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<document id="6D0EED12218D27440ABC7B0B731640C3" ID-DOI="10.1080/00222933.2017.1319519" ID-GBIF-Dataset="7dd54e06-5524-476c-8ef1-eefc5c571da3" ID-ISSN="1464-5262" ID-Zenodo-Dep="4746652" IM.bibliography_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.taxonomicNames_approvedBy="admin" IM.treatmentCitations_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1620669599041" checkinUser="carolina" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Vaz-de-Mello, Fernando Z." docDate="2017" docId="03EE87E3FFA1FFD3BBD6AF8CFB74FCA2" docLanguage="en" docName="JNATHIST.51.19-20.1035-1090.pdf" docOrigin="Journal of Natural History 51 (19 - 20)" docSource="http://dx.doi.org/10.1080/00222933.2017.1319519" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Aegidium minor Paul. R. Paulian 1984" docType="treatment" docVersion="7" lastPageNumber="1081" masterDocId="FFD7FF9BFF8FFFFCBA0DAF01FFF0FFB5" masterDocTitle="Revision of the mainland species of the Neotropical genus Aegidium Westwood (Coleoptera: Scarabaeidae: Orphninae)" masterLastPageNumber="1090" masterPageNumber="1035" pageNumber="1080" updateTime="1694885189776" updateUser="admin" zenodo-license-document="CLOSED">
<mods:mods id="7706295BA03D962A48DA9AE1D9A27B52" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1C9B4E26BB8FBC8D0CE6B774C7DC9E18">
<mods:title id="CF62C1D74E344AF5A9052DDD961DF36D">Revision of the mainland species of the Neotropical genus Aegidium Westwood (Coleoptera: Scarabaeidae: Orphninae)</mods:title>
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<mods:name id="0D169E09886A9F634E1329A670977CC6" type="personal">
<mods:role id="295E74F8BE13861D651D9297FA28C373">
<mods:roleTerm id="14782A60BCF91830284A78381B84B5EF">Author</mods:roleTerm>
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<mods:namePart id="A36AE4B665A3864067E315B6AE467EFA">Frolov, Andrey V.</mods:namePart>
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<mods:name id="109501035D76255A7E9B81FA53BAFED3" type="personal">
<mods:role id="A5CD40E0150D42ED163901062F71724F">
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<mods:namePart id="C1B055C81F7ED3F534381E8778250C74">Akhmetova, Lilia A.</mods:namePart>
</mods:name>
<mods:name id="7D1922E825916B83CBD57502741E3640" type="personal">
<mods:role id="B696CB752AC58D6E72E561533CA9151C">
<mods:roleTerm id="D5362B38009676C9EFDB0DF5489A9D58">Author</mods:roleTerm>
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<mods:namePart id="FF31E2716F00656C178BD15613A09398">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="F36FD4C7EFE6BBA96526F02A7A5D1C60">text</mods:typeOfResource>
<mods:relatedItem id="86A63B2B6F43786E6DB716DF23DF8BBA" type="host">
<mods:titleInfo id="29797F547959BA1648A0329082BF05F8">
<mods:title id="672774E4744AC17DF4DF42FB3086F5A7">Journal of Natural History</mods:title>
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<mods:part id="DEC14955FED752113916986B661AF62A">
<mods:date id="D2FBB140522B5B007AA1B1AB1C5995A2">2017</mods:date>
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<mods:number id="88EA495A4211DD4892B522F23D6698CC">2017-05-17</mods:number>
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<mods:detail id="DCBBF764D1F205837D941A68051132E4" type="volume">
<mods:number id="75054177D8F2AFC61D7567AC19E51BD0">51</mods:number>
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<mods:detail id="F898E399128968C8FF1A003AC0864F9A" type="issue">
<mods:number id="5ED9484C64546201CE113F3F6762008C">19 - 20</mods:number>
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<mods:start id="C6044ED1360E6137E1654325652AB7A9">1035</mods:start>
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<mods:url id="83A361BBD75F2E78458977F77BA168CC">http://dx.doi.org/10.1080/00222933.2017.1319519</mods:url>
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<mods:classification id="5B71CD47E12802F239898605C4FCF6EA">journal article</mods:classification>
<mods:identifier id="E1955CBECEBC990083348F5F29D9A796" type="DOI">10.1080/00222933.2017.1319519</mods:identifier>
<mods:identifier id="BFFCE533A3E4AD2A52EC478635EC3C16" type="GBIF-Dataset">7dd54e06-5524-476c-8ef1-eefc5c571da3</mods:identifier>
<mods:identifier id="2B9C1C87E228747A6234C9F852DF7E0C" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="9C59E884B05177FDCA5BC23EB8ACDC90" type="Zenodo-Dep">4746652</mods:identifier>
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<treatment id="03EE87E3FFA1FFD3BBD6AF8CFB74FCA2" ID-DOI="http://doi.org/10.5281/zenodo.6506528" ID-GBIF-Taxon="194877628" ID-Zenodo-Dep="6506528" LSID="urn:lsid:plazi:treatment:03EE87E3FFA1FFD3BBD6AF8CFB74FCA2" httpUri="http://treatment.plazi.org/id/03EE87E3FFA1FFD3BBD6AF8CFB74FCA2" lastPageId="47" lastPageNumber="1081" pageId="46" pageNumber="1080">
<subSubSection id="C35D657EFFA1FFD2BBD6AF8CFCB8FF13" box="[475,840,141,166]" pageId="46" pageNumber="1080" type="nomenclature">
<paragraph id="8BF836F5FFA1FFD2BBD6AF8CFCB8FF13" blockId="46.[475,840,141,201]" box="[475,840,141,166]" pageId="46" pageNumber="1080">
<heading id="D0B08199FFA1FFD2BBD6AF8CFCB8FF13" box="[475,840,141,166]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="1080" reason="2">
<taxonomicName id="4C474D76FFA1FFD2BBD6AF8CFCB8FF13" authority="Paulian, 1984" authorityName="Paul. R. Paulian" authorityYear="1984" box="[475,840,141,166]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="1080" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B933EAE7FFA1FFD2BBD6AF8CFD50FF13" bold="true" box="[475,672,141,166]" italics="true" pageId="46" pageNumber="1080">Aegidium minor</emphasis>
<bibRefCitation id="EFD64B04FFA1FFD2B8A7AF8CFCB8FF13" author="Paulian R" box="[682,840,141,166]" pageId="46" pageNumber="1080" pagination="65 - 92" refId="ref21167" refString="Paulian R. 1984. Les Orphnidae Americains (Coleopteres, Scarabaeoidea). Ann Soc Ent Fr (N S). 20: 65 - 92." type="journal article" year="1984">Paulian, 1984</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35D657EFFA1FFD2B839AFB1FD1FFF7C" box="[564,751,176,201]" pageId="46" pageNumber="1080" type="description">
<paragraph id="8BF836F5FFA1FFD2B839AFB1FD1FFF7C" blockId="46.[475,840,141,201]" box="[564,751,176,201]" pageId="46" pageNumber="1080">
(
<figureCitation id="137C2A70FFA1FFD2B831AFB1FD5DFF7C" box="[572,685,176,201]" captionStart="Figure 15" captionStartId="21.[160,225,1522,1545]" captionTargetBox="[163,1152,143,1491]" captionTargetId="figure-1@21.[163,1153,142,1491]" captionTargetPageId="21" captionText="Figure 15. Aegidium minor, holotype male (a, d, e), male (b, f), female (c). Habitus in dorsal view (a, c), parameres in ventral view (b), aedeagus in lateral view (d), parameres in dorsal view (e), spiculum gastrale (f), locality map (g)." figureDoi="http://doi.org/10.5281/zenodo.4746701" httpUri="https://zenodo.org/record/4746701/files/figure.png" pageId="46" pageNumber="1080">Figure 15</figureCitation>
(a–g))
</paragraph>
</subSubSection>
<subSubSection id="C35D657EFFA1FFD2BAADAFE4FDE4FF48" box="[160,532,229,253]" pageId="46" pageNumber="1080" type="reference_group">
<paragraph id="8BF836F5FFA1FFD2BAADAFE4FDE4FF48" blockId="46.[160,532,229,253]" box="[160,532,229,253]" pageId="46" pageNumber="1080">
<treatmentCitationGroup id="AB5711DBFFA1FFD2BAADAFE4FDE4FF48" box="[160,532,229,253]" pageId="46" pageNumber="1080">
<treatmentCitation id="0AE610E4FFA1FFD2BAADAFE4FDE4FF48" author="Paulian R" box="[160,532,229,253]" page="72" pageId="46" pageNumber="1080" year="1984">
<taxonomicName id="4C474D76FFA1FFD2BAADAFE4FDE4FF48" authority="Paulian 1984: 72" authorityName="Paul. R. Paulian" authorityPageNumber="72" authorityYear="1984" box="[160,532,229,253]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="1080" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B933EAE7FFA1FFD2BAADAFE4FEA0FF48" box="[160,336,229,253]" italics="true" pageId="46" pageNumber="1080">Aegidium minor</emphasis>
<bibRefCitation id="EFD64B04FFA1FFD2BB54AFE4FDE4FF48" author="Paulian R" box="[345,532,229,253]" pageId="46" pageNumber="1080" pagination="65 - 92" refId="ref21167" refString="Paulian R. 1984. Les Orphnidae Americains (Coleopteres, Scarabaeoidea). Ann Soc Ent Fr (N S). 20: 65 - 92." type="journal article" year="1984">Paulian 1984: 72</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35D657EFFA1FFD2BAADAE35FEF2FEC5" pageId="46" pageNumber="1080" type="materials_examined">
<paragraph id="8BF836F5FFA1FFD2BAADAE35FEC7FEF8" blockId="46.[160,311,308,368]" box="[160,311,308,333]" pageId="46" pageNumber="1080">
<emphasis id="B933EAE7FFA1FFD2BAADAE35FEC7FEF8" bold="true" box="[160,311,308,333]" italics="true" pageId="46" pageNumber="1080">Type locality</emphasis>
</paragraph>
<paragraph id="8BF836F5FFA1FFD2BAADAE59FEF2FEC5" blockId="46.[160,311,308,368]" box="[160,258,344,368]" pageId="46" pageNumber="1080">
<collectingCountry id="F3507665FFA1FFD2BAADAE59FF0EFEC5" box="[160,254,344,368]" name="Ecuador" pageId="46" pageNumber="1080">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D657EFFA1FFD2BAADAEA7FBCDFC90" pageId="46" pageNumber="1080" type="materials_examined">
<paragraph id="8BF836F5FFA1FFD2BAADAEA7FE32FE0A" blockId="46.[160,1155,422,517]" box="[160,450,422,447]" pageId="46" pageNumber="1080">
<emphasis id="B933EAE7FFA1FFD2BAADAEA7FE32FE0A" bold="true" box="[160,450,422,447]" italics="true" pageId="46" pageNumber="1080">Type material examined</emphasis>
</paragraph>
<paragraph id="8BF836F5FFA1FFD2BAADAECBFCC8FDB0" blockId="46.[160,1155,422,517]" pageId="46" pageNumber="1080">
<materialsCitation id="3B2F3CA8FFA1FFD2BAADAECBFCDCFDB0" ID-GBIF-Occurrence="3757961367" collectionCode="MNHN" country="Ecuador" determinerName="Paul. R. Paulian" location="Balzen" pageId="46" pageNumber="1080" specimenCount="1" typeStatus="holotype">
<typeStatus id="54FC8857FFA1FFD2BAADAECBFEF9FE57" box="[160,265,458,482]" pageId="46" pageNumber="1080">Holotype</typeStatus>
,
<specimenCount id="9D41FD7CFFA1FFD2BB13AECBFEA5FE57" box="[286,341,458,482]" pageId="46" pageNumber="1080" type="male">male</specimenCount>
at
<collectionCode id="ED56AE30FFA1FFD2BB82AECBFE29FE57" box="[399,473,458,482]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="46" pageNumber="1080">MNHN</collectionCode>
labeled ‘
<collectingCountry id="F3507665FFA1FFD2B858AECBFD43FE57" box="[597,691,458,482]" name="Ecuador" pageId="46" pageNumber="1080">Equador</collectingCountry>
<location id="8E98602EFFA1FFD2B8C8AECBFCFEFE57" LSID="urn:lsid:plazi:treatment:03EE87E3FFA1FFD3BBD6AF8CFB74FCA2:8E98602EFFA1FFD2B8C8AECBFCFEFE57" box="[709,782,458,482]" country="Ecuador" name="Balzen" pageId="46" pageNumber="1080">Balzen</location>
[III]/52559/Fry Coll. 1905.100./ Holotype/
<taxonomicName id="4C474D76FFA1FFD2BB1DAEECFD86FDB0" authority="Paul. R. Paulian" authorityName="Paul. R. Paulian" authorityYear="1984" box="[272,630,493,517]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="1080" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B933EAE7FFA1FFD2BB1DAEECFE30FDB0" box="[272,448,493,517]" italics="true" pageId="46" pageNumber="1080">Aegidium minor</emphasis>
<determinerName id="7E24DA31FFA1FFD2BBC4AEECFD86FDB0" box="[457,630,493,517]" pageId="46" pageNumber="1080">Paul. R. Paulian</determinerName>
</taxonomicName>
det./HOLOTYPE’
</materialsCitation>
.
</paragraph>
<paragraph id="8BF836F5FFA1FFD2BAADAD3DFDF5FDE0" blockId="46.[160,1157,572,805]" box="[160,517,572,597]" pageId="46" pageNumber="1080">
<emphasis id="B933EAE7FFA1FFD2BAADAD3DFDF5FDE0" bold="true" box="[160,517,572,597]" italics="true" pageId="46" pageNumber="1080">Additional material examined</emphasis>
</paragraph>
<paragraph id="8BF836F5FFA1FFD2BAADAD5EFBCDFC90" blockId="46.[160,1157,572,805]" pageId="46" pageNumber="1080">
<materialsCitation id="3B2F3CA8FFA1FFD2BAADAD5EFEFBFD2F" ID-GBIF-Occurrence="3757961365" collectingDate="1894-04-03" collectingDateMax="1985-07-31" collectingDateMin="1985-05" collectionCode="MNHN" collectorName="Balzapamba, M" country="Ecuador" elevation="2600" isTailEnumeration="true" location="Bolivar" municipality="Patricia Pilar" pageId="46" pageNumber="1080" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Cotopaxi">
<collectingCountry id="F3507665FFA1FFD2BAADAD5EFEE4FDC2" box="[160,276,607,631]" name="Ecuador" pageId="46" pageNumber="1080">ECUADOR</collectingCountry>
.
<location id="8E98602EFFA1FFD2BB2FAD5EFE81FDC2" LSID="urn:lsid:plazi:treatment:03EE87E3FFA1FFD3BBD6AF8CFB74FCA2:8E98602EFFA1FFD2BB2FAD5EFE81FDC2" box="[290,369,607,631]" country="Ecuador" municipality="Patricia Pilar" name="Bolivar" pageId="46" pageNumber="1080" stateProvince="Cotopaxi">Bolívar</location>
:
<specimenCount id="9D41FD7CFFA1FFD2BB73AD5EFE1BFDC2" box="[382,491,607,631]" pageId="46" pageNumber="1080" type="male">one male</specimenCount>
at
<collectionCode id="ED56AE30FFA1FFD2B815AD5EFD95FDC2" box="[536,613,607,631]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="46" pageNumber="1080">MNHN</collectionCode>
,
<collectorName id="26B25323FFA1FFD2B87FAD5EFCD3FDC2" box="[626,803,607,631]" pageId="46" pageNumber="1080">Balzapamba, M</collectorName>
. de
<location id="8E98602EFFA1FFD2B957AD5EFC42FDC2" LSID="urn:lsid:plazi:treatment:03EE87E3FFA1FFD3BBD6AF8CFB74FCA2:8E98602EFFA1FFD2B957AD5EFC42FDC2" box="[858,946,607,631]" country="Ecuador" municipality="Patricia Pilar" name="Mathan" pageId="46" pageNumber="1080" stateProvince="Cotopaxi">Mathan</location>
,
<date id="FFF91035FFA1FFD2B9CDAD5EFBA0FDC2" box="[960,1104,607,631]" pageId="46" pageNumber="1080" value="1894-04-03">
<collectingDate id="EFBDE9DDFFA1FFD2B9CDAD5EFBA0FDC2" box="[960,1104,607,631]" pageId="46" pageNumber="1080" value="1894-04-03">3 April 1894</collectingDate>
</date>
leg.
<collectingRegion id="4983F817FFA1FFD2BAADAD83FEF7FD2F" box="[160,263,642,666]" country="Ecuador" name="Cotopaxi" pageId="46" pageNumber="1080">Cotopaxi</collectingRegion>
:
</materialsCitation>
<materialsCitation id="3B2F3CA8FFA1FFD2BB1CAD83FB74FD2F" ID-GBIF-Occurrence="3757961424" box="[273,1156,642,666]" collectingDate="1983-02" collectingDateMax="1985-07-31" collectingDateMin="1985-05" collectionCode="MHNG" collectorName="G. Onore" country="Ecuador" elevation="2600" isTailEnumeration="true" location="Chugchilan" municipality="Patricia Pilar" pageId="46" pageNumber="1080" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Cotopaxi">
<specimenCount id="9D41FD7CFFA1FFD2BB1CAD83FE89FD2F" box="[273,377,642,666]" pageId="46" pageNumber="1080" type="male">one male</specimenCount>
at
<collectionCode id="ED56AE30FFA1FFD2BB91AD83FE18FD2F" box="[412,488,642,666]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="46" pageNumber="1080" type="Museum">MHNG</collectionCode>
,
<location id="8E98602EFFA1FFD2BBFFAD83FD83FD2F" LSID="urn:lsid:plazi:treatment:03EE87E3FFA1FFD3BBD6AF8CFB74FCA2:8E98602EFFA1FFD2BBFFAD83FD83FD2F" box="[498,627,642,666]" country="Ecuador" municipality="Patricia Pilar" name="Chugchilan" pageId="46" pageNumber="1080" stateProvince="Cotopaxi">Chugchilán</location>
,
<quantity id="4CBF9B10FFA1FFD2B870AD83FD23FD2F" box="[637,723,642,666]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="46" pageNumber="1080" unit="m" value="2600.0">
<elevation id="006AD1C6FFA1FFD2B870AD83FD23FD2F" box="[637,723,642,666]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="46" pageNumber="1080" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<date id="FFF91035FFA1FFD2B8D0AD83FC70FD2F" box="[733,896,642,666]" pageId="46" pageNumber="1080" value="1983-02">
<collectingDate id="EFBDE9DDFFA1FFD2B8D0AD83FC70FD2F" box="[733,896,642,666]" pageId="46" pageNumber="1080" value="1983-02">February 1983</collectingDate>
</date>
,
<collectorName id="26B25323FFA1FFD2B987AD83FC1EFD2F" box="[906,1006,642,666]" pageId="46" pageNumber="1080">G. Onore</collectorName>
leg. Los Ríos:
</materialsCitation>
<materialsCitation id="3B2F3CA8FFA1FFD2BAADADA4FE28FD6A" ID-GBIF-Occurrence="3757961415" collectingDate="1985-05" collectingDateMax="1985-07-31" collectingDateMin="1985-05" collectionCode="CMN" collectorName="Peck" country="Ecuador" elevation="230" isTailEnumeration="true" location="Palenque River" municipality="Patricia Pilar" pageId="46" pageNumber="1080" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Cotopaxi">
<specimenCount id="9D41FD7CFFA1FFD2BAADADA4FEFCFD08" box="[160,268,677,701]" pageId="46" pageNumber="1080" type="male">one male</specimenCount>
at
<collectionCode id="ED56AE30FFA1FFD2BB3AADA4FE80FD08" box="[311,368,677,701]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="46" pageNumber="1080" type="Museum">CMN</collectionCode>
,
<collectingMunicipality id="6B9CAC8FFFA1FFD2BB70ADA4FDFCFD08" box="[381,524,677,701]" pageId="46" pageNumber="1080">Patricia Pilar</collectingMunicipality>
,
<locationDeviation id="9A4A2FD7FFA1FFD2B817ADA4FCCBFD08" box="[538,827,677,701]" location="Santo Domingo" pageId="46" pageNumber="1080">
<quantity id="4CBF9B10FFA1FFD2B817ADA4FD93FD08" box="[538,611,677,701]" metricMagnitude="4" metricUnit="m" metricValue="4.7" pageId="46" pageNumber="1080" unit="km" value="47.0">47 km</quantity>
S Santo Domingo
</locationDeviation>
,
<location id="8E98602EFFA1FFD2B944ADA4FC06FD08" LSID="urn:lsid:plazi:treatment:03EE87E3FFA1FFD3BBD6AF8CFB74FCA2:8E98602EFFA1FFD2B944ADA4FC06FD08" box="[841,1014,677,701]" country="Ecuador" municipality="Patricia Pilar" name="Palenque River" pageId="46" pageNumber="1080" stateProvince="Cotopaxi">Palenque River</location>
,
<quantity id="4CBF9B10FFA1FFD2BE0EADA4FBBFFD08" box="[1027,1103,677,701]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="46" pageNumber="1080" unit="m" value="230.0">
<elevation id="006AD1C6FFA1FFD2BE0EADA4FBBFFD08" box="[1027,1103,677,701]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="46" pageNumber="1080" unit="m" value="230.0">230 m</elevation>
</quantity>
, FIT,
<date id="FFF91035FFA1FFD2BAADADC6FEB9FD6A" box="[160,329,711,735]" pageId="46" pageNumber="1080" value="1985-05" valueMax="1985-07-31" valueMin="1985-05-00">
<collectingDate id="EFBDE9DDFFA1FFD2BAADADC6FEB9FD6A" box="[160,329,711,735]" pageId="46" pageNumber="1080" value="1985-05" valueMax="1985-07-31" valueMin="1985-05-00">May–July 1985</collectingDate>
</date>
, S.
<collectorName id="26B25323FFA1FFD2BB63ADC6FE52FD6A" box="[366,418,711,735]" pageId="46" pageNumber="1080">Peck</collectorName>
leg.;
</materialsCitation>
<materialsCitation id="3B2F3CA8FFA1FFD2BBD2ADC6FF3EFC90" ID-GBIF-Occurrence="3757961313" collectingDate="1985-05-05" collectingDateMax="1985-07-25" collectingDateMin="1985-05-05" collectionCode="CMN" collectorName="Balzapamba, M" country="Ecuador" elevation="250" isTailEnumeration="true" location="Palenque River" municipality="Patricia Pilar" pageId="46" pageNumber="1080" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Cotopaxi">
<specimenCount id="9D41FD7CFFA1FFD2BBD2ADC6FD98FD6A" box="[479,616,711,735]" pageId="46" pageNumber="1080" type="female">two females</specimenCount>
at
<collectionCode id="ED56AE30FFA1FFD2B883ADC6FD37FD6A" box="[654,711,711,735]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="46" pageNumber="1080" type="Museum">CMN</collectionCode>
,
<collectingMunicipality id="6B9CAC8FFFA1FFD2B8DFADC6FCAFFD6A" box="[722,863,711,735]" pageId="46" pageNumber="1080">Patricia Pilar</collectingMunicipality>
,
<locationDeviation id="9A4A2FD7FFA1FFD2B967ADC6FB70FD6A" box="[874,1152,711,735]" location="Santo Domingo" pageId="46" pageNumber="1080">
<quantity id="4CBF9B10FFA1FFD2B967ADC6FC40FD6A" box="[874,944,711,735]" metricMagnitude="4" metricUnit="m" metricValue="4.7" pageId="46" pageNumber="1080" unit="km" value="47.0">47 km</quantity>
S Santo Domingo
</locationDeviation>
,
<location id="8E98602EFFA1FFD2BAADADEBFEBCFCB7" LSID="urn:lsid:plazi:treatment:03EE87E3FFA1FFD3BBD6AF8CFB74FCA2:8E98602EFFA1FFD2BAADADEBFEBCFCB7" box="[160,332,746,770]" country="Ecuador" municipality="Patricia Pilar" name="Palenque River" pageId="46" pageNumber="1080" stateProvince="Cotopaxi">Palenque River</location>
,
<quantity id="4CBF9B10FFA1FFD2BB54ADEBFE54FCB7" box="[345,420,746,770]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="1080" unit="m" value="250.0">
<elevation id="006AD1C6FFA1FFD2BB54ADEBFE54FCB7" box="[345,420,746,770]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="1080" unit="m" value="250.0">250 m</elevation>
</quantity>
, malaise-FIT rainforest,
<date id="FFF91035FFA1FFD2B8BEADEBFC6AFCB7" box="[691,922,746,770]" pageId="46" pageNumber="1080" value="1985-05-05" valueMax="1985-07-25" valueMin="1985-05-05">
<collectingDate id="EFBDE9DDFFA1FFD2B8BEADEBFC6AFCB7" box="[691,922,746,770]" pageId="46" pageNumber="1080" value="1985-05-05" valueMax="1985-07-25" valueMin="1985-05-05">5 May–25 July 1985</collectingDate>
</date>
, S.
<location id="8E98602EFFA1FFD2B9C9ADEBFC08FCB7" LSID="urn:lsid:plazi:treatment:03EE87E3FFA1FFD3BBD6AF8CFB74FCA2:8E98602EFFA1FFD2B9C9ADEBFC08FCB7" box="[964,1016,746,770]" country="Ecuador" municipality="Patricia Pilar" name="Peck" pageId="46" pageNumber="1080" stateProvince="Cotopaxi">Peck</location>
and J. Peck leg.;
</materialsCitation>
<materialsCitation id="3B2F3CA8FFA1FFD2BADBAC0CFBCAFC90" ID-GBIF-Occurrence="3757961419" box="[214,1082,781,805]" collectingDate="1983-02-16" collectingDateMax="1985-07-31" collectingDateMin="1985-05" collectionCode="CEMT" collectorName="Sandoval" country="Ecuador" elevation="2600" isTailEnumeration="true" location="Provincia de Los Rios" municipality="Patricia Pilar" pageId="46" pageNumber="1080" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Provincia de Los Rios">
<specimenCount id="9D41FD7CFFA1FFD2BADBAC0CFEB1FC90" box="[214,321,781,805]" pageId="46" pageNumber="1080" type="male">one male</specimenCount>
at
<collectionCode id="ED56AE30FFA1FFD2BB65AC0CFE5AFC90" box="[360,426,781,805]" pageId="46" pageNumber="1080">CEMT</collectionCode>
,
<collectingRegion id="4983F817FFA1FFD2BBBBAC0CFD5AFC90" box="[438,682,781,805]" country="Ecuador" name="Los Rios" pageId="46" pageNumber="1080">Provincia de Los Ríos</collectingRegion>
,
<date id="FFF91035FFA1FFD2B8B8AC0CFC8EFC90" box="[693,894,781,805]" pageId="46" pageNumber="1080" value="1983-02-16">
<collectingDate id="EFBDE9DDFFA1FFD2B8B8AC0CFC8EFC90" box="[693,894,781,805]" pageId="46" pageNumber="1080" value="1983-02-16">16 February 1983</collectingDate>
</date>
, S.
<collectorName id="26B25323FFA1FFD2B9A8AC0CFBFBFC90" box="[933,1035,781,805]" pageId="46" pageNumber="1080">Sandoval</collectorName>
leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D657EFFA1FFD2BAADAC5DFC2FFC4A" pageId="46" pageNumber="1080" type="diagnosis">
<paragraph id="8BF836F5FFA1FFD2BAADAC5DFEE4FCC0" blockId="46.[160,1156,860,1023]" box="[160,276,860,885]" pageId="46" pageNumber="1080">
<emphasis id="B933EAE7FFA1FFD2BAADAC5DFEE4FCC0" bold="true" box="[160,276,860,885]" italics="true" pageId="46" pageNumber="1080">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF836F5FFA1FFD2BAADAC7EFC2FFC4A" blockId="46.[160,1156,860,1023]" pageId="46" pageNumber="1080">
<taxonomicName id="4C474D76FFA1FFD2BAADAC7EFEBBFC22" authorityName="Paul. R. Paulian" authorityYear="1984" box="[160,331,895,919]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="1080" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B933EAE7FFA1FFD2BAADAC7EFEBBFC22" box="[160,331,895,919]" italics="true" pageId="46" pageNumber="1080">Aegidium minor</emphasis>
</taxonomicName>
is similar to
<emphasis id="B933EAE7FFA1FFD2BBDAAC7EFD97FC22" box="[471,615,895,919]" italics="true" pageId="46" pageNumber="1080">
Ae.
<taxonomicName id="4C474D76FFA1FFD2BBF2AC7EFD97FC22" box="[511,615,895,919]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="1080" phylum="Arthropoda" rank="species" species="cribratum">cribratum</taxonomicName>
</emphasis>
and
<emphasis id="B933EAE7FFA1FFD2B892AC7EFCC9FC22" box="[671,825,895,919]" italics="true" pageId="46" pageNumber="1080">
Ae.
<taxonomicName id="4C474D76FFA1FFD2B8CAAC7EFCC9FC22" authorityName="Preudhomme de Borre (Arrow" authorityYear="1912" box="[711,825,895,919]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="1080" phylum="Arthropoda" rank="species" species="asperatum">asperatum</taxonomicName>
</emphasis>
in the shape of the aedeagus but differs from them by the parameres having small sensory areas apicad of lateral teeth (
<figureCitation id="137C2A70FFA1FFD2BAAAACC4FEE3FC68" box="[167,275,965,989]" captionStart="Figure 16" captionStartId="22.[160,225,1510,1533]" captionTargetBox="[164,1152,143,1479]" captionTargetId="figure-1@22.[163,1153,142,1479]" captionTargetPageId="22" captionText="Figure 16. Aegidium asperatum, holotype male (a, c, d), male (b, e). Habitus in dorsal view (a, b), aedeagus in lateral view (c), parameres in dorsal view (d), spiculum gastrale (e), locality map (f)." figureDoi="http://doi.org/10.5281/zenodo.4746705" httpUri="https://zenodo.org/record/4746705/files/figure.png" pageId="46" pageNumber="1080">Figure 16</figureCitation>
(b)). From the former species it also differs in having more protruding anterior angles of frontoclypeus, and from the later by sparser punctation of elytra.
</paragraph>
</subSubSection>
<subSubSection id="C35D657EFFA1FFD3BAADAB37FC6CFD1B" lastPageId="47" lastPageNumber="1081" pageId="46" pageNumber="1080" type="description">
<paragraph id="8BF836F5FFA1FFD2BAADAB37FED6FBFA" blockId="46.[160,1156,1078,1727]" box="[160,294,1078,1103]" pageId="46" pageNumber="1080">
<emphasis id="B933EAE7FFA1FFD2BAADAB37FED6FBFA" bold="true" box="[160,294,1078,1103]" italics="true" pageId="46" pageNumber="1080">Description</emphasis>
</paragraph>
<paragraph id="8BF836F5FFA1FFD2BAADAB5BFB8EFBC7" blockId="46.[160,1156,1078,1727]" box="[160,1150,1114,1138]" pageId="46" pageNumber="1080">
Male (
<figureCitation id="137C2A70FFA1FFD2BAEBAB5BFEA2FBC7" box="[230,338,1114,1138]" captionStart="Figure 15" captionStartId="21.[160,225,1522,1545]" captionTargetBox="[163,1152,143,1491]" captionTargetId="figure-1@21.[163,1153,142,1491]" captionTargetPageId="21" captionText="Figure 15. Aegidium minor, holotype male (a, d, e), male (b, f), female (c). Habitus in dorsal view (a, c), parameres in ventral view (b), aedeagus in lateral view (d), parameres in dorsal view (e), spiculum gastrale (f), locality map (g)." figureDoi="http://doi.org/10.5281/zenodo.4746701" httpUri="https://zenodo.org/record/4746701/files/figure.png" pageId="46" pageNumber="1080">Figure 15</figureCitation>
(a,b,d–f)). Body length 12.0–17.0 mm. Colour uniformly blackish brown.
</paragraph>
<paragraph id="8BF836F5FFA1FFD2BACDAB7CFE46FB48" blockId="46.[160,1156,1078,1727]" pageId="46" pageNumber="1080">Anterior margin of frontoclypeus slightly convex in middle, slightly sinuate each side of medial convexity, with strongly protruding anterior angles, without distinct border or slightly upturned. Frontoclypeus punctate with rounded punctures separated by about 1–2 puncture diameters.</paragraph>
<paragraph id="8BF836F5FFA1FFD2BACDAA06FBD2FAAA" blockId="46.[160,1156,1078,1727]" box="[192,1058,1287,1311]" pageId="46" pageNumber="1080">Eyes relatively large: width about 1/7 distance between eyes in dorsal view.</paragraph>
<paragraph id="8BF836F5FFA1FFD2BACDAA2BFBBAF9A7" blockId="46.[160,1156,1078,1727]" pageId="46" pageNumber="1080">Pronotum with widely rounded lateral margins, slightly wider than elytra, 1.6 times wider than length, 0.55 times length of elytra. Posterior angles widely rounded, indistinct. Anterior margin bordered, border interrupted medially and not reaching anterior angles. Base of pronotum without furrow, punctate with a row of rounded punctures. Pronotal disc deeply excavated, with flattened area mediobasally. Lateral pronotal processes long, horn-shaped in lateral view, not protruding past lateral margin outline of pronotum in dorsal view. Medial excavation of pronotum punctate with rounded punctures, lateral processes smooth.</paragraph>
<paragraph id="8BF836F5FFA1FFD2BACDA91CFC80F980" blockId="46.[160,1156,1078,1727]" box="[192,880,1565,1589]" pageId="46" pageNumber="1080">Scutellum narrow subtriangular, about 1/11 length of elytra.</paragraph>
<paragraph id="8BF836F5FFA1FFD2BACDA93EFBCCF928" blockId="46.[160,1156,1078,1727]" pageId="46" pageNumber="1080">Elytra 1.2 times longer than wide, with humeral and apical umbones, widest in basal 1/3, and tapering apically in dorsal view. Elytral carinae feebly distinct. Elytra covered with relatively rounded punctures separated by 1–3 puncture diameters, becoming denser at apices.</paragraph>
<paragraph id="8BF836F5FFA1FFD2BACDA9A6FE92F90A" blockId="46.[160,1156,1078,1727]" box="[192,354,1703,1727]" pageId="46" pageNumber="1080">Macropterous.</paragraph>
<paragraph id="8BF836F5FFA0FFD3BACDAF8FFCE7FF7C" blockId="47.[160,1156,142,548]" pageId="47" pageNumber="1081">Mesotibiae without tuft of setae ventroapically. Inner apical spur of mesotibia shorter than basal mesotarsomere and not curved downwards.</paragraph>
<paragraph id="8BF836F5FFA0FFD3BACDAFD5FD06FE09" blockId="47.[160,1156,142,548]" pageId="47" pageNumber="1081">
Aedeagus with relatively long (0.55 times length of phallobase), tapering parameres (
<figureCitation id="137C2A70FFA0FFD3BAAAAFF7FEE7FEBB" box="[167,279,246,270]" captionStart="Figure 15" captionStartId="21.[160,225,1522,1545]" captionTargetBox="[163,1152,143,1491]" captionTargetId="figure-1@21.[163,1153,142,1491]" captionTargetPageId="21" captionText="Figure 15. Aegidium minor, holotype male (a, d, e), male (b, f), female (c). Habitus in dorsal view (a, c), parameres in ventral view (b), aedeagus in lateral view (d), parameres in dorsal view (e), spiculum gastrale (f), locality map (g)." figureDoi="http://doi.org/10.5281/zenodo.4746701" httpUri="https://zenodo.org/record/4746701/files/figure.png" pageId="47" pageNumber="1081">Figure 15</figureCitation>
(d)). Parameres almost as wide as apical part of phallobase in dorsal view (
<figureCitation id="137C2A70FFA0FFD3BAAAAE18FEE4FE84" box="[167,276,281,305]" captionStart="Figure 16" captionStartId="22.[160,225,1510,1533]" captionTargetBox="[164,1152,143,1479]" captionTargetId="figure-1@22.[163,1153,142,1479]" captionTargetPageId="22" captionText="Figure 16. Aegidium asperatum, holotype male (a, c, d), male (b, e). Habitus in dorsal view (a, b), aedeagus in lateral view (c), parameres in dorsal view (d), spiculum gastrale (e), locality map (f)." figureDoi="http://doi.org/10.5281/zenodo.4746705" httpUri="https://zenodo.org/record/4746705/files/figure.png" pageId="47" pageNumber="1081">Figure 16</figureCitation>
(e)). Ventrobasal transverse plate of parameres distinct. Lateral teeth of parameres feebly protruding past paramere outline in dorsal view (
<figureCitation id="137C2A70FFA0FFD3B952AE3DFC39FEE1" box="[863,969,316,340]" captionStart="Figure 15" captionStartId="21.[160,225,1522,1545]" captionTargetBox="[163,1152,143,1491]" captionTargetId="figure-1@21.[163,1153,142,1491]" captionTargetPageId="21" captionText="Figure 15. Aegidium minor, holotype male (a, d, e), male (b, f), female (c). Habitus in dorsal view (a, c), parameres in ventral view (b), aedeagus in lateral view (d), parameres in dorsal view (e), spiculum gastrale (f), locality map (g)." figureDoi="http://doi.org/10.5281/zenodo.4746701" httpUri="https://zenodo.org/record/4746701/files/figure.png" pageId="47" pageNumber="1081">Figure 15</figureCitation>
(e)). Lateral sides of parameres with small sensory areas apicad of lateral teeth (
<figureCitation id="137C2A70FFA0FFD3B961AE5FFC29FEC3" box="[876,985,350,374]" captionStart="Figure 15" captionStartId="21.[160,225,1522,1545]" captionTargetBox="[163,1152,143,1491]" captionTargetId="figure-1@21.[163,1153,142,1491]" captionTargetPageId="21" captionText="Figure 15. Aegidium minor, holotype male (a, d, e), male (b, f), female (c). Habitus in dorsal view (a, c), parameres in ventral view (b), aedeagus in lateral view (d), parameres in dorsal view (e), spiculum gastrale (f), locality map (g)." figureDoi="http://doi.org/10.5281/zenodo.4746701" httpUri="https://zenodo.org/record/4746701/files/figure.png" pageId="47" pageNumber="1081">Figure 15</figureCitation>
(b,d), arrowed). Endophallus without strongly sclerotised sclerites. Cranial part of spiculum gastrale narrow, tapering and rounded apically (
<figureCitation id="137C2A70FFA0FFD3B86BAEA5FD23FE09" box="[614,723,420,444]" captionStart="Figure 15" captionStartId="21.[160,225,1522,1545]" captionTargetBox="[163,1152,143,1491]" captionTargetId="figure-1@21.[163,1153,142,1491]" captionTargetPageId="21" captionText="Figure 15. Aegidium minor, holotype male (a, d, e), male (b, f), female (c). Habitus in dorsal view (a, c), parameres in ventral view (b), aedeagus in lateral view (d), parameres in dorsal view (e), spiculum gastrale (f), locality map (g)." figureDoi="http://doi.org/10.5281/zenodo.4746701" httpUri="https://zenodo.org/record/4746701/files/figure.png" pageId="47" pageNumber="1081">Figure 15</figureCitation>
(f)).
</paragraph>
<paragraph id="8BF836F5FFA0FFD3BACDAEC7FDF0FD91" blockId="47.[160,1156,142,548]" pageId="47" pageNumber="1081">
Female (
<figureCitation id="137C2A70FFA0FFD3BB28AEC7FE64FE6B" box="[293,404,454,478]" captionStart="Figure 15" captionStartId="21.[160,225,1522,1545]" captionTargetBox="[163,1152,143,1491]" captionTargetId="figure-1@21.[163,1153,142,1491]" captionTargetPageId="21" captionText="Figure 15. Aegidium minor, holotype male (a, d, e), male (b, f), female (c). Habitus in dorsal view (a, c), parameres in ventral view (b), aedeagus in lateral view (d), parameres in dorsal view (e), spiculum gastrale (f), locality map (g)." figureDoi="http://doi.org/10.5281/zenodo.4746701" httpUri="https://zenodo.org/record/4746701/files/figure.png" pageId="47" pageNumber="1081">Figure 15</figureCitation>
(c)) differs from male in having protibial spur, relatively smaller pronotum without armature and in absence of inner protibial tooth. Body length of examined specimens 13.0 mm.
</paragraph>
<paragraph id="8BF836F5FFA0FFD3BAADAD51FEFEFDDC" blockId="47.[160,1155,592,686]" box="[160,270,592,617]" pageId="47" pageNumber="1081">
<emphasis id="B933EAE7FFA0FFD3BAADAD51FEFEFDDC" bold="true" box="[160,270,592,617]" italics="true" pageId="47" pageNumber="1081">Variation</emphasis>
</paragraph>
<paragraph id="8BF836F5FFA0FFD3BAADAD75FC6CFD1B" blockId="47.[160,1155,592,686]" pageId="47" pageNumber="1081">Pronotum shape in males varies from fully developed as described above to ill-defined lateral pronotal processes and shallow central pronotal excavation.</paragraph>
</subSubSection>
<subSubSection id="C35D657EFFA0FFD3BAADADDAFB74FCA2" box="[160,1156,731,791]" pageId="47" pageNumber="1081" type="distribution">
<paragraph id="8BF836F5FFA0FFD3BAADADDAFEDCFD41" blockId="47.[160,1156,731,791]" box="[160,300,731,756]" pageId="47" pageNumber="1081">
<emphasis id="B933EAE7FFA0FFD3BAADADDAFEDCFD41" bold="true" box="[160,300,731,756]" italics="true" pageId="47" pageNumber="1081">Distribution</emphasis>
</paragraph>
<paragraph id="8BF836F5FFA0FFD3BAADADFEFB74FCA2" blockId="47.[160,1156,731,791]" box="[160,1156,767,791]" pageId="47" pageNumber="1081">
The species is known from a few localities in the western Ecuadorian Andes (
<figureCitation id="137C2A70FFA0FFD3B9FDADFEFBAAFCA2" box="[1008,1114,767,791]" captionStart="Figure 15" captionStartId="21.[160,225,1522,1545]" captionTargetBox="[163,1152,143,1491]" captionTargetId="figure-1@21.[163,1153,142,1491]" captionTargetPageId="21" captionText="Figure 15. Aegidium minor, holotype male (a, d, e), male (b, f), female (c). Habitus in dorsal view (a, c), parameres in ventral view (b), aedeagus in lateral view (d), parameres in dorsal view (e), spiculum gastrale (f), locality map (g)." figureDoi="http://doi.org/10.5281/zenodo.4746701" httpUri="https://zenodo.org/record/4746701/files/figure.png" pageId="47" pageNumber="1081">Figure 15</figureCitation>
(g)).
</paragraph>
</subSubSection>
</treatment>
</document>
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