233 lines
28 KiB
XML
233 lines
28 KiB
XML
<document id="8451D11D354720FA028E2B80C6C62A62" ID-DOI="10.11646/zootaxa.3990.3.1" ID-GBIF-Dataset="cf747765-b9bf-42ed-8479-35355e712db0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244301" ID-ZooBank="378C0359-E0E4-4CCC-821D-649144E37A63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461268581225" checkinUser="plazi" docAuthor="Toledo, Mario & Michat, Mariano C." docDate="2015" docId="03EE87FEFFE6FFE291A6FA08FCB8A49E" docLanguage="en" docName="zt03990p354.pdf" docOrigin="Zootaxa 3990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccomimus spangleri Toledo & Michat, 2015, sp.n." docType="treatment" docVersion="9" lastPageNumber="323" masterDocId="FFD7FF86FFF3FFF49131FFBDFFF8A03E" masterDocTitle="Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)" masterLastPageNumber="354" masterPageNumber="301" pageNumber="322" updateTime="1698611165842" updateUser="plazi">
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<mods:mods id="DD5C9F32E6973C58AA270F99CFEE5649" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C9A4042C3DEB5EC39D23A4D29A5ECE1B">
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<mods:title id="B06E67E0475D1C5130859DCFF0D5AAEB">Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="E63DB132D7FCEF9F41D4B03C95C7F5BF" type="personal">
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<mods:role id="5576EAAC38DAAEA65F5C61284F644D5E">
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<mods:roleTerm id="6442DCB41C1E2343775593AD37A43B1E">Author</mods:roleTerm>
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<mods:namePart id="78D0B8485A57A22317BC80ED381ECA90">Toledo, Mario</mods:namePart>
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</mods:name>
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<mods:name id="B85FA763C3825F11F7DDDE96FAD2E014" type="personal">
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<mods:role id="00C6DA6B848869CDDBEC9F80E2337AB1">
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<mods:roleTerm id="A751744C3F25B2DCE21AE81FCC1A85B1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3B33891AF775340DF01386C27C731EE7">Michat, Mariano C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="64E43D6E67A85CA346CF7DFC8DB73EA5">text</mods:typeOfResource>
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<mods:titleInfo id="B16977B112BF0463A83134E7C5AB55F2">
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<mods:title id="A748B466DE7EF72615A61F8E4C1C7E37">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7A9A27461B31B8DB3FF00BC0D3F062E1">
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<mods:date id="0AA96E5185D57D0E9191AFC334C4660C">2015</mods:date>
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<mods:detail id="C4F90566A865F65D20FD524C4D778337" type="volume">
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<mods:number id="D7EA0322EEC58F86CB315FBEA158561A">3990</mods:number>
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<mods:detail id="E29DD876C29540A489F7049B58FBD4B6" type="issue">
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<mods:number id="F8CC35695DB8EED35D227396580A1C0A">3</mods:number>
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<mods:extent id="3C755F94807D18AAFD570C172AAE86DB" unit="page">
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<mods:start id="E28779491DA4AE37161DAC2C8674E68A">301</mods:start>
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<mods:end id="330072815BB85782F44C172CFBB31DE2">354</mods:end>
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</mods:part>
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<mods:classification id="F81A9EF2008618AD6D1880893DE28E54">journal article</mods:classification>
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<mods:identifier id="DB7B36F17897373C414EF89D1A6EC012" type="DOI">10.11646/zootaxa.3990.3.1</mods:identifier>
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<mods:identifier id="638F5B0631F3802A6788AF64CEE86DF9" type="GBIF-Dataset">cf747765-b9bf-42ed-8479-35355e712db0</mods:identifier>
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<mods:identifier id="9E798C45EEA492C2775DBF7AF0ABA645" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0B38EDE408B5A067E6E279C98152016E" type="Zenodo-Dep">244301</mods:identifier>
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<mods:identifier id="FCFF0E9E5EA525C7EAFCF8BB1C9508BC" type="ZooBank">378C0359-E0E4-4CCC-821D-649144E37A63</mods:identifier>
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</mods:mods>
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<treatment id="03EE87FEFFE6FFE291A6FA08FCB8A49E" ID-DOI="http://doi.org/10.5281/zenodo.5692058" ID-GBIF-Taxon="119638442" ID-Zenodo-Dep="5692058" LSID="urn:lsid:plazi:treatment:03EE87FEFFE6FFE291A6FA08FCB8A49E" httpUri="http://treatment.plazi.org/id/03EE87FEFFE6FFE291A6FA08FCB8A49E" lastPageId="22" lastPageNumber="323" pageId="21" pageNumber="322">
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<subSubSection id="C35D6563FFE6FFE191A6FA08FE28A5CE" pageId="21" pageNumber="322" type="nomenclature">
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<paragraph id="8BF836E8FFE6FFE191A6FA08FE14A5F1" blockId="21.[151,492,1461,1520]" box="[151,492,1461,1487]" pageId="21" pageNumber="322">
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<heading id="D0B08184FFE6FFE191A6FA08FE14A5F1" bold="true" box="[151,492,1461,1487]" fontSize="11" level="1" pageId="21" pageNumber="322" reason="1">
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||
<emphasis id="B933EAFAFFE6FFE191A6FA08FE14A5F1" bold="true" box="[151,492,1461,1487]" pageId="21" pageNumber="322">
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||
<taxonomicName id="4C474D6BFFE6FFE191A6FA08FE51A5F1" ID-CoL="8LZSC" box="[151,425,1461,1487]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="spangleri" status="sp. nov.">
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<emphasis id="B933EAFAFFE6FFE191A6FA08FE51A5F1" bold="true" box="[151,425,1461,1487]" italics="true" pageId="21" pageNumber="322">Laccomimus spangleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2005781FFE6FFE19081FA08FE14A5F1" box="[432,492,1461,1487]" pageId="21" pageNumber="322" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF836E8FFE6FFE191A6FA6AFE28A5CE" blockId="21.[151,492,1461,1520]" box="[151,464,1495,1520]" pageId="21" pageNumber="322">
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(
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<figureCitation id="137C2A6DFFE6FFE191AEFA6AFF0EA5CE" box="[159,246,1495,1520]" captionStart="FIGURES 59 – 66" captionStartId="38.[151,264,1953,1975]" captionTargetBox="[161,1426,197,1927]" captionTargetId="figure@38.[151,1435,193,1932]" captionTargetPageId="38" captionText="FIGURES 59 – 66. Subschematic body shape and colour patterns in Laccomimus species: 59) L. distinctus; 60) L. malkini, a) typical specimen from Colombia, b) specimen from Piracicaba, Brazil; 61) L. variegatus (Perú); 62) L. amazonas; 63) L. pumilio (Florida, USA); 64) L. alvarengi, a – b) variation in colouration patterns; 65) L. bordoni; 66) L. spangleri." httpUri="https://zenodo.org/record/244311/files/figure.png" pageId="21" pageNumber="322">Figs 66</figureCitation>
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; 72; 104; 114; 128)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6563FFE6FFE191A6F9A4FCD3A76C" pageId="21" pageNumber="322" type="materials_examined">
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<paragraph id="8BF836E8FFE6FFE191A6F9A4FD89A60C" blockId="21.[151,1437,1561,1982]" box="[151,625,1561,1586]" pageId="21" pageNumber="322">
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<emphasis id="B933EAFAFFE6FFE191A6F9A4FECDA60C" bold="true" box="[151,309,1561,1586]" pageId="21" pageNumber="322">
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<typeStatus id="54FC884AFFE6FFE191A6F9A4FF2AA60C" box="[151,210,1561,1586]" pageId="21" pageNumber="322">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3507678FFE6FFE1900DF9A4FE6CA60C" box="[316,404,1561,1586]" name="Mexico" pageId="21" pageNumber="322">Mexico</collectingCountry>
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, Sinaloa, Mazatlan.
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</paragraph>
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<paragraph id="8BF836E8FFE6FFE191F6F980FECCA6FC" blockId="21.[151,1437,1561,1982]" pageId="21" pageNumber="322">
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<emphasis id="B933EAFAFFE6FFE191F6F980FE8DA668" bold="true" box="[199,373,1597,1622]" pageId="21" pageNumber="322">
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<typeStatus id="54FC884AFFE6FFE191F6F980FEFAA668" box="[199,258,1597,1622]" pageId="21" pageNumber="322">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54FC884AFFE6FFE1904DF980FE1DA668" box="[380,485,1597,1622]" pageId="21" pageNumber="322" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED56AE2DFFE6FFE19303F983FD72A668" LSID="urn:lsid:biocol.org:col:34871" box="[562,650,1598,1622]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="322">USNM</collectionCode>
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): dissected by us; median lobe of aedeagus and parameres included in
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<collectionCode id="ED56AE2DFFE6FFE191A6F9DFFF10A647" box="[151,232,1634,1657]" pageId="21" pageNumber="322">DMHF</collectionCode>
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on plastic label, pinned under the
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<typeStatus id="54FC884AFFE6FFE19358F9DCFD33A644" box="[617,715,1633,1658]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
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“
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<collectingCountry id="F3507678FFE6FFE193EDF9DCFCCEA644" box="[732,822,1633,1658]" name="Mexico" pageId="21" pageNumber="322">Mexico</collectingCountry>
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, Sinaloa, Mazatlan,
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<date id="FFF91028FFE6FFE1952BF9DCFB98A644" box="[1050,1120,1633,1658]" pageId="21" pageNumber="322" value="1917-07">
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<collectionCode id="ED56AE2DFFE6FFE1952BF9DCFBBBA644" box="[1050,1091,1633,1658]" pageId="21" pageNumber="322">VII</collectionCode>
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.17
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</date>
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–23.1963, P. & P. Spangler” [rectangular, white label, printed], “
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<typeStatus id="54FC884AFFE6FFE1930CF93BFD2CA6A0" box="[573,724,1670,1694]" pageId="21" pageNumber="322" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C474D6BFFE6FFE193D1F93BFC1DA6A3" box="[736,997,1669,1693]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="spangleri">
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<emphasis id="B933EAFAFFE6FFE193D1F93BFC1DA6A3" box="[736,997,1669,1693]" italics="true" pageId="21" pageNumber="322">Laccomimus spangleri</emphasis>
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</taxonomicName>
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Toledo & Michat” [red, rectangular, printed label].
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</paragraph>
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<paragraph id="8BF836E8FFE6FFE191F6F973FCD3A76C" blockId="21.[151,1437,1561,1982]" pageId="21" pageNumber="322">
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<typeStatus id="54FC884AFFE6FFE191F6F973FECCA6D8" box="[199,308,1742,1766]" pageId="21" pageNumber="322" type="paratype">Paratypes</typeStatus>
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(6 exs, all bearing red, printed label “
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<typeStatus id="54FC884AFFE6FFE19236F973FC6DA6D8" box="[775,917,1742,1766]" pageId="21" pageNumber="322" type="paratype">PARATYPE</typeStatus>
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<taxonomicName id="4C474D6BFFE6FFE19293F973FB5EA6DB" box="[930,1190,1741,1765]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="spangleri">
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<emphasis id="B933EAFAFFE6FFE19293F973FB5EA6DB" box="[930,1190,1741,1765]" italics="true" pageId="21" pageNumber="322">Laccomimus spangleri</emphasis>
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</taxonomicName>
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Toledo & Michat”):
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<emphasis id="B933EAFAFFE6FFE191A6F94CFF0FA734" bold="true" box="[151,247,1777,1802]" pageId="21" pageNumber="322">
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<collectingCountry id="F3507678FFE6FFE191A6F94CFF0BA734" box="[151,243,1777,1802]" name="Mexico" pageId="21" pageNumber="322">Mexico</collectingCountry>
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:
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</emphasis>
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same data as
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<typeStatus id="54FC884AFFE6FFE190AAF94CFDF8A734" box="[411,512,1777,1802]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
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; two specimens bearing the following additional label “Collected in blk litetrap” [smaller white, rectangular, printed label] (3 exs
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<collectionCode id="ED56AE2DFFE6FFE193FBF8ABFCD8A710" LSID="urn:lsid:biocol.org:col:34871" box="[714,800,1814,1838]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="322">USNM</collectionCode>
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).
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<emphasis id="B933EAFAFFE6FFE19205F8A8FC3AA710" bold="true" box="[820,962,1813,1838]" pageId="21" pageNumber="322">
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<collectingCountry id="F3507678FFE6FFE19205F8A8FC45A710" box="[820,957,1813,1838]" name="Costa Rica" pageId="21" pageNumber="322">Costa Rica</collectingCountry>
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:
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</emphasis>
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“16 Mi. S La Cruz,
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<collectionCode id="ED56AE2DFFE6FFE19584F8A8FB26A710" box="[1205,1246,1813,1838]" pageId="21" pageNumber="322">VII</collectionCode>
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.13.1965, Paul J. Spangler”, “
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<taxonomicName id="4C474D6BFFE6FFE19010F887FE56A76F" box="[289,430,1850,1873]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B933EAFAFFE6FFE19010F887FE56A76F" box="[289,430,1850,1873]" italics="true" pageId="21" pageNumber="322">Laccomimus</emphasis>
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</taxonomicName>
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det. PJ Spangler” (2 exs
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<collectionCode id="ED56AE2DFFE6FFE193F8F887FCE7A76C" LSID="urn:lsid:biocol.org:col:34871" box="[713,799,1850,1874]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="322">USNM</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6563FFE6FFE191F6F8E0FDEDA780" pageId="21" pageNumber="322" type="diagnosis">
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<paragraph id="8BF836E8FFE6FFE191F6F8E0FDEDA780" blockId="21.[151,1437,1561,1982]" pageId="21" pageNumber="322">
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<emphasis id="B933EAFAFFE6FFE191F6F8E0FEB9A748" bold="true" box="[199,321,1885,1910]" pageId="21" pageNumber="322">Diagnosis.</emphasis>
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Externally very similar to most of the species of the genus, but visibly smaller. Median lobe of aedeagus distinctive, distal portion straight, almost parallel-sided up to abruptly narrowed apex, ending in a small button-like expansion (
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<figureCitation id="137C2A6DFFE6FFE190ABF818FE03A780" box="[410,507,1957,1982]" captionStart="FIGURES 103 – 106" captionStartId="45.[151,264,1913,1935]" captionTargetBox="[158,1429,198,1873]" captionTargetId="figure@45.[151,1436,193,1892]" captionTargetPageId="45" captionText="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." httpUri="https://zenodo.org/record/244318/files/figure.png" pageId="21" pageNumber="322">Fig. 104</figureCitation>
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a).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6563FFE5FFE291F6FF2AFE61A341" pageId="22" pageNumber="323" type="description">
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||
<paragraph id="8BF836E8FFE5FFE291F6FF2AFDF6A0C9" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="323">
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<emphasis id="B933EAFAFFE5FFE291F6FF2AFEA1A08E" bold="true" box="[199,345,151,176]" pageId="22" pageNumber="323">Description.</emphasis>
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<emphasis id="B933EAFAFFE5FFE2906EFF25FE40A08E" box="[351,440,152,176]" italics="true" pageId="22" pageNumber="323">Habitus</emphasis>
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||
(
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<figureCitation id="137C2A6DFFE5FFE290F6FF2AFDE3A08E" box="[455,539,151,176]" captionStart="FIGURES 59 – 66" captionStartId="38.[151,264,1953,1975]" captionTargetBox="[161,1426,197,1927]" captionTargetId="figure@38.[151,1435,193,1932]" captionTargetPageId="38" captionText="FIGURES 59 – 66. Subschematic body shape and colour patterns in Laccomimus species: 59) L. distinctus; 60) L. malkini, a) typical specimen from Colombia, b) specimen from Piracicaba, Brazil; 61) L. variegatus (Perú); 62) L. amazonas; 63) L. pumilio (Florida, USA); 64) L. alvarengi, a – b) variation in colouration patterns; 65) L. bordoni; 66) L. spangleri." httpUri="https://zenodo.org/record/244311/files/figure.png" pageId="22" pageNumber="323">Fig. 66</figureCitation>
|
||
). TL:
|
||
<quantity id="4CBF9B0DFFE5FFE2936EFF25FD11A091" box="[607,745,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="22" pageNumber="323" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
|
||
; MW: 0.9–1.0 mm (
|
||
<typeStatus id="54FC884AFFE5FFE292FBFF2AFBCAA08E" box="[970,1074,151,176]" pageId="22" pageNumber="323" type="holotype">holotype</typeStatus>
|
||
: TL:
|
||
<quantity id="4CBF9B0DFFE5FFE2955CFF25FB3DA091" box="[1133,1221,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="22" pageNumber="323" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; MW: 1.0 mm); A- TL:
|
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<quantity id="4CBF9B0DFFE5FFE291FDFF01FED2A0EA" box="[204,298,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="22" pageNumber="323" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; A-MW:
|
||
<quantity id="4CBF9B0DFFE5FFE290ACFF01FE00A0EA" box="[413,504,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="323" unit="mm" value="1.1">1.1 mm</quantity>
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||
(NUM: 6 exs); smaller than any other
|
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<taxonomicName id="4C474D6BFFE5FFE292E1FF00FBA5A0EA" box="[976,1117,189,212]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B933EAFAFFE5FFE292E1FF00FBA5A0EA" box="[976,1117,189,212]" italics="true" pageId="22" pageNumber="323">Laccomimus</emphasis>
|
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</taxonomicName>
|
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species; body quite broad, regularly tapering to elytral apex.
|
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</paragraph>
|
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<paragraph id="8BF836E8FFE5FFE291F6FEB9FE28A192" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="323">
|
||
<emphasis id="B933EAFAFFE5FFE291F6FEB9FEEDA122" box="[199,277,260,284]" italics="true" pageId="22" pageNumber="323">Colour</emphasis>
|
||
. Head and pronotum paler than elytra. Head testaceous or reddish-brown, often darker between eyes; pronotum testaceous, with more or less defined dark, elongate marking along hind margin. Elytra chestnut-brown, with quite defined pale subbasal band, fragmented in irregular spots; subapical lateral yellow area reduced to a narrow lateral band or completely obsolete. Whole underside testaceous or pale-brown. Antennae and mouthparts testaceous. Legs testaceous.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE5FFE291F6FE05FE07A261" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="323">
|
||
<emphasis id="B933EAFAFFE5FFE291F6FE05FE26A1EE" box="[199,478,440,464]" italics="true" pageId="22" pageNumber="323">Structures and sculpture</emphasis>
|
||
. Head smooth, shiny, with fine microreticulation of rounded cells. Pronotum with microreticulation as on head, shiny, with faint iridescence. Surface with few scattered punctures. Elytra gradually tapering, with subapical sinuation obsolete; smooth, with poorly impressed reticulation but visible and quite intense iridescence. Prosternal process slightly elevated, almost acuminate at tip. Lobes of metacoxal processes regularly rounded in both sexes.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE5FFE291F6FDD1FBA7A309" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="323">
|
||
<emphasis id="B933EAFAFFE5FFE291F6FDD1FEF9A2BA" box="[199,257,620,644]" italics="true" pageId="22" pageNumber="323">Male</emphasis>
|
||
. Last abdominal ventrite tectiform, pointed, with anterolateral sides rectilinear. Outer claw of forelegs scimitar-shaped (
|
||
<figureCitation id="137C2A6DFFE5FFE2906BFD32FE49A296" box="[346,433,655,680]" captionStart="FIGURES 67 – 83" captionStartId="39.[151,264,1798,1820]" captionTargetBox="[159,1430,201,1748]" captionTargetId="figure@39.[151,1436,193,1777]" captionTargetPageId="39" captionText="FIGURES 67 – 83. Prosternal process and claws in Laccomimus species. 67 – 69) P rosternal process: 67) L. pumilio (Florida, USA); 68) L. pumilio (Cuba); 69) L. alvarengi (Perú). 70 – 83) M ale left protarsal claws: 70) L. bordoni (Paraguay); 71) L. malkini (Madre de Dios, Perú); 72) L. spangleri (holotype); 73) L. distinctus (holotype); 74) L. bolivari (Trinidad); 75) L. amazonas (holotype); 76) L. variegatus (Colonia Calleria, Perú); 77) L. alvarengi (holotype); 78) L. alvarengi (Loreto, Perú); 79) L. alvarengi (Guayaramerin, Bolivia); 80) L. alvarengi (Esmeraldas, Ecuador); 81) L. alvarengi (Albrook, Panamá); 82) L. pumilio (Cayo District, Belize); 83) L. pumilio (Cayamas, Cuba)." httpUri="https://zenodo.org/record/244312/files/figure.png" pageId="22" pageNumber="323">Fig. 72</figureCitation>
|
||
). Median lobe of aedeagus, in left lateral view (
|
||
<figureCitation id="137C2A6DFFE5FFE292D1FD32FBBCA296" box="[992,1092,655,680]" captionStart="FIGURES 103 – 106" captionStartId="45.[151,264,1913,1935]" captionTargetBox="[158,1429,198,1873]" captionTargetId="figure@45.[151,1436,193,1892]" captionTargetPageId="45" captionText="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." httpUri="https://zenodo.org/record/244318/files/figure.png" pageId="22" pageNumber="323">Fig. 104</figureCitation>
|
||
a), elongate, slender, base and distal portion forming an angle of about 100°; distal portion straight, almost parallel-sided up to abruptly narrowed apex, ending in a small button-like expansion. In dorsal view (
|
||
<figureCitation id="137C2A6DFFE5FFE2926EFD6AFC46A2CE" box="[863,958,727,752]" captionStart="FIGURES 107 – 115" captionStartId="46.[151,264,1951,1973]" captionTargetBox="[184,1390,196,1925]" captionTargetId="figure@46.[170,1417,193,1930]" captionTargetPageId="46" captionText="FIGURES 107 – 115. Median lobe of aedeagus in Laccomimus species, dorsal view: 107) L. distinctus; 108) L. malkini; 109) L. variegatus; 110) L. amazonas; 111) L. alvarengi; 112) L. improvidus; 113) L. bordoni; 114) L. spangleri; 115) L. bolivari." httpUri="https://zenodo.org/record/244319/files/figure.png" pageId="22" pageNumber="323">Fig. 114</figureCitation>
|
||
) straight and slender, with almost parallel sides, then gradually narrowed and turned to right; apex quite dilated; dorsal lamina covering distal half of ejaculator groove. Left paramere (
|
||
<figureCitation id="137C2A6DFFE5FFE29324FCA2FD8CA306" box="[533,628,799,824]" captionStart="FIGURES 103 – 106" captionStartId="45.[151,264,1913,1935]" captionTargetBox="[158,1429,198,1873]" captionTargetId="figure@45.[151,1436,193,1892]" captionTargetPageId="45" captionText="FIGURES 103 – 106. Male genitalia in Laccomimus species, left lateral view: 103) L. bordoni, a) median lobe, b) magnification of apex of median lobe, c) left paramere, showing apex in different positions, d) right paramere; 104) L. spangleri, a) median lobe, b) left paramere; c) right paramere; 105) L. bolivari (holotype), a) median lobe, b) magnification of apex of median lobe, c) left paramere, d) right paramere; 106) L. bolivari (Suriname), a) median lobe, b) magnification of apex of median lobe." httpUri="https://zenodo.org/record/244318/files/figure.png" pageId="22" pageNumber="323">Fig. 104</figureCitation>
|
||
b) sabre-shaped, gradually tapering to apex.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE5FFE291F6FCF9FE61A341" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="323">
|
||
<emphasis id="B933EAFAFFE5FFE291F6FCF9FEE3A362" box="[199,283,836,860]" italics="true" pageId="22" pageNumber="323">Female</emphasis>
|
||
. Not duller than male. Last abdominal ventrite subtruncate, with small medial notch at apex. Genital pieces as in
|
||
<taxonomicName id="4C474D6BFFE5FFE2902EFCD4FE69A3BE" box="[287,401,872,896]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE5FFE2902EFCD4FE69A3BE" box="[287,401,872,896]" italics="true" pageId="22" pageNumber="323">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6563FFE5FFE291F6FC31FD3BA39A" box="[199,707,908,933]" pageId="22" pageNumber="323" type="distribution">
|
||
<paragraph id="8BF836E8FFE5FFE291F6FC31FD3BA39A" blockId="22.[151,1437,151,1185]" box="[199,707,908,933]" pageId="22" pageNumber="323">
|
||
<emphasis id="B933EAFAFFE5FFE291F6FC31FE98A39B" bold="true" box="[199,352,908,933]" pageId="22" pageNumber="323">Distribution.</emphasis>
|
||
<collectingCountry id="F3507678FFE5FFE29056FC31FE47A39B" box="[359,447,908,933]" name="Mexico" pageId="22" pageNumber="323">Mexico</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507678FFE5FFE290FAFC31FDBCA39B" box="[459,580,908,933]" name="Costa Rica" pageId="22" pageNumber="323">Costa Rica</collectingCountry>
|
||
(
|
||
<figureCitation id="137C2A6DFFE5FFE29365FC31FD4EA39A" box="[596,694,908,933]" captionStart="FIGURES 126 – 129" captionStartId="49.[151,264,1953,1975]" captionTargetBox="[157,1432,197,1931]" captionTargetId="figure@49.[151,1436,193,1932]" captionTargetPageId="49" captionText="FIGURES 126 – 129. Distribution maps of Laccomimus species: 126) L. bordoni in South America; 127) L. bolivari (full circles), L. youngi (single open square in Brazil); 128) L. pumilio (open circles), L. alvarengi in Central America (single triangle in Panamá), L. spangleri (stars), L. bordoni in Central America (asterisks); 129) L. improvidus." httpUri="https://zenodo.org/record/244322/files/figure.png" pageId="22" pageNumber="323">Fig. 128</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6563FFE5FFE291F6FC12FBA1A40A" pageId="22" pageNumber="323" type="etymology">
|
||
<paragraph id="8BF836E8FFE5FFE291F6FC12FD6BA3D2" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="323">
|
||
<emphasis id="B933EAFAFFE5FFE291F6FC12FEB6A3F6" bold="true" box="[199,334,943,968]" pageId="22" pageNumber="323">Etymology.</emphasis>
|
||
Dedicated to the late Paul J. Spangler, who collected this species from both known localities. The specific epithet is a noun in the genitive case.
|
||
</paragraph>
|
||
<paragraph id="8BF836E8FFE5FFE291F6FC4AFBA1A40A" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="323">
|
||
<emphasis id="B933EAFAFFE5FFE291F6FC4AFE42A42E" bold="true" box="[199,442,1015,1040]" pageId="22" pageNumber="323">Notes on variability.</emphasis>
|
||
The only specimen seen from
|
||
<collectingCountry id="F3507678FFE5FFE2922DFC45FC60A42E" box="[796,920,1015,1040]" name="Costa Rica" pageId="22" pageNumber="323">Costa Rica</collectingCountry>
|
||
(the second one is known only from a single elytron) is the smallest known specimen of
|
||
<taxonomicName id="4C474D6BFFE5FFE2934FFBA0FCF3A40A" box="[638,779,1053,1076]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B933EAFAFFE5FFE2934FFBA0FCF3A40A" box="[638,779,1053,1076]" italics="true" pageId="22" pageNumber="323">Laccomimus</emphasis>
|
||
</taxonomicName>
|
||
(TL:
|
||
<quantity id="4CBF9B0DFFE5FFE2927CFBA1FC5EA40A" box="[845,934,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="323" unit="mm" value="1.8">1.8 mm</quantity>
|
||
; MW:
|
||
<quantity id="4CBF9B0DFFE5FFE292C2FBA1FBB6A40A" box="[1011,1102,1052,1076]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="323" unit="mm" value="0.9">0.9 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D6563FFE5FFE291F6FB82FCB8A49E" pageId="22" pageNumber="323" type="discussion">
|
||
<paragraph id="8BF836E8FFE5FFE291F6FB82FCB8A49E" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="323">
|
||
<emphasis id="B933EAFAFFE5FFE291F6FB82FEC3A466" bold="true" box="[199,315,1087,1112]" pageId="22" pageNumber="323">Remarks.</emphasis>
|
||
<taxonomicName id="4C474D6BFFE5FFE29072FBFDFDBBA466" box="[323,579,1088,1112]" class="Insecta" family="Dytiscidae" genus="Laccomimus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="B933EAFAFFE5FFE29072FBFDFDBBA466" box="[323,579,1088,1112]" italics="true" pageId="22" pageNumber="323">Laccomimus spangleri</emphasis>
|
||
</taxonomicName>
|
||
is close to
|
||
<taxonomicName id="4C474D6BFFE5FFE293F6FBFCFCC4A466" box="[711,828,1088,1112]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="pumilio">
|
||
<emphasis id="B933EAFAFFE5FFE293F6FBFCFCC4A466" box="[711,828,1088,1112]" italics="true" pageId="22" pageNumber="323">L. pumilio</emphasis>
|
||
</taxonomicName>
|
||
and similar species, sharing with them external male characters, similar prosternal and metacoxal processes, and similar female genitalia. It is, however, well characterised by aedeagal features and by its very small size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |