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<document id="DE6ADADCD16C5EFB08E8269CA0D7C3EB" ID-CLB-Dataset="27870" ID-DOI="10.11646/zootaxa.4527.1.11" ID-GBIF-Dataset="ac565270-3576-4125-b47f-13e57c3b17fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="2611919" ID-ZooBank="3B3B334F-5083-4F53-851A-F12DA6A91E18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553697984126" checkinUser="plazi" docAuthor="Komai, Furumi, Byun, Bong-Kyu &amp; Yamazaki, Saburo" docDate="2018" docId="1A17878BA37B7B7BFF69FBF78957FC99" docLanguage="en" docName="zootaxa.4527.1.11.pdf" docOrigin="Zootaxa 4527 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metacosma Kuznetzov 1985" docType="treatment" docVersion="8" lastPageNumber="133" masterDocId="E62EFFF3A37A7B79FFFEFFEF8B14FF8C" masterDocTitle="Remarks on the genus Metacosma Kuznetzov, 1985 (Lepidoptera: Tortricidae, Olethreutinae, Eucosmini), with description of a new species from Japan and Korea" masterLastPageNumber="139" masterPageNumber="131" pageNumber="132" updateTime="1698714385838" updateUser="ExternalLinkService">
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<mods:title id="3F0E0D3BB6E70AB95E1366CF2FB53E8E">Remarks on the genus Metacosma Kuznetzov, 1985 (Lepidoptera: Tortricidae, Olethreutinae, Eucosmini), with description of a new species from Japan and Korea</mods:title>
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<mods:namePart id="062D9EE79C4ACDCD1D1F56982702B5FC">Komai, Furumi</mods:namePart>
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<mods:namePart id="1DA1BBAC7CF35434049E4E3BD92EAEDF">Byun, Bong-Kyu</mods:namePart>
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<mods:title id="DA48231B2A5CB0CD0F114F9B14D8C925">Zootaxa</mods:title>
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<treatment id="1A17878BA37B7B7BFF69FBF78957FC99" ID-DOI="http://doi.org/10.5281/zenodo.6488447" ID-GBIF-Taxon="154507891" ID-Zenodo-Dep="6488447" LSID="urn:lsid:plazi:treatment:1A17878BA37B7B7BFF69FBF78957FC99" httpUri="http://treatment.plazi.org/id/1A17878BA37B7B7BFF69FBF78957FC99" lastPageId="2" lastPageNumber="133" pageId="1" pageNumber="132">
<subSubSection id="DAA46516A37B7B78FF69FBF78AE2FBBF" box="[151,502,1048,1075]" pageId="1" pageNumber="132" type="nomenclature">
<paragraph id="9201369DA37B7B78FF69FBF78AE2FBBF" blockId="1.[151,502,1048,1075]" box="[151,502,1048,1075]" pageId="1" pageNumber="132">
<heading id="C94981F1A37B7B78FF69FBF78AE2FBBF" bold="true" box="[151,502,1048,1075]" fontSize="11" level="1" pageId="1" pageNumber="132" reason="1">
<taxonomicName id="55BE4D1EA37B7B78FF69FBF78AE2FBBF" ID-CoL="92FFY" authority="Kuznetzov, 1985" authorityName="Kuznetzov" authorityYear="1985" box="[151,502,1048,1075]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA37B7B78FF69FBF78AE2FBBF" bold="true" box="[151,502,1048,1075]" pageId="1" pageNumber="132">
<emphasis id="A0CAEA8FA37B7B78FF69FBF78A35FBBE" bold="true" box="[151,289,1048,1074]" italics="true" pageId="1" pageNumber="132">Metacosma</emphasis>
<bibRefCitation id="F62F4B6CA37B7B78FED6FBF68AE2FBBF" author="Kuznetzov, V. I." box="[296,502,1049,1075]" pageId="1" pageNumber="132" pagination="3 - 11" refId="ref4543" refString="Kuznetzov, V. I. (1985) New representatives of the tribe Eucosmini (Lepidoptera, Tortricidae) of the USSR Asiatic part. Festnik Zoologii, 1985 (1), 3 - 11. [in Russian]" type="journal article" year="1985">Kuznetzov, 1985</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DAA46516A37B7B78FF69FB8E8FEEF935" pageId="1" pageNumber="132" type="materials_examined">
<paragraph id="9201369DA37B7B78FF69FB8E8F40FBFA" blockId="1.[151,1108,1120,1142]" box="[151,1108,1120,1142]" pageId="1" pageNumber="132">
<typeStatus id="4D05883FA37B7B78FF69FB8E8BDDFBFA" box="[151,201,1121,1142]" pageId="1" pageNumber="132">Type</typeStatus>
species:
<taxonomicName id="55BE4D1EA37B7B78FED8FB8E89D6FBFA" authority="Kuznetzov, 1985" authorityName="Kuznetzov" authorityYear="1985" box="[294,706,1120,1142]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="impolitana">
<emphasis id="A0CAEA8FA37B7B78FED8FB8E891BFBFA" box="[294,527,1120,1142]" italics="true" pageId="1" pageNumber="132">Metacosma impolitana</emphasis>
<bibRefCitation id="F62F4B6CA37B7B78FDEBFB8E89D6FBFA" author="Kuznetzov, V. I." box="[533,706,1120,1142]" pageId="1" pageNumber="132" pagination="3 - 11" refId="ref4543" refString="Kuznetzov, V. I. (1985) New representatives of the tribe Eucosmini (Lepidoptera, Tortricidae) of the USSR Asiatic part. Festnik Zoologii, 1985 (1), 3 - 11. [in Russian]" type="journal article" year="1985">Kuznetzov, 1985</bibRefCitation>
</taxonomicName>
, by original designation and monotypy.
</paragraph>
<paragraph id="9201369DA37B7B78FF69FB4D8FEEF935" blockId="1.[151,1437,1186,2009]" pageId="1" pageNumber="132">
<emphasis id="A0CAEA8FA37B7B78FF69FB4D896FFB37" bold="true" box="[151,635,1186,1211]" pageId="1" pageNumber="132">
Redesciption (excluding
<taxonomicName id="55BE4D1EA37B7B78FE3AFB4C8965FB37" authorityName="Kuznetzov" authorityYear="1988" box="[452,625,1187,1211]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="miratorana">
<emphasis id="A0CAEA8FA37B7B78FE3AFB4C8965FB37" bold="true" box="[452,625,1187,1211]" italics="true" pageId="1" pageNumber="132">M. miratorana</emphasis>
</taxonomicName>
)
</emphasis>
. Head: Antenna (
<figureCitation id="0A852A18A37B7B78FCB0FB4D88ABFB37" box="[846,959,1186,1211]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1940,1963]" captionTargetBox="[151,1437,844,1919]" captionTargetId="figure@2.[151,1437,844,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. Metacosma echinophora. 1. Adult moth, ♂, paratype, Yakushima Is., Japan. 2. Notch on base of male antenna. 3. Lateral view of male head. 4. Larval habitat, resin dome constructed among winter buds of Pinus thunbergii, Gifu Pref., Japan. 5. Detail of resin dome. 6. lateral view of mature larva (photo. Akihiko Miyano)." httpUri="https://zenodo.org/record/2611921/files/figure.png" pageId="1" pageNumber="132">Figs. 2, 3</figureCitation>
) in male with a notch near base. Labial palpus (
<figureCitation id="0A852A18A37B7B78FF0DFB298A2EFB53" box="[243,314,1222,1247]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1940,1963]" captionTargetBox="[151,1437,844,1919]" captionTargetId="figure@2.[151,1437,844,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. Metacosma echinophora. 1. Adult moth, ♂, paratype, Yakushima Is., Japan. 2. Notch on base of male antenna. 3. Lateral view of male head. 4. Larval habitat, resin dome constructed among winter buds of Pinus thunbergii, Gifu Pref., Japan. 5. Detail of resin dome. 6. lateral view of mature larva (photo. Akihiko Miyano)." httpUri="https://zenodo.org/record/2611921/files/figure.png" pageId="1" pageNumber="132">Fig. 3</figureCitation>
) with second segment expanded distally by scales, roughly triangular, ascending; third segment short, porrect. Thorax: Forewing (
<figureCitation id="0A852A18A37B7B78FDDFFB058965FA8F" box="[545,625,1258,1283]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1940,1963]" captionTargetBox="[151,1437,844,1919]" captionTargetId="figure@2.[151,1437,844,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. Metacosma echinophora. 1. Adult moth, ♂, paratype, Yakushima Is., Japan. 2. Notch on base of male antenna. 3. Lateral view of male head. 4. Larval habitat, resin dome constructed among winter buds of Pinus thunbergii, Gifu Pref., Japan. 5. Detail of resin dome. 6. lateral view of mature larva (photo. Akihiko Miyano)." httpUri="https://zenodo.org/record/2611921/files/figure.png" pageId="1" pageNumber="132">Figs. 1</figureCitation>
,
<figureCitation id="0A852A18A37B7B78FD7DFB048986FA8F" box="[643,658,1259,1283]" captionStart="FIGURES 7–9" captionStartId="3.[151,264,1135,1158]" captionTargetBox="[200,1402,281,1079]" captionTargetId="figure@3.[150,1436,193,1115]" captionTargetPageId="3" captionText="FIGURES 7–9. Metacosma echinophora, wing structure. 7. Wing venation, ♂, Gifu Pref., Japan, slide FK 1292. 8. Detail of anal fold on hindwing, Nagano Pref., Japan, slide FK 2142. 9. Wing venation, ♀, Nagano Pref., Japan, slide FK 2142." httpUri="https://zenodo.org/record/2611923/files/figure.png" pageId="1" pageNumber="132">7</figureCitation>
) long and narrow, without costal fold in male; M-stem and chorda absent; all veins present and separate beyond discal cell; R
<subScript id="0E3A34D8A37B7B78FCC8FAF2882AFAA7" attach="left" box="[822,830,1309,1323]" fontSize="6" pageId="1" pageNumber="132">3</subScript>
approximated to R
<subScript id="0E3A34D8A37B7B78FBE5FAF28F37FAA7" attach="left" box="[1051,1059,1309,1323]" fontSize="6" pageId="1" pageNumber="132">4</subScript>
at base. Hindwing (
<figureCitation id="0A852A18A37B7B78FAF4FAE18E4DFAAB" box="[1290,1369,1294,1319]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1940,1963]" captionTargetBox="[151,1437,844,1919]" captionTargetId="figure@2.[151,1437,844,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. Metacosma echinophora. 1. Adult moth, ♂, paratype, Yakushima Is., Japan. 2. Notch on base of male antenna. 3. Lateral view of male head. 4. Larval habitat, resin dome constructed among winter buds of Pinus thunbergii, Gifu Pref., Japan. 5. Detail of resin dome. 6. lateral view of mature larva (photo. Akihiko Miyano)." httpUri="https://zenodo.org/record/2611921/files/figure.png" pageId="1" pageNumber="132">Figs. 1</figureCitation>
,
<figureCitation id="0A852A18A37B7B78FA97FAE08E80FAAB" box="[1385,1428,1295,1319]" captionStart="FIGURES 7–9" captionStartId="3.[151,264,1135,1158]" captionTargetBox="[200,1402,281,1079]" captionTargetId="figure@3.[150,1436,193,1115]" captionTargetPageId="3" captionText="FIGURES 7–9. Metacosma echinophora, wing structure. 7. Wing venation, ♂, Gifu Pref., Japan, slide FK 1292. 8. Detail of anal fold on hindwing, Nagano Pref., Japan, slide FK 2142. 9. Wing venation, ♀, Nagano Pref., Japan, slide FK 2142." httpUri="https://zenodo.org/record/2611923/files/figure.png" pageId="1" pageNumber="132">7–9</figureCitation>
) almost equal to forewing in width, with sparse cubital pecten. Hindwing in male with anal margin produced into a free elongate lobe with the outer edge folded upward and embracing a scale pencil from the base of the anal area (
<figureCitation id="0A852A18A37B7B78FF61FA928BF8FA1A" box="[159,236,1405,1430]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1940,1963]" captionTargetBox="[151,1437,844,1919]" captionTargetId="figure@2.[151,1437,844,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. Metacosma echinophora. 1. Adult moth, ♂, paratype, Yakushima Is., Japan. 2. Notch on base of male antenna. 3. Lateral view of male head. 4. Larval habitat, resin dome constructed among winter buds of Pinus thunbergii, Gifu Pref., Japan. 5. Detail of resin dome. 6. lateral view of mature larva (photo. Akihiko Miyano)." httpUri="https://zenodo.org/record/2611921/files/figure.png" pageId="1" pageNumber="132">Figs. 1</figureCitation>
,
<figureCitation id="0A852A18A37B7B78FF07FA918A30FA1A" box="[249,292,1406,1430]" captionStart="FIGURES 7–9" captionStartId="3.[151,264,1135,1158]" captionTargetBox="[200,1402,281,1079]" captionTargetId="figure@3.[150,1436,193,1115]" captionTargetPageId="3" captionText="FIGURES 7–9. Metacosma echinophora, wing structure. 7. Wing venation, ♂, Gifu Pref., Japan, slide FK 1292. 8. Detail of anal fold on hindwing, Nagano Pref., Japan, slide FK 2142. 9. Wing venation, ♀, Nagano Pref., Japan, slide FK 2142." httpUri="https://zenodo.org/record/2611923/files/figure.png" pageId="1" pageNumber="132">7, 8</figureCitation>
); M
<subScript id="0E3A34D8A37B7B78FEADFA638A4FFA16" attach="left" box="[339,347,1420,1434]" fontSize="6" pageId="1" pageNumber="132">2</subScript>
and M
<subScript id="0E3A34D8A37B7B78FE57FA638AA5FA16" attach="left" box="[425,433,1420,1434]" fontSize="6" pageId="1" pageNumber="132">3</subScript>
nearly approximated at base and from lower angle of discal cell, CuA
<subScript id="0E3A34D8A37B7B78FB42FA638FD0FA16" attach="left" box="[1212,1220,1420,1434]" fontSize="6" pageId="1" pageNumber="132">1</subScript>
from lower margin of discal cell near angle. Abdomen: Hindmargin of S8 strongly bilobed in male (
<figureCitation id="0A852A18A37B7B78FBD6FA4B8F6EFA31" box="[1064,1146,1444,1469]" captionStart="FIGURES 10–13" captionStartId="5.[151,264,1911,1934]" captionTargetBox="[220,1325,193,1878]" captionTargetId="figure@5.[198,1388,193,1893]" captionTargetPageId="5" captionText="FIGURES 10–13. Metacosma echinophora, abdominal structure. 10. Male genitalia, paratype, Gifu Pref., Japan, slide FK 2137. 11. Postabdomen, ♂, paratype, Iriomote Is., Japan, slide FK 2133, unrolled preparation. 12. Female genitalia, paratype, Amami Is., Japan, slide FK 2136, ventral view. 13. Female genitalia, paratype, sterigma detail (arrow. entrance to ostium), paratype, Iriomote Is., Japan, slide FK 2182, dorsal view (T7 removed)." httpUri="https://zenodo.org/record/2611925/files/figure.png" pageId="1" pageNumber="132">Fig. 11</figureCitation>
). Male genitalia (
<figureCitation id="0A852A18A37B7B78FABCFA4B8E80FA31" box="[1346,1428,1444,1469]" captionStart="FIGURES 10–13" captionStartId="5.[151,264,1911,1934]" captionTargetBox="[220,1325,193,1878]" captionTargetId="figure@5.[198,1388,193,1893]" captionTargetPageId="5" captionText="FIGURES 10–13. Metacosma echinophora, abdominal structure. 10. Male genitalia, paratype, Gifu Pref., Japan, slide FK 2137. 11. Postabdomen, ♂, paratype, Iriomote Is., Japan, slide FK 2133, unrolled preparation. 12. Female genitalia, paratype, Amami Is., Japan, slide FK 2136, ventral view. 13. Female genitalia, paratype, sterigma detail (arrow. entrance to ostium), paratype, Iriomote Is., Japan, slide FK 2182, dorsal view (T7 removed)." httpUri="https://zenodo.org/record/2611925/files/figure.png" pageId="1" pageNumber="132">Fig. 10</figureCitation>
) with valva without basal process; cucullus with a bristled flap produced from dorsal margin near its apex, with one or two strong spines on the apex and three to six similar ones along the ventrodistal margin. Uncus a short process, often distally bifid. Socii elongate lobes with long setae, completely fused with diaphragma. Female genitalia (
<figureCitation id="0A852A18A37B7B78FF61F9DB8A3FF9C1" box="[159,299,1588,1613]" captionStart="FIGURES 10–13" captionStartId="5.[151,264,1911,1934]" captionTargetBox="[220,1325,193,1878]" captionTargetId="figure@5.[198,1388,193,1893]" captionTargetPageId="5" captionText="FIGURES 10–13. Metacosma echinophora, abdominal structure. 10. Male genitalia, paratype, Gifu Pref., Japan, slide FK 2137. 11. Postabdomen, ♂, paratype, Iriomote Is., Japan, slide FK 2133, unrolled preparation. 12. Female genitalia, paratype, Amami Is., Japan, slide FK 2136, ventral view. 13. Female genitalia, paratype, sterigma detail (arrow. entrance to ostium), paratype, Iriomote Is., Japan, slide FK 2182, dorsal view (T7 removed)." httpUri="https://zenodo.org/record/2611925/files/figure.png" pageId="1" pageNumber="132">Figs. 12, 13</figureCitation>
) with S7 heavily sclerotized, wide, divided into three areas: raised, semicircular median plate and variously shaped lateral plates. Sterigma funnel-shaped, fused with folded posterior apex of S7 (ostium bursae behind S7) (
<figureCitation id="0A852A18A37B7B78FED8F9938A69F919" box="[294,381,1660,1685]" captionStart="FIGURES 10–13" captionStartId="5.[151,264,1911,1934]" captionTargetBox="[220,1325,193,1878]" captionTargetId="figure@5.[198,1388,193,1893]" captionTargetPageId="5" captionText="FIGURES 10–13. Metacosma echinophora, abdominal structure. 10. Male genitalia, paratype, Gifu Pref., Japan, slide FK 2137. 11. Postabdomen, ♂, paratype, Iriomote Is., Japan, slide FK 2133, unrolled preparation. 12. Female genitalia, paratype, Amami Is., Japan, slide FK 2136, ventral view. 13. Female genitalia, paratype, sterigma detail (arrow. entrance to ostium), paratype, Iriomote Is., Japan, slide FK 2182, dorsal view (T7 removed)." httpUri="https://zenodo.org/record/2611925/files/figure.png" pageId="1" pageNumber="132">Fig. 13</figureCitation>
: arrow). Ductus bursae with long, anteriorly bifurcate sclerotization in anterior 3/4. Corpus bursae ovate or rounded, spinulose except around signum, with two, rather small horn-shaped signa.
</paragraph>
</subSubSection>
<subSubSection id="DAA46516A37B7B7BFF39F92B8957FC99" lastPageId="2" lastPageNumber="133" pageId="1" pageNumber="132" type="discussion">
<paragraph id="9201369DA37B7B78FF39F92B8BF4F81D" blockId="1.[151,1437,1186,2009]" pageId="1" pageNumber="132">
<emphasis id="A0CAEA8FA37B7B78FF39F92B8A2FF951" bold="true" box="[199,315,1732,1757]" pageId="1" pageNumber="132">Remarks.</emphasis>
<taxonomicName id="55BE4D1EA37B7B78FEBFF92A8AD6F950" authorityName="Kuznetzov" authorityYear="1985" box="[321,450,1733,1756]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA37B7B78FEBFF92A8AD6F950" box="[321,450,1733,1756]" italics="true" pageId="1" pageNumber="132">Metacosma</emphasis>
</taxonomicName>
is characterized by the male hindwing with a free elongate fold embracing a scale pencil from the base of the anal area (
<figureCitation id="0A852A18A37B7B78FE0FF907892AF88C" box="[497,574,1768,1793]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1940,1963]" captionTargetBox="[151,1437,844,1919]" captionTargetId="figure@2.[151,1437,844,1919]" captionTargetPageId="2" captionText="FIGURES 1–6. Metacosma echinophora. 1. Adult moth, ♂, paratype, Yakushima Is., Japan. 2. Notch on base of male antenna. 3. Lateral view of male head. 4. Larval habitat, resin dome constructed among winter buds of Pinus thunbergii, Gifu Pref., Japan. 5. Detail of resin dome. 6. lateral view of mature larva (photo. Akihiko Miyano)." httpUri="https://zenodo.org/record/2611921/files/figure.png" pageId="1" pageNumber="132">Figs. 1</figureCitation>
,
<figureCitation id="0A852A18A37B7B78FDB2F9068963F88C" box="[588,631,1768,1792]" captionStart="FIGURES 7–9" captionStartId="3.[151,264,1135,1158]" captionTargetBox="[200,1402,281,1079]" captionTargetId="figure@3.[150,1436,193,1115]" captionTargetPageId="3" captionText="FIGURES 7–9. Metacosma echinophora, wing structure. 7. Wing venation, ♂, Gifu Pref., Japan, slide FK 1292. 8. Detail of anal fold on hindwing, Nagano Pref., Japan, slide FK 2142. 9. Wing venation, ♀, Nagano Pref., Japan, slide FK 2142." httpUri="https://zenodo.org/record/2611923/files/figure.png" pageId="1" pageNumber="132">7, 8</figureCitation>
); the cucullus with a bristled flap produced from dorsal margin near the apex and with one or two strong spines on the apex and three to six similar ones along the ventrodistal margin (
<figureCitation id="0A852A18A37B7B78FA91F8E38BA2F8C4" captionStart="FIGURES 10–13" captionStartId="5.[151,264,1911,1934]" captionTargetBox="[220,1325,193,1878]" captionTargetId="figure@5.[198,1388,193,1893]" captionTargetPageId="5" captionText="FIGURES 10–13. Metacosma echinophora, abdominal structure. 10. Male genitalia, paratype, Gifu Pref., Japan, slide FK 2137. 11. Postabdomen, ♂, paratype, Iriomote Is., Japan, slide FK 2133, unrolled preparation. 12. Female genitalia, paratype, Amami Is., Japan, slide FK 2136, ventral view. 13. Female genitalia, paratype, sterigma detail (arrow. entrance to ostium), paratype, Iriomote Is., Japan, slide FK 2182, dorsal view (T7 removed)." httpUri="https://zenodo.org/record/2611925/files/figure.png" pageId="1" pageNumber="132">Fig. 10</figureCitation>
); and the seventh sternite of the female consisting of raised, semicircular median plate and variously shaped lateral plates (
<figureCitation id="0A852A18A37B7B78FEC2F8BB8ADFF8E1" box="[316,459,1876,1901]" captionStart="FIGURES 10–13" captionStartId="5.[151,264,1911,1934]" captionTargetBox="[220,1325,193,1878]" captionTargetId="figure@5.[198,1388,193,1893]" captionTargetPageId="5" captionText="FIGURES 10–13. Metacosma echinophora, abdominal structure. 10. Male genitalia, paratype, Gifu Pref., Japan, slide FK 2137. 11. Postabdomen, ♂, paratype, Iriomote Is., Japan, slide FK 2133, unrolled preparation. 12. Female genitalia, paratype, Amami Is., Japan, slide FK 2136, ventral view. 13. Female genitalia, paratype, sterigma detail (arrow. entrance to ostium), paratype, Iriomote Is., Japan, slide FK 2182, dorsal view (T7 removed)." httpUri="https://zenodo.org/record/2611925/files/figure.png" pageId="1" pageNumber="132">Figs. 12, 13</figureCitation>
). All these character states appear to represent synapomorphies for members of the genus.
</paragraph>
<paragraph id="9201369DA37B7B7BFF39F8738ADDFECC" blockId="1.[151,1437,1186,2009]" lastBlockId="2.[151,1437,151,789]" lastPageId="2" lastPageNumber="133" pageId="1" pageNumber="132">
The genus belongs to the
<taxonomicName id="55BE4D1EA37B7B78FE0DF873894BF838" authorityName="Stephens" authorityYear="1829" box="[499,607,1948,1972]" class="Insecta" family="Tortricidae" genus="Spilonota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA37B7B78FE0DF873894BF838" box="[499,607,1948,1972]" italics="true" pageId="1" pageNumber="132">Spilonota</emphasis>
</taxonomicName>
group, an informal genus group proposed by
<bibRefCitation id="F62F4B6CA37B7B78FB8DF8738E04F839" author="Horak, M." box="[1139,1296,1948,1973]" pageId="1" pageNumber="132" pagination="1 - 522" refId="ref4300" refString="Horak, M. (2006) Olethreutine moths of Australia (Lepidoptera: Tortricidae). Monographs on Australian Lepidoptera, 10, i-x + 1 - 522." type="journal article" year="2006">Horak (2006)</bibRefCitation>
, comprising
<taxonomicName id="55BE4D1EA37B7B78FF69F82E8AADF855" authority="Walsingham" authorityName="Walsingham" authorityYear="1907" box="[151,441,1984,2009]" class="Insecta" family="Tortricidae" genus="Eccoptocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA37B7B78FF69F82E8A31F854" box="[151,293,1985,2008]" italics="true" pageId="1" pageNumber="132">Eccoptocera</emphasis>
Walsingham
</taxonomicName>
,
<taxonomicName id="55BE4D1EA37B7B78FE3AF82F89C6F855" authority="Meyrick" authorityName="Meyrick" authorityYear="1882" box="[452,722,1984,2009]" class="Insecta" family="Tortricidae" genus="Hendecasticha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA37B7B78FE3AF82F897EF854" box="[452,618,1984,2008]" italics="true" pageId="1" pageNumber="132">Hendecasticha</emphasis>
Meyrick
</taxonomicName>
,
<taxonomicName id="55BE4D1EA37B7B78FD23F82E88D4F855" authority="Meyrick" authorityName="Meyrick" authorityYear="1911" box="[733,960,1984,2009]" class="Insecta" family="Tortricidae" genus="Hermenias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA37B7B78FD23F82E884CF854" box="[733,856,1985,2008]" italics="true" pageId="1" pageNumber="132">Hermenias</emphasis>
Meyrick
</taxonomicName>
,
<taxonomicName id="55BE4D1EA37B7B78FC35F82F8F8FF855" authority="Meyrick" authorityName="Meyrick" authorityYear="1882" box="[971,1179,1984,2009]" class="Insecta" family="Tortricidae" genus="Holocola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA37B7B78FC35F82F8F27F854" box="[971,1075,1984,2008]" italics="true" pageId="1" pageNumber="132">Holocola</emphasis>
Meyrick
</taxonomicName>
,
<taxonomicName id="55BE4D1EA37B7B78FB5BF82F8E8CF855" authority="Diakonoff" authorityName="Diakonoff" authorityYear="1984" box="[1189,1432,1984,2009]" class="Insecta" family="Tortricidae" genus="Lepidunca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA37B7B78FB5BF82F8E0FF854" box="[1189,1307,1984,2008]" italics="true" pageId="1" pageNumber="132">Lepidunca</emphasis>
Diakonoff
</taxonomicName>
,
<taxonomicName id="55BE4D1EA3787B7BFF69FF778A56FF3C" authority="Berg" authorityName="Berg" authorityYear="1899" box="[151,322,152,176]" class="Insecta" family="Tortricidae" genus="Parienia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFF69FF778BEEFF23" box="[151,250,152,175]" italics="true" pageId="2" pageNumber="133">Parienia</emphasis>
Berg
</taxonomicName>
,
<taxonomicName id="55BE4D1EA3787B7BFEA8FF778955FF3C" authority="Meyrick" authorityName="Meyrick" authorityYear="1882" box="[342,577,151,176]" class="Insecta" family="Tortricidae" genus="Protithona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFEA8FF778ADAFF3C" box="[342,462,152,176]" italics="true" pageId="2" pageNumber="133">Protithona</emphasis>
Meyrick
</taxonomicName>
,
<taxonomicName id="55BE4D1EA3787B7BFDABFF77882DFF3C" authority="Stephens" authorityName="Stephens" authorityYear="1829" box="[597,825,151,176]" class="Insecta" family="Tortricidae" genus="Spilonota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFDABFF7789D4FF3C" box="[597,704,152,176]" italics="true" pageId="2" pageNumber="133">Spilonota</emphasis>
Stephens
</taxonomicName>
,
<taxonomicName id="55BE4D1EA3787B7BFCB3FF778F44FF3C" authority="Meyrick" authorityName="Meyrick" authorityYear="1888" box="[845,1104,151,176]" class="Insecta" family="Tortricidae" genus="Strepsicrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFCB3FF7788CAFF23" box="[845,990,152,175]" italics="true" pageId="2" pageNumber="133">Strepsicrates</emphasis>
Meyrick
</taxonomicName>
, and
<taxonomicName id="55BE4D1EA3787B7BFB63FF778E89FF3C" authority="Diakonoff." authorityName="Diakonoff." box="[1181,1437,151,176]" class="Insecta" family="Tortricidae" genus="Xenosocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFB63FF778E06FF23" box="[1181,1298,152,175]" italics="true" pageId="2" pageNumber="133">Xenosocia</emphasis>
Diakonoff.
</taxonomicName>
<taxonomicName id="55BE4D1EA3787B7BFF69FF528A0DFF58" authorityName="Kuznetzov" authorityYear="1985" box="[151,281,189,212]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFF69FF528A0DFF58" box="[151,281,189,212]" italics="true" pageId="2" pageNumber="133">Metacosma</emphasis>
</taxonomicName>
shares with members of the
<taxonomicName id="55BE4D1EA3787B7BFD9BFF5389C5FF58" authorityName="Stephens" authorityYear="1829" box="[613,721,188,212]" class="Insecta" family="Tortricidae" genus="Spilonota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFD9BFF5389C5FF58" box="[613,721,188,212]" italics="true" pageId="2" pageNumber="133">Spilonota</emphasis>
</taxonomicName>
group the following derived traits presented by
<bibRefCitation id="F62F4B6CA3787B7BFB04FF538E8CFF59" author="Horak, M." box="[1274,1432,188,213]" pageId="2" pageNumber="133" pagination="1 - 522" refId="ref4300" refString="Horak, M. (2006) Olethreutine moths of Australia (Lepidoptera: Tortricidae). Monographs on Australian Lepidoptera, 10, i-x + 1 - 522." type="journal article" year="2006">Horak (2006)</bibRefCitation>
: antennal notch in male, extended anal region in male hindwing with upturned anal margin folded over long scale pencil from base, funnel-shaped sterigma fused with S7, and a tendency for signa to be reduced or lost and bursa to become strongly spinulose.
</paragraph>
<paragraph id="9201369DA3787B7BFF39FEA28881FDB0" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="133">
In addition to the above genera the North African
<taxonomicName id="55BE4D1EA3787B7BFCC5FEA38F7AFEE9" authority="Razowski" authorityName="Razowski" authorityYear="1961" box="[827,1134,332,357]" class="Insecta" family="Tortricidae" genus="Cirrilaspeyresia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFCC5FEA388E5FEE8" box="[827,1009,332,356]" italics="true" pageId="2" pageNumber="133">Cirrilaspeyresia</emphasis>
Razowski
</taxonomicName>
(
<typeStatus id="4D05883FA3787B7BFB7BFEA38FA3FEE8" box="[1157,1207,332,356]" pageId="2" pageNumber="133">type</typeStatus>
species:
<taxonomicName id="55BE4D1EA3787B7BFAD2FEA38A6AFE04" authority="Kennel" authorityName="Kennel" class="Insecta" family="Tortricidae" genus="Euxanthis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="imbecillana">
<emphasis id="A0CAEA8FA3787B7BFAD2FEA38A08FE04" italics="true" pageId="2" pageNumber="133">Euxanthis imbecillana</emphasis>
Kennel
</taxonomicName>
) may belong to the
<taxonomicName id="55BE4D1EA3787B7BFD74FE9F89E2FE04" authorityName="Stephens" authorityYear="1829" box="[650,758,368,392]" class="Insecta" family="Tortricidae" genus="Spilonota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFD74FE9F89E2FE04" box="[650,758,368,392]" italics="true" pageId="2" pageNumber="133">Spilonota</emphasis>
</taxonomicName>
group and may be most closely allied to
<taxonomicName id="55BE4D1EA3787B7BFAEAFE9E8E81FE04" authorityName="Kuznetzov" authorityYear="1985" box="[1300,1429,369,392]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFAEAFE9E8E81FE04" box="[1300,1429,369,392]" italics="true" pageId="2" pageNumber="133">Metacosma</emphasis>
</taxonomicName>
.
<taxonomicName id="55BE4D1EA3787B7BFF69FE7B8957FE21" authority="(Kennel)" authorityName="Kennel" baseAuthorityName="Kennel" box="[151,579,404,429]" class="Insecta" family="Tortricidae" genus="Cirrilaspeyresia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="imbecillana">
<emphasis id="A0CAEA8FA3787B7BFF69FE7B8ACDFE20" box="[151,473,404,428]" italics="true" pageId="2" pageNumber="133">Cirrilaspeyresia imbecillana</emphasis>
(Kennel)
</taxonomicName>
shares with the
<taxonomicName id="55BE4D1EA3787B7BFD07FE7B8871FE20" authorityName="Stephens" authorityYear="1829" box="[761,869,404,428]" class="Insecta" family="Tortricidae" genus="Spilonota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFD07FE7B8871FE20" box="[761,869,404,428]" italics="true" pageId="2" pageNumber="133">Spilonota</emphasis>
</taxonomicName>
group the tendency towards reduction of the signa (absent in
<taxonomicName id="55BE4D1EA3787B7BFEEAFE578AA8FE5C" box="[276,444,440,464]" class="Insecta" family="Tortricidae" genus="Cirrilaspeyresia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="imbecillana">
<emphasis id="A0CAEA8FA3787B7BFEEAFE578AA8FE5C" box="[276,444,440,464]" italics="true" pageId="2" pageNumber="133">C. imbecillana</emphasis>
</taxonomicName>
) and the funnel-shaped sterigma fused with S7 (
<bibRefCitation id="F62F4B6CA3787B7BFC02FE588E06FE5C" author="Razowski, J." box="[1020,1298,439,464]" pageId="2" pageNumber="133" pagination="661 - 697" refId="ref4779" refString="Razowski, J. (1961) Notes on some little known Tortricidae (Lepidoptera). Acta Zoologica Cracoviensia, 5 (14), 661 - 697." type="journal article" year="1961">Razowski 1961: fig. 25</bibRefCitation>
), but in the drawing of the adult head the antenna has no notch at its base (
<bibRefCitation id="F62F4B6CA3787B7BFC8CFE338FA7FE78" author="Razowski, J." box="[882,1203,476,501]" pageId="2" pageNumber="133" pagination="661 - 697" refId="ref4779" refString="Razowski, J. (1961) Notes on some little known Tortricidae (Lepidoptera). Acta Zoologica Cracoviensia, 5 (14), 661 - 697." type="journal article" year="1961">Razowski, 1961: 675, fig. 3</bibRefCitation>
), though there is no indication of its sex. Moreover, this species is similar to members of
<taxonomicName id="55BE4D1EA3787B7BFC67FDEE8F0EFD94" authorityName="Kuznetzov" authorityYear="1985" box="[921,1050,513,536]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFC67FDEE8F0EFD94" box="[921,1050,513,536]" italics="true" pageId="2" pageNumber="133">Metacosma</emphasis>
</taxonomicName>
in having the cucullus with strong spines along the outer and ventral margins (
<bibRefCitation id="F62F4B6CA3787B7BFD81FDCB8893FDB0" author="Razowski, J." box="[639,903,548,573]" pageId="2" pageNumber="133" pagination="661 - 697" refId="ref4779" refString="Razowski, J. (1961) Notes on some little known Tortricidae (Lepidoptera). Acta Zoologica Cracoviensia, 5 (14), 661 - 697." type="journal article" year="1961">Razowski 1961: fig. 11</bibRefCitation>
).
</paragraph>
<paragraph id="9201369DA3787B7BFF39FDA88957FC99" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="133">
Prior to this report, five
<taxonomicName id="55BE4D1EA3787B7BFE12FDA6897AFDEC" authorityName="Kuznetzov" authorityYear="1985" box="[492,622,585,608]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFE12FDA6897AFDEC" box="[492,622,585,608]" italics="true" pageId="2" pageNumber="133">Metacosma</emphasis>
</taxonomicName>
species were known mainly from North
<collectingCountry id="EAA9760DA3787B7BFBA9FDA78FD2FDEC" box="[1111,1222,583,608]" name="China" pageId="2" pageNumber="133">East Asia</collectingCountry>
(see the checklist below). However, the generic assignment of
<taxonomicName id="55BE4D1EA3787B7BFD77FD8288B9FD08" authority="Kuznetzov" authorityName="Kuznetzov" authorityYear="1988" box="[649,941,621,644]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="miratorana">
<emphasis id="A0CAEA8FA3787B7BFD77FD82883AFD08" box="[649,814,621,644]" italics="true" pageId="2" pageNumber="133">M. miratorana</emphasis>
Kuznetzov
</taxonomicName>
is questionable, as no synapomorphies have been identified that would support this assignment, though the species is known only from males. Furthermore,
<taxonomicName id="55BE4D1EA3787B7BFA83FD7E8A03FD40" authorityName="Kuznetzov" authorityYear="1988" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="miratorana">
<emphasis id="A0CAEA8FA3787B7BFA83FD7E8A03FD40" italics="true" pageId="2" pageNumber="133">M. miratorana</emphasis>
</taxonomicName>
differs from the other species of
<taxonomicName id="55BE4D1EA3787B7BFD5CFD5A8830FD40" authorityName="Kuznetzov" authorityYear="1985" box="[674,804,693,716]" class="Insecta" family="Tortricidae" genus="Metacosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFD5CFD5A8830FD40" box="[674,804,693,716]" italics="true" pageId="2" pageNumber="133">Metacosma</emphasis>
</taxonomicName>
in possessing semioval socii and long basal processes (
<bibRefCitation id="F62F4B6CA3787B7BFF60FD378AE0FD7C" author="Kuznetzov, V. I." box="[158,500,727,752]" pageId="2" pageNumber="133" pagination="615 - 631" refId="ref4585" refString="Kuznetzov, V. I. (1988) New species of tortricid moths of the subfamily Olethreutinae (Lepidoptera, Tortricidae) of the fauna of North Vietnam. Entomologicheskoe Obozrenie, 67, 615 - 631. [in Russian]" type="journal article" year="1988">Kuznetzov 1988: 629, fig. 21</bibRefCitation>
). Superficially it is reminiscent of some species of
<taxonomicName id="55BE4D1EA3787B7BFBBCFD378E24FD7C" authority="Meyrick" authorityName="Meyrick" authorityYear="1895" box="[1090,1328,727,752]" class="Insecta" family="Tortricidae" genus="Gypsonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A0CAEA8FA3787B7BFBBCFD378FDCFD63" box="[1090,1224,728,751]" italics="true" pageId="2" pageNumber="133">Gypsonoma</emphasis>
Meyrick
</taxonomicName>
(see also
<bibRefCitation id="F62F4B6CA3787B7BFF69FD138A7CFC98" author="Nedoshivina, S. V." box="[151,360,764,789]" pageId="2" pageNumber="133" pagination="335 - 347" refId="ref4692" refString="Nedoshivina, S. V. (2010) A catalogue of type specimens of the Tortricidae described by V. I. Kuznetzov from Vietnam and deposited in the Zoological Institute, St. Petersburg. Atalanta, 41 (3 / 4), 335 - 347, pls. 7 - 9." type="journal article" year="2010">Nedoshivina 2010</bibRefCitation>
: 343, pl. 9, fig. 67).
</paragraph>
</subSubSection>
</treatment>
</document>
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