319 lines
34 KiB
XML
319 lines
34 KiB
XML
<document id="22ADE97BABEE4F75958CB7AD46A25D43" ID-DOI="10.1649/828.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="10105234" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630173236196" checkinUser="tatiana" docAuthor="Morón, Miguel-Angel & Riley, Edward G." docDate="2005" docId="196C87F3FF8FFFDC7A6EABD4FE56B2BC" docLanguage="en" docName="828.1.pdf.imf" docOrigin="The Coleopterists Bulletin 59 (3)" docSource="http://dx.doi.org/10.1649/828.1" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Phyllophaga (Phyllophaga) cruces Moron and Riley, new species" docType="treatment" docVersion="4" lastPageNumber="399" masterDocId="E555FF8BFF89FFD47B6FA94AFFD0B057" masterDocTitle="Four New Species of Phyllophaga from Mexico and Central America (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="399" masterPageNumber="391" pageNumber="397" updateTime="1699632931453" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BB0CC2B9A5D9F3AA8AAF1FFCB65F4E9C">
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<mods:title id="37042964977A0D02942CB6B9577848B6">Four New Species of Phyllophaga from Mexico and Central America (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:namePart id="1E2DFEF55272327EFB3A7F13C66DCA0D">Morón, Miguel-Angel</mods:namePart>
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<mods:roleTerm id="1EB558A55F174B9C8B0B2088021A10D2">Author</mods:roleTerm>
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<mods:namePart id="E78E3DD53215022C63F98B57A353A15A">Riley, Edward G.</mods:namePart>
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<mods:typeOfResource id="396D4AF14320910B1C38CB45D57D3F1B">text</mods:typeOfResource>
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<mods:title id="539523076A230D3C20BC09D607FBDB8E">The Coleopterists Bulletin</mods:title>
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<mods:part id="7799B404BCF546DC5C08AACD53C2178B">
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<mods:date id="72605EAEF30B7DC86F2F0FA11DD717E6">2005</mods:date>
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<mods:number id="ED9EE40A2536428EE6A212352761FF46">2005-09-30</mods:number>
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<mods:number id="2B64F01F7806F11C2C8E9A32D75925A7">59</mods:number>
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<mods:url id="8B7ED19C2F039A566A45A06C388938CD">http://dx.doi.org/10.1649/828.1</mods:url>
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<mods:classification id="366D0B18C2E1B180922051634BF66E1A">journal article</mods:classification>
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<mods:identifier id="55537A05B9C2681DFAC641621DD139EA" type="DOI">10.1649/828.1</mods:identifier>
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<mods:identifier id="6E56D7532FCB950CBE57EDE5ED206263" type="ISSN">1938-4394</mods:identifier>
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<treatment id="196C87F3FF8FFFDC7A6EABD4FE56B2BC" ID-DOI="http://doi.org/10.5281/zenodo.5411956" ID-Zenodo-Dep="5411956" LSID="urn:lsid:plazi:treatment:196C87F3FF8FFFDC7A6EABD4FE56B2BC" httpUri="http://treatment.plazi.org/id/196C87F3FF8FFFDC7A6EABD4FE56B2BC" lastPageId="8" lastPageNumber="399" pageId="6" pageNumber="397">
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<subSubSection id="D9DF656EFF8FFFD27A6EABD4FC4AB2E4" box="[257,922,670,691]" pageId="6" pageNumber="397" type="nomenclature">
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<paragraph id="917A36E5FF8FFFD27A6EABD4FC4AB2E4" blockId="6.[257,922,670,718]" box="[257,922,670,691]" pageId="6" pageNumber="397">
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<heading id="CA328189FF8FFFD27A6EABD4FC4AB2E4" box="[257,922,670,691]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="397" reason="2">
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<emphasis id="A3B1EAF7FF8FFFD27A6EABD4FC4AB2E4" bold="true" box="[257,922,670,691]" pageId="6" pageNumber="397">
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<taxonomicName id="56C54D66FF8FFFD27A6EABD4FCC7B2E4" authority="Moron and Riley" authorityName="Moron and Riley" box="[257,791,670,691]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="397" phylum="Arthropoda" rank="species" species="cruces" status="sp. nov." subGenus="Phyllophaga">
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<emphasis id="A3B1EAF7FF8FFFD27A6EABD4FEAEB2E4" bold="true" box="[257,382,670,691]" italics="true" pageId="6" pageNumber="397">Phyllophaga</emphasis>
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(
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<emphasis id="A3B1EAF7FF8FFFD27AE1ABD4FDDBB2E4" bold="true" box="[398,523,670,691]" italics="true" pageId="6" pageNumber="397">Phyllophaga</emphasis>
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)
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<emphasis id="A3B1EAF7FF8FFFD27974ABD4FD8BB2E4" bold="true" box="[539,603,670,691]" italics="true" pageId="6" pageNumber="397">cruces</emphasis>
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Morón and Riley
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</taxonomicName>
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,
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<taxonomicNameLabel id="B882578CFF8FFFD2784CABD4FC4AB2E4" box="[803,922,670,691]" pageId="6" pageNumber="397" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D9DF656EFF8FFFDC7962ABF3FC76B080" lastPageId="8" lastPageNumber="399" pageId="6" pageNumber="397" type="description">
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<paragraph id="917A36E5FF8FFFD27962ABF3FD5FB299" blockId="6.[257,922,670,718]" box="[525,655,697,718]" pageId="6" pageNumber="397">
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(
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<figureCitation id="09FE2A60FF8FFFD2797BABF3FD58B299" box="[532,648,697,718]" captionStart="Figs" captionStartId="7.[169,208,1115,1134]" captionTargetBox="[153,975,195,1094]" captionTargetId="figure-183@7.[149,975,194,1094]" captionTargetPageId="7" captionText="Figs. 19–24. Phyllophaga cruces. 19) Inner protarsal claw, male; 20) genital capsule, lateral view; 21) female genital plates, ventral view; 22) parameres, distal view; 23) parameres, ventral view; 24) tectum, phallobase and parameres, dorsal view. Scale lines: Fig. 19 ¼ 0.5 mm, Figs. 20–24 ¼ 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10105242" httpUri="https://zenodo.org/record/10105242/files/figure.png" pageId="6" pageNumber="397">Figs. 19–24</figureCitation>
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)
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</paragraph>
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<paragraph id="917A36E5FF8FFFD27BAAABA8FDE2B3B3" blockId="6.[175,1007,738,1570]" pageId="6" pageNumber="397">
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<emphasis id="A3B1EAF7FF8FFFD27BAAABA8FE46B2A2" bold="true" box="[197,406,738,757]" pageId="6" pageNumber="397">
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Description.
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<typeStatus id="4E7E8847FF8FFFD27A52ABA8FE41B2A2" box="[317,401,738,757]" pageId="6" pageNumber="397" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male.Body and legs shiny reddish brown,elytra shiny yellowish brown. Clypeus glabrous, 2.6
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<emphasis id="A3B1EAF7FF8FFFD27A08ABB3FEA9B35B" bold="true" box="[359,377,761,780]" pageId="6" pageNumber="397">3</emphasis>
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wider than long, anterior border strongly elevated, anterior margin nearly straight, disk surface wide and deeply concave, with sparse, shallow round punctures. Frontoclypeal suture clearly impressed and closely sinuate on shallow middle depression. Frons 2.3
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<emphasis id="A3B1EAF7FF8FFFD27812AA63FC5FB36B" bold="true" box="[893,911,809,828]" pageId="6" pageNumber="397">3</emphasis>
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wider than long, widely convex, with irregular round punctuation, and some short erect setae on disk. Antenna 10-segmented, with three segmented club, lamellae 1.7
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<emphasis id="A3B1EAF7FF8FFFD279F8AA13FD79B33B" bold="true" box="[663,681,857,876]" pageId="6" pageNumber="397">3</emphasis>
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longer than length of preceding six segments combined; segments 3 to 6 progressively longer, with rounded or conical prominences on anterior sides; segment 7 wider than long, with acute prominence on anterior side. Frons 2.3
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<emphasis id="A3B1EAF7FF8FFFD278C6AAC3FC6BB3CB" bold="true" box="[937,955,905,924]" pageId="6" pageNumber="397">3</emphasis>
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wider than dorsal diameter of eye.Eye canthus long and narrow, with 14 setae.Labrum reniform,shallowly concave, with scattered slender setae along borders. Mentum nearly flat, impunctate, with scarce, slender setae at sides, anterior border straight.
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</paragraph>
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<paragraph id="917A36E5FF8FFFD27BA8AAA3FDE6B564" blockId="6.[175,1007,738,1570]" pageId="6" pageNumber="397">
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Pronotum 1.6
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<emphasis id="A3B1EAF7FF8FFFD27A56AAA2FE9BB3AC" bold="true" box="[313,331,1000,1019]" pageId="6" pageNumber="397">3</emphasis>
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wider than long and 2.8
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<emphasis id="A3B1EAF7FF8FFFD27975AAA2FDFCB3AC" bold="true" box="[538,556,1000,1019]" pageId="6" pageNumber="397">3</emphasis>
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wider than frons. Pronotal disk glabrous, with deep, round punctures irregularly separated by 1–4 diameters; lateral borders widely curved, lateral marginal bead crenulate, mainly on posterior half, with scarce slender, long setae; basal bead indicated by punctures on middle third; anterior angles obtuse, rounded; posterior angles nearly straight, weakly prominent. Scutellum 1.4
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<emphasis id="A3B1EAF7FF8FFFD27943AD02FDEEB40C" bold="true" box="[556,574,1096,1115]" pageId="6" pageNumber="397">3</emphasis>
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wider than long, with 8 punctures irregularly distributed, and anterior border widely sinuate. Elytron 2.4
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<emphasis id="A3B1EAF7FF8FFFD279CAAD2AFD67B424" bold="true" box="[677,695,1120,1139]" pageId="6" pageNumber="397">3</emphasis>
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longer than wide, glabrous, densely and irregularly punctate; epipleural border progressively narrowed toward apex, with erect, slender setae; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Pterosternum with many yellowish, long setae. Visible abdominal sternites 2 to 4 widely depressed at middle with wide, shallow longitudinal furrow; sternite 5 slightly concave on anterior central border, shiny, with scattered setae; anal plate large, shiny, shallowly concave, with slender setae toward sides and posterior border. Propygidium shiny, sparsely punctate. Pygidium shiny, widely convex, with scarce, shallow punctures, irregularly distributed; apical margin with 28 slender setae; basal margin effaced at middle.
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</paragraph>
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<paragraph id="917A36E5FF8FFFD37BA8AC72FDE5B51F" blockId="6.[175,1007,738,1570]" lastBlockId="7.[148,978,1238,1570]" lastPageId="7" lastPageNumber="398" pageId="6" pageNumber="397">
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Protibia shorter than protarsus (0.7:1), with two large teeth and one basal small tooth on external border, preapical spur acute, straight, nearly as long as 2nd protarsomere. Mesotibia with one oblique, well marked, setiferous carina on external side; upper apical spur with acute apex, 1.2
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<emphasis id="A3B1EAF7FF8FFFD278B2AC2DFC3FB52D" bold="true" box="[989,1007,1383,1402]" pageId="6" pageNumber="397">3</emphasis>
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longer than lower acute spur. Metatibia slightly shorter than metatarsus (0.8:1), with one oblique setiferous carina on external side; upper apical spur articulated, curved, sharply pointed, longer than basal metatarsomere (1.4:1), and 1.3 longer than lower spur; lower apical spur articulated, curved, apex acute. Protarsomeres 1–4 semicylindrical, elongate, with subapical, inner, rounded projection. Meso- and metatarsomeres semicylindrical, elongate, each with enlarged apex, some setae apically, and two fine, keels with scarce setae along ventral side. Tarsal claws dentate, with ventral tooth narrow, acute, located toward base (
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<figureCitation id="09FE2A60FF8FFFD2791BAF45FD67B675" box="[628,695,1551,1570]" captionStart="Figs" captionStartId="7.[169,208,1115,1134]" captionTargetBox="[153,975,195,1094]" captionTargetId="figure-183@7.[149,975,194,1094]" captionTargetPageId="7" captionText="Figs. 19–24. Phyllophaga cruces. 19) Inner protarsal claw, male; 20) genital capsule, lateral view; 21) female genital plates, ventral view; 22) parameres, distal view; 23) parameres, ventral view; 24) tectum, phallobase and parameres, dorsal view. Scale lines: Fig. 19 ¼ 0.5 mm, Figs. 20–24 ¼ 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10105242" httpUri="https://zenodo.org/record/10105242/files/figure.png" pageId="6" pageNumber="397">Fig. 19</figureCitation>
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). Genital capsule with long, wide parameres dorsally and ventrally fused at phallobase, apex enlarged, narrowed, acute, divergent, and curved downward; ventro-basal border with 2 narrow, curved projections (
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<figureCitation id="09FE2A60FF8EFFD37858ADA7FC18B557" box="[823,968,1261,1280]" captionStart="Figs" captionStartId="7.[169,208,1115,1134]" captionTargetBox="[153,975,195,1094]" captionTargetId="figure-183@7.[149,975,194,1094]" captionTargetPageId="7" captionText="Figs. 19–24. Phyllophaga cruces. 19) Inner protarsal claw, male; 20) genital capsule, lateral view; 21) female genital plates, ventral view; 22) parameres, distal view; 23) parameres, ventral view; 24) tectum, phallobase and parameres, dorsal view. Scale lines: Fig. 19 ¼ 0.5 mm, Figs. 20–24 ¼ 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10105242" httpUri="https://zenodo.org/record/10105242/files/figure.png" pageId="7" pageNumber="398">Figs. 20, 22–24</figureCitation>
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). Aedeagus with sclerotized, tube like support and preapical curved spine. Tectum wide, weakly bicallose. Length of genital capsule from apex of parameres to border of basal piece:
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<quantity id="563D9B00FF8EFFD37807AC57FC4CB567" box="[872,924,1309,1328]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="398" unit="mm" value="6.0">6 mm</quantity>
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. Total body length: 19.0 mm. Humeral width:
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||
<quantity id="563D9B00FF8EFFD37A82AC7FFDE1B51F" box="[493,561,1333,1352]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="7" pageNumber="398" unit="mm" value="8.1">8.1 mm</quantity>
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.
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</paragraph>
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<caption id="C5BA666DFF8EFFD37BC6AD11FECCB4E1" ID-DOI="http://doi.org/10.5281/zenodo.10105242" ID-Zenodo-Dep="10105242" httpUri="https://zenodo.org/record/10105242/files/figure.png" pageId="7" pageNumber="398" startId="7.[169,208,1115,1134]" targetBox="[153,975,195,1094]" targetPageId="7">
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<paragraph id="917A36E5FF8EFFD37BC6AD11FECCB4E1" blockId="7.[148,978,1115,1207]" pageId="7" pageNumber="398">
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<emphasis id="A3B1EAF7FF8EFFD37BC6AD11FEC5B439" bold="true" box="[169,277,1115,1134]" pageId="7" pageNumber="398">Figs. 19–24.</emphasis>
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<taxonomicName id="56C54D66FF8EFFD37A5FAD11FE09B439" box="[304,473,1115,1134]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="398" phylum="Arthropoda" rank="species" species="cruces">
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||
<emphasis id="A3B1EAF7FF8EFFD37A5FAD11FE09B439" box="[304,473,1115,1134]" italics="true" pageId="7" pageNumber="398">Phyllophaga cruces</emphasis>
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</taxonomicName>
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||
.
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||
<emphasis id="A3B1EAF7FF8EFFD37A8BAD11FDD0B439" bold="true" box="[484,512,1115,1134]" pageId="7" pageNumber="398">19)</emphasis>
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Inner protarsal claw, male;
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<emphasis id="A3B1EAF7FF8EFFD3799FAD11FCDDB439" bold="true" box="[752,781,1115,1134]" pageId="7" pageNumber="398">20)</emphasis>
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||
genital capsule, lateral view;
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||
<emphasis id="A3B1EAF7FF8EFFD37BA5AD39FF36B4D1" bold="true" box="[202,230,1139,1158]" pageId="7" pageNumber="398">21)</emphasis>
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female genital plates, ventral view;
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||
<emphasis id="A3B1EAF7FF8EFFD3794EAD39FDEDB4D1" bold="true" box="[545,573,1139,1158]" pageId="7" pageNumber="398">22)</emphasis>
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parameres, distal view;
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<emphasis id="A3B1EAF7FF8EFFD3787EAD39FCFDB4D1" bold="true" box="[785,813,1139,1158]" pageId="7" pageNumber="398">23)</emphasis>
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parameres, ventral view;
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<emphasis id="A3B1EAF7FF8EFFD37BA5ADC1FF37B4C9" bold="true" box="[202,231,1163,1182]" pageId="7" pageNumber="398">24)</emphasis>
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tectum, phallobase and parameres, dorsal view. Scale lines: Fig. 19
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<emphasis id="A3B1EAF7FF8EFFD3782FADC0FC80B4C8" bold="true" box="[832,848,1162,1183]" italics="true" pageId="7" pageNumber="398">¼</emphasis>
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0.5 mm, Figs. 20–24
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||
<emphasis id="A3B1EAF7FF8EFFD37BA1ADE8FF0EB4E0" bold="true" box="[206,222,1186,1207]" italics="true" pageId="7" pageNumber="398">¼</emphasis>
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1 mm.
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||
</paragraph>
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||
</caption>
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||
<paragraph id="917A36E5FF8EFFD37BC6AC1AFDE5B675" blockId="7.[148,978,1238,1570]" pageId="7" pageNumber="398">
|
||
<emphasis id="A3B1EAF7FF8EFFD37BC6AC1AFF2BB534" bold="true" box="[169,251,1360,1379]" pageId="7" pageNumber="398">
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<typeStatus id="4E7E8847FF8EFFD37BC6AC1AFF27B534" box="[169,247,1360,1379]" pageId="7" pageNumber="398" type="allotype">Allotype</typeStatus>
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.
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</emphasis>
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Female. Similar to male except as follows: anterior border of clypeus briefly elevated; antennal club as long as preceding six segments combined; pronotal punctation deeper. Visible abdominal sternites 2 to 5 convex, with scattered, setiferous punctures, more abundant on sternite 5; anal plate large, widely convex, with scattered slender setae. Both apical spurs of metatibia widened, curved, with rounded apices. Tarsal claws dentate, with lower tooth long, acute, located near middle of ventral border. Ventral genital plates well sclerotized, nearly symmetrical, with rounded lobes on posterior border; dorsal genital plates fused laterally with ventral plates, with wide, rounded apices and some short setae on distal borders (
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<figureCitation id="09FE2A60FF8EFFD3783CACBDFC43B65D" box="[851,915,1527,1546]" captionStart="Figs" captionStartId="7.[169,208,1115,1134]" captionTargetBox="[153,975,195,1094]" captionTargetId="figure-183@7.[149,975,194,1094]" captionTargetPageId="7" captionText="Figs. 19–24. Phyllophaga cruces. 19) Inner protarsal claw, male; 20) genital capsule, lateral view; 21) female genital plates, ventral view; 22) parameres, distal view; 23) parameres, ventral view; 24) tectum, phallobase and parameres, dorsal view. Scale lines: Fig. 19 ¼ 0.5 mm, Figs. 20–24 ¼ 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10105242" httpUri="https://zenodo.org/record/10105242/files/figure.png" pageId="7" pageNumber="398">Fig. 21</figureCitation>
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). Total body length:
|
||
<quantity id="563D9B00FF8EFFD37A69AF45FE86B675" box="[262,342,1551,1570]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" pageId="7" pageNumber="398" unit="mm" value="20.1">20.1 mm</quantity>
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. Humeral width: 9.0 mm.
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</paragraph>
|
||
<paragraph id="917A36E5FF81FFDC7BA8A988FC76B080" blockId="8.[175,1005,194,747]" box="[199,934,194,215]" pageId="8" pageNumber="399">
|
||
<emphasis id="A3B1EAF7FF81FFDC7BA8A988FEFFB080" bold="true" box="[199,303,194,215]" pageId="8" pageNumber="399">Variation.</emphasis>
|
||
Total body length:
|
||
<quantity id="563D9B00FF81FFDC7A98A988FD55B080" box="[503,645,194,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.9700000000000002" metricValueMax="2.0300000000000002" metricValueMin="1.9100000000000001" pageId="8" pageNumber="399" unit="mm" value="19.7" valueMax="20.3" valueMin="19.1">19.1–20.3 mm</quantity>
|
||
, humeral width:
|
||
<quantity id="563D9B00FF81FFDC7845A988FC72B080" box="[810,930,194,215]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.2" metricValueMin="8.2" pageId="8" pageNumber="399" unit="mm" value="8.7" valueMax="9.2" valueMin="8.2">8.2–9.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9DF656EFF81FFDC7BA8A997FE39B1B6" pageId="8" pageNumber="399" type="materials_examined">
|
||
<paragraph id="917A36E5FF81FFDC7BA8A997FE9DB115" blockId="8.[175,1005,194,747]" pageId="8" pageNumber="399">
|
||
<emphasis id="A3B1EAF7FF81FFDC7BA8A997FE99B0A5" bold="true" box="[199,329,221,242]" pageId="8" pageNumber="399">Type Series.</emphasis>
|
||
(Described from
|
||
<specimenCount id="87C3FD6CFF81FFDC796FA997FD9DB0A5" box="[512,589,221,242]" pageId="8" pageNumber="399" type="male">3 males</specimenCount>
|
||
and
|
||
<specimenCount id="87C3FD6CFF81FFDC79EAA997FD38B0A5" box="[645,744,221,242]" pageId="8" pageNumber="399" type="female">5 females</specimenCount>
|
||
).
|
||
<materialsCitation id="21AD3CB8FF81FFDC7992A997FC39B15A" collectingDate="2003-04-20" collectionCode="INBio" collectorName="E. G. Riley" country="Costa Rica" location="Estacion Biologica Las Cruces" pageId="8" pageNumber="399" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||
<typeStatus id="4E7E8847FF81FFDC7992A997FC85B0A5" box="[765,853,221,242]" pageId="8" pageNumber="399" type="holotype">Holotype</typeStatus>
|
||
male;
|
||
<collectingCountry id="E9D27675FF81FFDC78F1A997FF3BB15A" name="Costa Rica" pageId="8" pageNumber="399">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="5301F807FF81FFDC7B9BA9B2FE8EB15A" box="[244,350,248,269]" country="Costa Rica" name="Puntarenas" pageId="8" pageNumber="399">Puntarenas</collectingRegion>
|
||
,
|
||
<location id="941A603EFF81FFDC7A07A9B2FD5DB15A" LSID="urn:lsid:plazi:treatment:196C87F3FF8FFFDC7A6EABD4FE56B2BC:941A603EFF81FFDC7A07A9B2FD5DB15A" box="[360,653,248,269]" country="Costa Rica" name="Estacion Biologica Las Cruces" pageId="8" pageNumber="399" stateProvince="Puntarenas">Estación Biologica Las Cruces</location>
|
||
, 17/
|
||
<date id="E57B1025FF81FFDC79DBA9B2FCF8B15A" box="[692,808,248,269]" pageId="8" pageNumber="399" value="2003-04-20">
|
||
<collectingDate id="F53FE9CDFF81FFDC79DBA9B2FCF8B15A" box="[692,808,248,269]" pageId="8" pageNumber="399" value="2003-04-20">20-IV-2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="3C305333FF81FFDC785EA9B2FC45B15A" box="[817,917,248,269]" pageId="8" pageNumber="399">E.G. Riley</collectorName>
|
||
(
|
||
<collectionCode id="F7D4AE20FF81FFDC78CDA9B2FC32B15A" box="[930,994,248,269]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="8" pageNumber="399">INBio</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="21AD3CB8FF81FFDC7BC0A858FE99B115" collectingDate="2003-04-20" collectionCode="TAMU, INBio, MXAL" collectorName="E. G. Riley" country="Costa Rica" location="Estacion Biologica Las Cruces" pageId="8" pageNumber="399" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="allotype">
|
||
<typeStatus id="4E7E8847FF81FFDC7BC0A858FED4B170" box="[175,260,274,295]" pageId="8" pageNumber="399" type="allotype">Allotype</typeStatus>
|
||
,
|
||
<specimenCount id="87C3FD6CFF81FFDC7A79A858FE8EB170" box="[278,350,274,295]" pageId="8" pageNumber="399" type="male">2 male</specimenCount>
|
||
and
|
||
<specimenCount id="87C3FD6CFF81FFDC7AF4A858FE25B170" box="[411,501,274,295]" pageId="8" pageNumber="399" type="female">4 female</specimenCount>
|
||
<typeStatus id="4E7E8847FF81FFDC796CA858FD8EB170" box="[515,606,274,295]" pageId="8" pageNumber="399" type="paratype">paratypes</typeStatus>
|
||
with same data as holotype (
|
||
<collectionCode id="F7D4AE20FF81FFDC78F0A858FC39B170" box="[927,1001,274,295]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;M University" pageId="8" pageNumber="399" type="Herbarium">TAMU</collectionCode>
|
||
,
|
||
<collectionCode id="F7D4AE20FF81FFDC7BC0A867FF3DB115" box="[175,237,301,322]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="8" pageNumber="399">INBio</collectionCode>
|
||
,
|
||
<collectionCode id="F7D4AE20FF81FFDC7B97A867FE92B115" box="[248,322,301,322]" pageId="8" pageNumber="399">MXAL</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="917A36E5FF81FFDC7BA8A80DFD62B120" blockId="8.[175,1005,194,747]" pageId="8" pageNumber="399">
|
||
<emphasis id="A3B1EAF7FF81FFDC7BA8A80DFEB1B10B" bold="true" box="[199,353,327,348]" pageId="8" pageNumber="399">Type Locality.</emphasis>
|
||
<materialsCitation id="21AD3CB8FF81FFDC7A02A80DFD7EB120" country="Costa Rica" latitude="8.785833" location="Estacion Biologica Las Cruces" longLatPrecision="21" longitude="-8.952499" municipality="Province" pageId="8" pageNumber="399" specimenCount="1" stateProvince="Puntarenas">
|
||
<location id="941A603EFF81FFDC7A02A80DFD73B10B" LSID="urn:lsid:plazi:treatment:196C87F3FF8FFFDC7A6EABD4FE56B2BC:941A603EFF81FFDC7A02A80DFD73B10B" box="[365,675,327,348]" country="Costa Rica" latitude="8.785833" longLatPrecision="21" longitude="-8.952499" municipality="Province" name="Estacion Biologica Las Cruces" pageId="8" pageNumber="399" stateProvince="Puntarenas">Estacion Biologica Las Cruces</location>
|
||
,
|
||
<locationDeviation id="80C82FC7FF81FFDC79DCA80DFCB9B10B" box="[691,873,327,348]" location="San Vito" pageId="8" pageNumber="399">
|
||
<quantity id="563D9B00FF81FFDC79DCA80DFD39B10B" box="[691,745,327,348]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="399" unit="km" value="7.0">7 km</quantity>
|
||
S San Vito
|
||
</locationDeviation>
|
||
,
|
||
<collectingMunicipality id="711EAC9FFF81FFDC7816A80DFC1DB10B" box="[889,973,327,348]" pageId="8" pageNumber="399">Province</collectingMunicipality>
|
||
of
|
||
<collectingRegion id="5301F807FF81FFDC7BC0A828FEC9B120" box="[175,281,354,375]" country="Costa Rica" name="Puntarenas" pageId="8" pageNumber="399">Puntarenas</collectingRegion>
|
||
,
|
||
<collectingCountry id="E9D27675FF81FFDC7A4AA828FE5EB120" box="[293,398,354,375]" name="Costa Rica" pageId="8" pageNumber="399">Costa Rica</collectingCountry>
|
||
(approx. 08
|
||
<geoCoordinate id="F4F15022FF81FFDC7969A82BFD92B120" box="[518,578,353,375]" degrees="8" direction="north" minutes="47" orientation="latitude" pageId="8" pageNumber="399" precision="15" seconds="9" value="8.785833">
|
||
<emphasis id="A3B1EAF7FF81FFDC7969A82BFDDEB120" bold="true" box="[518,526,353,375]" pageId="8" pageNumber="399">8</emphasis>
|
||
47
|
||
<emphasis id="A3B1EAF7FF81FFDC7949A82BFDFEB120" bold="true" box="[550,558,353,375]" pageId="8" pageNumber="399">9</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
; 82
|
||
<geoCoordinate id="F4F15022FF81FFDC790AA82BFD77B120" box="[613,679,353,375]" degrees="8" direction="west" minutes="57" orientation="longitude" pageId="8" pageNumber="399" precision="15" seconds="9" value="-8.952499">
|
||
<emphasis id="A3B1EAF7FF81FFDC790AA82BFDBDB120" bold="true" box="[613,621,353,375]" pageId="8" pageNumber="399">8</emphasis>
|
||
57
|
||
<emphasis id="A3B1EAF7FF81FFDC79EAA82BFD5DB120" bold="true" box="[645,653,353,375]" pageId="8" pageNumber="399">9</emphasis>
|
||
W
|
||
</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="917A36E5FF81FFDC7BA8A836FE39B1B6" blockId="8.[175,1005,194,747]" pageId="8" pageNumber="399">
|
||
<emphasis id="A3B1EAF7FF81FFDC7BA8A836FEB9B1C6" bold="true" box="[199,361,380,401]" pageId="8" pageNumber="399">Biological Data.</emphasis>
|
||
This species inhabits the tropical forest located at
|
||
<quantity id="563D9B00FF81FFDC7823A836FC49B1C6" box="[844,921,380,401]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="399" unit="m" value="1200.0">1,200 m</quantity>
|
||
altitude. All specimens were collected during April. Other species of
|
||
<emphasis id="A3B1EAF7FF81FFDC7997A8DDFCA1B1FB" box="[760,881,407,428]" italics="true" pageId="8" pageNumber="399">Phyllophaga</emphasis>
|
||
flying at the same time and place were:
|
||
<taxonomicName id="56C54D66FF81FFDC7ADAA8F8FD1CB191" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[437,716,433,455]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="chiriquina">
|
||
<emphasis id="A3B1EAF7FF81FFDC7ADAA8F8FE13B190" box="[437,451,434,455]" italics="true" pageId="8" pageNumber="399">P</emphasis>
|
||
. (
|
||
<emphasis id="A3B1EAF7FF81FFDC7AB7A8F8FDDEB190" box="[472,526,434,455]" italics="true" pageId="8" pageNumber="399">s. str.</emphasis>
|
||
)
|
||
<emphasis id="A3B1EAF7FF81FFDC7973A8F8FDAFB190" box="[540,639,434,455]" italics="true" pageId="8" pageNumber="399">chiriquina</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="A3B1EAF7FF81FFDC79B8A8F8FD35B190" box="[727,741,434,455]" italics="true" pageId="8" pageNumber="399">P</emphasis>
|
||
. (
|
||
<emphasis id="A3B1EAF7FF81FFDC7995A8F8FCE0B190" box="[762,816,434,455]" italics="true" pageId="8" pageNumber="399">s. str.</emphasis>
|
||
)
|
||
<emphasis id="A3B1EAF7FF81FFDC7851A8F8FC4EB190" box="[830,926,434,455]" italics="true" pageId="8" pageNumber="399">chorotega</emphasis>
|
||
Morón, and
|
||
<emphasis id="A3B1EAF7FF81FFDC7BB5A886FF38B1B6" box="[218,232,460,481]" italics="true" pageId="8" pageNumber="399">P</emphasis>
|
||
. (
|
||
<emphasis id="A3B1EAF7FF81FFDC7B91A886FEE4B1B6" box="[254,308,460,481]" italics="true" pageId="8" pageNumber="399">s. str.</emphasis>
|
||
)
|
||
<emphasis id="A3B1EAF7FF81FFDC7A2CA886FE46B1B6" box="[323,406,460,481]" italics="true" pageId="8" pageNumber="399">gigantea</emphasis>
|
||
(Bates).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9DF656EFF81FFDC7BA8A8ACFD81B2E1" pageId="8" pageNumber="399" type="discussion">
|
||
<paragraph id="917A36E5FF81FFDC7BA8A8ACFD81B2E1" blockId="8.[175,1005,194,747]" pageId="8" pageNumber="399">
|
||
<emphasis id="A3B1EAF7FF81FFDC7BA8A8ACFEFAB1AC" bold="true" box="[199,298,486,507]" pageId="8" pageNumber="399">Remarks.</emphasis>
|
||
<taxonomicName id="56C54D66FF81FFDC7A59A8ADFE28B1AB" box="[310,504,487,508]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="cruces">
|
||
<emphasis id="A3B1EAF7FF81FFDC7A59A8ADFE28B1AB" box="[310,504,487,508]" italics="true" pageId="8" pageNumber="399">Phyllophaga cruces</emphasis>
|
||
</taxonomicName>
|
||
does not belong to any species group defined by
|
||
<bibRefCitation id="F5544B14FF81FFDC7BC0AB4BFEE7B241" author="Moron" box="[175,311,513,534]" pageId="8" pageNumber="399" refId="ref5144" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica. Instituto de Ecologia, Mexico. 341 pp." type="book" year="1986">Morón (1986)</bibRefCitation>
|
||
or Solís and Morón (1998). It is similar to
|
||
<taxonomicName id="56C54D66FF81FFDC79B0AB4BFCBEB241" authority="Moser" authorityName="Moser" box="[735,878,513,534]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="nuda">
|
||
<emphasis id="A3B1EAF7FF81FFDC79B0AB4BFCFBB241" box="[735,811,513,534]" italics="true" pageId="8" pageNumber="399">P. nuda</emphasis>
|
||
Moser
|
||
</taxonomicName>
|
||
from British
|
||
<collectingCountry id="E9D27675FF81FFDC7BC0AB56FF2BB266" box="[175,251,540,561]" name="Guyana" pageId="8" pageNumber="399">Guyana</collectingCountry>
|
||
, and
|
||
<collectingRegion id="5301F807FF81FFDC7A56AB56FE52B266" box="[313,386,540,561]" country="Venezuela" name="Bolivar" pageId="8" pageNumber="399">Bolivar</collectingRegion>
|
||
,
|
||
<collectingCountry id="E9D27675FF81FFDC7AFDAB56FE28B266" box="[402,504,540,561]" name="Venezuela" pageId="8" pageNumber="399">Venezuela</collectingCountry>
|
||
, but the projection of the parameres are broadly rounded, not extended and curved apically as in
|
||
<taxonomicName id="56C54D66FF81FFDC7916AB7CFD13B21C" box="[633,707,566,587]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="nuda">
|
||
<emphasis id="A3B1EAF7FF81FFDC7916AB7CFD13B21C" box="[633,707,566,587]" italics="true" pageId="8" pageNumber="399">P. nuda</emphasis>
|
||
</taxonomicName>
|
||
. Also, it is externally similar to
|
||
<taxonomicName id="56C54D66FF81FFDC7BC0AB1BFEB5B231" authority="Saylor" authorityName="Saylor" authorityYear="1935" box="[175,357,593,614]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="vandykei">
|
||
<emphasis id="A3B1EAF7FF81FFDC7BC0AB1BFECFB231" box="[175,287,593,614]" italics="true" pageId="8" pageNumber="399">P. vandykei</emphasis>
|
||
Saylor
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="E9D27675FF81FFDC7ACEAB1BFE3EB231" box="[417,494,593,614]" name="Panama" pageId="8" pageNumber="399">Panama</collectingCountry>
|
||
, but the projections of the ventro-basal border of the parameres of the new species are much longer than the rounded distal projection of the parameres of
|
||
<taxonomicName id="56C54D66FF81FFDC7A5DABCCFE72B2CC" authorityName="Saylor" authorityYear="1935" box="[306,418,646,667]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="vandykei">
|
||
<emphasis id="A3B1EAF7FF81FFDC7A5DABCCFE72B2CC" box="[306,418,646,667]" italics="true" pageId="8" pageNumber="399">P. vandykei</emphasis>
|
||
</taxonomicName>
|
||
. The male mesotarsal claws are asymmetrical in
|
||
<taxonomicName id="56C54D66FF81FFDC7812ABCCFC3CB2CC" authorityName="Saylor" authorityYear="1935" box="[893,1004,646,667]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="vandykei">
|
||
<emphasis id="A3B1EAF7FF81FFDC7812ABCCFC3CB2CC" box="[893,1004,646,667]" italics="true" pageId="8" pageNumber="399">P. vandykei</emphasis>
|
||
</taxonomicName>
|
||
whereas they are symmetrical in
|
||
<taxonomicName id="56C54D66FF81FFDC7A9EABEBFD9BB2E1" box="[497,587,673,694]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="cruces">
|
||
<emphasis id="A3B1EAF7FF81FFDC7A9EABEBFD9BB2E1" box="[497,587,673,694]" italics="true" pageId="8" pageNumber="399">P. cruces</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9DF656EFF81FFDC7BA8ABF1FE56B2BC" pageId="8" pageNumber="399" type="etymology">
|
||
<paragraph id="917A36E5FF81FFDC7BA8ABF1FE56B2BC" blockId="8.[175,1005,194,747]" pageId="8" pageNumber="399">
|
||
<emphasis id="A3B1EAF7FF81FFDC7BA8ABF1FEEBB287" bold="true" box="[199,315,699,720]" pageId="8" pageNumber="399">Etymology.</emphasis>
|
||
Specific epithet derived from the name of
|
||
<typeStatus id="4E7E8847FF81FFDC7865ABF1FCE3B287" box="[778,819,699,720]" pageId="8" pageNumber="399">type</typeStatus>
|
||
locality, Estación Biologica Las Cruces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |