320 lines
40 KiB
XML
320 lines
40 KiB
XML
<document id="6C7447D836208F062B4CEB82153595C9" ID-CLB-Dataset="72886" ID-DOI="10.11646/zootaxa.5150.3.6" ID-GBIF-Dataset="c8c16c97-1eae-42d2-9ab3-750dc14ea775" ID-ISSN="1175-5326" ID-Zenodo-Dep="6623125" ID-ZooBank="D59052B5-85CD-466F-B4EB-812226DC913E" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1654676678377" checkinUser="plazi" docAuthor="Shamshev, Igor V. & Perkovsky, Evgeny E." docDate="2022" docId="03EE87B7D14951390AB6579937E02E76" docLanguage="en" docName="zootaxa.5150.3.6.pdf" docOrigin="Zootaxa 5150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microphor Macquart" docType="treatment" docVersion="7" lastPageNumber="417" masterDocId="FFD7FFCFD14F513F0A21551736482909" masterDocTitle="A review of fossil taxa of Microphorinae (Diptera, Dolichopodidae sensu lato), with redescription of the Eocene genus Meghyperiella Meunier" masterLastPageNumber="427" masterPageNumber="411" pageNumber="417" updateTime="1731614897330" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="E62173D99F73A964EAAEF37606273DDB">A review of fossil taxa of Microphorinae (Diptera, Dolichopodidae sensu lato), with redescription of the Eocene genus Meghyperiella Meunier</mods:title>
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<mods:namePart id="3F1696A015FC95C7E2A27354B10008F9">Shamshev, Igor V.</mods:namePart>
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<mods:affiliation id="697B7B3602D1D1E3FC42FA3F489F4156">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:namePart id="172A9C7EFBFCAB045429D76ECB420D98">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="B3D2975A80A49DFB5A7CC323EEA4A969">Schmalhausen Institute of Zoology NAS of Ukraine, vul. B. Khmelnytskogo, 15, Kiev, 01030 Ukraine. & Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117997, Russia</mods:affiliation>
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<mods:title id="E1AF39C492C289ACE3BF899DDFD7C423">Zootaxa</mods:title>
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<mods:date id="4820101980E403DE5CCF2D9E5B70F7EB">2022</mods:date>
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<treatment id="03EE87B7D14951390AB6579937E02E76" ID-GBIF-Taxon="195913585" LSID="urn:lsid:plazi:treatment:03EE87B7D14951390AB6579937E02E76" httpUri="http://treatment.plazi.org/id/03EE87B7D14951390AB6579937E02E76" lastPageNumber="417" pageId="6" pageNumber="417">
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<paragraph id="8BF836A1D14951390AB6579937D62BA0" blockId="6.[151,414,654,681]" box="[151,414,654,681]" pageId="6" pageNumber="417">
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<heading id="D0B081CDD14951390AB6579937D62BA0" bold="true" box="[151,414,654,681]" fontSize="11" level="1" pageId="6" pageNumber="417" reason="1">
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<taxonomicName id="4C474D22D14951390AB6579937D62BA0" ID-CoL="5S7S" ID-ENA="2498632" authority="Macquart" authorityName="Macquart" box="[151,414,654,681]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAB3D14951390AB6579937D62BA0" bold="true" box="[151,414,654,681]" pageId="6" pageNumber="417">
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<emphasis id="B933EAB3D14951390AB6579937512BA0" bold="true" box="[151,281,654,681]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
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Macquart
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D652AD14951390AB657C437002ABD" pageId="6" pageNumber="417" type="reference_group">
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<paragraph id="8BF836A1D14951390AB657C434982A03" blockId="6.[151,1436,723,839]" pageId="6" pageNumber="417">
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<treatmentCitationGroup id="AB57118FD14951390AB657C437B92BE2" box="[151,497,723,747]" pageId="6" pageNumber="417">
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<treatmentCitation id="0AE610B0D14951390AB657C437A52BE2" author="Macquart, J." box="[151,493,723,747]" page="139" pageId="6" pageNumber="417" year="1827">
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<taxonomicName id="4C474D22D14951390AB657C437A52BE2" ID-CoL="5S7S" authority="Macquart, 1827: 139" authorityName="Macquart" authorityPageNumber="139" authorityYear="1827" box="[151,493,723,747]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAB3D14951390AB657C4374C2BE2" box="[151,260,723,747]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
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<bibRefCitation id="EFD64B50D14951390B2F57C337A52BE2" author="Macquart, J." box="[270,493,723,747]" pageId="6" pageNumber="417" refId="ref15120" refString="Macquart, J. (1827) Insectes dipteres du nord de la France. Platypezines, dolichopodes, empides, hybotides. Imprimerie de Leleux, Lille, 159 pp. https: // doi. org / 10.5962 / bhl. title. 148911" type="book" year="1827">Macquart, 1827: 139</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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<typeStatus id="54FC8803D14951390BDB57C334642BE5" box="[506,556,724,748]" pageId="6" pageNumber="417">Type</typeStatus>
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species:
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<taxonomicName id="4C474D22D149513908B357C4325B2BE2" ID-CoL="433C4" authority="Macquart, 1827" authorityName="Macquart" authorityYear="1827" box="[658,1043,723,747]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="velutinus">
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<emphasis id="B933EAB3D149513908B357C4352C2BE2" box="[658,868,723,747]" italics="true" pageId="6" pageNumber="417">Microphor velutinus</emphasis>
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<bibRefCitation id="EFD64B50D1495139094E57C3325B2BE2" author="Macquart, J." box="[879,1043,723,747]" pageId="6" pageNumber="417" refId="ref15120" refString="Macquart, J. (1827) Insectes dipteres du nord de la France. Platypezines, dolichopodes, empides, hybotides. Imprimerie de Leleux, Lille, 159 pp. https: // doi. org / 10.5962 / bhl. title. 148911" type="book" year="1827">Macquart, 1827</bibRefCitation>
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</taxonomicName>
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(=
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<taxonomicName id="4C474D22D14951390E1F57C437452A03" ID-CoL="433BL" authority="(Meigen, 1804))" authorityName="" baseAuthorityName="Meigen" baseAuthorityYear="1804" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="holosericeus">
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<emphasis id="B933EAB3D14951390E1F57C4337E2BE2" box="[1086,1334,723,747]" italics="true" pageId="6" pageNumber="417">Microphor holosericeus</emphasis>
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(Meigen, 1804))
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</taxonomicName>
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, by subsequent designation (
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<bibRefCitation id="EFD64B50D1495139081157E5348D2A03" author="Rondani, C." box="[560,709,754,778]" pageId="6" pageNumber="417" pagination="2" refId="ref15970" refString="Rondani, C. (1856) Dipterologiae Italicae prodromus. Vol. 1. Genera Italica ordinis Dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. A. Storschi, Parmae [Parma], 228 + [2] pp. https: // doi. org / 10.5962 / bhl. title. 8160" type="journal article" year="1856">Rondani 1856</bibRefCitation>
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).
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||
</paragraph>
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<paragraph id="8BF836A1D14951390AB6560734BF2A21" blockId="6.[151,1436,723,839]" box="[151,759,784,809]" pageId="6" pageNumber="417">
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<treatmentCitationGroup id="AB57118FD14951390AB6560734BF2A21" box="[151,759,784,809]" pageId="6" pageNumber="417">
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<treatmentCitation id="0AE610B0D14951390AB6560737B02A21" author="Macquart, J." box="[151,504,784,809]" page="345" pageId="6" pageNumber="417" year="1834">
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<taxonomicName id="4C474D22D14951390AB6560737B02A21" ID-CoL="BYSS6" authority="Macquart, 1834: 345" authorityName="Macquart" authorityPageNumber="345" authorityYear="1834" box="[151,504,784,809]" class="Insecta" family="Dolichopodidae" genus="Microphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B933EAB3D14951390AB6560737522A21" box="[151,282,784,808]" italics="true" pageId="6" pageNumber="417">Microphorus</emphasis>
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<bibRefCitation id="EFD64B50D14951390B00560637B02A21" author="Macquart, J." box="[289,504,784,809]" pageId="6" pageNumber="417" pagination="1 - 578" refId="ref15166" refString="Macquart, J. (1834) Histoire naturelle des Insectes. Dipteres. Tome premier. In: Roret, N. E. (Ed.), Collection des suites a Buffon, Paris, 1, pp. 1 - 578, 12 pls. https: // doi. org / 10.5962 / bhl. title. 14274" type="journal article" year="1834">Macquart, 1834: 345</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
. Unjustified emendation.
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BF836A1D14951390AB65638331C2A4E" blockId="6.[151,1436,723,839]" box="[151,1364,815,839]" pageId="6" pageNumber="417">
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<treatmentCitationGroup id="AB57118FD14951390AB6563837B52A4E" box="[151,509,815,839]" pageId="6" pageNumber="417">
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<treatmentCitation id="0AE610B0D14951390AB6563837B12A4E" author="Zetterstedt, J. W." box="[151,505,815,839]" page="253" pageId="6" pageNumber="417" year="1842">
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<taxonomicName id="4C474D22D14951390AB6563837B12A4E" ID-CoL="5S7T" authority="Zetterstedt, 1842: 253" authorityName="Zetterstedt" authorityPageNumber="253" authorityYear="1842" box="[151,505,815,839]" class="Insecta" family="Athericidae" genus="Microphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAB3D14951390AB6563837582A4E" box="[151,272,815,839]" italics="true" pageId="6" pageNumber="417">Microphora</emphasis>
|
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<bibRefCitation id="EFD64B50D14951390B36563837B12A4E" author="Zetterstedt, J. W." box="[279,505,815,839]" pageId="6" pageNumber="417" refId="ref16776" refString="Zetterstedt, J. W. (1842) Diptera scandinaviae disposa et descripta. Vol. 1. ex Officina Lundbergiana, sumtibus auctoris, Lundae [Lund], 140 pp." type="book" year="1842">Zetterstedt, 1842: 253</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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||
Unjustified emendation or error, not
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<treatmentCitation id="0AE610B0D14951390956563832F82A4E" author="Krober, O." box="[887,1200,815,839]" page="245" pageId="6" pageNumber="417" year="1912">
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<taxonomicName id="4C474D22D14951390956563832F82A4E" ID-CoL="5S7T" authority="Krober, 1912: 245" authorityName="Krober" authorityPageNumber="245" authorityYear="1912" box="[887,1200,815,839]" class="Insecta" family="Athericidae" genus="Microphora" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAB3D14951390956563835B82A4E" box="[887,1008,815,839]" italics="true" pageId="6" pageNumber="417">Microphora</emphasis>
|
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<bibRefCitation id="EFD64B50D149513909D7563832F82A4E" author="Krober, O." box="[1014,1200,815,839]" pageId="6" pageNumber="417" pagination="242 - 256" refId="ref14649" refString="Krober, O. (1912) Die Thereviden der indo-australischen region (Dipt.). Entomologische Mitteilungen, 1, 242 - 256." type="journal article" year="1912">Kröber, 1912: 245</bibRefCitation>
|
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</taxonomicName>
|
||
</treatmentCitation>
|
||
(=
|
||
<taxonomicName id="4C474D22D14951390EF2563833022A4E" ID-CoL="H6L" box="[1235,1354,815,839]" class="Insecta" family="Therevidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="family">Therevidae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14951390AB6566237002ABD" blockId="6.[151,1437,885,1919]" pageId="6" pageNumber="417">
|
||
<emphasis id="B933EAB3D14951390AB6566237ED2A86" bold="true" box="[151,421,885,911]" pageId="6" pageNumber="417">Included fossil species.</emphasis>
|
||
<taxonomicName id="4C474D22D14951390B8D5662350E2A86" ID-CoL="8CVGL" authority="(Meunier, 1908)" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[428,838,885,911]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="rusticus">
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||
<emphasis id="B933EAB3D14951390B8D566234CC2A86" box="[428,644,885,911]" italics="true" pageId="6" pageNumber="417">Microphor rusticus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD64B50D149513908B3566235752A86" author="Meunier, F." box="[658,829,885,911]" pageId="6" pageNumber="417" pagination="81 - 135" refId="ref15542" refString="Meunier, F. (1908) Monographie des Empidae de l'ambre de la Baltique et catalogue bibliographique complet sur les Dipteres fossiles de cette resine. Annales des Sciences Naturelles, Zoologie, Series 9, 7, 81 - 135." type="journal article" year="1908">Meunier, 1908</bibRefCitation>
|
||
)
|
||
</taxonomicName>
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||
: 105 (
|
||
<taxonomicName id="4C474D22D149513909AB5662326B2A86" ID-CoL="6PMV" authorityName="Zetterstedt" authorityYear="1837" box="[906,1059,885,911]" class="Insecta" family="Empididae" genus="Phyllodromia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B933EAB3D149513909AB5662326B2A86" box="[906,1059,885,911]" italics="true" pageId="6" pageNumber="417">Phyllodromia</emphasis>
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||
</taxonomicName>
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||
, male and female) (Baltic region, Upper Eocene).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D652AD14951390AE656AA37E02E76" pageId="6" pageNumber="417" type="discussion">
|
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<paragraph id="8BF836A1D14951390AE656AA32592D16" blockId="6.[151,1437,885,1919]" pageId="6" pageNumber="417">
|
||
<emphasis id="B933EAB3D14951390AE656AA37732ADE" bold="true" box="[199,315,957,983]" pageId="6" pageNumber="417">Remarks.</emphasis>
|
||
<taxonomicName id="4C474D22D14951390B6356AA37F22ADE" box="[322,442,957,983]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B933EAB3D14951390B6356AA37F22ADE" box="[322,442,957,983]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
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remains a broadly defined group with not quite clear phylogenetic background. The very recently published revision of the North American species provides the first provisional evidence of monophyly of this genus, especially with respect to
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<taxonomicName id="4C474D22D14951390818511232452D16" authority="(Brooks & Cumming 2022)" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[569,1037,1029,1055]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="417" phylum="Ascomycota" rank="genus">
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||
<emphasis id="B933EAB3D14951390818511234852D16" box="[569,717,1029,1055]" italics="true" pageId="6" pageNumber="417">Schistostoma</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD64B50D149513908FA5112324E2D16" author="Brooks, S. E. & Cumming, J. M." box="[731,1030,1029,1055]" pageId="6" pageNumber="417" pagination="1 - 141" refId="ref13522" refString="Brooks, S. E. & Cumming, J. M. (2022) Revision of the Nearctic Microphorinae (Diptera: Dolichopodidae sensu lato). Zootaxa, 5095 (1), 1 - 141. https: // doi. org / 10.11646 / zootaxa. 5095.1.1" type="journal article" year="2022">Brooks & Cumming 2022</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14951390AE6513E32D82D82" blockId="6.[151,1437,885,1919]" pageId="6" pageNumber="417">
|
||
Currently,
|
||
<taxonomicName id="4C474D22D14951390B60513E37F12D4A" box="[321,441,1065,1091]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390B60513E37F12D4A" box="[321,441,1065,1091]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
|
||
comprises 16 known extant and one extinct species. The recent species are distributed mostly in the Holarctic, one species was described from mountains of South
|
||
<collectingCountry id="F3507631D14951390E2E515A321A2D6E" box="[1039,1106,1101,1127]" name="China" pageId="6" pageNumber="417">China</collectingCountry>
|
||
and one unnamed species is known from South
|
||
<collectingCountry id="F3507631D14951390B51516637E52D82" box="[368,429,1137,1163]" name="Chile" pageId="6" pageNumber="417">Chile</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFD64B50D14951390B9A516634072D82" author="Chvala, M." box="[443,591,1137,1163]" pageId="6" pageNumber="417" pagination="19 - 26" refId="ref13737" refString="Chvala, M. (1991) A new species of the Holarctic genus Schistostoma Becker (Diptera: Microphoridae) from South Africa, and first record of the family from South America. Annals of the Natal Museum, 32, 19 - 26." type="journal article" year="1991">Chvála 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD64B50D1495139087B516635182D82" author="Saigusa, T. & Yang, D." box="[602,848,1137,1164]" pageId="6" pageNumber="417" pagination="519 - 543" refId="ref16030" refString="Saigusa, T. & Yang, D. (2003) Empididae (Diptera) from Funiu Mountains, Henan, China (I). Studia dipterologica, 9 (2), 519 - 543. [2002]" type="journal article" year="2003">Saigusa & Yang 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD64B50D1495139097D516632CC2D82" author="Brooks, S. E. & Cumming, J. M." box="[860,1156,1137,1163]" pageId="6" pageNumber="417" pagination="1 - 141" refId="ref13522" refString="Brooks, S. E. & Cumming, J. M. (2022) Revision of the Nearctic Microphorinae (Diptera: Dolichopodidae sensu lato). Zootaxa, 5095 (1), 1 - 141. https: // doi. org / 10.11646 / zootaxa. 5095.1.1" type="journal article" year="2022">Brooks & Cumming 2022</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14951390AE6518237E02E76" blockId="6.[151,1437,885,1919]" pageId="6" pageNumber="417">
|
||
The single fossil species of
|
||
<taxonomicName id="4C474D22D14951390820518234312DA6" box="[513,633,1173,1199]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390820518234312DA6" box="[513,633,1173,1199]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EFD64B50D1495139096A518235B42DA6" author="Meunier, F." box="[843,1020,1173,1199]" pageId="6" pageNumber="417" pagination="81 - 135" refId="ref15542" refString="Meunier, F. (1908) Monographie des Empidae de l'ambre de la Baltique et catalogue bibliographique complet sur les Dipteres fossiles de cette resine. Annales des Sciences Naturelles, Zoologie, Series 9, 7, 81 - 135." type="journal article" year="1908">Meunier (1908)</bibRefCitation>
|
||
from Baltic amber recognized in the genus
|
||
<taxonomicName id="4C474D22D14951390AC151AE34582DDD" authorityName="Zetterstedt" authorityYear="1837" box="[224,528,1209,1236]" class="Insecta" family="Empididae" genus="Phyllodromia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390AC151AE37312DDA" box="[224,377,1209,1235]" italics="true" pageId="6" pageNumber="417">Phyllodromia</emphasis>
|
||
(Empididae)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFD64B50D1495139083B51AE34F72DDA" author="Hennig, W." box="[538,703,1209,1235]" pageId="6" pageNumber="417" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971)</bibRefCitation>
|
||
examined the type series of
|
||
<taxonomicName id="4C474D22D14951390E2051AD32C82DDA" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[1025,1152,1210,1236]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="rusticus">
|
||
<emphasis id="B933EAB3D14951390E2051AD32502DDA" box="[1025,1048,1210,1235]" italics="true" pageId="6" pageNumber="417">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14951390E0651AD32C82DDA" box="[1063,1152,1210,1235]" italics="true" pageId="6" pageNumber="417">rusticus</emphasis>
|
||
</taxonomicName>
|
||
and indicated the correct systematic position of this species (also designated the
|
||
<typeStatus id="54FC8803D1495139092351CA35262DF1" box="[770,878,1245,1272]" pageId="6" pageNumber="417" type="lectotype">lectotype</typeStatus>
|
||
). However, he did not provide a full redescription of
|
||
<taxonomicName id="4C474D22D14951390A975015377C2C12" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[182,308,1282,1308]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="rusticus">
|
||
<emphasis id="B933EAB3D14951390A97501536852C12" box="[182,205,1282,1307]" italics="true" pageId="6" pageNumber="417">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14951390AFA5015377C2C12" box="[219,308,1282,1307]" italics="true" pageId="6" pageNumber="417">rusticus</emphasis>
|
||
</taxonomicName>
|
||
but included figures of the female habitus as well as the male head, wing and postabdomen (
|
||
<bibRefCitation id="EFD64B50D14951390F695016369B2C36" author="Hennig, W." pageId="6" pageNumber="417" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig 1971</bibRefCitation>
|
||
: figs 26–29). It should be noted that
|
||
<taxonomicName id="4C474D22D1495139085B503134B12C36" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[634,761,1318,1344]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="rusticus">
|
||
<emphasis id="B933EAB3D1495139085B503134D92C36" box="[634,657,1318,1343]" italics="true" pageId="6" pageNumber="417">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14951390881503134B12C36" box="[672,761,1318,1343]" italics="true" pageId="6" pageNumber="417">rusticus</emphasis>
|
||
</taxonomicName>
|
||
may actually be a species of
|
||
<taxonomicName id="4C474D22D14951390E6D503232A82C36" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[1100,1248,1317,1343]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="417" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14951390E6D503232A82C36" box="[1100,1248,1317,1343]" italics="true" pageId="6" pageNumber="417">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
because
|
||
<bibRefCitation id="EFD64B50D14951390F6B503236AA2C6A" author="Hennig, W." pageId="6" pageNumber="417" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971)</bibRefCitation>
|
||
did not separate
|
||
<taxonomicName id="4C474D22D14951390B83505E34522C6A" box="[418,538,1353,1379]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390B83505E34522C6A" box="[418,538,1353,1379]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14951390873505E34AE2C6A" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[594,742,1353,1379]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="417" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14951390873505E34AE2C6A" box="[594,742,1353,1379]" italics="true" pageId="6" pageNumber="417">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
. As recently re-defined,
|
||
<taxonomicName id="4C474D22D149513909DA505E32C72C6A" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[1019,1167,1353,1379]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="417" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D149513909DA505E32C72C6A" box="[1019,1167,1353,1379]" italics="true" pageId="6" pageNumber="417">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C474D22D14951390F05505E33D42C6A" box="[1316,1436,1353,1379]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390F05505E33D42C6A" box="[1316,1436,1353,1379]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
|
||
primarily by medial hypandrial prolongation of the male genitalia (
|
||
<bibRefCitation id="EFD64B50D149513909A6507A32142C8E" author="Brooks, S. E. & Cumming, J. M. & Grimaldi, D. A." box="[903,1116,1389,1416]" pageId="6" pageNumber="417" pagination="121 - 131" refId="ref13574" refString="Brooks, S. E., Cumming, J. M. & Grimaldi, D. A. (2019) Remarkable new fossil species of Schistostoma Becker (Diptera: Dolichopodidae: Microphorinae) from mid-Cretaceous Burmese amber. Zootaxa, 4624 (1), 121 - 131. https: // doi. org / 10.11646 / zootaxa. 4624.1.8" type="journal article" year="2019">
|
||
Brooks
|
||
<emphasis id="B933EAB3D149513909C05079325B2C8E" box="[993,1043,1389,1415]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD64B50D14951390E49507A33D92C8E" author="Brooks, S. E. & Cumming, J. M." box="[1128,1425,1389,1415]" pageId="6" pageNumber="417" pagination="1 - 141" refId="ref13522" refString="Brooks, S. E. & Cumming, J. M. (2022) Revision of the Nearctic Microphorinae (Diptera: Dolichopodidae sensu lato). Zootaxa, 5095 (1), 1 - 141. https: // doi. org / 10.11646 / zootaxa. 5095.1.1" type="journal article" year="2022">Brooks & Cumming 2022</bibRefCitation>
|
||
). However, the structure of the hypandrium is invisible on Hennig’s figure 28 (illustrated from the left side and obscured by pregenital segments). In addition, the female of
|
||
<taxonomicName id="4C474D22D1495139090750A135ED2CC6" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[806,933,1462,1488]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="rusticus">
|
||
<emphasis id="B933EAB3D1495139090750A135752CC6" box="[806,829,1462,1487]" italics="true" pageId="6" pageNumber="417">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1495139096D50A135ED2CC6" box="[844,933,1462,1487]" italics="true" pageId="6" pageNumber="417">rusticus</emphasis>
|
||
</taxonomicName>
|
||
apparently has at least six visible abdominal segments (
|
||
<bibRefCitation id="EFD64B50D14951390B2F50CE37EE2CFA" author="Hennig, W." box="[270,422,1497,1523]" pageId="6" pageNumber="417" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig 1971</bibRefCitation>
|
||
: fig. 26) that matches with another possible apomorphic feature of
|
||
<taxonomicName id="4C474D22D14951390EBB50CE33662CFA" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[1178,1326,1497,1523]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="417" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14951390EBB50CE33662CFA" box="[1178,1326,1497,1523]" italics="true" pageId="6" pageNumber="417">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B933EAB3D14951390F1E50CD33CF2CFA" box="[1343,1415,1498,1523]" italics="true" pageId="6" pageNumber="417">versus</emphasis>
|
||
5 exposed segments in
|
||
<taxonomicName id="4C474D22D14951390BAB50EA344A2F1E" box="[394,514,1533,1559]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390BAB50EA344A2F1E" box="[394,514,1533,1559]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFD64B50D1495139083A50EA35022F1E" author="Brooks, S. E. & Cumming, J. M." box="[539,842,1533,1559]" pageId="6" pageNumber="417" pagination="1 - 141" refId="ref13522" refString="Brooks, S. E. & Cumming, J. M. (2022) Revision of the Nearctic Microphorinae (Diptera: Dolichopodidae sensu lato). Zootaxa, 5095 (1), 1 - 141. https: // doi. org / 10.11646 / zootaxa. 5095.1.1" type="journal article" year="2022">Brooks & Cumming 2022</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C474D22D1495139097C50EA327F2F1E" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[861,1079,1533,1559]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="rusticus">
|
||
<emphasis id="B933EAB3D1495139097C50EA327F2F1E" box="[861,1079,1533,1559]" italics="true" pageId="6" pageNumber="417">Microphor rusticus</emphasis>
|
||
</taxonomicName>
|
||
has two pairs of scutellar setae (
|
||
<bibRefCitation id="EFD64B50D14951390ABE5336377D2F32" author="Hennig, W." box="[159,309,1569,1595]" pageId="6" pageNumber="417" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig 1971</bibRefCitation>
|
||
: fig. 26) that is usually present in
|
||
<taxonomicName id="4C474D22D14951390889533635742F32" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[680,828,1569,1595]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="417" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14951390889533635742F32" box="[680,828,1569,1595]" italics="true" pageId="6" pageNumber="417">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B933EAB3D1495139096A533535DB2F32" box="[843,915,1570,1595]" italics="true" pageId="6" pageNumber="417">versus</emphasis>
|
||
3–4 pairs in
|
||
<taxonomicName id="4C474D22D14951390E00533632D12F32" box="[1057,1177,1569,1595]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390E00533632D12F32" box="[1057,1177,1569,1595]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
|
||
), however, this character is somewhat variable (
|
||
<bibRefCitation id="EFD64B50D14951390B9C535234BC2F56" author="Shamshev, I. V. & Sinclair, B. J." box="[445,756,1605,1631]" pageId="6" pageNumber="417" pagination="335 - 346" refId="ref16072" refString="Shamshev, I. V. & Sinclair, B. J. (2006) The genus Schistostoma Becker from southern Africa, with an evaluation of its generic status (Diptera: Dolichopodidae s. l.: Microphorinae). African Invertebrates, 47, 335 - 346." type="journal article" year="2006">Shamshev & Sinclair 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD64B50D14951390921535232662F56" author="Brooks, S. E. & Cumming, J. M." box="[768,1070,1605,1631]" pageId="6" pageNumber="417" pagination="1 - 141" refId="ref13522" refString="Brooks, S. E. & Cumming, J. M. (2022) Revision of the Nearctic Microphorinae (Diptera: Dolichopodidae sensu lato). Zootaxa, 5095 (1), 1 - 141. https: // doi. org / 10.11646 / zootaxa. 5095.1.1" type="journal article" year="2022">Brooks & Cumming 2022</bibRefCitation>
|
||
). It seems that
|
||
<bibRefCitation id="EFD64B50D14951390EFA535233DF2F69" author="Hennig, W." box="[1243,1431,1605,1632]" pageId="6" pageNumber="417" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971: 5</bibRefCitation>
|
||
, fig. 4) did not illustrate correctly the stylus segmentation in
|
||
<taxonomicName id="4C474D22D14951390963537D35892F8A" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[834,961,1642,1668]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="rusticus">
|
||
<emphasis id="B933EAB3D14951390963537D35112F8A" box="[834,857,1642,1667]" italics="true" pageId="6" pageNumber="417">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14951390949537D35892F8A" box="[872,961,1642,1667]" italics="true" pageId="6" pageNumber="417">rusticus</emphasis>
|
||
</taxonomicName>
|
||
. Actually, the basal article of the stylus is much shorter and hardly distinguishable, contrary to
|
||
<bibRefCitation id="EFD64B50D149513908C2539A35E52FAE" author="Ulrich, H." box="[739,941,1677,1703]" pageId="6" pageNumber="417" pagination="187 - 216" refId="ref16641" refString="Ulrich, H. (1991) Two new genera of parathalassiine-like flies from South Africa (Diptera: Empidoidea). Bonner zoologische Beitrage, 42, 187 - 216." type="journal article" year="1991">Ulrich (1991: 212</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD64B50D14951390999539A32612FA1" author="Ulrich, H." box="[952,1065,1677,1704]" pageId="6" pageNumber="417" pagination="233 - 243" refId="ref16673" refString="Ulrich, H. (2004) Phylogenetic considerations about an early colonization of the sea coasts by Dolichopodidae (Diptera). Studia dipterologica, 11, 233 - 243." type="journal article" year="2004">2004: 235</bibRefCitation>
|
||
) who considered the basal article to be absent in this species.
|
||
<taxonomicName id="4C474D22D14951390BF753A634E72FC2" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[470,687,1713,1739]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="rusticus">
|
||
<emphasis id="B933EAB3D14951390BF753A634E72FC2" box="[470,687,1713,1739]" italics="true" pageId="6" pageNumber="417">Microphor rusticus</emphasis>
|
||
</taxonomicName>
|
||
is probably present in
|
||
<collectingRegion id="4983F843D1495139099553A535B72FC5" box="[948,1023,1714,1740]" country="Ukraine" name="Rivne" pageId="6" pageNumber="417">Rovno</collectingRegion>
|
||
amber. We have examined two very well-preserved inclusions that are apparently a mated male and female (Shamshev & Perkovsky, unpub. data). In addition, an undescribed species with an incomplete vein M
|
||
<subScript id="17C334E4D14951390939521B35682E13" attach="left" box="[792,800,1804,1818]" fontSize="6" pageId="6" pageNumber="417">2</subScript>
|
||
found in
|
||
<collectingRegion id="4983F843D149513909AB53ED359D2E1D" box="[906,981,1786,1812]" country="Ukraine" name="Rivne" pageId="6" pageNumber="417">Rovno</collectingRegion>
|
||
amber, which has been mentioned under
|
||
<taxonomicName id="4C474D22D14951390AB6520A37722E3E" authorityName="Meunier" authorityYear="1908" box="[151,314,1821,1847]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390AB6520A37722E3E" box="[151,314,1821,1847]" italics="true" pageId="6" pageNumber="417">Meghyperiella</emphasis>
|
||
</taxonomicName>
|
||
, probably is another representative of
|
||
<taxonomicName id="4C474D22D149513908C6520A35172E3E" box="[743,863,1821,1847]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D149513908C6520A35172E3E" box="[743,863,1821,1847]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
|
||
. Although
|
||
<taxonomicName id="4C474D22D149513909FD520A321C2E3E" box="[988,1108,1821,1847]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D149513909FD520A321C2E3E" box="[988,1108,1821,1847]" italics="true" pageId="6" pageNumber="417">Microphor</emphasis>
|
||
</taxonomicName>
|
||
is known from the Eocene, it must exist at least in the mid-Cretaceous because both its closest allies (i.e.,
|
||
<taxonomicName id="4C474D22D149513909CD525632C82E52" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[1004,1152,1857,1883]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="6" pageNumber="417" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D149513909CD525632C82E52" box="[1004,1152,1857,1883]" italics="true" pageId="6" pageNumber="417">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14951390E975256331D2E52" authorityName="Hennig" authorityYear="1971" box="[1206,1365,1857,1883]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14951390E975256331D2E52" box="[1206,1365,1857,1883]" italics="true" pageId="6" pageNumber="417">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
) were recorded from that time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |