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<document id="B61D54D21E680E9EAA249C58BB913610" ID-CLB-Dataset="8243" ID-DOI="10.11646/zootaxa.4923.1.1" ID-GBIF-Dataset="0a027439-c601-4533-bd03-f0e5d4442801" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496757" ID-ZooBank="41D057A7-F855-47C2-A598-8000CFD83A5B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612332986149" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2021" docId="1A3236786202223009D9FCA2FA5F6EED" docLanguage="en" docName="zootaxa.4923.1.1.pdf" docOrigin="Zootaxa 4923 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lepanus carbinensis Gunter &amp; Weir 2021, new species" docType="key" docVersion="10" lastPageNumber="45" masterDocId="E60B4E00622E221C094EFFBDFFC26960" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species" masterLastPageNumber="66" masterPageNumber="1" pageNumber="45" updateTime="1698926692322" updateUser="ExternalLinkService">
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<mods:title id="28BC3DC41EC0E93881DFF6A632FB96EC">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species</mods:title>
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<mods:namePart id="0FFCF62D4139E4F967E3BD74732428A6">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="A82F637B3EF840C0E97EF17F1C9B938D">Weir, Thomas A.</mods:namePart>
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<subSubSection id="DA81D4E56202223009D9FCA2FD7D6A59" box="[151,703,799,825]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="9224876E6202223009D9FCA2FD7D6A59" blockId="44.[151,705,799,969]" box="[151,703,799,825]" pageId="44" pageNumber="45">
<taxonomicName id="559BFCED6202223009D9FCA2FDEC6A59" ID-CoL="7WWVP" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[151,558,799,825]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="carbinensis" status="sp. nov.">
<emphasis id="A0EF5B7C6202223009D9FCA2FEBC6A59" box="[151,382,799,825]" italics="true" pageId="44" pageNumber="45">Lepanus carbinensis</emphasis>
Gunter &amp; Weir
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,
<taxonomicNameLabel id="BBDCE607620222300B77FCA2FD7D6A59" box="[569,703,799,825]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
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<paragraph id="9224876E6202223009D9FCFEFE3F6A3D" blockId="44.[151,705,799,969]" box="[151,509,834,861]" pageId="44" pageNumber="45">
<taxonomicName id="559BFCED6202223009D9FCFEFE3F6A3D" ID-CoL="6P88C" authority="(Gillet, 1925)" baseAuthorityName="Gillet" baseAuthorityYear="1925" box="[151,509,834,861]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="dichrous">
<emphasis id="A0EF5B7C6202223009D9FCFEFE9C6A3D" box="[151,350,835,861]" italics="true" pageId="44" pageNumber="45">Lepanus dichrous</emphasis>
(
<bibRefCitation id="F60AFA9F620222300823FCFEFE376A3D" author="Gillet" box="[365,501,834,861]" firstAuthor="Gillet" pageId="44" pageNumber="45" pagination="1 - 16" refId="ref76399" refString="Gillet, J. J. E. (1925) Results of Dr. E. Mj oberg's Swedish Scientific expedition to Australia, 1910 - 1913. 40. Scarabaeidae: Geo- trupinae et Coprinae. Arkiv for zoologi, 17, 1 - 16." type="journal article" year="1925">Gillet, 1925</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="9224876E6202223009D9FCDAFDB76AE1" blockId="44.[151,705,799,969]" box="[151,629,871,897]" pageId="44" pageNumber="45">
<taxonomicName id="559BFCED6202223009D9FCDAFE266AE1" ID-CoL="7WWVR" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[151,484,871,897]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="kulki" status="sp. nov.">
<emphasis id="A0EF5B7C6202223009D9FCDAFEF16AE1" box="[151,307,871,897]" italics="true" pageId="44" pageNumber="45">Lepanus kulki</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="BBDCE6076202223008A0FCDAFDB76AE1" box="[494,629,871,897]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</paragraph>
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<taxonomicName id="559BFCED6202223009D9FC36FDC76AC5" ID-CoL="3T8V5" authority="Matthews, 1974" authorityName=", Matthews" authorityYear="1974" box="[151,517,907,933]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="A0EF5B7C6202223009D9FC36FE8B6AC5" box="[151,329,907,933]" italics="true" pageId="44" pageNumber="45">Lepanus nitidus</emphasis>
<bibRefCitation id="F60AFA9F62022230081EFC36FDC76AC5" author="Matthews" box="[336,517,907,933]" firstAuthor="Matthews" pageId="44" pageNumber="45" pagination="1 - 211" refId="ref77152" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211. https: // doi. org / 10.1071 / AJZS 024" type="journal article" year="1974">Matthews, 1974</bibRefCitation>
</taxonomicName>
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<paragraph id="9224876E6202223009D9FC12FD036AA9" blockId="44.[151,705,799,969]" box="[151,705,943,969]" pageId="44" pageNumber="45">
<taxonomicName id="559BFCED6202223009D9FC12FDF26AA9" ID-CoL="7WWVV" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[151,560,943,969]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="vangerweni" status="sp. nov.">
<emphasis id="A0EF5B7C6202223009D9FC12FE426AA8" box="[151,384,943,968]" italics="true" pageId="44" pageNumber="45">Lepanus vangerweni</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="BBDCE607620222300B74FC12FD036AA9" box="[570,705,943,969]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</paragraph>
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<paragraph id="9224876E6202223009D9FBA6FCCD6D55" blockId="44.[151,783,1050,1077]" box="[151,783,1050,1077]" pageId="44" pageNumber="45">
<emphasis id="A0EF5B7C6202223009D9FBA6FCCD6D55" bold="true" box="[151,783,1050,1077]" pageId="44" pageNumber="45">
Key to species of the
<taxonomicName id="559BFCED6202223008D3FBA6FD9D6D55" ID-CoL="3T8V5" authorityName=", Matthews" authorityYear="1974" box="[413,607,1050,1077]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="A0EF5B7C6202223008D3FBA6FD9D6D55" bold="true" box="[413,607,1050,1077]" italics="true" pageId="44" pageNumber="45">Lepanus nitidus</emphasis>
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species group
</emphasis>
</paragraph>
<key id="E60AD05F6202223009D9FBDDFA5F6EED" pageId="44" pageNumber="45">
<keyStep id="296F9FCB6202223009D9FBDDFA5E6DAC" pageId="44" pageNumber="45">
<paragraph id="9224876E6202223009D9FBDDFA5E6DF2" blockId="44.[151,1437,1120,1933]" pageId="44" pageNumber="45">
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1 Ridge on ventral side of protibia apically more or less joined to the second tooth by a fine carina (
<figureCitation id="0AA09BEB620222300DCAFBDDFB1F6D16" box="[1156,1245,1120,1142]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13G</figureCitation>
); aedeagus with left paramere excised apically...............................................................................
<emphasis id="A0EF5B7C620222300CDEFBC0FA5E6DF2" bold="true" box="[1424,1436,1149,1170]" pageId="44" pageNumber="45">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="9224876E6202223009D9FB26FA5E6DAC" blockId="44.[151,1437,1120,1933]" pageId="44" pageNumber="45">
<keyLead id="296A245B6202223009D9FB26FA5E6DAC" pageId="44" pageNumber="45">
Ridge on ventral side of protibia not joined to the second tooth by a fine carina (
<figureCitation id="0AA09BEB620222300A97FB27FBF06DD0" box="[985,1074,1178,1200]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13H</figureCitation>
); aedeagus with left paramere excised apically or not........................................................................................
<emphasis id="A0EF5B7C620222300CDEFB0AFA5E6DAC" bold="true" box="[1424,1436,1207,1228]" pageId="44" pageNumber="45">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296F9FCB6202223009D9FB68FA5F6CD8" pageId="44" pageNumber="45">
<paragraph id="9224876E6202223009D9FB68FA5E6C22" blockId="44.[151,1437,1120,1933]" pageId="44" pageNumber="45">
<keyLead id="296A245B6202223009D9FB68FA5E6C22" pageId="44" pageNumber="45">
2 Lateral lobe of metaventrite with inner border beside mesocoxa anteriorly widened (
<figureCitation id="0AA09BEB620222300D43FB68FBA96D8B" box="[1037,1131,1237,1259]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13M</figureCitation>
); male medial lobe of metaventrite with a very small tubercle located approximately halfway between mesocoxae and metacoxae (
<figureCitation id="0AA09BEB620222300DEFFB4FFB326C68" box="[1185,1264,1266,1288]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13I</figureCitation>
); male protibia as in
<figureCitation id="0AA09BEB6202223009B4FAADFE906C45" box="[250,338,1296,1318]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13A</figureCitation>
; aedeagus as in
<figureCitation id="0AA09BEB6202223008A5FAADFD806C46" box="[491,578,1296,1318]" captionStart="FIGURE 16" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[246,1337,193,1906]" captionTargetId="figure-19@61.[234,1353,181,1919]" captionTargetPageId="61" captionText="FIGURE 16. Left, dorsal and right view of aedeagi of species within the Lepanus nitidus species group. A, Lepanus carbinensis; B, Lepanus dichrous; C, Lepanus kulki; D, Lepanus nitidus; E, Lepanus vangerweni. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496791" httpUri="https://zenodo.org/record/4496791/files/figure.png" pageId="44" pageNumber="45">Fig. 16D</figureCitation>
; generally larger species
<quantity id="55632A8B620222300A7DFAADFC6B6C45" box="[819,937,1296,1318]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.4" metricValueMin="3.6" pageId="44" pageNumber="45" unit="mm" value="4.0" valueMax="4.4" valueMin="3.6">3.64.4 mm</quantity>
; widely distributed in
<collectingRegion id="505F498C620222300D30FAADFB2D6C46" box="[1150,1263,1296,1318]" country="Australia" name="Queensland" pageId="44" pageNumber="45">Queensland</collectingRegion>
Wet Tropics from south of
<location id="9744D1B5620222300878FA90FE586C23" LSID="urn:lsid:plazi:treatment:1A3236786202223009D9FCA2FA5F6EED:9744D1B5620222300878FA90FE586C23" box="[310,410,1325,1347]" name="Cooktown" pageId="44" pageNumber="45" stateProvince="Queensland">Cooktown</location>
to Cardwell Range..........................................
<taxonomicName id="559BFCED620222300D18FA90FA5E6C22" ID-CoL="3T8V5" authority="Matthews, 1974" authorityName=", Matthews" authorityYear="1974" box="[1110,1436,1324,1346]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="A0EF5B7C620222300D18FA90FA5E6C22" bold="true" box="[1110,1436,1324,1346]" pageId="44" pageNumber="45">
<emphasis id="A0EF5B7C620222300D18FA90FB376C22" bold="true" box="[1110,1269,1324,1346]" italics="true" pageId="44" pageNumber="45">Lepanus nitidus</emphasis>
<bibRefCitation id="F60AFA9F620222300DB5FA90FA5E6C22" author="Matthews" box="[1275,1436,1325,1346]" firstAuthor="Matthews" pageId="44" pageNumber="45" pagination="1 - 211" refId="ref77152" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211. https: // doi. org / 10.1071 / AJZS 024" type="journal article" year="1974">Matthews, 1974</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="9224876E6202223009D9FAF6FA5F6CD8" blockId="44.[151,1437,1120,1933]" pageId="44" pageNumber="45">
<keyLead id="296A245B6202223009D9FAF6FA5F6CD8" pageId="44" pageNumber="45">
<collectingCounty id="7B45FFE26202223009ACFAF7FEE46C00" box="[226,294,1354,1376]" pageId="44" pageNumber="45">Lateral</collectingCounty>
lobe of metaventrite with inner border beside mesocoxa not anteriorly widened, of same width throughout (
<figureCitation id="0AA09BEB620222300C79FAF7FA526C00" box="[1335,1424,1354,1376]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13N</figureCitation>
); male medial lobe of metaventrite without very small tubercle; male protibia as in
<figureCitation id="0AA09BEB620222300AB3FAD5FB966C1D" box="[1021,1108,1384,1406]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13C</figureCitation>
; aedeagus as in
<figureCitation id="0AA09BEB620222300DBDFAD5FA896C1E" box="[1267,1355,1384,1406]" captionStart="FIGURE 16" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[246,1337,193,1906]" captionTargetId="figure-19@61.[234,1353,181,1919]" captionTargetPageId="61" captionText="FIGURE 16. Left, dorsal and right view of aedeagi of species within the Lepanus nitidus species group. A, Lepanus carbinensis; B, Lepanus dichrous; C, Lepanus kulki; D, Lepanus nitidus; E, Lepanus vangerweni. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496791" httpUri="https://zenodo.org/record/4496791/files/figure.png" pageId="44" pageNumber="45">Fig. 16C</figureCitation>
; smaller species
<quantity id="55632A8B620222300860FA38FEB96CFA" box="[302,379,1413,1435]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="44" pageNumber="45" unit="mm" value="3.5">3.5 mm</quantity>
;
<collectingRegion id="505F498C6202223008CBFA38FE346CFB" box="[389,502,1413,1435]" country="Australia" name="Queensland" pageId="44" pageNumber="45">Queensland</collectingRegion>
<collectingMunicipality id="72401D146202223008B3FA38FD0E6CFB" box="[509,716,1413,1435]" pageId="44" pageNumber="45">northern Wet Tropics</collectingMunicipality>
, apparently restricted to the
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and
<location id="9744D1B5620222300DC6FA38FAD56CFB" LSID="urn:lsid:plazi:treatment:1A3236786202223009D9FCA2FA5F6EED:9744D1B5620222300DC6FA38FAD56CFB" box="[1160,1303,1413,1435]" county="Lateral" municipality="northern Wet Tropics" name="Thornton Peak" pageId="44" pageNumber="45" stateProvince="Queensland">Thornton Peak</location>
blocks above
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<elevation id="19B6605D6202223009ACFA1FFEDD6CD8" box="[226,287,1442,1464]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="44" pageNumber="45" unit="m" value="700.0">700 m</elevation>
</quantity>
.
</specimenCount>
..........................................................
<emphasis id="A0EF5B7C620222300AA2FA1EFA5F6CD8" bold="true" box="[1004,1437,1442,1464]" pageId="44" pageNumber="45">
<taxonomicName id="559BFCED620222300AA2FA1EFADE6CD8" ID-CoL="7WWVR" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[1004,1308,1442,1464]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="kulki" status="sp. nov.">
<emphasis id="A0EF5B7C620222300AA2FA1EFBB86CD8" bold="true" box="[1004,1146,1442,1464]" italics="true" pageId="44" pageNumber="45">Lepanus kulki</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="BBDCE607620222300C68FA1FFA5F6CD8" box="[1318,1437,1442,1464]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296F9FCB6202223009D9FA7DFA5E6FC2" pageId="44" pageNumber="45">
<paragraph id="9224876E6202223009D9FA7DFA5F6F4D" blockId="44.[151,1437,1120,1933]" pageId="44" pageNumber="45">
<keyLead id="296A245B6202223009D9FA7DFA5F6F4D" pageId="44" pageNumber="45">
3
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of elytra nitid, with stria 1 and usually stria 2 superficial, appearing as a single line (
<figureCitation id="0AA09BEB620222300D3FFA7DFB056CB6" box="[1137,1223,1472,1494]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13L</figureCitation>
); metaventrite smooth and nitid between metacoxae, male with a very small tubercle located closer to metacoxae than mesocoxae (
<figureCitation id="0AA09BEB620222300DB6FA60FA8B6C93" box="[1272,1353,1501,1523]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13J</figureCitation>
), female without tubercle or depression; male protibia as in
<figureCitation id="0AA09BEB620222300B95FA47FCF46F6F" box="[731,822,1530,1552]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13D</figureCitation>
; aedeagus as in
<figureCitation id="0AA09BEB620222300A97FA47FBF36F70" box="[985,1073,1530,1552]" captionStart="FIGURE 16" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[246,1337,193,1906]" captionTargetId="figure-19@61.[234,1353,181,1919]" captionTargetPageId="61" captionText="FIGURE 16. Left, dorsal and right view of aedeagi of species within the Lepanus nitidus species group. A, Lepanus carbinensis; B, Lepanus dichrous; C, Lepanus kulki; D, Lepanus nitidus; E, Lepanus vangerweni. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496791" httpUri="https://zenodo.org/record/4496791/files/figure.png" pageId="44" pageNumber="45">Fig. 16B</figureCitation>
, with left paramere excised apically;
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;
<collectingRegion id="505F498C62022230082CF9A5FE116F4E" box="[354,467,1560,1582]" country="Australia" name="Queensland" pageId="44" pageNumber="45">Queensland</collectingRegion>
Wet Tropics from Lamb Range to Ravenshoe..................
<taxonomicName id="559BFCED620222300D11F9A5FA5F6F4D" ID-CoL="6P88C" authority="(Gillet, 1925)" baseAuthorityName="Gillet" baseAuthorityYear="1925" box="[1119,1437,1559,1581]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="dichrous">
<emphasis id="A0EF5B7C620222300D11F9A5FA5F6F4D" bold="true" box="[1119,1437,1559,1581]" pageId="44" pageNumber="45">
<emphasis id="A0EF5B7C620222300D11F9A5FAD36F4D" bold="true" box="[1119,1297,1559,1581]" italics="true" pageId="44" pageNumber="45">Lepanus dichrous</emphasis>
(
<bibRefCitation id="F60AFA9F620222300C50F9AAFA576F4D" author="Gillet" box="[1310,1429,1559,1581]" firstAuthor="Gillet" pageId="44" pageNumber="45" pagination="1 - 16" refId="ref76399" refString="Gillet, J. J. E. (1925) Results of Dr. E. Mj oberg's Swedish Scientific expedition to Australia, 1910 - 1913. 40. Scarabaeidae: Geo- trupinae et Coprinae. Arkiv for zoologi, 17, 1 - 16." type="journal article" year="1925">Gillet, 1925</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="9224876E6202223009D9F988FA5E6FC2" blockId="44.[151,1437,1120,1933]" pageId="44" pageNumber="45">
<keyLead id="296A245B6202223009D9F988FA5E6FC2" pageId="44" pageNumber="45">
Apical area of elytra usually very finely reticulate, with stria 1 and usually stria 2 slightly impressed, appearing as a double line (
<figureCitation id="0AA09BEB62022230085DF9EFFEAF6F08" box="[275,365,1618,1640]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13O</figureCitation>
); metaventrite between metacoxae more-or-less reticulate, male with a medial longitudinal depression between mesocoxae and metacoxae (
<figureCitation id="0AA09BEB6202223008A2F9CDFD866FE6" box="[492,580,1648,1670]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13K</figureCitation>
), female with a small depression and medial line between metacoxae; aedeagus as in
<figureCitation id="0AA09BEB620222300C23F9CDFEE46FC2" captionStart="FIGURE 16" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[246,1337,193,1906]" captionTargetId="figure-19@61.[234,1353,181,1919]" captionTargetPageId="61" captionText="FIGURE 16. Left, dorsal and right view of aedeagi of species within the Lepanus nitidus species group. A, Lepanus carbinensis; B, Lepanus dichrous; C, Lepanus kulki; D, Lepanus nitidus; E, Lepanus vangerweni. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496791" httpUri="https://zenodo.org/record/4496791/files/figure.png" pageId="44" pageNumber="45">Figs. 16A, E</figureCitation>
with left paramere not excised apically, somewhat pointed; larger species 4.0
<quantity id="55632A8B620222300D54F930FBA16FC2" box="[1050,1123,1677,1699]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="44" pageNumber="45" unit="mm" value="5.2">5.2 mm</quantity>
.........................
<emphasis id="A0EF5B7C620222300CDEF930FA5E6FC2" bold="true" box="[1424,1436,1677,1698]" pageId="44" pageNumber="45">4</emphasis>
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</paragraph>
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<paragraph id="9224876E6202223009D9F917FA5F6E78" blockId="44.[151,1437,1120,1933]" pageId="44" pageNumber="45">
<keyLead id="296A245B6202223009D9F917FA5F6E78" pageId="44" pageNumber="45">
4
<collectingCounty id="7B45FFE26202223009ACF917FE9F6FA0" box="[226,349,1706,1728]" pageId="44" pageNumber="45">Metaventrite</collectingCounty>
with punctures on medial lobe minute, much smaller than those on the lateral lobe (
<figureCitation id="0AA09BEB620222300DCDF917FB1B6FA0" box="[1155,1241,1706,1728]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13P</figureCitation>
); male protibia as in
<figureCitation id="0AA09BEB6202223009ACF975FEF56FBD" box="[226,311,1736,1758]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13E</figureCitation>
; pygidial reticulation usually confined to central and basal areas; aedeagus as in
<figureCitation id="0AA09BEB620222300D73F975FB506FBE" box="[1085,1170,1736,1758]" captionStart="FIGURE 16" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[246,1337,193,1906]" captionTargetId="figure-19@61.[234,1353,181,1919]" captionTargetPageId="61" captionText="FIGURE 16. Left, dorsal and right view of aedeagi of species within the Lepanus nitidus species group. A, Lepanus carbinensis; B, Lepanus dichrous; C, Lepanus kulki; D, Lepanus nitidus; E, Lepanus vangerweni. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496791" httpUri="https://zenodo.org/record/4496791/files/figure.png" pageId="44" pageNumber="45">Fig. 16E</figureCitation>
, with apically deflexed part of right paramere rounded;
<collectingRegion id="505F498C6202223008A4F958FD996F9B" box="[490,603,1765,1787]" country="Australia" name="Queensland" pageId="44" pageNumber="45">Queensland</collectingRegion>
<collectingMunicipality id="72401D14620222300B2FF958FD166F9B" box="[609,724,1765,1787]" pageId="44" pageNumber="45">Wet Tropics</collectingMunicipality>
from
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to
<location id="9744D1B5620222300AE7F958FBCF6F9B" LSID="urn:lsid:plazi:treatment:1A3236786202223009D9FCA2FA5F6EED:9744D1B5620222300AE7F958FBCF6F9B" box="[937,1037,1765,1787]" county="Metaventrite" municipality="Wet Tropics" name="Cardstone" pageId="44" pageNumber="45" stateProvince="Queensland">Cardstone</location>
, above
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<quantity id="55632A8B620222300D18F958FB516F9A" box="[1110,1171,1765,1787]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="44" pageNumber="45" unit="m" value="750.0">
<elevation id="19B6605D620222300D18F958FB516F9A" box="[1110,1171,1765,1787]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="44" pageNumber="45" unit="m" value="750.0">750 m</elevation>
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.
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................................................................................
<emphasis id="A0EF5B7C620222300AE2F8BEFA5F6E78" bold="true" box="[940,1437,1794,1816]" pageId="44" pageNumber="45">
<taxonomicName id="559BFCED620222300AE2F8BEFADE6E78" ID-CoL="7WWVV" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[940,1308,1794,1816]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="vangerweni" status="sp. nov.">
<emphasis id="A0EF5B7C620222300AE2F8BEFBB86E77" bold="true" box="[940,1146,1794,1816]" italics="true" pageId="44" pageNumber="45">Lepanus vangerweni</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="BBDCE607620222300C68F8BFFA5F6E78" box="[1318,1437,1794,1816]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="9224876E6202223009D9F89DFA5F6EED" blockId="44.[151,1437,1120,1933]" pageId="44" pageNumber="45">
<keyLead id="296A245B6202223009D9F89DFA5F6EED" pageId="44" pageNumber="45">
Metaventrite with punctures on medial lobe larger, about the same size as those on the lateral lobe (
<figureCitation id="0AA09BEB620222300DDEF89DFB2B6E56" box="[1168,1257,1824,1846]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig. 13Q</figureCitation>
); male protibiae as in
<figureCitation id="0AA09BEB6202223009B7F880FE856E33" box="[249,327,1853,1875]" captionStart="FIGURE 13" captionStartId="57.[151,250,1614,1638]" captionTargetBox="[293,1291,189,1590]" captionTargetId="figure-19@57.[293,1295,181,1590]" captionTargetPageId="57" captionText="FIGURE 13. Some diagnostic characters of the Lepanus nitidus species group. A, Dorsal view of male protibia of Lepanus nitidus; B, dorsal view of female protibia of Lepanus nitidus; C, dorsal view of male protibia of Lepanus kulki; D, dorsal view of male protibia of Lepanus dichrous; E, dorsal view of male protibia of Lepanus vangerweni; F, dorsal view of male protibia of Lepanus carbinensis; G, ventral view of male protibia of Lepanus nitidus, grey arrow points to diagnostic carina; H, ventral view of male protibia of Lepanus dichrous showing lack of carina; I, ventral view of metaventrite of Lepanus nitidus, showing tubercle (grey arrow); J, ventral view of metaventrite of Lepanus dichrous, showing tubercle (grey arrow); K, ventrolateral view of metaventrite of Lepanus vangerweni, showing medial longitudinal depression; L, apical view of elytra of Lepanus dichrous, showing stria 1 appearing superficial towards the apex (grey arrow); M, lateral lobe of metaventrite of Lepanus nitidus, grey arrow points to apically widened inner border; N, lateral lobe of metaventrite of Lepanus kulki; O, apical view of elytra of Lepanus carbinensis, showing stria 1 visibly impressed towards the apex (grey arrow); P, metaventrite of Lepanus vangerweni showing punctures on medial lobe much smaller than those on lateral lobe; Q, metaventrite of Lepanus carbinensis showing punctures on medial lobe same size as those on lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.4496785" httpUri="https://zenodo.org/record/4496785/files/figure.png" pageId="44" pageNumber="45">Fig.13F</figureCitation>
; pygidium usually nearly completely reticulate; aedeagus as in
<figureCitation id="0AA09BEB620222300AD0F880FC336E33" box="[926,1009,1853,1875]" captionStart="FIGURE 16" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[246,1337,193,1906]" captionTargetId="figure-19@61.[234,1353,181,1919]" captionTargetPageId="61" captionText="FIGURE 16. Left, dorsal and right view of aedeagi of species within the Lepanus nitidus species group. A, Lepanus carbinensis; B, Lepanus dichrous; C, Lepanus kulki; D, Lepanus nitidus; E, Lepanus vangerweni. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496791" httpUri="https://zenodo.org/record/4496791/files/figure.png" pageId="44" pageNumber="45">Fig. 16A</figureCitation>
with apically deflexed part of right paramere angulate;
<collectingRegion id="505F498C62022230080EF8E7FE736E10" box="[320,433,1882,1904]" country="Australia" name="Queensland" pageId="44" pageNumber="45">Queensland</collectingRegion>
Wet Tropics, apparently restricted to the
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......................................................................................
<emphasis id="A0EF5B7C620222300AE3F8C5FA5F6EED" bold="true" box="[941,1437,1911,1933]" pageId="44" pageNumber="45">
<taxonomicName id="559BFCED620222300AE3F8C5FADE6EED" ID-CoL="7WWVP" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[941,1308,1911,1933]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="carbinensis" status="sp. nov.">
<emphasis id="A0EF5B7C620222300AE3F8C5FBB86EED" bold="true" box="[941,1146,1911,1933]" italics="true" pageId="44" pageNumber="45">Lepanus carbinensis</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
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</emphasis>
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</treatment>
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