1998</collectingDate>
</date>
,
<collectorName id="3F6EE2B86206223408ACF8BDFDAA6E7A" box="[482,616,1792,1818]" pageId="40" pageNumber="41">G. Williams</collectorName>
(25–65523
<specimenCount id="849D4CE7620622340BA2F8BDFC466E7A" box="[748,900,1792,1818]" pageId="40" pageNumber="41" type="female">1 specimen ♀</specimenCount>
in
<collectionCode id="F48A1FAB620622340AEBF8BDFC296E7A" box="[933,1003,1792,1818]" pageId="40" pageNumber="41">GWC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22F38D33620622340AB3F8BDFD2F6E02" ID-GBIF-Occurrence="3034608881" collectingDate="1980-11-17" collectingDateMax="1980-11-20" collectingDateMin="1980-11-17" collectionCode="ANIC, GWC" collectorName="G. Williams &amp; T. Williams" country="Gibraltar" latitude="-31.716667" location="Vale Road" longLatPrecision="20" longitude="152.5" municipality="Lansdowne State Forest" pageId="40" pageNumber="41" specimenCount="3" specimenCount-female="2" typeStatus="paratype">
Langley
<location id="9744D1B5620622340D10F8BDFB166E7A" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0:9744D1B5620622340D10F8BDFB166E7A" box="[1118,1236,1792,1818]" country="Gibraltar" latitude="-31.716667" longLatPrecision="20" longitude="152.5" municipality="Lansdowne State Forest" name="Vale Road" pageId="40" pageNumber="41">Vale Road</location>
,
<collectingMunicipality id="72401D14620622340D93F8BDFF1F6E5E" pageId="40" pageNumber="41">Lansdowne State Forest</collectingMunicipality>
[
<geoCoordinate id="F7AFE1A96206223409A2F899FEB76E5E" box="[236,373,1828,1854]" degrees="31" direction="south" minutes="43" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="-31.716667">31°43’00”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A96206223408CEF89EFDDB6E5E" box="[384,537,1827,1854]" degrees="152" direction="east" minutes="30" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="152.5">152°30’00”E</geoCoordinate>
],
<date id="E625A1AE620622340B62F899FD126E5E" box="[556,720,1828,1854]" pageId="40" pageNumber="41" value="1980-11-17" valueMax="1980-11-20" valueMin="1980-11-17">
<collectingDate id="F6615846620622340B62F899FD126E5E" box="[556,720,1828,1854]" pageId="40" pageNumber="41" value="1980-11-17" valueMax="1980-11-20" valueMin="1980-11-17">17–20.xi.1980</collectingDate>
</date>
,
<collectorName id="3F6EE2B8620622340B92F899FCA66E5E" box="[732,868,1828,1854]" pageId="40" pageNumber="41">G. Williams</collectorName>
&amp;
<collectorName id="3F6EE2B8620622340AC9F899FBC86E5E" box="[903,1034,1828,1854]" pageId="40" pageNumber="41">T. Williams</collectorName>
(25-054071
<specimenCount id="849D4CE7620622340DD5F899FAD86E5E" box="[1179,1306,1828,1854]" pageId="40" pageNumber="41" type="generic">1 specimen</specimenCount>
♂, 1 specimen
<specimenCount id="849D4CE7620622340981F8F5FF266E02" box="[207,228,1864,1890]" pageId="40" pageNumber="41" type="female">♀</specimenCount>
in
<collectionCode id="F48A1FAB620622340846F8F5FE8D6E02" box="[264,335,1864,1890]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="40" pageNumber="41">ANIC</collectionCode>
, 25-058087
<specimenCount id="849D4CE7620622340893F8F5FDB86E02" box="[477,634,1864,1890]" pageId="40" pageNumber="41" type="female">1 specimen ♀</specimenCount>
in
<collectionCode id="F48A1FAB620622340BD1F8F5FD276E02" box="[671,741,1864,1890]" pageId="40" pageNumber="41">GWC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22F38D33620622340BB4F8F5FCA66EE6" ID-GBIF-Occurrence="3034608950" collectingDate="1980-10-25" collectionCode="GWC" collectorName="G. Williams &amp; T. Williams" country="Gibraltar" latitude="-31.866667" location="Lorien" longLatPrecision="20" longitude="152.48334" municipality="Lansdowne" pageId="40" pageNumber="41" specimenCount="1" typeStatus="paratype">
<location id="9744D1B5620622340BB4F8F5FC8A6E02" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0:9744D1B5620622340BB4F8F5FC8A6E02" box="[762,840,1864,1890]" country="Gibraltar" latitude="-31.866667" longLatPrecision="20" longitude="152.48334" municipality="Lansdowne" name="Lorien" pageId="40" pageNumber="41">Lorien</location>
,
<collectingMunicipality id="72401D14620622340A1DF8F5FC166E02" box="[851,980,1864,1890]" pageId="40" pageNumber="41">Lansdowne</collectingMunicipality>
[
<geoCoordinate id="F7AFE1A9620622340AADF8F5FBAF6E03" box="[995,1133,1864,1891]" degrees="31" direction="south" minutes="52" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="-31.866667">31°52’00”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A9620622340D37F8FAFAD06E02" box="[1145,1298,1863,1890]" degrees="152" direction="east" minutes="29" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="152.48334">152°29’00”E</geoCoordinate>
],
<date id="E625A1AE620622340C6BF8F5FA5A6E03" box="[1317,1432,1864,1891]" pageId="40" pageNumber="41" value="1980-10-25">
<collectingDate id="F6615846620622340C6BF8F5FA5A6E03" box="[1317,1432,1864,1891]" pageId="40" pageNumber="41" value="1980-10-25">25.x.1980</collectingDate>
</date>
,
<collectorName id="3F6EE2B86206223409D9F8D1FEDD6EE6" box="[151,287,1900,1926]" pageId="40" pageNumber="41">G. Williams</collectorName>
&amp;
<collectorName id="3F6EE2B862062234080DF8D1FE076EE6" box="[323,453,1900,1926]" pageId="40" pageNumber="41">T. Williams</collectorName>
(25-058086
<specimenCount id="849D4CE7620622340B18F8D1FD176EE6" box="[598,725,1900,1926]" pageId="40" pageNumber="41" type="generic">1 specimen</specimenCount>
♂ in
<collectionCode id="F48A1FAB620622340A58F8D1FC9E6EE6" box="[790,860,1900,1926]" pageId="40" pageNumber="41">GWC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22F38D33620622340A21F8D1FC236ECA" ID-GBIF-Occurrence="3034608897" collectingDate="1979-10-27" collectionCode="ANIC" collectorName="G. Williams &amp; T. Williams" country="Gibraltar" latitude="-34.066666" location="Royal National Park" longLatPrecision="19" longitude="151.06667" pageId="40" pageNumber="41" specimenCount="1" typeStatus="paratype">
<location id="9744D1B5620622340A21F8D1FB966EE6" LSID="urn:lsid:plazi:treatment:1A3236786206223609D9FAC9FA836ED0:9744D1B5620622340A21F8D1FB966EE6" box="[879,1108,1900,1926]" country="Gibraltar" latitude="-34.066666" longLatPrecision="19" longitude="151.06667" name="Royal National Park" pageId="40" pageNumber="41">Royal National Park</location>
[
<geoCoordinate id="F7AFE1A9620622340D2DF8D1FB2E6EE6" box="[1123,1260,1900,1926]" degrees="34" direction="south" minutes="04" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="-34.066666">34°04’00”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A9620622340DB6F8D6FA536EE6" box="[1272,1425,1899,1926]" degrees="151" direction="east" minutes="04" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="00" value="151.06667">151°04’00”E</geoCoordinate>
],
<date id="E625A1AE6206223409D9F82DFECB6ECA" box="[151,265,1936,1962]" pageId="40" pageNumber="41" value="1979-10-27">
<collectingDate id="F66158466206223409D9F82DFECB6ECA" box="[151,265,1936,1962]" pageId="40" pageNumber="41" value="1979-10-27">27.x.1979</collectingDate>
</date>
,
<collectorName id="3F6EE2B862062234085BF82DFE5F6ECA" box="[277,413,1936,1962]" pageId="40" pageNumber="41">G. Williams</collectorName>
&amp;
<collectorName id="3F6EE2B862062234088EF82DFD806ECA" box="[448,578,1936,1962]" pageId="40" pageNumber="41">T. Williams</collectorName>
(25-058085
<specimenCount id="849D4CE7620622340B9CF82DFC936ECA" box="[722,849,1936,1962]" pageId="40" pageNumber="41" type="generic">1 specimen</specimenCount>
♂ in
<collectionCode id="F48A1FAB620622340AC1F82DFC1B6ECA" box="[911,985,1936,1962]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="40" pageNumber="41">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="C6E4D7E66207223509D9FB6AFE4D6C57" ID-DOI="http://doi.org/10.5281/zenodo.4496779" ID-Zenodo-Dep="4496779" httpUri="https://zenodo.org/record/4496779/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,1239,1263]" targetBox="[158,1430,191,1207]" targetPageId="41">
<paragraph id="9224876E6207223509D9FB6AFE4D6C57" blockId="41.[151,1436,1239,1335]" pageId="41" pageNumber="42">
<emphasis id="A0EF5B7C6207223509D9FB6AFEDD6D8F" bold="true" box="[151,287,1239,1263]" pageId="41" pageNumber="42">FIGURE 10.</emphasis>
Pygidial configuration of species within the
<taxonomicName id="559BFCED620722350BAFFB65FC426D8F" authorityName="Weir &amp; Monteith" authorityYear="2010" box="[737,896,1240,1263]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="A0EF5B7C620722350BAFFB65FC426D8F" box="[737,896,1240,1263]" italics="true" pageId="41" pageNumber="42">Lepanus storeyi</emphasis>
</taxonomicName>
species group.
<emphasis id="A0EF5B7C620722350D55FB6AFBEF6D8F" bold="true" box="[1051,1069,1239,1263]" pageId="41" pageNumber="42">A</emphasis>
,
<taxonomicName id="559BFCED620722350D77FB65FB256D8F" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[1081,1255,1240,1263]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="meierae">
<emphasis id="A0EF5B7C620722350D77FB65FB256D8F" box="[1081,1255,1240,1263]" italics="true" pageId="41" pageNumber="42">Lepanus meierae</emphasis>
</taxonomicName>
male; B,
<taxonomicName id="559BFCED620722350C08FB65FF2B6C73" authorityName="Gunter &amp; Weir" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="meierae">
<emphasis id="A0EF5B7C620722350C08FB65FF2B6C73" italics="true" pageId="41" pageNumber="42">Lepanus meierae</emphasis>
</taxonomicName>
female;
<emphasis id="A0EF5B7C62072235080EFB46FE906C73" bold="true" box="[320,338,1275,1299]" pageId="41" pageNumber="42">C</emphasis>
,
<taxonomicName id="559BFCED620722350811FB41FE3D6C73" authorityName="Weir &amp; Monteith" authorityYear="2010" box="[351,511,1276,1299]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="A0EF5B7C620722350811FB41FE3D6C73" box="[351,511,1276,1299]" italics="true" pageId="41" pageNumber="42">Lepanus storeyi</emphasis>
</taxonomicName>
male;
<emphasis id="A0EF5B7C620722350B0DFB46FD996C73" bold="true" box="[579,603,1275,1299]" pageId="41" pageNumber="42">D,</emphasis>
<taxonomicName id="559BFCED620722350B2FFB41FCC36C73" authorityName="Weir &amp; Monteith" authorityYear="2010" box="[609,769,1276,1299]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="A0EF5B7C620722350B2FFB41FCC36C73" box="[609,769,1276,1299]" italics="true" pageId="41" pageNumber="42">Lepanus storeyi</emphasis>
</taxonomicName>
female;
<emphasis id="A0EF5B7C620722350A16FB46FCAB6C73" bold="true" box="[856,873,1275,1299]" pageId="41" pageNumber="42">E</emphasis>
,
<taxonomicName id="559BFCED620722350A3BFB41FBEC6C73" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[885,1070,1275,1299]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="A0EF5B7C620722350A3BFB41FBEC6C73" box="[885,1070,1275,1299]" italics="true" pageId="41" pageNumber="42">Lepanus williamsi</emphasis>
</taxonomicName>
male. All specimens in comparative scale. Scale bar 0.5 mm.
</paragraph>
</caption>
<subSubSection id="DA81D4E5620722360989FAD8FE556FD4" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="9224876E620722350989FAD8FEBC6CC4" blockId="41.[151,1437,1381,2020]" pageId="41" pageNumber="42">
<emphasis id="A0EF5B7C620722350989FAD8FE9B6C1F" bold="true" box="[199,345,1381,1407]" pageId="41" pageNumber="42">Description.</emphasis>
Head, pronotum and elytra brown, humeral and apical areas with ill-defined orange colouration. Antennal clubs grey.
</paragraph>
<paragraph id="9224876E620722350989FA10FB296CA8" blockId="41.[151,1437,1381,2020]" box="[199,1259,1453,1480]" pageId="41" pageNumber="42">
Total length:
<quantity id="55632A8B620722350812FA10FE246CA8" box="[348,486,1453,1480]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="41" pageNumber="42" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mm</quantity>
.
<typeStatus id="4D2039CC6207223508BFFA10FD9B6CA7" box="[497,601,1453,1479]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
measurements: total length
<quantity id="55632A8B620722350ADBFA10FC2F6CA8" box="[917,1005,1453,1480]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="41" pageNumber="42" unit="mm" value="3.5">3.5 mm</quantity>
, elytral width
<quantity id="55632A8B620722350DC0FA13FB256CA8" box="[1166,1255,1454,1480]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="41" pageNumber="42" unit="mm" value="2.2">2.2 mm</quantity>
.
</paragraph>
<paragraph id="9224876E620722350989FA6CFF776F54" blockId="41.[151,1437,1381,2020]" pageId="41" pageNumber="42">
<emphasis id="A0EF5B7C620722350989FA6CFECA6C8B" box="[199,264,1489,1515]" italics="true" pageId="41" pageNumber="42">Male.</emphasis>
Head: Width to length ratio 56: 40. Surface smooth and nitid, with fine punctation, becoming very fine anteriorly. Dorsal part of eyes narrow, separated by an interocular space approximately 22–23 times eye width (45: 2).
</paragraph>
<paragraph id="9224876E620722350989F980FD846FA4" blockId="41.[151,1437,1381,2020]" pageId="41" pageNumber="42">Prothorax: Pronotum anterior angles obtuse. Surface smooth, nitid, finely punctate. A row of slightly elongate punctures present along middle two thirds of hind margin. Hypomeral stria about half the length of hypomeron and very close to lateral margin which almost appears double at that point. Hypomeral surface very finely reticulate. Pronotum width to length ratio 47: 27.</paragraph>
<paragraph id="9224876E620722350989F970FE796E4F" blockId="41.[151,1437,1381,2020]" pageId="41" pageNumber="42">Elytra: Surface reticulate with very fine superficial striae. Striae 1–4 equal width and appearing double at least in part. Striae 6 extending to the elytral base. Epipleura reticulate. Ratio of length of the elytra along suture to maximum elytral width 58: 60.</paragraph>
<paragraph id="9224876E620722350989F887FED66EFB" blockId="41.[151,1437,1381,2020]" pageId="41" pageNumber="42">Legs: Protibia with 2 teeth on outer edge, front edge deeply angulate between outer tooth and apical digit. Apical digit parallel sided and obliquely truncate, with a sharp inner angle. Mesotibia with a brush of setae apically on inner edge.</paragraph>
<paragraph id="9224876E620722350989F818FB746E84" blockId="41.[151,1437,1381,2020]" pageId="41" pageNumber="42">Abdomen: Pygidium smooth, nitid, without a medial depression, with a transverse sinuate fold curving away from base. Abdominal ventrites finely reticulate for full width. Ventrite 6 very finely punctate.</paragraph>
<caption id="C6E4D7E66204223609D9FA22FBB06CBB" ID-DOI="http://doi.org/10.5281/zenodo.4496781" ID-Zenodo-Dep="4496781" httpUri="https://zenodo.org/record/4496781/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1439,1463]" targetBox="[159,1423,189,1407]" targetPageId="42">
<paragraph id="9224876E6204223609D9FA22FBB06CBB" blockId="42.[151,1436,1439,1500]" pageId="42" pageNumber="43">
<emphasis id="A0EF5B7C6204223609D9FA22FEDD6CD7" bold="true" box="[151,287,1439,1463]" pageId="42" pageNumber="43">FIGURE 11.</emphasis>
Left, dorsal and right view of aedeagi of species within the
<taxonomicName id="559BFCED620422360AC9FA1DFBEA6CD7" authorityName="Weir &amp; Monteith" authorityYear="2010" box="[903,1064,1440,1463]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="A0EF5B7C620422360AC9FA1DFBEA6CD7" box="[903,1064,1440,1463]" italics="true" pageId="42" pageNumber="43">Lepanus storeyi</emphasis>
</taxonomicName>
species group.
<emphasis id="A0EF5B7C620422360D89FA22FB1B6CD7" bold="true" box="[1223,1241,1439,1463]" pageId="42" pageNumber="43">A</emphasis>
,
<taxonomicName id="559BFCED620422360DA9FA1DFA546CD7" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[1255,1430,1440,1463]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="meierae">
<emphasis id="A0EF5B7C620422360DA9FA1DFA546CD7" box="[1255,1430,1440,1463]" italics="true" pageId="42" pageNumber="43">Lepanus meierae</emphasis>
</taxonomicName>
;
<emphasis id="A0EF5B7C6204223609D9FA79FF6A6CBC" bold="true" box="[151,168,1476,1500]" pageId="42" pageNumber="43">B</emphasis>
,
<taxonomicName id="559BFCED6204223609FBFA79FE976CBB" authorityName="Weir &amp; Monteith" authorityYear="2010" box="[181,341,1476,1499]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="A0EF5B7C6204223609FBFA79FE976CBB" box="[181,341,1476,1499]" italics="true" pageId="42" pageNumber="43">Lepanus storeyi</emphasis>
</taxonomicName>
;
<emphasis id="A0EF5B7C62042236082CFA7EFEB66CBB" bold="true" box="[354,372,1475,1499]" pageId="42" pageNumber="43">C</emphasis>
,
<taxonomicName id="559BFCED6204223608CFFA79FDF86CBB" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[385,570,1475,1499]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="A0EF5B7C6204223608CFFA79FDF86CBB" box="[385,570,1475,1499]" italics="true" pageId="42" pageNumber="43">Lepanus williamsi</emphasis>
</taxonomicName>
. All specimens in comparative scale. Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="9224876E620422360989F9B7FDB96F0C" blockId="42.[151,1437,1546,1968]" pageId="42" pageNumber="43">Pterothorax: Medial lobe of metaventrite very finely punctate and broadly margined between mesocoxae. Lateral lobe of metaventrite punctate and setose. Mesoventrite smooth with a few very fine punctures. Mesepimeron smooth. Metanepisternum finely reticulate.</paragraph>
<paragraph id="9224876E620422360989F9CBFE556FD4" blockId="42.[151,1437,1546,1968]" pageId="42" pageNumber="43">
<emphasis id="A0EF5B7C620422360989F9CBFEE16FF0" box="[199,291,1654,1680]" italics="true" pageId="42" pageNumber="43">Female.</emphasis>
Protibia with a short apical digit giving rise to a long apical spur. Mesotibia without a brush of setae apically on inner edge.
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E5620422360989F903FCDB6F9C" pageId="42" pageNumber="43" type="etymology">
<paragraph id="9224876E620422360989F903FCDB6F9C" blockId="42.[151,1437,1546,1968]" pageId="42" pageNumber="43">
<emphasis id="A0EF5B7C620422360989F903FE8C6FB8" bold="true" box="[199,334,1726,1752]" pageId="42" pageNumber="43">Etymology.</emphasis>
This species is named for Geoff Williams who has collected almost all specimens known to date, as well as many other
<taxonomicName id="559BFCED6204223608DFF95FFE326F9B" box="[401,496,1762,1787]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="A0EF5B7C6204223608DFF95FFE326F9B" box="[401,496,1762,1787]" italics="true" pageId="42" pageNumber="43">Lepanus</emphasis>
</taxonomicName>
examined in this revision.
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E5620422360989F8B8FE506E08" pageId="42" pageNumber="43" type="distribution">
<paragraph id="9224876E620422360989F8B8FE506E08" blockId="42.[151,1437,1546,1968]" pageId="42" pageNumber="43">
<emphasis id="A0EF5B7C620422360989F8B8FEA36E7F" bold="true" box="[199,353,1797,1823]" pageId="42" pageNumber="43">Distribution.</emphasis>
Known from subtropical rainforests and wet sclerophyll forests of eastern
<collectingRegion id="505F498C620422360DD5F8BBFAA26E40" box="[1179,1376,1798,1824]" country="Australia" name="New South Wales" pageId="42" pageNumber="43">New South Wales</collectingRegion>
from Port Macquarie to south of Sydney in the
<collectingRegion id="505F498C620422360B36F897FC816E24" box="[632,835,1834,1860]" country="Australia" name="New South Wales" pageId="42" pageNumber="43">New South Wales</collectingRegion>
North
<collectingRegion id="505F498C620422360ADBF897FC176E24" box="[917,981,1834,1860]" country="New Zealand" name="West Coast" pageId="42" pageNumber="43">Coast</collectingRegion>
(NNC) and Sydney Basin (SYB) IBRA bioregions (
<figureCitation id="0AA09BEB620422360853F8F3FE476E08" box="[285,389,1870,1896]" captionStart="FIGURE 12" captionStartId="43.[151,250,1221,1245]" captionTargetBox="[155,1427,186,1191]" captionTargetId="figure-19@43.[151,1436,181,1197]" captionTargetPageId="43" captionText="FIGURE 12. A, Map of known records of Lepanus meierae, Lepanus storeyi, and Lepanus williamsi, B, predicted distribution of Lepanus meierae." figureDoi="http://doi.org/10.5281/zenodo.4496783" httpUri="https://zenodo.org/record/4496783/files/figure.png" pageId="42" pageNumber="43">Fig. 12A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E5620422360989F8CCFA836ED0" pageId="42" pageNumber="43" type="discussion">
<paragraph id="9224876E620422360989F8CCFA836ED0" blockId="42.[151,1437,1546,1968]" pageId="42" pageNumber="43">
<emphasis id="A0EF5B7C620422360989F8CCFE8D6EEB" bold="true" box="[199,335,1905,1931]" pageId="42" pageNumber="43">Comments.</emphasis>
This species has been referred to to under the informal name of “
<taxonomicName id="559BFCED620422360D6AF8CFFB0C6EEC" authority="NSW" authorityName="NSW" box="[1060,1230,1906,1932]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="A0EF5B7C620422360D6AF8CFFB416EEB" box="[1060,1155,1906,1931]" italics="true" pageId="42" pageNumber="43">Lepanus</emphasis>
NSW
</taxonomicName>
3”. The majority of specimens has been collected at faeces or in faeces baited pitfall traps but it has also collected in leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>
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