treatments-xml/data/03/F5/31/03F5311F9475850852AD0E2FE596FD68.xml
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<document id="25AA43BA3C7C075D25C215874AFC9D0D" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627012910610" checkinUser="felipe" docAuthor="Bruce, A. J." docDate="2008" docId="03F5311F9475850852AD0E2FE596FD68" docLanguage="en" docName="zt01815p024.pdf" docOrigin="Zootaxa 1815" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pseudoclimenes holthuisi Bruce 2008, sp. nov." docType="treatment" docVersion="2" lastPageNumber="23" masterDocId="FFCC49679465851E523A0F18E468FFD2" masterDocTitle="Palaemonoid shrimps from the Australian north west shelf" masterLastPageNumber="24" masterPageNumber="1" pageNumber="17" updateTime="1699014906470" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B1469430A9AAE671D1FA5B543F20AA15">Palaemonoid shrimps from the Australian north west shelf</mods:title>
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<mods:namePart id="FADAD8D890D3E3D8CCDBEFB3ECDC6917">Bruce, A. J.</mods:namePart>
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<mods:date id="6481720FBCDF2667192C3752B23A59C7">2008</mods:date>
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<mods:number id="E3A0401D313D1C77474C78A0E0FFF983">2008-07-04</mods:number>
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<treatment id="03F5311F9475850852AD0E2FE596FD68" LSID="urn:lsid:plazi:treatment:03F5311F9475850852AD0E2FE596FD68" httpUri="http://treatment.plazi.org/id/03F5311F9475850852AD0E2FE596FD68" lastPageId="22" lastPageNumber="23" pageId="16" pageNumber="17">
<subSubSection id="C346D3829475850E52AD0E2FE658FE83" box="[151,560,311,337]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BE380099475850E52AD0E2FE658FE83" blockId="16.[151,560,311,378]" box="[151,560,311,337]" pageId="16" pageNumber="17">
<heading id="D0AB37659475850E52AD0E2FE658FE83" bold="true" box="[151,560,311,337]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B9285C1B9475850E52AD0E2FE658FE83" bold="true" box="[151,560,311,337]" pageId="16" pageNumber="17">
<taxonomicName id="4C5CFB8A9475850E52AD0E2FE5A4FE83" ID-CoL="098fe727-025d-4292-b687-c030a170a838" authority="Bruce, 2008" authorityName="Bruce" authorityYear="2008" box="[151,460,311,337]" class="Malacostraca" family="Palaemonidae" genus="Pseudoclimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="holthuisi" status="sp. nov.">
<emphasis id="B9285C1B9475850E52AD0E2FE5A4FE83" bold="true" box="[151,460,311,337]" italics="true" pageId="16" pageNumber="17">Pseudoclimenes holthuisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE1609475850E53E90E2FE658FE83" box="[467,560,311,337]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C346D3829475850E52AD0E78E546FEA8" box="[151,302,352,378]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BE380099475850E52AD0E78E546FEA8" blockId="16.[151,560,311,378]" box="[151,302,352,378]" pageId="16" pageNumber="17">
(
<figureCitation id="13679C8C9475850E52A50E78E54DFEA8" box="[159,293,352,378]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStartId-0="17.[151,255,1571,1595]" captionStartId-1="18.[154,258,1680,1704]" captionStartId-2="19.[151,255,1499,1523]" captionStartId-3="20.[151,255,1831,1855]" captionTargetBox-0="[151,1419,446,1536]" captionTargetBox-1="[163,1413,350,1634]" captionTargetBox-2="[184,1406,364,1467]" captionTargetBox-3="[151,1409,360,1771]" captionTargetId-0="figure-248@17.[151,1436,434,1548]" captionTargetId-1="figure-191@18.[160,1426,350,1652]" captionTargetId-2="figure-272@19.[180,1406,354,1475]" captionTargetId-3="figure-138@20.[151,1436,314,1807]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="FIGURE 10. Pseudoclimenes holthuisi gen. nov., sp. nov., holotype female, NMV J55446. Scale bar in millimetres." captionText-1="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." captionText-2="FIGURE 12. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath." captionText-3="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="16" pageNumber="17">Figs 1013</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C346D3829475850E52AD0EB7E5E0FDC8" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BE380099475850E52AD0EB7E5E0FDC8" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52AD0EB7E5EDFE1B" bold="true" box="[151,389,431,457]" pageId="16" pageNumber="17">Material examined</emphasis>
.
<specimenCount id="9D5A4B809475850E53AC0EA8E5ADFE11" box="[406,453,431,458]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
<typeStatus id="54E73EAB9475850E53F10EA8E65CFE18" box="[459,564,432,458]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
,
<materialsCitation id="3B348A549475850E50790EA8E5ECFDC8" accessionNumber="J55446" collectingDate="2005-12-02" collectionCode="NMV" collectorName="Cruise SS &amp; Sherman &amp; G. C. Poore" elevation="114" latitude="-29.814722" location="Jurien Bay" longLatPrecision="20" longitude="114.43194" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D5A4B809475850E50790EA8E618FE11" box="[579,624,431,458]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
<typeStatus id="54E73EAB9475850E504D0EA8E6B7FE18" box="[631,735,432,458]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
, CLs 1.85, 1.6,
<collectorName id="26A9E5DF9475850E519A0EA8E07DFE18" box="[928,1045,432,458]" pageId="16" pageNumber="17">Cruise SS</collectorName>
2005/10 083,
<location id="8E83D6D29475850E56870EA8E156FE18" LSID="urn:lsid:plazi:treatment:03F5311F9475850852AD0E2FE596FD68:8E83D6D29475850E56870EA8E156FE18" box="[1213,1342,432,458]" latitude="-29.814722" longLatPrecision="20" longitude="114.43194" name="Jurien Bay" pageId="16" pageNumber="17">Jurien Bay</location>
,
<geoCoordinate id="EE68E6CE9475850E57760EA8E4B9FE20" degrees="29" direction="south" minutes="48" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="25" value="-29.806944">29°48 25”S</geoCoordinate>
<geoCoordinate id="EE68E6CE9475850E52E40EC0E51FFE20" box="[222,375,472,498]" degrees="114" direction="east" minutes="25" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="52" value="114.43111">114°2552”E</geoCoordinate>
to
<geoCoordinate id="EE68E6CE9475850E539D0EC0E65AFE20" box="[423,562,472,498]" degrees="29" direction="south" minutes="48" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="53" value="-29.814722">29°4853”S</geoCoordinate>
<geoCoordinate id="EE68E6CE9475850E50040EC0E6B1FE20" box="[574,729,472,498]" degrees="114" direction="east" minutes="25" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="55" value="114.43194">114°2555”E</geoCoordinate>
,
<quantity id="4CA42DEC9475850E50D10EC0E701FE20" box="[747,873,472,498]" metricMagnitude="2" metricUnit="m" metricValue="1.135" metricValueMax="1.14" metricValueMin="1.13" pageId="16" pageNumber="17" unit="m" value="113.5" valueMax="114.0" valueMin="113.0">
<elevation id="0071673A9475850E50D10EC0E701FE20" box="[747,873,472,498]" metricMagnitude="2" metricUnit="m" metricValue="1.135" metricValueMax="1.14" metricValueMin="1.13" pageId="16" pageNumber="17" unit="m" value="113.5" valueMax="114.0" valueMin="113.0">113114m</elevation>
</quantity>
,
<collectorName id="26A9E5DF9475850E51400EC0E78AFE20" box="[890,994,472,498]" pageId="16" pageNumber="17">Sherman</collectorName>
sled, coll.
<collectorName id="26A9E5DF9475850E564A0EC0E090FE20" box="[1136,1272,472,498]" pageId="16" pageNumber="17">G.C. Poore</collectorName>
,
<date id="FFE2A6C99475850E57330EC0E4BCFDC8" pageId="16" pageNumber="17" value="2005-12-02">
<collectingDate id="EFA65F219475850E57330EC0E4BCFDC8" pageId="16" pageNumber="17" value="2005-12-02">2 December 2005</collectingDate>
</date>
,
<collectionCode id="ED4D18CC9475850E52DA0D18E54CFDC8" box="[224,292,512,538]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="16" pageNumber="17" type="Museum">NMV</collectionCode>
<accessionNumber id="940F1DEA9475850E53160D18E5ECFDC8" box="[300,388,512,538]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J55446" pageId="16" pageNumber="17" type="EnaNcbi">J55446</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C346D3829475850E52FF0D3FE67DFD90" box="[197,533,551,578]" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BE380099475850E52FF0D3FE67DFD90" blockId="16.[151,1436,431,2018]" box="[197,533,551,578]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0D3FE52DFD93" bold="true" box="[197,325,551,577]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C346D3829475850852FF0D57E02FFF60" lastPageId="22" lastPageNumber="23" pageId="16" pageNumber="17" type="description">
<paragraph id="8BE380099475850E52FF0D57E036FDB8" blockId="16.[151,1436,431,2018]" box="[197,1118,591,618]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0D57E53EFDBB" bold="true" box="[197,342,591,617]" pageId="16" pageNumber="17">Description</emphasis>
. A slenderly built pontoniine shrimp of subcylindrical body form.
</paragraph>
<paragraph id="8BE380099475850E52FF0D60E06AFCD8" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0D60E540FD43" box="[197,296,632,657]" italics="true" pageId="16" pageNumber="17">Rostrum</emphasis>
(
<figureCitation id="13679C8C9475850E53040D60E5C2FD40" box="[318,426,632,658]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="16" pageNumber="17">Fig. 11B</figureCitation>
) slender, compressed, reaching to about distal margin of intermediate segment of antennular peduncle, 0.7 of CL, dorsal carina distinct, with 78 small acute teeth, first two postorbital, lateral and ventral carinae obsolescent, ventral margin with numerous short plumose median setae, two small acute teeth distally, with additional minute preterminal ventral tooth in
<typeStatus id="54E73EAB9475850E51A90DE8E795FCD8" box="[915,1021,752,778]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BE380099475850E52FF0C00E617FC50" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0C00E550FCE3" box="[197,312,792,817]" italics="true" pageId="16" pageNumber="17">Carapace</emphasis>
(
<figureCitation id="13679C8C9475850E53730C00E5DBFCE0" box="[329,435,792,818]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="16" pageNumber="17">Fig. 11A</figureCitation>
) smooth, glabrous, without supraorbital, epigastric, or hepatic spines, orbits obsolescent, inferior orbital angle (
<figureCitation id="13679C8C9475850E53DC0C58E639FC88" box="[486,593,832,858]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="16" pageNumber="17">Fig. 11C</figureCitation>
) broadly rounded, without ventral flange, antennal spine robust, well below inferior orbital angle, submarginal.
</paragraph>
<paragraph id="8BE380099475850E52FF0C88E08CFC00" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0C88E55BFC7B" box="[197,307,912,937]" italics="true" pageId="16" pageNumber="17">Abdomen</emphasis>
with third tergite not produced or carinate, sixth segment about 1.6 of CL, 0.9 of telson length, twice as long as deep, posteroventral angle minutely acute, posterolateral angle broadly acute.
</paragraph>
<paragraph id="8BE380099475850E52FF0CF8E5A9FAC0" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0CF8E554FC2B" box="[197,316,992,1017]" italics="true" pageId="16" pageNumber="17">Antennule</emphasis>
(
<figureCitation id="13679C8C9475850E536B0CF8E5D6FC28" box="[337,446,992,1018]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="16" pageNumber="17">Fig. 11D</figureCitation>
) proximal segment of peduncle (
<figureCitation id="13679C8C9475850E51630CF8E7ABFC28" box="[857,963,992,1018]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="16" pageNumber="17">Fig. 11E</figureCitation>
) about 1.6 times longer than maximal width, tapering distally, medial margin straight, sparsely setose distally, with small acute ventromedial tooth at about half length, lateral margin convergent distally, concave, with small rounded distal lobe with slender divergent acute tooth laterally (
<figureCitation id="13679C8C9475850E50390B40E607FBA0" box="[515,623,1112,1138]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="16" pageNumber="17">Fig. 13A</figureCitation>
), reaching well beyond adjacent lobe, statocyst normal, with granular statolith, stylocerite acute, reaching to about half segment length, intermediate and distal segments normal, combined length about 0.5 of proximal segment length, upper flagellum biramous, proximal three segments fused, upper ramus with four segments, about nine groups of aesthetascs, with long terminal group, lower ramus with ten segments.
</paragraph>
<paragraph id="8BE380099475850E52FF0A38E1FDFA60" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0A38E54EFAEB" box="[197,294,1312,1337]" italics="true" pageId="16" pageNumber="17">Antenna</emphasis>
(
<figureCitation id="13679C8C9475850E53060A38E5CDFAE8" box="[316,421,1312,1338]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="16" pageNumber="17">Fig. 11F</figureCitation>
) normal, basicerite with small acute lateral tooth, carpocerite short, about 0.27 of scaphocerite length, subcylindrical, 2.3 times longer than wide, flagellum well developed; scaphocerite well exceeding antennular peduncle, 3.0 times longer than width, lateral margin straight, with acute distolateral tooth (
<figureCitation id="13679C8C9475850E52D90A80E55DFA60" box="[227,309,1432,1458]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="16" pageNumber="17">Fig. 13</figureCitation>
AB), 0.08 of scaphocerite length, about at 0.8 of scaphocerite length, well exceeded by lamella.
</paragraph>
<paragraph id="8BE380099475850E52FF0AD8E069FA08" blockId="16.[151,1436,431,2018]" box="[197,1025,1472,1498]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0AD8E5CBFA0B" box="[197,419,1472,1497]" italics="true" pageId="16" pageNumber="17">Ophthalmic somite</emphasis>
without median process or accessory pigment spot.
</paragraph>
<paragraph id="8BE380099475850E52FF0AF0E0F4F9F8" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0AF0E499F9D3" box="[197,241,1512,1537]" italics="true" pageId="16" pageNumber="17">Eye</emphasis>
(
<figureCitation id="13679C8C9475850E533F0AF0E519F9D0" box="[261,369,1512,1538]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="16" pageNumber="17">Fig. 11G</figureCitation>
) with well pigmented hemispherical cornea, about 0.25 of CL, subequal to stalk width, without accessory pigment spot, stalk subcylindrical, about 1.4 times longer than width.
</paragraph>
<paragraph id="8BE380099475850E52FF0920E101F980" blockId="16.[151,1436,431,2018]" box="[197,1385,1592,1618]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0920E55AF983" box="[197,306,1592,1617]" italics="true" pageId="16" pageNumber="17">Epistome</emphasis>
unarmed. Paragnaths (
<figureCitation id="13679C8C9475850E507A0920E6C2F980" box="[576,682,1592,1618]" captionStart="FIGURE 12" captionStartId="19.[151,255,1499,1523]" captionTargetBox="[184,1406,364,1467]" captionTargetId="figure-272@19.[180,1406,354,1475]" captionTargetPageId="19" captionText="FIGURE 12. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath." pageId="16" pageNumber="17">Fig. 12G</figureCitation>
) with alae feebly bilobed, corpus with deep median groove.
</paragraph>
<paragraph id="8BE380099475850E52FF0978E518F920" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0978E55DF9AB" box="[197,309,1632,1657]" italics="true" pageId="16" pageNumber="17">Mandible</emphasis>
(
<figureCitation id="13679C8C9475850E53700978E5D0F9A8" box="[330,440,1632,1658]" captionStart="FIGURE 12" captionStartId="19.[151,255,1499,1523]" captionTargetBox="[184,1406,364,1467]" captionTargetId="figure-272@19.[180,1406,354,1475]" captionTargetPageId="19" captionText="FIGURE 12. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath." pageId="16" pageNumber="17">Fig. 12A</figureCitation>
) corpus moderately robust, without palp; incisor process (
<figureCitation id="13679C8C9475850E56BA0978E086F9A8" box="[1152,1262,1632,1658]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="16" pageNumber="17">Fig. 13D</figureCitation>
) slender, with three acute teeth distally, lateral tooth largest, central tooth smallest; molar process (
<figureCitation id="13679C8C9475850E564F0990E0B6F970" box="[1141,1246,1672,1698]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="16" pageNumber="17">Fig. 13C</figureCitation>
) subcylindrical, distally truncate, with five marginal teeth. Posterior tooth large and projecting, with small adjacent tufts of short simple setae.
</paragraph>
<paragraph id="8BE380099475850E52FF0818E61FF840" blockId="16.[151,1436,431,2018]" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF0818E55FF8CB" box="[197,311,1792,1817]" italics="true" pageId="16" pageNumber="17">Maxillula</emphasis>
(
<figureCitation id="13679C8C9475850E537D0818E5D9F8C8" box="[327,433,1792,1818]" captionStart="FIGURE 12" captionStartId="19.[151,255,1499,1523]" captionTargetBox="[184,1406,364,1467]" captionTargetId="figure-272@19.[180,1406,354,1475]" captionTargetPageId="19" captionText="FIGURE 12. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath." pageId="16" pageNumber="17">Fig. 12B</figureCitation>
) of normal form, palp (
<figureCitation id="13679C8C9475850E50FC0818E745F8C8" box="[710,813,1792,1818]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="16" pageNumber="17">Fig. 13E</figureCitation>
) feebly bilobed, lower lobe with small ventral tubercle with minute seta; upper lacinia (
<figureCitation id="13679C8C9475850E50020830E6F5F890" box="[568,669,1832,1858]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="16" pageNumber="17">Fig. 13F</figureCitation>
) tapering distally, rounded, with six short stout simple spines and several spiniform setae; lower lacinia subcylindrical, tapering distally, sparsely setose, with 23 short spiniform terminal setae and few simple setae
</paragraph>
<paragraph id="8BE380099475850F52FF08B8E06BFF08" blockId="16.[151,1436,431,2018]" lastBlockId="17.[151,1436,152,378]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B9285C1B9475850E52FF08B8E548F86B" box="[197,288,1952,1977]" italics="true" pageId="16" pageNumber="17">Maxilla</emphasis>
(
<figureCitation id="13679C8C9475850E530808B8E5F4F868" box="[306,412,1952,1978]" captionStart="FIGURE 12" captionStartId="19.[151,255,1499,1523]" captionTargetBox="[184,1406,364,1467]" captionTargetId="figure-272@19.[180,1406,354,1475]" captionTargetPageId="19" captionText="FIGURE 12. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath." pageId="16" pageNumber="17">Fig. 12C</figureCitation>
) with simple elongate non-setiferous palp; basal endite bilobed, lobes short, subequal, each with about six slender simple seta; coxal endite obsolete, medial margin rounded, non-setose; scaphog- nathite about 3.0 times longer than wide, anterior lobe 1.2 times longer than basal width, distally rounded, medially not emarginate, posterior lobe narrow, 0.8 of anterior lobe length.
</paragraph>
<paragraph id="8BE380099474850F52FF0FF0E614FEA8" blockId="17.[151,1436,152,378]" pageId="17" pageNumber="18">
<emphasis id="B9285C1B9474850F52FF0FF0E5EBFED3" box="[197,387,232,257]" italics="true" pageId="17" pageNumber="18">First maxilliped</emphasis>
(
<figureCitation id="13679C8C9474850F53AD0FF0E66BFED0" box="[407,515,232,258]" captionStart="FIGURE 12" captionStartId="19.[151,255,1499,1523]" captionTargetBox="[184,1406,364,1467]" captionTargetId="figure-272@19.[180,1406,354,1475]" captionTargetPageId="19" captionText="FIGURE 12. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath." pageId="17" pageNumber="18">Fig. 12D</figureCitation>
) with simple elongate palp with preterminal plumose seta; basal endite well developed, rounded, sparsely setose, with long simple setae, coxal endite obsolescent, convex, with single short seta only; exopod well developed with slender flagellum with four plumose terminal setae, caridean lobe normal, narrow, epipod lost in dissection.
</paragraph>
<caption id="DF23D0819474850F52AD093BE11EF9E9" box="[151,1398,1571,1595]" pageId="17" pageNumber="18" startId="17.[151,255,1571,1595]" targetBox="[151,1419,446,1536]" targetPageId="17">
<paragraph id="8BE380099474850F52AD093BE11EF9E9" blockId="17.[151,1398,1571,1595]" box="[151,1398,1571,1595]" pageId="17" pageNumber="18">
<emphasis id="B9285C1B9474850F52AD093BE54EF9E9" bold="true" box="[151,294,1571,1595]" pageId="17" pageNumber="18">FIGURE 10.</emphasis>
<taxonomicName id="4C5CFB8A9474850F5317093CE654F9E9" authority="Bruce, 2008" authorityName="Bruce" authorityYear="2008" box="[301,572,1572,1595]" class="Malacostraca" family="Palaemonidae" genus="Pseudoclimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="holthuisi" status="gen. nov., sp. nov.">
<emphasis id="B9285C1B9474850F5317093CE654F9E9" box="[301,572,1572,1595]" italics="true" pageId="17" pageNumber="18">Pseudoclimenes holthuisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE1609474850F507E093BE762F9E9" box="[580,778,1571,1595]" pageId="17" pageNumber="18" rank="species">
<emphasis id="B9285C1B9474850F507E093BE6C0F9E9" bold="true" box="[580,680,1571,1595]" pageId="17" pageNumber="18">gen. nov.</emphasis>
,
<emphasis id="B9285C1B9474850F508F093BE762F9E9" bold="true" box="[693,778,1571,1595]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype female, NMV
<accessionNumber id="940F1DEA9474850F562C093CE00EF9E9" box="[1046,1126,1572,1595]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J55446" pageId="17" pageNumber="18" type="EnaNcbi">J55446</accessionNumber>
. Scale bar in millimetres.
</paragraph>
</caption>
<paragraph id="8BE380099474850F52FC0977E485F8FB" blockId="17.[151,1436,1647,2033]" pageId="17" pageNumber="18">
<emphasis id="B9285C1B9474850F52FC0977E5F5F95A" box="[198,413,1647,1672]" italics="true" pageId="17" pageNumber="18">Second maxilliped</emphasis>
(
<figureCitation id="13679C8C9474850F53970977E67CF95B" box="[429,532,1647,1673]" captionStart="FIGURE 12" captionStartId="19.[151,255,1499,1523]" captionTargetBox="[184,1406,364,1467]" captionTargetId="figure-272@19.[180,1406,354,1475]" captionTargetPageId="19" captionText="FIGURE 12. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath." pageId="17" pageNumber="18">Fig. 12E</figureCitation>
) of normal form, endopod with dactylar segment about 4.0 times longer than wide, medial margin straight with numerous denticulate spines, propod with anteromedial margin rounded, small, with few spiniform setae, carpus, ischiomerus and basis normal, exopod well developed with slender flagellum with four plumose terminal setae, coxa with small subrectangular epipod laterally, without podobranch,
</paragraph>
<paragraph id="8BE380099474850C52FF082FE713FEF8" blockId="17.[151,1436,1647,2033]" lastBlockId="18.[151,1436,152,298]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B9285C1B9474850F52FF082FE5E1F882" box="[197,393,1847,1872]" italics="true" pageId="17" pageNumber="18">Third maxilliped</emphasis>
(
<figureCitation id="13679C8C9474850F53A0082FE668F883" box="[410,512,1847,1873]" captionStart="FIGURE 12" captionStartId="19.[151,255,1499,1523]" captionTargetBox="[184,1406,364,1467]" captionTargetId="figure-272@19.[180,1406,354,1475]" captionTargetPageId="19" captionText="FIGURE 12. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath." pageId="17" pageNumber="18">Fig. 12F</figureCitation>
) of normal form, endopod exceeding carpocerite by terminal segment, ischiomerus feebly demarkated from basis by small margin notch, uniform, about 4.0 times longer than wide, laterally glabrous, medially with few simple setae, proximal third of medial margin with row of seven short submarginal plumose setae; penultimate segment about 0.75 of ischiomeral length, subcylindrical, 4.0 times longer than width, with pair of spiniform setae distomedially, three similar setae proximally; terminal segment about 0.8 of penultimate segment length, 4.0 times longer than proximal width, tapering distally, medial margin with five pairs of denticulate spines and similar terminal spines; exopod with slender flagellum, reaching to distal margin of ischiomerus, with four plumose terminal setae; coxa with small rounded lateral plate, medial margin not produced, non-setose; without arthrobranch.
</paragraph>
<caption id="DF23D0819477850C52A00988E6BFF93F" pageId="18" pageNumber="19" startId="18.[154,258,1680,1704]" targetBox="[163,1413,350,1634]" targetPageId="18">
<paragraph id="8BE380099477850C52A00988E6BFF93F" blockId="18.[151,1437,1680,1773]" pageId="18" pageNumber="19">
<emphasis id="B9285C1B9477850C52A00988E542F97A" bold="true" box="[154,298,1680,1704]" pageId="18" pageNumber="19">FIGURE 11.</emphasis>
<taxonomicName id="4C5CFB8A9477850C53080989E62CF97A" authority="Bruce, 2008" authorityName="Bruce" authorityYear="2008" box="[306,580,1681,1704]" class="Malacostraca" family="Palaemonidae" genus="Pseudoclimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="holthuisi" status="gen. nov., sp. nov.">
<emphasis id="B9285C1B9477850C53080989E62CF97A" box="[306,580,1681,1704]" italics="true" pageId="18" pageNumber="19">Pseudoclimenes holthuisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE1609477850C50770988E772F97A" box="[589,794,1680,1704]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B9285C1B9477850C50770988E6DBF97A" bold="true" box="[589,691,1680,1704]" pageId="18" pageNumber="19">gen. nov.</emphasis>
,
<emphasis id="B9285C1B9477850C50F80988E772F97A" bold="true" box="[706,794,1680,1704]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, NMV
<accessionNumber id="940F1DEA9477850C51E10988E043F975" box="[987,1067,1680,1703]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J55446" pageId="18" pageNumber="19" type="EnaNcbi">J55446</accessionNumber>
. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod.
</paragraph>
</caption>
<paragraph id="8BE380099477850C52FC0839E75DF8B1" blockId="18.[151,1436,1825,2011]" pageId="18" pageNumber="19">
<emphasis id="B9285C1B9477850C52FC0839E5F3F8E8" box="[198,411,1825,1850]" italics="true" pageId="18" pageNumber="19">Thoracic sternites</emphasis>
unarmed, third with broad transverse ridge, fourth sternite without finger-like median process, with low transverse ridge without median notch.
</paragraph>
<paragraph id="8BE380099477850D52FF0869E6D4FEF8" blockId="18.[151,1436,1825,2011]" lastBlockId="19.[151,1437,152,298]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B9285C1B9477850C52FF0869E516F858" box="[197,382,1905,1930]" italics="true" pageId="18" pageNumber="19">First pereiopod</emphasis>
(
<figureCitation id="13679C8C9477850C53B50869E593F859" box="[399,507,1905,1931]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="18" pageNumber="19">Fig. 13G</figureCitation>
) with chela (
<figureCitation id="13679C8C9477850C50A10869E76FF859" box="[667,775,1905,1931]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="18" pageNumber="19">Fig. 13H</figureCitation>
) about 0.4 of CL, palm subcylindrical, about 1.7 times longer than deep, with three rows of short cleaning setae proximoventrally, fingers subequal to palm length, similar, sparsely setose, slender with small acute hooked tips, narrow sharp entire lamina over distal half of cutting edge; carpus short, stout, 0.79 of chela length, 3.4 times longer than distal width, tapering proximally, with several distoventral cleaning setae; merus subequal to chela length, uniform, about 6.3 times longer than wide; ischium normal, about half merus length; basis and coxa normal, coxa with small rounded ventromedial process with four short simple spiniform setae.
</paragraph>
<caption id="DF23D0819476850D52AD0AC3E78EF9C7" pageId="19" pageNumber="20" startId="19.[151,255,1499,1523]" targetBox="[184,1406,364,1467]" targetPageId="19">
<paragraph id="8BE380099476850D52AD0AC3E78EF9C7" blockId="19.[151,1436,1499,1557]" pageId="19" pageNumber="20">
<emphasis id="B9285C1B9476850D52AD0AC3E54FFA21" bold="true" box="[151,295,1499,1523]" pageId="19" pageNumber="20">FIGURE 12.</emphasis>
<taxonomicName id="4C5CFB8A9476850D53140AC4E655FA21" authority="Bruce, 2008" authorityName="Bruce" authorityYear="2008" box="[302,573,1500,1523]" class="Malacostraca" family="Palaemonidae" genus="Pseudoclimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="holthuisi" status="gen. nov., sp. nov.">
<emphasis id="B9285C1B9476850D53140AC4E655FA21" box="[302,573,1500,1523]" italics="true" pageId="19" pageNumber="20">Pseudoclimenes holthuisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE1609476850D507E0AC3E764FA21" box="[580,780,1499,1523]" pageId="19" pageNumber="20" rank="species">
<emphasis id="B9285C1B9476850D507E0AC3E6C1FA21" bold="true" box="[580,681,1499,1523]" pageId="19" pageNumber="20">gen. nov.</emphasis>
,
<emphasis id="B9285C1B9476850D508C0AC3E764FA21" bold="true" box="[694,780,1499,1523]" pageId="19" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, NMV
<accessionNumber id="940F1DEA9476850D51F20AC4E070FA21" box="[968,1048,1500,1523]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J55446" pageId="19" pageNumber="20" type="EnaNcbi">J55446</accessionNumber>
. A, mandible. B, maxillula. C, maxilla. D, first maxilliped. E, second maxilliped. F, third maxilliped. G, paragnath.
</paragraph>
</caption>
<paragraph id="8BE380099476850D52FC0951E6D6F881" blockId="19.[151,1436,1609,2035]" pageId="19" pageNumber="20">
<emphasis id="B9285C1B9476850D52FC0951E5F3F9B0" box="[198,411,1609,1634]" italics="true" pageId="19" pageNumber="20">Second pereiopod</emphasis>
(
<figureCitation id="13679C8C9476850D53950951E67AF9B1" box="[431,530,1609,1635]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="19" pageNumber="20">Fig. 13I</figureCitation>
) small, slender; chela (
<figureCitation id="13679C8C9476850D510B0951E7FEF9B1" box="[817,918,1609,1635]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="19" pageNumber="20">Fig. 13J</figureCitation>
) about 0.8 times CL, palm subcylindrical, smooth, about 4.0 times linger than central width, fingers (
<figureCitation id="13679C8C9476850D51530969E7BFF959" box="[873,983,1649,1675]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="19" pageNumber="20">Fig. 13K</figureCitation>
) 0.85 of palm length, dactyl slender, sparsely setose, 6.2 times longer than basal width, with acute hooked tip, cutting edge unarmed, sharp, entire over distal two thirds, fixed finger similar, tip minutely notched; carpus about 0.6 of palm length, unarmed, 2.5 times longer than distal width, tapering proximally, 1.25 times palm length, uniform, 6.5 times longer than width, unarmed; ischium about 0.8 of merus length, 5.0 times longer than distal width, tapering slightly proximally, basis and coxa without special features.
</paragraph>
<paragraph id="8BE380099476850A52FF0879E0BDFF08" blockId="19.[151,1436,1609,2035]" lastBlockId="20.[151,1435,152,258]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="B9285C1B9476850D52FF0879E5EDF8A8" box="[197,389,1889,1914]" italics="true" pageId="19" pageNumber="20">Third pereiopod</emphasis>
of normal form, dactyl (
<figureCitation id="13679C8C9476850D508B0879E74AF8A9" box="[689,802,1889,1915]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="19" pageNumber="20">Fig. 13M</figureCitation>
) curved, biunguiculate, about 0.26 of propod length, unguis (
<figureCitation id="13679C8C9476850D52CE0891E537F871" box="[244,351,1929,1955]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="19" pageNumber="20">Fig. 13N</figureCitation>
) clearly demarkated, slender, about 4.0 times longer than oblique basal width, 0.63 of corpus length, corpus 3.5 times longer than basal width, dorsal margin concave, ventral margin straight with slightly preterminal slender straight accessory spine, about 0.25 of unguis length; propod (
<figureCitation id="13679C8C9476850D565008C1E0D5F821" box="[1130,1213,2009,2035]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="19" pageNumber="20">Fig. 13</figureCitation>
LM) about 0.57 of CL, 10.5 times longer than wide, with single or paired distoventral spine, slightly shorter than basal width of dactylar corpus, with four similar ventral spines. Fourth and fifth pereiopods similar to third.
</paragraph>
<paragraph id="8BE380099471850A52FF0FF0E15DFED0" blockId="20.[151,1435,152,258]" box="[197,1333,232,258]" pageId="20" pageNumber="21">
<emphasis id="B9285C1B9471850A52FF0FF0E55AFED3" box="[197,306,232,257]" italics="true" pageId="20" pageNumber="21">Pleopods</emphasis>
without special features, endopods of second pleopods without appendices masculinae.
</paragraph>
<caption id="DF23D0819471850A52AD083FE4B0F81B" pageId="20" pageNumber="21" startId="20.[151,255,1831,1855]" targetBox="[151,1409,360,1771]" targetPageId="20">
<paragraph id="8BE380099471850A52AD083FE4B0F81B" blockId="20.[151,1436,1831,1993]" pageId="20" pageNumber="21">
<emphasis id="B9285C1B9471850A52AD083FE54FF8ED" bold="true" box="[151,295,1831,1855]" pageId="20" pageNumber="21">FIGURE 13.</emphasis>
<taxonomicName id="4C5CFB8A9471850A530A0830E628F8ED" authority="Bruce, 2008" authorityName="Bruce" authorityYear="2008" box="[304,576,1832,1855]" class="Malacostraca" family="Palaemonidae" genus="Pseudoclimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="holthuisi" status="gen. nov., sp. nov.">
<emphasis id="B9285C1B9471850A530A0830E628F8ED" box="[304,576,1832,1855]" italics="true" pageId="20" pageNumber="21">Pseudoclimenes holthuisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE1609471850A5073083FE77BF8ED" box="[585,787,1831,1855]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B9285C1B9471850A5073083FE6C6F8ED" bold="true" box="[585,686,1831,1855]" pageId="20" pageNumber="21">gen. nov.</emphasis>
,
<emphasis id="B9285C1B9471850A5086083FE77BF8ED" bold="true" box="[700,787,1831,1855]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, NMV
<accessionNumber id="940F1DEA9471850A51E8083FE04AF8EC" box="[978,1058,1831,1854]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J55446" pageId="20" pageNumber="21" type="EnaNcbi">J55446</accessionNumber>
. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle.
</paragraph>
</caption>
<caption id="DF23D0819470850B52AD0935E681F958" pageId="21" pageNumber="22" startId="21.[151,255,1581,1605]" targetBox="[206,1384,184,1557]" targetPageId="21">
<paragraph id="8BE380099470850B52AD0935E681F958" blockId="21.[151,1436,1581,1674]" pageId="21" pageNumber="22">
<emphasis id="B9285C1B9470850B52AD0935E54EF997" bold="true" box="[151,294,1581,1605]" pageId="21" pageNumber="22">FIGURE 14.</emphasis>
A,
<taxonomicName id="4C5CFB8A9470850B53760936E62AF997" authority="Bruce, 2008" authorityName="Bruce" authorityYear="2008" box="[332,578,1582,1605]" class="Malacostraca" family="Palaemonidae" genus="Apopontonia" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="seticauda" status="sp. nov.">
<emphasis id="B9285C1B9470850B53760936E62AF997" box="[332,578,1582,1605]" italics="true" pageId="21" pageNumber="22">Apopontonia seticauda</emphasis>
</taxonomicName>
<emphasis id="B9285C1B9470850B50720935E6F5F997" bold="true" box="[584,669,1581,1605]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="A21BE1609470850B50720935E6F5F997" box="[584,669,1581,1605]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NMV
<accessionNumber id="940F1DEA9470850B51630936E7C1F997" box="[857,937,1582,1605]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J56496" pageId="21" pageNumber="22" type="EnaNcbi">J56496</accessionNumber>
. B,
<taxonomicName id="4C5CFB8A9470850B51EB0936E1F0F997" authority="Li &amp; Bruce, 2006" authorityName="Li &amp; Bruce" authorityYear="2006" box="[977,1432,1582,1605]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="paralcocki">
<emphasis id="B9285C1B9470850B51EB0936E0BDF997" box="[977,1237,1582,1605]" italics="true" pageId="21" pageNumber="22">Periclimenes paralcocki</emphasis>
<bibRefCitation id="EFCDFDF89470850B56E10936E1F0F997" author="Li, X. &amp; A. J. Bruce" box="[1243,1432,1582,1605]" pageId="21" pageNumber="22" pagination="611 - 738" refId="ref11845" refString="Li, X., &amp; A. J. Bruce. (2006). Further Indo-West Pacific palaemonoid shrimps (Crustacea: Decapoda: Palaemonoidea), principally from the New Caledonian region. Journal of Natural History, 40 (11 - 12), 611 - 738, figs 1 - 31." type="journal article" year="2006">Li &amp; Bruce, 2006</bibRefCitation>
</taxonomicName>
, female, NMV
<accessionNumber id="940F1DEA9470850B53000948E5E3F9B5" box="[314,395,1616,1639]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J56595" pageId="21" pageNumber="22" type="EnaNcbi">J56595</accessionNumber>
. C,
<taxonomicName id="4C5CFB8A9470850B53860949E708F9B5" authority="Bruce, 1991" authorityName="Bruce" authorityYear="1991" box="[444,864,1616,1640]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuirostris">
<emphasis id="B9285C1B9470850B53860949E6A4F9BA" box="[444,716,1617,1640]" italics="true" pageId="21" pageNumber="22">Periclimenes tenuirostris</emphasis>
Bruce, 1991
</taxonomicName>
, ovigerous female, QM W 28404. D,
<taxonomicName id="4C5CFB8A9470850B572B0949E5E8F958" authority="Kubo, 1951" authorityName="Kubo" authorityYear="1951" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="tosaensis">
<emphasis id="B9285C1B9470850B572B0949E491F958" italics="true" pageId="21" pageNumber="22">Periclimenes tosaensis</emphasis>
<bibRefCitation id="EFCDFDF89470850B52C5096BE5E8F958" author="Kubo, I." box="[255,384,1651,1674]" pageId="21" pageNumber="22" pagination="259 - 289" refId="ref11767" refString="Kubo, I. (1951). Some macrurous decapod crustacea found in Japanese waters, with descriptions of four new species. Journal of the Tokyo University of Fisheries, 38, 259 - 289, figs. 1 - 16." type="journal article" year="1951">Kubo, 1951</bibRefCitation>
</taxonomicName>
, ovigerous female, NMV
<accessionNumber id="940F1DEA9470850B50AF096BE68DF958" box="[661,741,1651,1674]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J56497" pageId="21" pageNumber="22" type="EnaNcbi">J56497</accessionNumber>
.
</paragraph>
</caption>
<paragraph id="8BE380099470850B52FC09A3E5AFF84F" blockId="21.[151,1436,1723,2029]" pageId="21" pageNumber="22">
<emphasis id="B9285C1B9470850B52FC09A3E579F906" box="[198,273,1723,1748]" italics="true" pageId="21" pageNumber="22">Telson</emphasis>
(
<figureCitation id="13679C8C9470850B531F09A3E5FAF907" box="[293,402,1723,1749]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="21" pageNumber="22">Fig. 11H</figureCitation>
) about 0.75 of CL, subequal to sixth segment length, 3.0 times longer than anterior width, lateral margins straight, convergent, with two pairs of submarginal dorsal spines, about 0.06 of telson length, at 0.39 and 0.68 of telson length, posterior margin about 0.4 of anterior width (
<figureCitation id="13679C8C9470850B56B00813E080F8F7" box="[1162,1256,1803,1829]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="21" pageNumber="22">Fig. 11I</figureCitation>
) angular, without median point, with three pairs of spines, lateral spines about 0.8 of dorsal spine length, 0.28 of intermediate spine length, intermediate spines about 0.17 of telson length, submedian spines non-setulose, 0.5 of intermediate spine length.
</paragraph>
<paragraph id="8BE380099470850852FF08B3E02FFF60" blockId="21.[151,1436,1723,2029]" lastBlockId="22.[151,1436,152,698]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="B9285C1B9470850B52FF08B3E577F816" box="[197,287,1963,1988]" italics="true" pageId="21" pageNumber="22">Uropod</emphasis>
(
<figureCitation id="13679C8C9470850B530B08B3E5FBF817" box="[305,403,1963,1989]" captionStart="FIGURE 11" captionStartId="18.[154,258,1680,1704]" captionTargetBox="[163,1413,350,1634]" captionTargetId="figure-191@18.[160,1426,350,1652]" captionTargetPageId="18" captionText="FIGURE 11. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, carapace and rostrum. B, rostrum. C, inferior orbital angle and antennal spine. D, antennule. E, same, proximal segment. F, antenna. G, eye. H, telson. I, same, posterior spines, dorsal spine inset. J, uropod." pageId="21" pageNumber="22">Fig. 11J</figureCitation>
) with protopodite unarmed; exopod slightly exceeding endopod, 3.5 times longer than wide, lateral margin straight, with slender distolateral tooth and spine (
<figureCitation id="13679C8C9470850B51CB08CBE009F83F" box="[1009,1121,2003,2029]" captionStart="FIGURE 13" captionStartId="20.[151,255,1831,1855]" captionTargetBox="[151,1409,360,1771]" captionTargetId="figure-138@20.[151,1436,314,1807]" captionTargetPageId="20" captionText="FIGURE 13. Pseudoclimenes holthuisi gen. nov., sp. nov., paratype, NMV J55446. A, antennule, distolateral angle of proximal segment. B, scaphocerite, distolateral spine. C, mandible, molar process. D, same, incisor process. E, maxillula, palp. F, same, upper lacinia. G, first pereiopod. H, same, chela. I, second pereiopod. J, same, chela. K, same, fingers. L, third pereiopod, propod and dactyl. M, same, distal propod and dactyl. N, same, unguis. O, exopod of uropod, distolateral angle." pageId="21" pageNumber="22">Fig. 13O</figureCitation>
), with sparse submarginal ventral setae; endopod about 0.85 of exopod length, 3.6 times longer than width.
</paragraph>
</subSubSection>
<subSubSection id="C346D3829473850852FF0FA7E11CFEF8" pageId="22" pageNumber="23" type="etymology">
<paragraph id="8BE380099473850852FF0FA7E11CFEF8" blockId="22.[151,1436,152,698]" pageId="22" pageNumber="23">
<emphasis id="B9285C1B9473850852FF0FA7E53AFF0B" bold="true" box="[197,338,191,217]" pageId="22" pageNumber="23">Etymology.</emphasis>
Named after Professor Lipke Bijdeley Holthuis (
<date id="FFE2A6C994738508518E0FD8E00AFF08" box="[948,1122,192,218]" pageId="22" pageNumber="23" value="1921-04-21">21 April 1921</date>
<date id="FFE2A6C99473850856480FD8E14AFF08" box="[1138,1314,192,218]" pageId="22" pageNumber="23" value="2008-03-08">8 March 2008</date>
), news of whose death was received today, in recognition of his outstanding contribution to carcinology in general and the study of the
<taxonomicName id="4C5CFB8A9473850853690E08E5B7FEF8" authorityName="Kingsley" authorityYear="1878" box="[339,479,272,298]" class="Malacostraca" family="Pontoniinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="family">Pontoniinae</taxonomicName>
in particular, and his never failing assistance to the author over half a century.
</paragraph>
</subSubSection>
<subSubSection id="C346D3829473850852FF0E2FE75BFEA8" pageId="22" pageNumber="23" type="description">
<paragraph id="8BE380099473850852FF0E2FE75BFEA8" blockId="22.[151,1436,152,698]" pageId="22" pageNumber="23">
<emphasis id="B9285C1B9473850852FF0E2FE5ECFE83" bold="true" box="[197,388,311,337]" pageId="22" pageNumber="23">Measurements.</emphasis>
<typeStatus id="54E73EAB9473850853AA0E20E595FE80" box="[400,509,312,338]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
female, postorbital carapace length, 1.85; carapace and rostrum, 3.45; total body length (approx.), 10.4, second pereiopod chela, 2.0.
</paragraph>
</subSubSection>
<subSubSection id="C346D3829473850852FF0E9FE77BFE70" box="[197,787,391,418]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BE380099473850852FF0E9FE77BFE70" blockId="22.[151,1436,152,698]" box="[197,787,391,418]" pageId="22" pageNumber="23">
<emphasis id="B9285C1B9473850852FF0E9FE50EFE73" bold="true" box="[197,358,391,417]" pageId="22" pageNumber="23">Distribution.</emphasis>
Known only from the
<typeStatus id="54E73EAB94738508504B0E90E6CBFE70" box="[625,675,392,418]" pageId="22" pageNumber="23">type</typeStatus>
material.
</paragraph>
</subSubSection>
<subSubSection id="C346D3829473850852FF0EB7E596FD68" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BE380099473850852FF0EB7E596FD68" blockId="22.[151,1436,152,698]" pageId="22" pageNumber="23">
<emphasis id="B9285C1B9473850852FF0EB7E556FE1B" bold="true" box="[197,318,431,457]" pageId="22" pageNumber="23">Remarks.</emphasis>
Apart from the lack of a hepatic spine,
<taxonomicName id="4C5CFB8A94738508512D0EA8E02AFE1B" authorityName="Bruce" authorityYear="2008" box="[791,1090,432,457]" class="Malacostraca" family="Palaemonidae" genus="Pseudoclimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B9285C1B94738508512D0EA8E02AFE1B" box="[791,1090,432,457]" italics="true" pageId="22" pageNumber="23">Pseudoclimenes holthuisi</emphasis>
</taxonomicName>
bears a close resemblance to species of the
<taxonomicName id="4C5CFB8A9473850853730EC0E63EFE23" authorityName="Kemp" authorityYear="1922" box="[329,598,472,497]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B9285C1B9473850853730EC0E63EFE23" box="[329,598,472,497]" italics="true" pageId="22" pageNumber="23">Periclimenes obscurus</emphasis>
</taxonomicName>
species group Bruce (1987; in press, 2008), particularly
<taxonomicName id="4C5CFB8A9473850857280EC0E1F3FE23" authorityName="Kemp" authorityYear="1922" box="[1298,1435,472,497]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B9285C1B9473850857280EC0E1F3FE23" box="[1298,1435,472,497]" italics="true" pageId="22" pageNumber="23">P. obscurus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCDFDF89473850852A50D18E55BFDC8" author="Kemp, S." box="[159,307,512,538]" pageId="22" pageNumber="23" pagination="113 - 288" refId="ref11722" refString="Kemp, S. (1922). Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum. 24, 113 - 288, figs. 1 - 105, pls. 3 - 9." type="journal article" year="1922">Kemp, 1922</bibRefCitation>
, fig. 15). The ambulatory dactyls in the
<taxonomicName id="4C5CFB8A94738508513C0D18E718FDCB" authorityName="Kemp" authorityYear="1922" box="[774,880,512,537]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B9285C1B94738508513C0D18E718FDCB" box="[774,880,512,537]" italics="true" pageId="22" pageNumber="23">obscurus</emphasis>
</taxonomicName>
group are generally similar, being simply biunguiculate. The accessory spines are usually slender, about half the unguis length and originate immediately adjacent to the base of the unguis, projecting more or less in the line of the unguis. In
<taxonomicName id="4C5CFB8A9473850856AB0D48E4A9FD43" authorityName="Bruce" authorityYear="2008" class="Malacostraca" family="Palaemonidae" genus="Pseudoclimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B9285C1B9473850856AB0D48E4A9FD43" italics="true" pageId="22" pageNumber="23">Pseudoclimenes holthuisi</emphasis>
</taxonomicName>
, the slender accessory spine is slightly remote from the base of the unguis and projects at a divergent angle to the axis of the unguis.
</paragraph>
</subSubSection>
</treatment>
</document>