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<document id="18444F72C6A102EFC35CF21733D538F8" ID-CLB-Dataset="72886" ID-DOI="10.11646/zootaxa.5150.3.6" ID-GBIF-Dataset="c8c16c97-1eae-42d2-9ab3-750dc14ea775" ID-ISSN="1175-5326" ID-Zenodo-Dep="6623125" ID-ZooBank="D59052B5-85CD-466F-B4EB-812226DC913E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654676678377" checkinUser="plazi" docAuthor="Shamshev, Igor V. &amp; Perkovsky, Evgeny E." docDate="2022" docId="03EE87B7D14C51390AB65095379E2B34" docLanguage="en" docName="zootaxa.5150.3.6.pdf" docOrigin="Zootaxa 5150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Meghyperiella porphyropsoides Meunier 1908" docType="treatment" docVersion="7" lastPageNumber="417" masterDocId="FFD7FFCFD14F513F0A21551736482909" masterDocTitle="A review of fossil taxa of Microphorinae (Diptera, Dolichopodidae sensu lato), with redescription of the Eocene genus Meghyperiella Meunier" masterLastPageNumber="427" masterPageNumber="411" pageNumber="414" updateTime="1699339181600" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D1F61883A2D96B808A575FAA875D9738" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EF7E8EC48EDC1130E89A1E0A0C991942">A review of fossil taxa of Microphorinae (Diptera, Dolichopodidae sensu lato), with redescription of the Eocene genus Meghyperiella Meunier</mods:title>
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<mods:name id="7B3FE38FE9B291DC65500FB6B4113F9C" type="personal">
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<mods:namePart id="418362B65821ABDB6FD39923F01FFE72">Shamshev, Igor V.</mods:namePart>
<mods:affiliation id="E61880B67B8703C06A32B293EA83482E">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:namePart id="0BF9B46FEFDB1C3DB1D5255E5406BAF3">Perkovsky, Evgeny E.</mods:namePart>
<mods:affiliation id="F51A0B7BCDDD69FFCCE146E7034AED68">Schmalhausen Institute of Zoology NAS of Ukraine, vul. B. Khmelnytskogo, 15, Kiev, 01030 Ukraine. &amp; Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117997, Russia</mods:affiliation>
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<mods:title id="5E57B2270745AA8DE87379D386A9A242">Zootaxa</mods:title>
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<mods:date id="D90522B342150DEC594F4C91ECA38CEA">2022</mods:date>
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<mods:number id="8A88BC77DD4CC5E5F50D2C1387074368">2022-06-08</mods:number>
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<mods:number id="AD20E7FAF99735EDDEB2FD194D74A074">5150</mods:number>
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<treatment id="03EE87B7D14C51390AB65095379E2B34" ID-DOI="http://doi.org/10.5281/zenodo.6645946" ID-GBIF-Taxon="195913582" ID-Zenodo-Dep="6645946" LSID="urn:lsid:plazi:treatment:03EE87B7D14C51390AB65095379E2B34" httpUri="http://treatment.plazi.org/id/03EE87B7D14C51390AB65095379E2B34" lastPageId="6" lastPageNumber="417" pageId="3" pageNumber="414">
<subSubSection id="C35D652AD14C513C0AB6509534CF2C95" box="[151,647,1410,1437]" pageId="3" pageNumber="414" type="nomenclature">
<paragraph id="8BF836A1D14C513C0AB6509534CF2C95" blockId="3.[151,647,1410,1473]" box="[151,647,1410,1437]" pageId="3" pageNumber="414">
<heading id="D0B081CDD14C513C0AB6509534CF2C95" bold="true" box="[151,647,1410,1437]" fontSize="11" level="1" pageId="3" pageNumber="414" reason="1">
<taxonomicName id="4C474D22D14C513C0AB6509534CF2C95" ID-CoL="8CS9P" authority="Meunier" authorityName="Meunier" authorityYear="1908" box="[151,647,1410,1437]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14C513C0AB6509534CF2C95" bold="true" box="[151,647,1410,1437]" pageId="3" pageNumber="414">
<emphasis id="B933EAB3D14C513C0AB65095345C2C94" bold="true" box="[151,532,1410,1437]" italics="true" pageId="3" pageNumber="414">Meghyperiella porphyropsoides</emphasis>
Meunier
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35D652AD14C513C0AB650B037442CC8" box="[151,268,1447,1473]" pageId="3" pageNumber="414" type="description">
<paragraph id="8BF836A1D14C513C0AB650B037442CC8" blockId="3.[151,647,1410,1473]" box="[151,268,1447,1473]" pageId="3" pageNumber="414">
(
<figureCitation id="137C2A24D14C513C0ABE50B0374B2CC8" box="[159,259,1447,1473]" captionStart="FIGURES 1, 2" captionStartId="5.[151,264,1893,1917]" captionTargetBox="[227,1360,180,1865]" captionTargetId="figure-19@5.[227,1361,179,1866]" captionTargetPageId="5" captionText="FIGURES 1, 2. Meghyperiella porphyropsoides Meunier, male, Rovno amber. 1. Habitus, right lateral view; 2. Wing and terminalia. Abbreviations: CuA+CuP —anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; epand—epandrium; hypd lb—apical lobe of hypandrium; M1, M2, M4—medial veins; ph—phallus; R1, R2+3, R 4+5 —radial veins." figureDoi="http://doi.org/10.5281/zenodo.6623130" httpUri="https://zenodo.org/record/6623130/files/figure.png" pageId="3" pageNumber="414">Figs 1, 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35D652AD14C513C0AB650FC33212E17" pageId="3" pageNumber="414" type="reference_group">
<paragraph id="8BF836A1D14C513C0AB650FC334E2F0A" blockId="3.[151,1286,1515,1570]" box="[151,1286,1515,1540]" pageId="3" pageNumber="414">
<treatmentCitationGroup id="AB57118FD14C513C0AB650FC334E2F0A" box="[151,1286,1515,1540]" pageId="3" pageNumber="414">
<treatmentCitation id="0AE610B0D14C513C0AB650FC34E12F0A" author="Meunier, F." box="[151,681,1515,1539]" page="112" pageId="3" pageNumber="414" year="1908">
<taxonomicName id="4C474D22D14C513C0AB650FC34E12F0A" ID-CoL="8CS9P" authority="Meunier, 1908: 112" authorityName="Meunier" authorityPageNumber="112" authorityYear="1908" box="[151,681,1515,1539]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14C513C0AB650FC37922F0A" box="[151,474,1515,1539]" italics="true" pageId="3" pageNumber="414">Meghyperiella porphyropsoides</emphasis>
<bibRefCitation id="EFD64B50D14C513C0BC050FC34E12F0A" author="Meunier, F." box="[481,681,1515,1539]" pageId="3" pageNumber="414" pagination="81 - 135" refId="ref15542" refString="Meunier, F. (1908) Monographie des Empidae de l'ambre de la Baltique et catalogue bibliographique complet sur les Dipteres fossiles de cette resine. Annales des Sciences Naturelles, Zoologie, Series 9, 7, 81 - 135." type="journal article" year="1908">Meunier, 1908: 112</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 8, figs 5–7.
<typeStatus id="54FC8803D14C513C096D50FB35362F0D" box="[844,894,1516,1540]" pageId="3" pageNumber="414">Type</typeStatus>
locality: Baltic region, Upper Eocene.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF836A1D14C513C0AB6531D35ED2F2B" blockId="3.[151,1286,1515,1570]" box="[151,933,1546,1570]" pageId="3" pageNumber="414">
<treatmentCitationGroup id="AB57118FD14C513C0AB6531D35ED2F2B" box="[151,933,1546,1570]" pageId="3" pageNumber="414">
<emphasis id="B933EAB3D14C513C0AB6531D37552F2B" box="[151,285,1546,1570]" italics="true" pageId="3" pageNumber="414">Meghyerellia</emphasis>
<treatmentCitation id="0AE610B0D14C513C0B05531D34E32F2B" author="Keilbach, R." box="[292,683,1546,1570]" page="369" pageId="3" pageNumber="414" year="1982">
<taxonomicName id="4C474D22D14C513C0B05531D34E32F2B" ID-CoL="8CS9P" authority=": Keilbach, 1982: 369" authorityName=": Keilbach" authorityPageNumber="369" authorityYear="1982" box="[292,683,1546,1570]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14C513C0B05531D37852F2B" box="[292,461,1546,1570]" italics="true" pageId="3" pageNumber="414">porphyropsoides</emphasis>
:
<bibRefCitation id="EFD64B50D14C513C0BFB531D34E32F2B" author="Keilbach, R." box="[474,683,1546,1570]" pageId="3" pageNumber="414" pagination="129 - 286" refId="ref14602" refString="Keilbach, R. (1982). Bibliographie und Liste der Arten tierischer Einschlusse in fossilen Harzen sowie ihrer Aufbewahrungsorte. Deutsche entomologische Zeitschrift, Neue Folge 29, Heft 1 - 3 &amp; 4 - 5, 129 - 286 &amp; 301 - 391." type="journal article" year="1982">Keilbach, 1982: 369</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, error (
<treatmentCitation id="0AE610B0D14C513C08D5531D35D12F2B" author="Spahr, U." box="[756,921,1546,1570]" page="45" pageId="3" pageNumber="414" year="1985">
<bibRefCitation id="EFD64B50D14C513C08D5531D35D12F2B" author="Spahr, U." box="[756,921,1546,1570]" pageId="3" pageNumber="414" pagination="1 - 146" refId="ref16471" refString="Spahr, U. (1985) Erganzungen und Berichtigungen zu R. Keilbach's Bibliographie und Liste der Bernsteinfossilien - Ordnung Diptera. Stutgarter Beitrage zur Naturkunde, Series B, 111, 1 - 146." type="journal article" year="1985">Spahr, 1985: 45</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF836A1D14C513C0AB6534733212E17" blockId="3.[151,1437,1616,2003]" pageId="3" pageNumber="414">
<emphasis id="B933EAB3D14C513C0AB6534737C42F63" bold="true" box="[151,396,1616,1642]" pageId="3" pageNumber="414">Notes on type series.</emphasis>
<bibRefCitation id="EFD64B50D14C513C0BB25347340C2F63" author="Meunier, F." box="[403,580,1616,1642]" pageId="3" pageNumber="414" pagination="81 - 135" refId="ref15542" refString="Meunier, F. (1908) Monographie des Empidae de l'ambre de la Baltique et catalogue bibliographique complet sur les Dipteres fossiles de cette resine. Annales des Sciences Naturelles, Zoologie, Series 9, 7, 81 - 135." type="journal article" year="1908">Meunier (1908)</bibRefCitation>
mentioned
<specimenCount id="9D41FD28D14C513C08EB534635742F63" box="[714,828,1616,1643]" pageId="3" pageNumber="414" type="male">two males</specimenCount>
and
<specimenCount id="9D41FD28D14C513C0955534635BC2F63" box="[884,1012,1616,1643]" pageId="3" pageNumber="414" type="female">one female</specimenCount>
. However,
<bibRefCitation id="EFD64B50D14C513C0E52534733402F63" author="Ulrich, H." box="[1139,1288,1616,1642]" pageId="3" pageNumber="414" pagination="233 - 243" refId="ref16673" refString="Ulrich, H. (2004) Phylogenetic considerations about an early colonization of the sea coasts by Dolichopodidae (Diptera). Studia dipterologica, 11, 233 - 243." type="journal article" year="2004">Ulrich (2004</bibRefCitation>
; unpublished notes on Meunier’s types) found only
<specimenCount id="9D41FD28D14C513C0864536235412F87" box="[581,777,1652,1679]" pageId="3" pageNumber="414" type="male" typeStatus="syntype">one syntype male</specimenCount>
in GZG (
<bibRefCitation id="EFD64B50D14C513C0957536332192F87" author="Meunier, F." box="[886,1105,1652,1678]" pageId="3" pageNumber="414" pagination="81 - 135" refId="ref15542" refString="Meunier, F. (1908) Monographie des Empidae de l'ambre de la Baltique et catalogue bibliographique complet sur les Dipteres fossiles de cette resine. Annales des Sciences Naturelles, Zoologie, Series 9, 7, 81 - 135." type="journal article" year="1908">Meunier 1908: 112</bibRefCitation>
, no. 5323, R. Klebs’s Collection), and the remaining
<specimenCount id="9D41FD28D14C513C0B86538E34712FBA" box="[423,569,1689,1715]" pageId="3" pageNumber="414" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
are probably lost. We were only able to examine photographs of this inclusion (should be the
<typeStatus id="54FC8803D14C513C0B6153AB37E42FDE" box="[320,428,1724,1751]" pageId="3" pageNumber="414" type="lectotype">lectotype</typeStatus>
). The amber piece has Meunier’s no. 5323 (new no. GZG.BST.00176). In addition, there is a Museum label with hand-written specific name (Z5323,
<taxonomicName id="4C474D22D14C513C091E53F732EC2FF3" ID-CoL="8CS9P" authorityName=": Keilbach" authorityYear="1982" box="[831,1188,1760,1786]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14C513C091E53F732EC2FF3" box="[831,1188,1760,1786]" italics="true" pageId="3" pageNumber="414">Meghyperiella porphyropsoides</emphasis>
</taxonomicName>
<specimenCount id="9D41FD28D14C513C0E8D53F632892FF2" box="[1196,1217,1761,1787]" pageId="3" pageNumber="414" type="male">♂</specimenCount>
) and Hans Ulrich’s hand-written label (Z5323,
<taxonomicName id="4C474D22D14C513C0BEA5213343C2E17" ID-CoL="9G27N" authorityName="Collin" authorityYear="1960" box="[459,628,1796,1822]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="subFamily" subFamily="Microphorinae">Microphorinae</taxonomicName>
, M2 reduziert, H. Ulrich 2001). The specimen is in good condition.
</paragraph>
</subSubSection>
<subSubSection id="C35D652AD14C513C0AE6523F34912EDA" pageId="3" pageNumber="414" type="materials_examined">
<paragraph id="8BF836A1D14C513C0AE6523F37412E83" blockId="3.[151,1437,1616,2003]" pageId="3" pageNumber="414">
<emphasis id="B933EAB3D14C513C0AE6523F37F92E4B" bold="true" box="[199,433,1832,1858]" pageId="3" pageNumber="414">Material examined.</emphasis>
<materialsCitation id="3B2F3CFCD14C513C0B99523F355D2E4B" ID-GBIF-Occurrence="3808178302" box="[440,789,1832,1859]" country="Ukraine" location="Rovno" pageId="3" pageNumber="414" specimenCount="1" stateProvince="Rivne">
<collectingCountry id="F3507631D14C513C0B99523F345D2E4B" box="[440,533,1832,1858]" name="Ukraine" pageId="3" pageNumber="414">Ukraine</collectingCountry>
,
<collectingRegion id="4983F843D14C513C0800523E34242E4A" box="[545,620,1833,1859]" country="Ukraine" name="Rivne" pageId="3" pageNumber="414">Rovno</collectingRegion>
amber: OL-39
</materialsCitation>
,
<materialsCitation id="3B2F3CFCD14C513C0900523F344F2E6F" ID-GBIF-Occurrence="3808178303" collectionCode="SIZK" country="Ukraine" county="Zhitomir Region" location="Upper Eocene" municipality="Olevsk" pageId="3" pageNumber="414" specimenCount="1" specimenCount-male="1" stateProvince="Rivne">
<collectingCountry id="F3507631D14C513C0900523F35362E4B" box="[801,894,1832,1858]" name="Ukraine" pageId="3" pageNumber="414">Ukraine</collectingCountry>
,
<collectingCounty id="62994E2DD14C513C09A8523F32002E4B" box="[905,1096,1832,1858]" pageId="3" pageNumber="414">Zhitomir Region</collectingCounty>
,
<collectingMunicipality id="6B9CACDBD14C513C0E72523F32ED2E4B" box="[1107,1189,1832,1858]" pageId="3" pageNumber="414">Olevsk</collectingMunicipality>
,
<collectingRegion id="4983F843D14C513C0E90523E32B42E4A" box="[1201,1276,1833,1859]" country="Ukraine" name="Rivne" pageId="3" pageNumber="414">Rovno</collectingRegion>
amber,
<location id="8E98607AD14C513C0F77523E36A12E6E" LSID="urn:lsid:plazi:treatment:03EE87B7D14C51390AB65095379E2B34:8E98607AD14C513C0F77523E36A12E6E" country="Ukraine" county="Zhitomir Region" municipality="Olevsk" name="Upper Eocene" pageId="3" pageNumber="414" stateProvince="Rivne">Upper Eocene</location>
(
<specimenCount id="9D41FD28D14C513C0ADC525B37602E6E" box="[253,296,1868,1895]" pageId="3" pageNumber="414" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED56AE64D14C513C0B15525B373F2E6F" box="[308,375,1868,1894]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="3" pageNumber="414">SIZK</collectionCode>
); LKV-1012
</materialsCitation>
,
<materialsCitation id="3B2F3CFCD14C513C0832525B374D2E83" ID-GBIF-Occurrence="3808178304" collectionCode="SIZK" country="Ukraine" location="Upper Eocene" municipality="Veselukha river" pageId="3" pageNumber="414" specimenCount="2" specimenCount-male="2" stateProvince="Rivne">
<collectingCountry id="F3507631D14C513C0832525B34382E6F" box="[531,624,1868,1894]" name="Ukraine" pageId="3" pageNumber="414">Ukraine</collectingCountry>
,
<collectingRegion id="4983F843D14C513C085D525A356A2E6F" box="[636,802,1868,1895]" country="Ukraine" name="Rivne" pageId="3" pageNumber="414">Rovno Region</collectingRegion>
,
<collectingMunicipality id="6B9CACDBD14C513C090C525B35952E6F" box="[813,989,1868,1894]" pageId="3" pageNumber="414">Veselukha river</collectingMunicipality>
valley,
<collectingRegion id="4983F843D14C513C0E16525A32CA2E6E" box="[1079,1154,1869,1895]" country="Ukraine" name="Rivne" pageId="3" pageNumber="414">Rovno</collectingRegion>
amber,
<location id="8E98607AD14C513C0EFC525A33352E6E" LSID="urn:lsid:plazi:treatment:03EE87B7D14C51390AB65095379E2B34:8E98607AD14C513C0EFC525A33352E6E" box="[1245,1405,1869,1895]" country="Ukraine" municipality="Veselukha river" name="Upper Eocene" pageId="3" pageNumber="414" stateProvince="Rivne">Upper Eocene</location>
(
<specimenCount id="9D41FD28D14C513C0FAC525B36E62E82" pageId="3" pageNumber="414" type="male">2 ♂</specimenCount>
,
<collectionCode id="ED56AE64D14C513C0A9B526736B52E83" box="[186,253,1904,1930]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="3" pageNumber="414">SIZK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF836A1D14C513C0AE6528234912EDA" blockId="3.[151,1437,1616,2003]" pageId="3" pageNumber="414">
In addition, we have examined photographs of the male and female specimens from the personal collection of Hans Ulrich (housed at
<collectionCode id="ED56AE64D14C513C0B8052AE37B92EDA" box="[417,497,1977,2003]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="414" type="Museum">ZFMK</collectionCode>
) (
<bibRefCitation id="EFD64B50D14C513C082652AF34842EDA" author="Ulrich, H." box="[519,716,1976,2003]" pageId="3" pageNumber="414" pagination="233 - 243" refId="ref16673" refString="Ulrich, H. (2004) Phylogenetic considerations about an early colonization of the sea coasts by Dolichopodidae (Diptera). Studia dipterologica, 11, 233 - 243." type="journal article" year="2004">Ulrich 2004: 235</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35D652AD14B513B0AE6558132B129DD" pageId="4" pageNumber="415" type="diagnosis">
<paragraph id="8BF836A1D14B513B0AE6558132B129DD" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE65581370A29B9" bold="true" box="[199,322,150,176]" pageId="4" pageNumber="415">Diagnosis.</emphasis>
Small (body nearly
<quantity id="4CBF9B44D14B513B08045581342F29B8" box="[549,615,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="415" unit="mm" value="2.0">2 mm</quantity>
) blackish flies; wing with two M veins beyond the discal cell (apical section of M
<subScript id="17C334E4D14B513B0B2055DA374129D2" attach="left" box="[257,265,205,219]" fontSize="6" pageId="4" pageNumber="415">2</subScript>
absent); vein closing discal cell (i.e., base of M
<subScript id="17C334E4D14B513B090155DA356029D2" attach="left" box="[800,808,205,219]" fontSize="6" pageId="4" pageNumber="415">2</subScript>
+ dm-m crossvein) smoothly undulating.
</paragraph>
</subSubSection>
<subSubSection id="C35D652AD14B513B0AE655C933162C05" pageId="4" pageNumber="415" type="description">
<paragraph id="8BF836A1D14B513B0AE655C935FE28D8" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE655C937F229F1" bold="true" box="[199,442,222,248]" pageId="4" pageNumber="415">Redescription. Male</emphasis>
(
<figureCitation id="137C2A24D14B513B0BEB55C9344429F1" box="[458,524,222,248]" captionStart="FIGURES 1, 2" captionStartId="5.[151,264,1893,1917]" captionTargetBox="[227,1360,180,1865]" captionTargetId="figure-19@5.[227,1361,179,1866]" captionTargetPageId="5" captionText="FIGURES 1, 2. Meghyperiella porphyropsoides Meunier, male, Rovno amber. 1. Habitus, right lateral view; 2. Wing and terminalia. Abbreviations: CuA+CuP —anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; epand—epandrium; hypd lb—apical lobe of hypandrium; M1, M2, M4—medial veins; ph—phallus; R1, R2+3, R 4+5 —radial veins." figureDoi="http://doi.org/10.5281/zenodo.6623130" httpUri="https://zenodo.org/record/6623130/files/figure.png" pageId="4" pageNumber="415">Fig. 1</figureCitation>
). Blackish. Body length
<quantity id="4CBF9B44D14B513B090855C935FB29F0" box="[809,947,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="4" pageNumber="415" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
, wing length
<quantity id="4CBF9B44D14B513B0E6F55C932E029F0" box="[1102,1192,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="415" unit="mm" value="1.5">1.5 mm</quantity>
.
<emphasis id="B933EAB3D14B513B0E9255C932B829F1" box="[1203,1264,222,248]" italics="true" pageId="4" pageNumber="415">Head</emphasis>
. Frons entirely obliterated by eyes. Face and clypeus recessed in oral cavity. Face broad, subrectangular. Ocellar triangle with 2 stronger, moderately long, almost parallel, proclinate ocellars and minute postocellars. Occiput with series of 8–9 visible postocular setae per side. Postgena covered with setae. Antenna with scape short, about half length of pedicel; pedicel about 2X longer than wide, with several setae; postpedicel subtriangular; gradually narrowing nearly beyond basal fourth to third, 3.5–4X longer than wide; stylus arista-like, terminal, about 2.3–2.5X longer than postpedicel, 2-articled, basal article short. Palpus with scattered short setae.
</paragraph>
<paragraph id="8BF836A1D14B513B0AE654CD37412BC5" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE654CD375028FD" box="[199,280,474,500]" italics="true" pageId="4" pageNumber="415">Thorax</emphasis>
. Proepisternum with 1 short seta on upper part. Postpronotal lobe with 1 strong, long and 1 short, weak seta. Mesonotum with 1 moderately long presutural supra-alar (with several shorter supplementary setae); 2 long notopleurals (with some additional short weak setae); 1 long postalar; scutellum with 6 setae (middle pair very long, lateral pairs short); rows of acrostichal and dorsocentral setae separated by bare space; acrostichals short, quadriserial, lacking on prescutellar depression; dorsocentrals uniserial, numerous, apparently 11–12 setae per row, short and closely spaced anteriorly, becoming longer posteriorly, 2 posterior-most setae (prescutellars) strong and widely separated.
</paragraph>
<paragraph id="8BF836A1D14B513B0AE657C032EF2AAD" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE657C036B42BF9" box="[199,252,727,752]" italics="true" pageId="4" pageNumber="415">Legs</emphasis>
. Coxae with scattered setae of different lengths. Fore and mid femora with complete rows of 8–9 short to moderately long (2–3 subapical setae on fore femur) posterodorsal setae, some slightly prominent setae anterodorsally; hind femur with complete row of moderately long setae dorsally (slightly shorter than femur width) and complete row of mostly short anteroventral setae (3–4 subapical setae longer). Fore and mid tibiae with somewhat longer setulae dorsally; hind tibia with more or less distinct row of short setae dorsally. Tarsomeres of all legs covered with simple setulae. Tarsal claws, pulvilli and empodium normally developed on all legs.
</paragraph>
<paragraph id="8BF836A1D14B513B0AE656B8334F2AC1" blockId="4.[151,1437,150,1977]" box="[199,1287,942,968]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE656B837492AC1" box="[199,257,943,968]" italics="true" pageId="4" pageNumber="415">Wing</emphasis>
membrane uniformly faintly infuscate. Calypter brownish, brown fringed. Halter brownish.
</paragraph>
<paragraph id="8BF836A1D14B513B0AE656C5342C2D19" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE656C537792AE5" box="[199,305,978,1004]" italics="true" pageId="4" pageNumber="415">Abdomen</emphasis>
with tergites 1–6 bearing numerous moderately long, uniform setae, sternites 1–6 with similar sparser setae; segments 7–8 obscured from view.
</paragraph>
<paragraph id="8BF836A1D14B513B0AE6510D37352DCD" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE6510D37092D3D" box="[199,321,1050,1076]" italics="true" pageId="4" pageNumber="415">Terminalia</emphasis>
small. Right epandrial lamella narrowed toward apex, with some hardly distinguishable, short apical lobes (surstyli?). Left epandrial lamella obscured from view. Hypandrium with basal part convex, broad apical lobes of subequal lengths; probably bearing 3 long setae on each side. Postgonites obscured from view. Phallus tubular, with tip not narrowed, phallic process (if developed) obscured from view. Hypoproct (if developed) and cercus obscured from view.
</paragraph>
<paragraph id="8BF836A1D14B513B0AE651D933162C05" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE651D937562DE1" bold="true" box="[199,286,1230,1256]" pageId="4" pageNumber="415">Female</emphasis>
. Similar to male except for following characters: head dichoptic; eyes smaller; dorsal and ventral ommatidia similar in size. Mesoscutum with slightly less pronounced arch. Wing with anal lobe slightly smaller.
</paragraph>
</subSubSection>
<subSubSection id="C35D652AD14B51390AE65001379E2B34" lastPageId="6" lastPageNumber="417" pageId="4" pageNumber="415" type="discussion">
<paragraph id="8BF836A1D14B513B0AE6500132242FB5" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
<emphasis id="B933EAB3D14B513B0AE6500137732C39" bold="true" box="[199,315,1302,1328]" pageId="4" pageNumber="415">Remarks.</emphasis>
Amongst the genera placed in the “
<taxonomicName id="4C474D22D14B513B08F2500135372C39" authorityName="Collin" authorityYear="1960" box="[723,895,1302,1328]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="subFamily" subFamily="Microphorinae">Microphorinae</taxonomicName>
” lineage of
<taxonomicName id="4C474D22D14B513B0E2B500132F52C39" authorityName="Latreille" authorityYear="1809" box="[1034,1213,1302,1328]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
<emphasis id="B933EAB3D14B513B0EE4500033702C39" box="[1221,1336,1302,1328]" italics="true" pageId="4" pageNumber="415">
<taxonomicName id="4C474D22D14B513B0EE45000334B2C39" box="[1221,1283,1303,1328]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
lato
</emphasis>
,
<taxonomicName id="4C474D22D14B513B0F66500136A62C5D" authorityName="Meunier" authorityYear="1908" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B0F66500136A62C5D" italics="true" pageId="4" pageNumber="415">Meghyperiella</emphasis>
</taxonomicName>
should be compared primarily with
<taxonomicName id="4C474D22D14B513B08AF502D35072C5D" box="[654,847,1338,1364]" class="Insecta" family="Dolichopodidae" genus="Pristinmicrophor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B08AF502D35072C5D" box="[654,847,1338,1364]" italics="true" pageId="4" pageNumber="415">Pristinmicrophor</emphasis>
</taxonomicName>
because they both possess a wing with only two M veins beyond the discal cell (apical section of M
<subScript id="17C334E4D14B513B089F5066348E2C76" attach="left" box="[702,710,1393,1407]" fontSize="6" pageId="4" pageNumber="415">2</subScript>
absent).
<taxonomicName id="4C474D22D14B513B090F504935A72C71" box="[814,1007,1374,1400]" class="Insecta" family="Dolichopodidae" genus="Pristinmicrophor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B090F504935A72C71" box="[814,1007,1374,1400]" italics="true" pageId="4" pageNumber="415">Pristinmicrophor</emphasis>
</taxonomicName>
(with single species
<taxonomicName id="4C474D22D14B513B0EFE504837F92C94" authority="Tang, Shi, Wang &amp; Yang" authorityName="Tang, Shi, Wang &amp; Yang" authorityYear="2019" class="Insecta" family="Dolichopodidae" genus="Pristinmicrophor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="hukawngensis">
<emphasis id="B933EAB3D14B513B0EFE504832B82C71" box="[1247,1264,1375,1400]" italics="true" pageId="4" pageNumber="415">P</emphasis>
.
<emphasis id="B933EAB3D14B513B0EDE504933D42C71" box="[1279,1436,1374,1400]" italics="true" pageId="4" pageNumber="415">hukawngensis</emphasis>
Tang, Shi, Wang &amp; Yang
</taxonomicName>
) was described very recently from mid-Cretaceous amber of northern
<collectingCountry id="F3507631D14B513B0EEA509433702C94" box="[1227,1336,1411,1437]" name="Myanmar" pageId="4" pageNumber="415">Myanmar</collectingCountry>
(
<bibRefCitation id="EFD64B50D14B513B0F66509536B12CC9" author="Tang, C. &amp; Shi, C. &amp; Wang, S. &amp; Yang, D." pageId="4" pageNumber="415" pagination="1 - 4" refId="ref16508" refString="Tang, C. - f., Shi, C., Wang, S. &amp; Yang, D. (2019) The first report of Dolichopodidae from mid-Cretaceous amber of northern Myanmar. Cretaceous Research, 104, 1 - 4. https: // doi. org / 10.1016 / j. cretres. 2019.07.009" type="journal article" year="2019">
Tang
<emphasis id="B933EAB3D14B513B0FA9509436E52CC9" italics="true" pageId="4" pageNumber="415">et al</emphasis>
. 2019
</bibRefCitation>
). However,
<taxonomicName id="4C474D22D14B513B0BA050B134AF2CC9" authorityName="Tang, Shi, Wang &amp; Yang" authorityYear="2019" box="[385,743,1446,1472]" class="Insecta" family="Dolichopodidae" genus="Pristinmicrophor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="hukawngensis">
<emphasis id="B933EAB3D14B513B0BA050B134AF2CC9" box="[385,743,1446,1472]" italics="true" pageId="4" pageNumber="415">Pristinmicrophor hukawngensis</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName id="4C474D22D14B513B0E5350B136952CED" authorityName=": Keilbach" authorityYear="1982" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14B513B0E5350B136952CED" italics="true" pageId="4" pageNumber="415">Meghyperiella porphyropsoides</emphasis>
</taxonomicName>
by several distinctive characters and these species are undoubtedly non-congeneric (see below for additional discussion). According to the original description,
<taxonomicName id="4C474D22D14B513B08F550F835D92F01" authorityName="Tang, Shi, Wang &amp; Yang" authorityYear="2019" box="[724,913,1518,1545]" class="Insecta" family="Dolichopodidae" genus="Pristinmicrophor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="hukawngensis">
<emphasis id="B933EAB3D14B513B08F550F834AD2F01" box="[724,741,1519,1544]" italics="true" pageId="4" pageNumber="415">P</emphasis>
.
<emphasis id="B933EAB3D14B513B08D550F935D92F01" box="[756,913,1518,1544]" italics="true" pageId="4" pageNumber="415">hukawngensis</emphasis>
</taxonomicName>
has a wing with very narrow, almost parallelsided cell dm (
<emphasis id="B933EAB3D14B513B0B1B530437CA2F25" box="[314,386,1555,1580]" italics="true" pageId="4" pageNumber="415">versus</emphasis>
cell dm broadened toward apex in
<taxonomicName id="4C474D22D14B513B0927530435AE2F25" authorityName=": Keilbach" authorityYear="1982" box="[774,998,1554,1581]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14B513B0927530435552F25" box="[774,797,1555,1580]" italics="true" pageId="4" pageNumber="415">M</emphasis>
.
<emphasis id="B933EAB3D14B513B090A530535AE2F25" box="[811,998,1554,1580]" italics="true" pageId="4" pageNumber="415">porphyropsoides</emphasis>
</taxonomicName>
) and dichoptic eyes in the male (
<emphasis id="B933EAB3D14B513B0F74530433D52F25" box="[1365,1437,1555,1580]" italics="true" pageId="4" pageNumber="415">versus</emphasis>
holoptic in
<taxonomicName id="4C474D22D14B513B0B37532037BE2F59" authorityName=": Keilbach" authorityYear="1982" box="[278,502,1590,1617]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14B513B0B37532037652F59" box="[278,301,1591,1616]" italics="true" pageId="4" pageNumber="415">M</emphasis>
.
<emphasis id="B933EAB3D14B513B0B1A532137BE2F59" box="[315,502,1590,1616]" italics="true" pageId="4" pageNumber="415">porphyropsoides</emphasis>
</taxonomicName>
).
<taxonomicName id="4C474D22D14B513B082C532134FE2F59" authorityName="Negrobov" authorityYear="1978" box="[525,694,1590,1616]" class="Insecta" family="Dolichopodidae" genus="Archichrysotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B082C532134FE2F59" box="[525,694,1590,1616]" italics="true" pageId="4" pageNumber="415">Archichrysotus</emphasis>
</taxonomicName>
is another “microphorine”-like genus possessing a wing with only two M veins beyond the discal cell (
<bibRefCitation id="EFD64B50D14B513B080E534D34AB2F7D" author="Negrobov, O. P." box="[559,739,1626,1652]" pageId="4" pageNumber="415" pagination="81 - 90" refId="ref15585" refString="Negrobov, O. P. (1978) Flies of the superfamily Empidoidea (Diptera) from Cretaceous retinite in northern Siberia. Paleontologicheskii Zhurnal, 1978 (2), 81 - 90. [in Russian]" type="journal article" year="1978">Negrobov 1978</bibRefCitation>
; Cumming &amp;
<bibRefCitation id="EFD64B50D14B513B09A5534D32642F7D" author="Grimaldi, D. A. &amp; Cumming, J. M." box="[900,1068,1626,1652]" pageId="4" pageNumber="415" pagination="1 - 124" refId="ref14337" refString="Grimaldi, D. A. &amp; Cumming, J. M. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 1 - 124." type="journal article" year="1999">Grimaldi 1999</bibRefCitation>
). However, species of this group share one-articulated arista-like stylus (
<emphasis id="B933EAB3D14B513B0870536834D12F91" box="[593,665,1663,1688]" italics="true" pageId="4" pageNumber="415">versus</emphasis>
two-articulated in
<taxonomicName id="4C474D22D14B513B094D536832042F91" authorityName=": Keilbach" authorityYear="1982" box="[876,1100,1662,1689]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14B513B094D536835CB2F91" box="[876,899,1663,1688]" italics="true" pageId="4" pageNumber="415">M</emphasis>
.
<emphasis id="B933EAB3D14B513B09B0536932042F91" box="[913,1100,1662,1688]" italics="true" pageId="4" pageNumber="415">porphyropsoides</emphasis>
</taxonomicName>
) and they are classified in the “
<taxonomicName id="4C474D22D14B513B0A8053B537132FB5" authorityName="Chvála" authorityYear="1981" box="[161,347,1698,1724]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="subFamily" subFamily="Parathalassiinae">Parathalassiinae</taxonomicName>
” lineage (
<bibRefCitation id="EFD64B50D14B513B0BEA53B534BC2FB5" author="Cumming, J. M. &amp; Brooks, S. E." box="[459,756,1698,1724]" pageId="4" pageNumber="415" pagination="41 - 54" refId="ref13849" refString="Cumming, J. M. &amp; Brooks, S. E. (2002) Electrophorella, a new genus of parathalassiine flies from Baltic amber, with a cladistic analysis of the Microphorinae + Dolichopodidae lineage (Diptera: Empidoidea). Studia dipterologica, 9 (1), 41 - 54." type="journal article" year="2002">Cumming &amp; Brooks 2002</bibRefCitation>
; 2019), which is followed herein.
</paragraph>
<paragraph id="8BF836A1D14B513B0AE653D135B42E44" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
Actually,
<taxonomicName id="4C474D22D14B513B0B1253D1379E2FE9" authorityName="Meunier" authorityYear="1908" box="[307,470,1734,1760]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B0B1253D1379E2FE9" box="[307,470,1734,1760]" italics="true" pageId="4" pageNumber="415">Meghyperiella</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C474D22D14B513B08A053D134B12FE9" box="[641,761,1734,1760]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B08A053D134B12FE9" box="[641,761,1734,1760]" italics="true" pageId="4" pageNumber="415">Microphor</emphasis>
</taxonomicName>
genus group and provisionally this genus is most closely related to
<taxonomicName id="4C474D22D14B513B0ACE53FD372F2E0D" box="[239,359,1770,1796]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B0ACE53FD372F2E0D" box="[239,359,1770,1796]" italics="true" pageId="4" pageNumber="415">Microphor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C474D22D14B513B0BBD53FD34732E0D" authorityName="Hennig" authorityYear="1971" box="[412,571,1770,1796]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B0BBD53FD34732E0D" box="[412,571,1770,1796]" italics="true" pageId="4" pageNumber="415">Microphorites</emphasis>
</taxonomicName>
primarily because they share the female syntergite 9+10 with acanthophorous setae (
<bibRefCitation id="EFD64B50D14B513B0AFF521934422E21" author="Cumming, J. M. &amp; Brooks, S. E." box="[222,522,1806,1832]" pageId="4" pageNumber="415" pagination="41 - 54" refId="ref13849" refString="Cumming, J. M. &amp; Brooks, S. E. (2002) Electrophorella, a new genus of parathalassiine flies from Baltic amber, with a cladistic analysis of the Microphorinae + Dolichopodidae lineage (Diptera: Empidoidea). Studia dipterologica, 9 (1), 41 - 54." type="journal article" year="2002">Cumming &amp; Brooks 2002</bibRefCitation>
). A unique feature of
<taxonomicName id="4C474D22D14B513B08DD5219322A2E21" authorityName=": Keilbach" authorityYear="1982" box="[764,1122,1806,1832]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14B513B08DD5219322A2E21" box="[764,1122,1806,1832]" italics="true" pageId="4" pageNumber="415">Meghyperiella porphyropsoides</emphasis>
</taxonomicName>
among all microphorines is the smoothly undulating vein closing cell dm (base of M
<subScript id="17C334E4D14B513B0930525235512E5A" attach="left" box="[785,793,1861,1875]" fontSize="6" pageId="4" pageNumber="415">2</subScript>
+ dm-m crossvein).
</paragraph>
<paragraph id="8BF836A1D14B513B0AE65240351B2EB1" blockId="4.[151,1437,150,1977]" pageId="4" pageNumber="415">
It is interesting to note that
<bibRefCitation id="EFD64B50D14B513B0BDC524134872E79" author="Collin, J. E." box="[509,719,1878,1904]" pageId="4" pageNumber="415" refId="ref13821" refString="Collin, J. E. (1961) Empididae. In: British Flies. Vol. 6. University Press, Cambridge, 782 pp." type="book" year="1961">Collin (1961: 320)</bibRefCitation>
mentioned female specimens of
<taxonomicName id="4C474D22D14B513B0E63524133D42E78" authority="Macquart" authorityName="Macquart" authorityYear="1827" box="[1090,1436,1878,1905]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="B933EAB3D14B513B0E63524133612E79" box="[1090,1321,1878,1904]" italics="true" pageId="4" pageNumber="415">Microphor crassipes</emphasis>
Macquart
</taxonomicName>
with “very much abbreviated” apical section of M
<subScript id="17C334E4D14B513B08E5529A34842E92" attach="left" box="[708,716,1933,1947]" fontSize="6" pageId="4" pageNumber="415">2</subScript>
. In addition, we have seen a female of an undescribed species of probably
<taxonomicName id="4C474D22D14B513B0B23528937322EB1" box="[258,378,1950,1976]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAB3D14B513B0B23528937322EB1" box="[258,378,1950,1976]" italics="true" pageId="4" pageNumber="415">Microphor</emphasis>
</taxonomicName>
from
<collectingRegion id="4983F843D14B513B0B9F528834412EB0" box="[446,521,1951,1977]" country="Ukraine" name="Rivne" pageId="4" pageNumber="415">Rovno</collectingRegion>
amber with similar venation.
</paragraph>
<caption id="DF386629D14A513A0AB6527234702EE0" ID-DOI="http://doi.org/10.5281/zenodo.6623130" ID-Zenodo-Dep="6623130" httpUri="https://zenodo.org/record/6623130/files/figure.png" pageId="5" pageNumber="416" startId="5.[151,264,1893,1917]" targetBox="[227,1360,180,1865]" targetPageId="5">
<paragraph id="8BF836A1D14A513A0AB6527234702EE0" blockId="5.[151,1437,1893,2032]" pageId="5" pageNumber="416">
<emphasis id="B933EAB3D14A513A0AB6527237092E74" bold="true" box="[151,321,1893,1917]" pageId="5" pageNumber="416">FIGURES 1, 2.</emphasis>
<taxonomicName id="4C474D22D14A513A0B6B527234BB2E74" authority="Meunier" authorityName="Meunier" authorityYear="1908" box="[330,755,1893,1917]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="416" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14A513A0B6B527234D92E74" box="[330,657,1893,1917]" italics="true" pageId="5" pageNumber="416">Meghyperiella porphyropsoides</emphasis>
Meunier
</taxonomicName>
, male, Rovno amber.
<emphasis id="B933EAB3D14A513A09FA527235A72E74" bold="true" box="[987,1007,1893,1917]" pageId="5" pageNumber="416">1.</emphasis>
Habitus, right lateral view;
<emphasis id="B933EAB3D14A513A0F3A527233662E74" bold="true" box="[1307,1326,1893,1917]" pageId="5" pageNumber="416">2.</emphasis>
Wing and terminalia. Abbreviations: CuA+CuP —anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; epand—epandrium; hypd lb—apical lobe of hypandrium; M
<subScript id="17C334E4D14A513A0EB652AA32D62EC2" attach="left" box="[1175,1182,1981,1995]" fontSize="6" pageId="5" pageNumber="416">1</subScript>
, M
<subScript id="17C334E4D14A513A0EE552AA32832EC2" attach="left" box="[1220,1227,1981,1995]" fontSize="6" pageId="5" pageNumber="416">2</subScript>
, M
<subScript id="17C334E4D14A513A0ED052AA32B02EC2" attach="left" box="[1265,1272,1981,1995]" fontSize="6" pageId="5" pageNumber="416">4</subScript>
—medial veins; ph—phallus; R
<subScript id="17C334E4D14A513A0B1052F637702EE6" attach="left" box="[305,312,2017,2031]" fontSize="6" pageId="5" pageNumber="416">1</subScript>
, R
<subScript id="17C334E4D14A513A0B7752F637262EE6" attach="left" box="[342,366,2017,2031]" fontSize="6" pageId="5" pageNumber="416">2+3</subScript>
, R
<subScript id="17C334E4D14A513A0BAA52F537EA2EF9" attach="left" box="[395,418,2018,2032]" fontSize="6" pageId="5" pageNumber="416">4+5</subScript>
—radial veins.
</paragraph>
</caption>
<paragraph id="8BF836A1D14951390AE65580379E2B34" blockId="6.[151,1437,151,573]" pageId="6" pageNumber="417">
It seems that
<taxonomicName id="4C474D22D14951390B405580348F29B8" authorityName="Meunier" authorityYear="1908" box="[353,711,151,177]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D14951390B405580348F29B8" box="[353,711,151,177]" italics="true" pageId="6" pageNumber="417">Meghyperiella porphyropsoides</emphasis>
</taxonomicName>
was not an uncommon species in the Upper Eocene on the territory of the modern Baltic region and
<collectingCountry id="F3507631D1495139087F55AC34F429DC" box="[606,700,187,213]" name="Ukraine" pageId="6" pageNumber="417">Ukraine</collectingCountry>
.
<taxonomicName id="4C474D22D149513908E855AC327A29DC" authorityName="Meunier" authorityYear="1908" box="[713,1074,187,213]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="porphyropsoides">
<emphasis id="B933EAB3D149513908E855AC327A29DC" box="[713,1074,187,213]" italics="true" pageId="6" pageNumber="417">Meghyperiella porphyropsoides</emphasis>
</taxonomicName>
is the first amber brachyceran, reported from Zhitomir Region of
<collectingCountry id="F3507631D1495139080555C8343629F0" box="[548,638,223,249]" name="Ukraine" pageId="6" pageNumber="417">Ukraine</collectingCountry>
(one rhagionid was determined earlier by B.B. Rohdendorf to family level). Fourteen amber species have been reported from this region (
<bibRefCitation id="EFD64B50D149513909B7541432CC2814" author="Legalov, A. &amp; Nazarenko, V. Yu. &amp; Perkovsky, E. E." box="[918,1156,259,285]" pageId="6" pageNumber="417" pagination="95 - 100" refId="ref14854" refString="Legalov, A. A, Nazarenko, V. Yu. &amp; Perkovsky, E. E. (2021 b) A new species of the genus Dorytomus Germar, 1817 (Coleoptera, Curculionidae) from Rovno amber. Zootaxa, 5006 (1), 95 - 100. https: // doi. org / 10.11646 / zootaxa. 5006.1.12" type="journal article" year="2021">
Legalov
<emphasis id="B933EAB3D149513909DA541432642814" box="[1019,1068,259,285]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
. 2021b
</bibRefCitation>
;
<bibRefCitation id="EFD64B50D14951390EAE541433D82814" author="Radchenko, A. G. &amp; Perkovsky, E. E. &amp; Vasilenko, D. V." box="[1167,1424,259,285]" pageId="6" pageNumber="417" pagination="237 - 251" refId="ref15909" refString="Radchenko, A. G., Perkovsky, E. E. &amp; Vasilenko, D. V. (2021) Formica species (Hymenoptera, Formicidae, Formicinae) in late Eocene Rovno amber. Journal of Hymenoptera Research, 82, 237 - 251. https: // doi. org / 10.3897 / jhr. 82.64599" type="journal article" year="2021">
Radchenko
<emphasis id="B933EAB3D14951390F355414330E2814" box="[1300,1350,259,285]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
. 2021
</bibRefCitation>
), including new ant species, a new anthribid (
<bibRefCitation id="EFD64B50D1495139085C5430352E2848" author="Legalov, A. A. &amp; Nazarenko, V. Yu. &amp; Perkovsky, E. E." box="[637,870,295,321]" pageId="6" pageNumber="417" pagination="1 - 7" refId="ref14785" refString="Legalov, A. A., Nazarenko, V. Yu. &amp; Perkovsky, E. E. (2021 a) A new species of the genus Glaesotropis Gratshev &amp; Zherikhin, 1995 (Coleoptera, Anthribidae) from Rovno amber. Fossil Record, 24, 1 - 7. https: // doi. org / 10.5194 / fr- 24 - 1 - 2021" type="journal article" year="2021">
Legalov
<emphasis id="B933EAB3D149513908C0543035592848" box="[737,785,295,321]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
. 2021a
</bibRefCitation>
), a new curculionid (
<bibRefCitation id="EFD64B50D14951390E6F543033632848" author="Legalov, A. A. &amp; Nazarenko, V. Yu. &amp; Vasilenko, D. V. &amp; Perkovsky, E. E." box="[1102,1323,295,321]" pageId="6" pageNumber="417" pagination="379 - 386" refId="ref14919" refString="Legalov, A. A., Nazarenko, V. Yu., Vasilenko, D. V. &amp; Perkovsky, E. E. (2022) Ceutorhynchus Germar (Coleoptera, Curculionidae) as proxy for Eocene core Brassicaceae: first record of the genus from Rovno amber. Journal of Paleontology, 96 (2), 379 - 386. https: // doi. org / 10.1017 / jpa. 2021.82" type="journal article" year="2022">
Legalov
<emphasis id="B933EAB3D14951390E90543032AA2848" box="[1201,1250,295,321]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
. 2022
</bibRefCitation>
), two new genera and four new species of gall midges (
<bibRefCitation id="EFD64B50D149513908A4545C3581286C" author="Fedotova, Z. A. &amp; Perkovsky, E. E." box="[645,969,330,357]" pageId="6" pageNumber="417" pagination="270 - 278" refId="ref14144" refString="Fedotova, Z. A. &amp; Perkovsky, E. E. (2015) New gall midges (Diptera, Cecidomyiidae, Stomatosematidi, Brachineuridi) from the Late Eocene amber of Gulyanka (Zhitomir Region, Ukraine). Paleontological Journal, 49, 270 - 278. https: // doi. org / 10.1134 / S 0031030115030041" type="journal article" year="2015">Fedotova &amp; Perkovsky 2015</bibRefCitation>
,
<bibRefCitation id="EFD64B50D149513909F2545C3247286C" author="Fedotova, Z. A. &amp; Perkovsky, E. E." box="[979,1039,331,357]" pageId="6" pageNumber="417" pagination="23 - 30" refId="ref14206" refString="Fedotova, Z. A. &amp; Perkovsky, E. E. (2017) New genus and species of gall midges (Diptera, Cecidomyiidae, Porricondylinae, Holoneurini) from the Late Eocene amber of Olevsk (Zhitomir Region, Ukraine). Vestnik Zoologii, 51 (1), 23 - 30. https: // doi. org / 10.1515 / vzoo- 2017 - 0004" type="journal article" year="2017">2017</bibRefCitation>
), a new micro-caddisfly (Melnitsky
<emphasis id="B933EAB3D14951390AB6547836802880" box="[151,200,367,393]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
. 2021); an additional new cicadellid genus and species is described from closely connected fauna of Perebrody (
<bibRefCitation id="EFD64B50D14951390ABE5484373728A4" author="Dietrich, C. H. &amp; Dmitriev, D. A. &amp; Perkovsky, E. E." box="[159,383,403,429]" pageId="6" pageNumber="417" pagination="445 - 452" refId="ref14042" refString="Dietrich, C. H., Dmitriev, D. A. &amp; Perkovsky, E. E. (2021) A new leafhopper (Hemiptera: Cicadellidae) from late Eocene Rovno amber described based on the adult and associated last-instar nymph from Perebrody. Palaeoentomology, 4 (5), 445 - 452. https: // doi. org / 10.11646 / palaeoentomology. 4.5.10" type="journal article" year="2021">
Dietrich
<emphasis id="B933EAB3D14951390B255484377D28A4" box="[260,309,403,429]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
. 2021
</bibRefCitation>
). Veselukha river valley (
<bibRefCitation id="EFD64B50D149513908BD5484324D28A4" author="Lyubarsky, G. Yu. &amp; Perkovsky, E. E." box="[668,1029,403,429]" pageId="6" pageNumber="417" pagination="25 - 35" refId="ref15061" refString="Lyubarsky, G. Yu. &amp; Perkovsky, E. E. (2020) First Rovno amber species of the genus Telmatophilus (Coleoptera: Clavicornia: Cryptophagidae) from Veselukha floodplain, Invertebrate Zoology, 17 (1), 25 - 35. https: // doi. org / 10.15298 / invertzool. 17.1.03" type="journal article" year="2020">Lyubarsky and Perkovsky 2020</bibRefCitation>
;
<bibRefCitation id="EFD64B50D14951390E315484335B28A4" author="Melnitsky, S. I. &amp; Ivanov, V. D. &amp; Perkovsky, E. E." box="[1040,1299,403,429]" pageId="6" pageNumber="417" pagination="106 - 109" refId="ref15481" refString="Melnitsky, S. I., Ivanov, V. D. &amp; Perkovsky, E. E. (2021 b) A new species of Plectrocnemia (Trichoptera: Polycentropodidae) from Rovno amber. Zootaxa, 5006 (1), 106 - 109. https: // doi. org / 10.11646 / zootaxa. 5006.1.14" type="journal article" year="2021">
Melnitsky
<emphasis id="B933EAB3D14951390EA85484328A28A4" box="[1161,1218,403,429]" italics="true" pageId="6" pageNumber="417">et al.</emphasis>
2021b
</bibRefCitation>
;
<bibRefCitation id="EFD64B50D14951390F3F5484369828D8" author="Gilka, W. &amp; Harbach, R. E. &amp; Perkovsky, E. E." pageId="6" pageNumber="417" pagination="257 - 270" refId="ref14277" refString="Gilka, W., Harbach, R. E. &amp; Perkovsky, E. E. (2021) Mosquitoes (Diptera: Culicidae) in Eocene amber from the Rovno region, Ukraine. Zootaxa, 5016 (2), 257 - 270. https: // doi. org / 10.11646 / zootaxa. 5016.2.6" type="journal article" year="2021">
Giłka
<emphasis id="B933EAB3D14951390F45548433D528A4" box="[1380,1437,403,429]" italics="true" pageId="6" pageNumber="417">et al.</emphasis>
2021
</bibRefCitation>
) as well as Voronki (
<bibRefCitation id="EFD64B50D14951390B9B54A034EB28D8" author="Simutnik, S. A. &amp; Perkovsky, E. E. &amp; Khomych, M. R. &amp; Vasilenko, D. V." box="[442,675,439,465]" pageId="6" pageNumber="417" pagination="85 - 102" refId="ref16181" refString="Simutnik, S. A., Perkovsky, E. E., Khomych, M. R. &amp; Vasilenko, D. V. (2021) First record of the Sulia glaesaria Simutnik, 2015 (Hymenoptera, Chalcidoidea, Encyrtidae) from Rovno amber. Journal of Hymenoptera Research, 88, 85 - 102. https: // doi. org / 10.3897 / jhr. 88.75941" type="journal article" year="2021">
Simutnik
<emphasis id="B933EAB3D1495139080854A0341228D8" box="[553,602,439,465]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
. 2021
</bibRefCitation>
;
<bibRefCitation id="EFD64B50D1495139088C54A035EE28D8" author="Yamamoto, S. &amp; Nazarenko, V. Y. &amp; Vasilenko, D. V. &amp; Perkovsky, E. E." box="[685,934,439,465]" pageId="6" pageNumber="417" pagination="65 - 74" refId="ref16705" refString="Yamamoto, S., Nazarenko, V. Y., Vasilenko, D. V. &amp; Perkovsky, E. E. (2022) First fossil species of ship-timber beetles (Coleoptera: Lymexylidae) from Eocene Rovno amber (Ukraine). Fossil Record, 25 (1), 65 - 74. https: // doi. org / 10.3897 / fr. 25.81054" type="journal article" year="2022">
Yamamoto
<emphasis id="B933EAB3D1495139090F54A0352E28D8" box="[814,870,439,465]" italics="true" pageId="6" pageNumber="417">et al.</emphasis>
2022
</bibRefCitation>
) and Velyki Telkovichi (
<bibRefCitation id="EFD64B50D14951390E9954A033DD28D8" author="Legalov, A. A. &amp; Nazarenko, V. Yu. &amp; Vasilenko, D. V. &amp; Perkovsky, E. E." box="[1208,1429,439,465]" pageId="6" pageNumber="417" pagination="379 - 386" refId="ref14919" refString="Legalov, A. A., Nazarenko, V. Yu., Vasilenko, D. V. &amp; Perkovsky, E. E. (2022) Ceutorhynchus Germar (Coleoptera, Curculionidae) as proxy for Eocene core Brassicaceae: first record of the genus from Rovno amber. Journal of Paleontology, 96 (2), 379 - 386. https: // doi. org / 10.1017 / jpa. 2021.82" type="journal article" year="2022">
Legalov
<emphasis id="B933EAB3D14951390F3D54A0331C28D8" box="[1308,1364,439,465]" italics="true" pageId="6" pageNumber="417">et al.</emphasis>
2022
</bibRefCitation>
) is one of the most important amber-bearing sites of the Varash district of
<collectingRegion id="4983F843D149513909EC54CC323C28FC" box="[973,1140,475,501]" country="Ukraine" name="Rivne" pageId="6" pageNumber="417">Rovno Region</collectingRegion>
. Two males were cut from a piece obtained from the private collection of M.R. Khomych (
<collectingRegion id="4983F843D1495139094654E835FD2B10" box="[871,949,511,537]" country="Ukraine" name="Rivne" pageId="6" pageNumber="417">Rovno</collectingRegion>
) that comprises four additional males, supposedly of the same species.
</paragraph>
</subSubSection>
</treatment>
</document>
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