333 lines
34 KiB
XML
333 lines
34 KiB
XML
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<mods:title id="FA9C90CF61C94AAA9D6CF1B7C8CF4F7D">Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)</mods:title>
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<mods:namePart id="29270F6969030C3FD64A5947680E03F5">Toledo, Mario</mods:namePart>
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<mods:namePart id="E9C908A7951F3BD3DB6CCA55916C7938">Michat, Mariano C.</mods:namePart>
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<mods:title id="E6D7588DC7BCC3DE1892E33B8076F7FE">Zootaxa</mods:title>
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<mods:date id="86D32E229A7DC6CBA7984F01CD86B5AC">2015</mods:date>
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<treatment id="03EE87FEFFFBFFFD91A6FDF5FA64A1A6" ID-DOI="http://doi.org/10.5281/zenodo.5692038" ID-GBIF-Taxon="119638427" ID-Zenodo-Dep="5692038" LSID="urn:lsid:plazi:treatment:03EE87FEFFFBFFFD91A6FDF5FA64A1A6" httpUri="http://treatment.plazi.org/id/03EE87FEFFFBFFFD91A6FDF5FA64A1A6" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
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<paragraph id="8BF836E8FFFBFFFC91A6FDF5FD83A25C" blockId="8.[151,635,584,610]" box="[151,635,584,610]" pageId="8" pageNumber="309">
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<heading id="D0B08184FFFBFFFC91A6FDF5FD83A25C" bold="true" box="[151,635,584,610]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
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<emphasis id="B933EAFAFFFBFFFC91A6FDF5FD83A25C" bold="true" box="[151,635,584,610]" pageId="8" pageNumber="309">
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Key to genera of the tribe
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<taxonomicName id="4C474D6BFFFBFFFC90EFFDF5FD83A25C" ID-CoL="KX5" box="[478,635,584,610]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF836E8FFFBFFFC91A6FD33FAA1A3F9" blockId="8.[151,1436,654,967]" pageId="8" pageNumber="309">
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Several authors provided keys for
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<taxonomicName id="4C474D6BFFFBFFFC931DFD33FD33A299" box="[556,715,654,679]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subFamily" subFamily="Laccophilinae">Laccophilinae</taxonomicName>
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genera, often associated with description of new genera (e.g.
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<bibRefCitation id="EFD64B19FFFBFFFC91A6FD0FFE6AA2F5" author="Balfour-Browne" box="[151,402,690,715]" pageId="8" pageNumber="309" refString="Balfour-Browne, J. (1939) A contribution to the study of the Dytiscidae - I. (Coleoptera, Adephaga). Annals and Magazine of Natural History, Series 11, 3, 97 - 114." type="journal article" year="1939">Balfour-Browne 1939</bibRefCitation>
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,
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<bibRefCitation id="EFD64B19FFFBFFFC90AEFD0FFDC9A2F5" author="Steiner" box="[415,561,690,715]" pageId="8" pageNumber="309" refString="Steiner Jr., W. E. (1981) A new genus and a new species of laccophiline water beetle from Ecuador (Coleoptera: Dytiscidae). Pan-Pacific Entomologist, 57, 251 - 259." type="journal article" year="1981">Steiner 1981</bibRefCitation>
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,
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<bibRefCitation id="EFD64B19FFFBFFFC930EFD0FFCFBA2F5" author="Brancucci" box="[575,771,690,715]" pageId="8" pageNumber="309" refString="Brancucci, M. (1983 a) A new genus of the subfamily Laccophilinae (Coleoptera, Dytiscidae). Aquatic Insects, 5, 351 - 254. http: // dx. doi. org / 10.1080 / 01650428309361151" type="journal volume" year="1983" yearSuffix="a">Brancucci 1983a</bibRefCitation>
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). The most recent key was provided by
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<bibRefCitation id="EFD64B19FFFBFFFC95E2FD0FFA6FA2F5" author="Pederzani" box="[1235,1431,690,715]" pageId="8" pageNumber="309" refString="Pederzani, F. (1995) Keys to the identification of the genera and subgenera of adult Dytiscidae (sensu lato) of the world (Coleoptera, Dytiscidae). Atti dell'Accademia Roveretana degli Agiati, a. 244 (1994), Series VII, IVB, 5 - 83." type="journal article" year="1995">Pederzani (1995)</bibRefCitation>
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, including 11 genera and two subgenera as the author did not consider
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<taxonomicName id="4C474D6BFFFBFFFC92DAFD6BFB0EA2D1" authority="Crotch, 1873" authorityName="Crotch" authorityYear="1873" box="[1003,1270,726,751]" class="Insecta" family="Dytiscidae" genus="Agabetes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAFAFFFBFFFC92DAFD6BFBAAA2D0" box="[1003,1106,726,750]" italics="true" pageId="8" pageNumber="309">Agabetes</emphasis>
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Crotch, 1873
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</taxonomicName>
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a member of
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<taxonomicName id="4C474D6BFFFBFFFC91A6FD47FECEA32D" box="[151,310,762,787]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subFamily" subFamily="Laccophilinae">Laccophilinae</taxonomicName>
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(
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<bibRefCitation id="EFD64B19FFFBFFFC9075FD47FDFEA32D" author="Burmeister" box="[324,518,762,787]" pageId="8" pageNumber="309" refString="Burmeister, E. - G. (1990) The systematic position of the genus Agabetes Crotch within Dytiscidae (Coleoptera, Adephaga). Quaestiones Entomologicae, 26, 221 - 238." type="journal article" year="1990">Burmeister 1990</bibRefCitation>
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;
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<bibRefCitation id="EFD64B19FFFBFFFC9320FD47FD21A32D" author="Alarie" box="[529,729,762,787]" pageId="8" pageNumber="309" refString="Alarie, Y., Spangler, P. J. & Steiner, W. E. Jr. (2002) Larval morphology of Agabetes Crotch (Coleoptera: Adephaga: Dytiscidae): the hypothesis of sister-group relationship with the subfamily Laccophilinae revisited. The Coleopterists Bulletin, 56, 547 - 567. http: // dx. doi. org / 10.1649 / 0010 - 065 X (2002) 056 [0547: LMOACC] 2.0. CO; 2" type="journal article" year="2002">
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Alarie
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<emphasis id="B933EAFAFFFBFFFC936FFD46FD6EA32D" box="[606,662,763,787]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
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2002
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</bibRefCitation>
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), and
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<taxonomicName id="4C474D6BFFFBFFFC922CFD46FBA2A32D" authority="Guignot, 1937" authorityName="Guignot" authorityYear="1937" box="[797,1114,762,787]" class="Insecta" family="Dytiscidae" genus="Philaccolilus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAFAFFFBFFFC922CFD46FC49A32D" box="[797,945,763,787]" italics="true" pageId="8" pageNumber="309">Philaccolilus</emphasis>
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Guignot, 1937
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</taxonomicName>
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had not yet been elevated to generic rank (Balke
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<emphasis id="B933EAFAFFFBFFFC90B2FCA2FE46A308" box="[387,446,798,822]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
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2000). Two years later, the reliability of the bifid metatibial spurs as a character that unequivocally defines
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<taxonomicName id="4C474D6BFFFBFFFC90A9FCFEFDD9A365" box="[408,545,835,859]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAFAFFFBFFFC90A9FCFEFDD9A365" box="[408,545,835,859]" italics="true" pageId="8" pageNumber="309">Laccophilus</emphasis>
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</taxonomicName>
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was discussed (
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<bibRefCitation id="EFD64B19FFFBFFFC93EAFCFFFC5CA365" author="Balke" box="[731,932,834,859]" pageId="8" pageNumber="309" refString="Balke, M., Larson, D. J. & Hendrich, L. (1997) A review of the New Guinean species of Laccophilus Leach 1815 with notes on regional melanism (Coleoptera, Dytiscidae). Tropical Zoology, 10, 293 - 320. http: // dx. doi. org / 10.1080 / 03946975.1997.10539344" type="journal article" year="1997">
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Balke
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<emphasis id="B933EAFAFFFBFFFC9217FCFEFC99A365" box="[806,865,835,859]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
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1997
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</bibRefCitation>
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), after the discovery of a species from New
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<collectingCountry id="F3507678FFFBFFFC91A6FCDBFF10A341" box="[151,232,870,895]" name="Guinea-Bissau" pageId="8" pageNumber="309">Guinea</collectingCountry>
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which has simple spurs. In this vein the authors suggested that
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<taxonomicName id="4C474D6BFFFBFFFC9530FCDBFB80A340" box="[1025,1144,870,894]" class="Insecta" family="Dytiscidae" genus="Philodytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAFAFFFBFFFC9530FCDBFB80A340" box="[1025,1144,870,894]" italics="true" pageId="8" pageNumber="309">Philodytes</emphasis>
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</taxonomicName>
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is a junior synonym of
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<taxonomicName id="4C474D6BFFFBFFFC91A6FC36FED8A39D" box="[151,288,907,931]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAFAFFFBFFFC91A6FC36FED8A39D" box="[151,288,907,931]" italics="true" pageId="8" pageNumber="309">Laccophilus</emphasis>
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</taxonomicName>
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, although this synonymy is not currently supported (
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<bibRefCitation id="EFD64B19FFFBFFFC92AAFC37FB8CA39D" author="Miller" box="[923,1140,906,931]" pageId="8" pageNumber="309" refString="Miller, K. B., Alarie, Y., Wolfe, W. G. & Whiting, M. F. (2005) Associations of insect life stages using DNA sequences: the larva of Phylodytes umbrinus (Motschulsky) (Coleoptera: Dytiscidae). Systematic Entomology, 30, 499 - 509. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2005.00320. x" type="journal article" year="2005">
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Miller
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<emphasis id="B933EAFAFFFBFFFC92DEFC36FBDDA39D" box="[1007,1061,907,931]" italics="true" pageId="8" pageNumber="309">et al</emphasis>
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. 2005
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</bibRefCitation>
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;
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<bibRefCitation id="EFD64B19FFFBFFFC95B4FC37FADCA39D" author="Nilsson" box="[1157,1316,906,931]" pageId="8" pageNumber="309" refString="Nilsson, A. N. (2015) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2015. Umea: distributed electronically as a PDF file by the author, 298 pp. Available from: http: // www 2. emg. umu. se / projects / biginst / andersn / World catalogue of Dytiscidae 2015. pdf (accessed 2 June 2015)" type="book" year="2015">Nilsson 2015</bibRefCitation>
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). All this, together with the new genus described here, makes it necessary to develop a new version of Pederzani’s key:
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6563FFFBFFFD91A6FC4BFA64A1A6" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309" type="key">
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<keyStep id="30B32E4DFFFBFFFC91A6FC4BFA63A417" pageId="8" pageNumber="309">
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<paragraph id="8BF836E8FFFBFFFC91A6FC4BFA63A432" blockId="8.[151,1436,1014,2033]" box="[151,1435,1014,1036]" pageId="8" pageNumber="309">
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<keyLead id="30B695DDFFFBFFFC91A6FC4BFA63A432" box="[151,1435,1014,1036]" pageId="8" pageNumber="309">
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1. Metatibiae with a single apical spur. Antennal segments enlarged and flattened in male. Neotropical............
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<taxonomicName id="4C474D6BFFFBFFFC9404FC4BFA63A432" ID-CoL="5XL2" box="[1333,1435,1014,1036]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAFAFFFBFFFC9404FC4BFA63A432" bold="true" box="[1333,1435,1014,1036]" italics="true" pageId="8" pageNumber="309">Napodytes</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BF836E8FFFBFFFC91A6FBA9FA63A417" blockId="8.[151,1436,1014,2033]" box="[151,1435,1044,1065]" pageId="8" pageNumber="309">
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<keyLead id="30B695DDFFFBFFFC91A6FBA9FA63A417" box="[151,1435,1044,1065]" pageId="8" pageNumber="309">- Metatibiae with 2 apical spurs. Antennal segments never enlarged or flattened in male............................... 2</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30B32E4DFFFBFFFC91A6FB8CFA63A4A0" pageId="8" pageNumber="309">
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<paragraph id="8BF836E8FFFBFFFC91A6FB8CFA63A45D" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
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<keyLead id="30B695DDFFFBFFFC91A6FB8CFA63A45D" pageId="8" pageNumber="309">2. Suture between elytron and epipleuron visible dorsally (= “lateral sides bordered”: in dorsal view elytra visibly outlined by a fine border along outer lateral side)....................................................................... 3</keyLead>
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</paragraph>
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<paragraph id="8BF836E8FFFBFFFC91A6FBD1FA63A4A0" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
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||
<keyLead id="30B695DDFFFBFFFC91A6FBD1FA63A4A0" pageId="8" pageNumber="309">- Suture between elytron and epipleuron not visible dorsally (= “lateral sides not bordered”: in dorsal view the elytra do not appear to be bordered)................................................................................. 5</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30B32E4DFFFBFFFC91A6FB1BFA63A52D" pageId="8" pageNumber="309">
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||
<paragraph id="8BF836E8FFFBFFFC91A6FB1BFA64A4E7" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
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||
<keyLead id="30B695DDFFFBFFFC91A6FB1BFA64A4E7" pageId="8" pageNumber="309">
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||
3.
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<collectingCountry id="F3507678FFFBFFFC91D0FB1BFEE9A485" box="[225,273,1190,1211]" name="India" pageId="8" pageNumber="309">Hind</collectingCountry>
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angles of pronotum rounded; base of pronotum projecting hindwards in a distinct angle at middle; prosternal process not extending beyond mesocoxae. Neotropical........................................
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<emphasis id="B933EAFAFFFBFFFC9554FB7EFA64A4E7" bold="true" box="[1125,1436,1219,1241]" pageId="8" pageNumber="309">
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||
<taxonomicName id="4C474D6BFFFBFFFC9554FB7EFAAEA4E7" ID-CoL="3RLZL" box="[1125,1366,1219,1241]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="phalacroides">
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<emphasis id="B933EAFAFFFBFFFC9554FB7EFAAEA4E7" bold="true" box="[1125,1366,1219,1241]" italics="true" pageId="8" pageNumber="309">Laccodytes phalacroides</emphasis>
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</taxonomicName>
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-group
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8BF836E8FFFBFFFC91A6FB5CFA63A52D" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
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<keyLead id="30B695DDFFFBFFFC91A6FB5CFA63A52D" pageId="8" pageNumber="309">
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-
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<collectingCountry id="F3507678FFFBFFFC91D0FB5CFEE9A4C8" box="[225,273,1249,1270]" name="India" pageId="8" pageNumber="309">Hind</collectingCountry>
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angles of pronotum extending posteriorly into acuminate spine; base of pronotum straight or almost so; prosternal pro- cess long, extending beyond mesocoxae................................................................... 4
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30B32E4DFFFBFFFC91A6FAA1FA64A5B7" pageId="8" pageNumber="309">
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<paragraph id="8BF836E8FFFBFFFC91A6FAA1FA63A570" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
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<keyLead id="30B695DDFFFBFFFC91A6FAA1FA63A570" pageId="8" pageNumber="309">
|
||
4. Prosternal process three-pointed; posterolateral lobes of metatarsi very short, almost lacking; apex of elytra rounded. Afrotro- pical and Oriental...........................................................................
|
||
<taxonomicName id="4C474D6BFFFBFFFC9428FA85FA63A570" ID-CoL="639Q" box="[1305,1435,1336,1358]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFBFFFC9428FA85FA63A570" bold="true" box="[1305,1435,1336,1358]" italics="true" pageId="8" pageNumber="309">Neptosternus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFBFFFC91A6FAEBFA64A5B7" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6FAEBFA64A5B7" pageId="8" pageNumber="309">
|
||
- Prosternal process single-pointed; posterolateral lobes of metatarsi longer, well developed; apex of elytra truncate. Neotropical..............................................................................
|
||
<emphasis id="B933EAFAFFFBFFFC95BCFACEFA64A5B7" bold="true" box="[1165,1436,1395,1417]" pageId="8" pageNumber="309">
|
||
<taxonomicName id="4C474D6BFFFBFFFC95BCFACEFAAEA5B7" ID-CoL="3RLZF" box="[1165,1366,1395,1417]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="B933EAFAFFFBFFFC95BCFACEFAAEA5B7" bold="true" box="[1165,1366,1395,1417]" italics="true" pageId="8" pageNumber="309">Laccodytes apalodes</emphasis>
|
||
</taxonomicName>
|
||
-group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFBFFFC91A6FA2CFA63A5DF" pageId="8" pageNumber="309">
|
||
<paragraph id="8BF836E8FFFBFFFC91A6FA2CFA64A5FA" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6FA2CFA64A5FA" pageId="8" pageNumber="309">
|
||
5. Metatibial spurs bifid apically (except
|
||
<taxonomicName id="4C474D6BFFFBFFFC9365FA2CFC10A598" authority="Balke, Larson & Hendrich, 1997" authorityName="Balke, Larson & Hendrich" authorityYear="1997" box="[596,1000,1425,1446]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="bapak">
|
||
<emphasis id="B933EAFAFFFBFFFC9365FA2CFD50A598" box="[596,680,1425,1446]" italics="true" pageId="8" pageNumber="309">L. bapak</emphasis>
|
||
<bibRefCitation id="EFD64B19FFFBFFFC939CFA2CFC10A598" author="Balke" box="[685,1000,1425,1446]" pageId="8" pageNumber="309" refString="Balke, M., Larson, D. J. & Hendrich, L. (1997) A review of the New Guinean species of Laccophilus Leach 1815 with notes on regional melanism (Coleoptera, Dytiscidae). Tropical Zoology, 10, 293 - 320. http: // dx. doi. org / 10.1080 / 03946975.1997.10539344" type="journal article" year="1997">Balke, Larson & Hendrich, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3507678FFFBFFFC9513FA2CFB18A598" box="[1058,1248,1425,1446]" name="Papua New Guinea" pageId="8" pageNumber="309">Papua New Guinea</collectingCountry>
|
||
). Worldwide.................................................................................................
|
||
<taxonomicName id="4C474D6BFFFBFFFC9412FA13FA64A5FA" ID-CoL="59RR" box="[1315,1436,1454,1476]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFBFFFC9412FA13FA64A5FA" bold="true" box="[1315,1436,1454,1476]" italics="true" pageId="8" pageNumber="309">Laccophilus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFBFFFC91A6FA71FA63A5DF" blockId="8.[151,1436,1014,2033]" box="[151,1435,1484,1505]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6FA71FA63A5DF" box="[151,1435,1484,1505]" pageId="8" pageNumber="309">- Metatibial spurs simple, apically acute..................................................................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFBFFFC91A6FA54FA63A625" pageId="8" pageNumber="309">
|
||
<paragraph id="8BF836E8FFFBFFFC91A6FA54FA63A5C0" blockId="8.[151,1436,1014,2033]" box="[151,1435,1513,1534]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6FA54FA63A5C0" box="[151,1435,1513,1534]" pageId="8" pageNumber="309">6. Base of pronotum projecting hindwards in a distinct angle at middle............................................ 7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F9BBFA63A625" blockId="8.[151,1436,1014,2033]" box="[151,1435,1542,1563]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F9BBFA63A625" box="[151,1435,1542,1563]" pageId="8" pageNumber="309">- Base of pronotum straight or almost so.................................................................... 11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFBFFFC91A6F999FA63A668" pageId="8" pageNumber="309">
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F999FA63A607" blockId="8.[151,1436,1014,2033]" box="[151,1435,1572,1593]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F999FA63A607" box="[151,1435,1572,1593]" pageId="8" pageNumber="309">7. Prosternal process laterally compressed behind procoxae...................................................... 8</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F9FCFA63A668" blockId="8.[151,1436,1014,2033]" box="[151,1435,1601,1622]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F9FCFA63A668" box="[151,1435,1601,1622]" pageId="8" pageNumber="309">- Prosternal process fairly broad behind procoxae............................................................. 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFBFFFC91A6F9E3FA63A6F2" pageId="8" pageNumber="309">
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F9E3FA64A6AF" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F9E3FA64A6AF" pageId="8" pageNumber="309">
|
||
8. Length not exceeding
|
||
<quantity id="4CBF9B0DFFFBFFFC9087F9E3FE08A64D" box="[438,496,1630,1651]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="309" unit="mm" value="5.0">5 mm</quantity>
|
||
; pro- and mesofemora and tibiae densely punctate; prosternal process markedly carinate. Tibet....................................................
|
||
<taxonomicName id="4C474D6BFFFBFFFC920DF9C6FC48A6AF" ID-CoL="59RT" box="[828,944,1659,1681]" class="Insecta" family="Dytiscidae" genus="Laccoporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFBFFFC920DF9C6FC48A6AF" bold="true" box="[828,944,1659,1681]" italics="true" pageId="8" pageNumber="309">Laccoporus</emphasis>
|
||
</taxonomicName>
|
||
(re-description and illustration in
|
||
<bibRefCitation id="EFD64B19FFFBFFFC95C6F9C1FA6DA6AF" author="Zhao" box="[1271,1429,1660,1681]" pageId="8" pageNumber="309" refString="Zhao, S., Jia, F. & Balke, M. (2011) Rhantus fengi sp. n. from Xizang, China, and notes on Laccoporus nigritulus (Gschwendtner) (Coleoptera, Dytiscidae). ZooKeys, 94, 61 - 71." type="journal article" year="2011">
|
||
Zhao
|
||
<emphasis id="B933EAFAFFFBFFFC941EF9C1FA98A6AF" box="[1327,1376,1660,1681]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F924FA63A6F2" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F924FA63A6F2" pageId="8" pageNumber="309">
|
||
- Length exceeding
|
||
<quantity id="4CBF9B0DFFFBFFFC90A3F924FE34A690" box="[402,460,1689,1710]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="309" unit="mm" value="5.0">5 mm</quantity>
|
||
; pro- and mesofemora and tibiae never densely punctate; prosternal process slightly carinate. Africa.............................................................................................
|
||
<taxonomicName id="4C474D6BFFFBFFFC9404F90BFA63A6F2" ID-CoL="6NGQ" box="[1333,1435,1718,1740]" class="Insecta" family="Dytiscidae" genus="Philodytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFBFFFC9404F90BFA63A6F2" bold="true" box="[1333,1435,1718,1740]" italics="true" pageId="8" pageNumber="309">Philodytes</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFBFFFC91A6F969FA64A77F" pageId="8" pageNumber="309">
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F969FA63A738" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F969FA63A738" pageId="8" pageNumber="309">
|
||
9. Metacoxal lines straight and fairly parallel; body short and broad, strongly narrowed caudally; prosternal process triangular behind procoxae, with acute apex. East Palearctic..............................................
|
||
<taxonomicName id="4C474D6BFFFBFFFC95D0F94DFA63A738" ID-CoL="62W6K" box="[1249,1435,1776,1798]" class="Insecta" family="Dytiscidae" genus="Japanolaccophilus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFBFFFC95D0F94DFA63A738" bold="true" box="[1249,1435,1776,1798]" italics="true" pageId="8" pageNumber="309">Japanolaccophilus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F8B3FA64A77F" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F8B3FA64A77F" pageId="8" pageNumber="309">- Metacoxal lines sinuate and strongly convergent anteriorly; body elongate, more gently narrowed caudally; prosternal process broadly triangular to cordiform behind procoxae, with apex rounded to acute.................................... 10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFBFFFC91A6F8F4FA63A788" pageId="8" pageNumber="309">
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F8F4FA63A742" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F8F4FA63A742" pageId="8" pageNumber="309">
|
||
10. Mesotibial spurs not longer than segments 1–2 of mesotarsi combined; elytra covered by numerous, more or less impressed punctures. Oriental..........................................................................
|
||
<taxonomicName id="4C474D6BFFFBFFFC9426F8DBFA63A742" ID-CoL="59S8" box="[1303,1435,1894,1916]" class="Insecta" family="Dytiscidae" genus="Laccosternus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFBFFFC9426F8DBFA63A742" bold="true" box="[1303,1435,1894,1916]" italics="true" pageId="8" pageNumber="309">Laccosternus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F839FA63A788" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F839FA63A788" pageId="8" pageNumber="309">
|
||
- Mesotibial spurs longer than segments 1–4 of mesotarsi combined; elytra lacking punctures, at most bearing very few, small, hardly visible punctures. Neotropical and SE Nearctic...............................................
|
||
<taxonomicName id="4C474D6BFFFBFFFC942CF81DFA63A788" ID-CoL="8LY8Y" box="[1309,1435,1952,1974]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B933EAFAFFFBFFFC942CF81DFA63A788" bold="true" box="[1309,1435,1952,1974]" italics="true" pageId="8" pageNumber="309">Laccomimus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFBFFFD91A6F803FA64A092" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
|
||
<paragraph id="8BF836E8FFFBFFFC91A6F803FA64A7CF" blockId="8.[151,1436,1014,2033]" pageId="8" pageNumber="309">
|
||
<keyLead id="30B695DDFFFBFFFC91A6F803FA64A7CF" pageId="8" pageNumber="309">
|
||
11. Reticulation on elytra consisting of longitudinal or broad polygonal meshes; metacoxal lines fairly parallel. Afrotropical...............................................................................................
|
||
<taxonomicName id="4C474D6BFFFBFFFC9411F866FA64A7CF" ID-CoL="Q38" box="[1312,1436,2011,2033]" class="Insecta" family="Dytiscidae" genus="Africophilus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFBFFFC9411F866FA64A7CF" bold="true" box="[1312,1436,2011,2033]" italics="true" pageId="8" pageNumber="309">Africophilus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFAFFFD91A6FF2AFA64A092" blockId="9.[151,1436,151,408]" box="[151,1436,151,172]" pageId="9" pageNumber="310">
|
||
<keyLead id="30B695DDFFFAFFFD91A6FF2AFA64A092" box="[151,1436,151,172]" pageId="9" pageNumber="310">- Reticulation on elytra consisting of small, transversal meshes; metacoxal lines more or less convergent anteriorly....... 12</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFAFFFD91A6FF08FA63A11C" pageId="9" pageNumber="310">
|
||
<paragraph id="8BF836E8FFFAFFFD91A6FF08FA63A0D6" blockId="9.[151,1436,151,408]" pageId="9" pageNumber="310">
|
||
<keyLead id="30B695DDFFFAFFFD91A6FF08FA63A0D6" pageId="9" pageNumber="310">
|
||
12. Space between metacoxal lines strongly rugose-punctate; metacoxal processes with round meshes and distally bilobed.
|
||
<collectingCountry id="F3507678FFFAFFFD9462FF08FF01A0D9" name="Australia" pageId="9" pageNumber="310">Australia</collectingCountry>
|
||
.......................................................................................
|
||
<taxonomicName id="4C474D6BFFFAFFFD9424FF6FFA63A0D6" ID-CoL="365J" box="[1301,1435,210,232]" class="Insecta" family="Dytiscidae" genus="Australphilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFAFFFD9424FF6FFA63A0D6" bold="true" box="[1301,1435,210,232]" italics="true" pageId="9" pageNumber="310">Australphilus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFAFFFD91A6FF52FA63A11C" blockId="9.[151,1436,151,408]" pageId="9" pageNumber="310">
|
||
<keyLead id="30B695DDFFFAFFFD91A6FF52FA63A11C" pageId="9" pageNumber="310">- Space between metacoxal lines somewhat smooth; metacoxal processes reticulate with transversal meshes and distally not bilobed............................................................................................. 13</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B32E4DFFFAFFFD91A6FE97FA64A1A6" pageId="9" pageNumber="310">
|
||
<paragraph id="8BF836E8FFFAFFFD91A6FE97FA64A163" blockId="9.[151,1436,151,408]" pageId="9" pageNumber="310">
|
||
<keyLead id="30B695DDFFFAFFFD91A6FE97FA64A163" pageId="9" pageNumber="310">
|
||
13. Metacoxa with stridulatory file; posterolatral lobes of metatarsi short; ventrite
|
||
<quantity id="4CBF9B0DFFFAFFFD92E0FE97FC0EA101" box="[977,1014,298,319]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="310" unit="in" value="6.0">6 in</quantity>
|
||
females rounded apically. Afrotropical...............................................................................................
|
||
<taxonomicName id="4C474D6BFFFAFFFD9419FEFAFA64A163" ID-CoL="63N6W" box="[1320,1436,327,349]" class="Insecta" family="Dytiscidae" genus="Philaccolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFAFFFD9419FEFAFA64A163" bold="true" box="[1320,1436,327,349]" italics="true" pageId="9" pageNumber="310">Philaccolus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFFAFFFD91A6FED8FA64A1A6" blockId="9.[151,1436,151,408]" pageId="9" pageNumber="310">
|
||
<keyLead id="30B695DDFFFAFFFD91A6FED8FA64A1A6" pageId="9" pageNumber="310">
|
||
- Metacoxa lacking stridulatory file; posterolateral lobes of metatarsi very long; ventrite
|
||
<quantity id="4CBF9B0DFFFAFFFD9518FED8FBB7A144" box="[1065,1103,357,378]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="310" unit="in" value="6.0">6 in</quantity>
|
||
females with an apical lobe, more or less developed.
|
||
<collectingCountry id="F3507678FFFAFFFD90A2FE3FFDB7A1A9" box="[403,591,386,407]" name="Papua New Guinea" pageId="9" pageNumber="310">Papua New Guinea</collectingCountry>
|
||
............................................................
|
||
<taxonomicName id="4C474D6BFFFAFFFD942AFE3FFA64A1A6" ID-CoL="63N79" box="[1307,1436,386,408]" class="Insecta" family="Dytiscidae" genus="Philaccolilus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAFAFFFAFFFD942AFE3FFA64A1A6" bold="true" box="[1307,1436,386,408]" italics="true" pageId="9" pageNumber="310">Philaccolilus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |