897 lines
109 KiB
XML
897 lines
109 KiB
XML
<document id="13AABDB51DBD2CE001AC2E49D12F8835" ID-CLB-Dataset="72886" ID-DOI="10.11646/zootaxa.5150.3.6" ID-GBIF-Dataset="c8c16c97-1eae-42d2-9ab3-750dc14ea775" ID-ISSN="1175-5326" ID-Zenodo-Dep="6623125" ID-ZooBank="D59052B5-85CD-466F-B4EB-812226DC913E" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1654676678377" checkinUser="plazi" docAuthor="Shamshev, Igor V. & Perkovsky, Evgeny E." docDate="2022" docId="03EE87B7D14851360AB6577732E82AC0" docLanguage="en" docName="zootaxa.5150.3.6.pdf" docOrigin="Zootaxa 5150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microphorites Hennig 1971" docType="treatment" docVersion="8" lastPageNumber="420" masterDocId="FFD7FFCFD14F513F0A21551736482909" masterDocTitle="A review of fossil taxa of Microphorinae (Diptera, Dolichopodidae sensu lato), with redescription of the Eocene genus Meghyperiella Meunier" masterLastPageNumber="427" masterPageNumber="411" pageNumber="418" updateTime="1731614897330" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="18CD90FB553BCF7C513C30B79979C728">A review of fossil taxa of Microphorinae (Diptera, Dolichopodidae sensu lato), with redescription of the Eocene genus Meghyperiella Meunier</mods:title>
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<mods:namePart id="1B0BE98F575C0C07D9E0D4FE8B99BDFD">Shamshev, Igor V.</mods:namePart>
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<mods:affiliation id="F3B0D9BDE7E81AF881B3B4603AF133CB">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:namePart id="35676D1CBC235589151FD53E1AC89188">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="994F0DECC7B45D7BDAD4DDE1FAD6AB56">Schmalhausen Institute of Zoology NAS of Ukraine, vul. B. Khmelnytskogo, 15, Kiev, 01030 Ukraine. & Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117997, Russia</mods:affiliation>
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<mods:title id="84CC49AD06CFC68943123CB3A7FA5C7F">Zootaxa</mods:title>
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<mods:part id="B5665245FC3171830D8E7306E001470C">
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<mods:date id="F241D8F7DE5E6655BEAADE5F87607DC4">2022</mods:date>
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<mods:number id="F18C277D56091B4707B207EAC3099AD7">2022-06-08</mods:number>
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<treatment id="03EE87B7D14851360AB6577732E82AC0" ID-DOI="http://doi.org/10.5281/zenodo.6646000" ID-GBIF-Taxon="195913586" ID-Zenodo-Dep="6646000" LSID="urn:lsid:plazi:treatment:03EE87B7D14851360AB6577732E82AC0" httpUri="http://treatment.plazi.org/id/03EE87B7D14851360AB6577732E82AC0" lastPageId="9" lastPageNumber="420" pageId="7" pageNumber="418">
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<subSubSection id="C35D652AD14851380AB6577737ED2B73" box="[151,421,608,635]" pageId="7" pageNumber="418" type="nomenclature">
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<paragraph id="8BF836A1D14851380AB6577737ED2B73" blockId="7.[151,421,608,635]" box="[151,421,608,635]" pageId="7" pageNumber="418">
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<heading id="D0B081CDD14851380AB6577737ED2B73" bold="true" box="[151,421,608,635]" fontSize="11" level="1" pageId="7" pageNumber="418" reason="1">
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<taxonomicName id="4C474D22D14851380AB6577737ED2B73" ID-CoL="89QQL" authority="Hennig" authorityName="Hennig" authorityYear="1971" box="[151,421,608,635]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
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<emphasis id="B933EAB3D14851380AB6577737ED2B73" bold="true" box="[151,421,608,635]" pageId="7" pageNumber="418">
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<emphasis id="B933EAB3D14851380AB65777370A2B72" bold="true" box="[151,322,608,635]" italics="true" pageId="7" pageNumber="418">Microphorites</emphasis>
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Hennig
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||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C35D652AD14851380AB657B2342A2D2F" pageId="7" pageNumber="418" type="reference_group">
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<paragraph id="8BF836A1D14851380AB657B232DB2BB4" blockId="7.[151,1171,677,701]" box="[151,1171,677,701]" pageId="7" pageNumber="418">
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<treatmentCitationGroup id="AB57118FD14851380AB657B237AD2BB4" box="[151,485,677,701]" pageId="7" pageNumber="418">
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<treatmentCitation id="0AE610B0D14851380AB657B237A92BB4" author="Hennig, W." box="[151,481,677,701]" page="16" pageId="7" pageNumber="418" year="1971">
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<taxonomicName id="4C474D22D14851380AB657B237A92BB4" ID-CoL="89QQL" authority="Hennig, 1971: 16" authorityName="Hennig" authorityPageNumber="16" authorityYear="1971" box="[151,481,677,701]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14851380AB657B2376F2BB4" box="[151,295,677,701]" italics="true" pageId="7" pageNumber="418">Microphorites</emphasis>
|
||
<bibRefCitation id="EFD64B50D14851380B0F57B237A92BB4" author="Hennig, W." box="[302,481,677,701]" pageId="7" pageNumber="418" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig, 1971: 16</bibRefCitation>
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||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
<typeStatus id="54FC8803D14851380BCB57B234542BB4" box="[490,540,677,701]" pageId="7" pageNumber="418">Type</typeStatus>
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||
species:
|
||
<taxonomicName id="4C474D22D1485138085857B235B42BB4" ID-CoL="8CVGQ" authority="Hennig, 1971" authorityName="Hennig" authorityYear="1971" box="[633,1020,677,701]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="extinctus">
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||
<emphasis id="B933EAB3D1485138085857B235222BB4" box="[633,874,677,701]" italics="true" pageId="7" pageNumber="418">Microphorites extinctus</emphasis>
|
||
<bibRefCitation id="EFD64B50D1485138095157B235B42BB4" author="Hennig, W." box="[880,1020,677,701]" pageId="7" pageNumber="418" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
, by monotypy.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14851380AB657FC342A2D2F" blockId="7.[151,1437,747,2034]" pageId="7" pageNumber="418">
|
||
<emphasis id="B933EAB3D14851380AB657FC37122A0C" bold="true" box="[151,346,747,773]" pageId="7" pageNumber="418">Included species</emphasis>
|
||
(alphabetically).
|
||
<taxonomicName id="4C474D22D1485138083E57FC331D2A0C" ID-CoL="8CVGP" authority="Nel, Perrichot, Daugeron & Neraudeau, 2004: 24" authorityName="Nel, Perrichot, Daugeron & Neraudeau" authorityPageNumber="24" authorityYear="2004" box="[543,1365,747,774]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="deploegi">
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||
<emphasis id="B933EAB3D1485138083E57FC356E2A0C" box="[543,806,747,773]" italics="true" pageId="7" pageNumber="418">Microphorites deploegi</emphasis>
|
||
<bibRefCitation id="EFD64B50D1485138090C57FC331D2A0C" author="Nel, A. & Perrichot, V. & Daugeron, C. & Neraudeau, D." box="[813,1365,747,774]" pageId="7" pageNumber="418" pagination="23 - 29" refId="ref15681" refString="Nel, A., Perrichot, V., Daugeron, C. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France, New Series, 40, 23 - 29. https: // doi. org / 10.1080 / 00379271.2004.10697401" type="journal article" year="2004">Nel, Perrichot, Daugeron & Néraudeau, 2004: 24</bibRefCitation>
|
||
</taxonomicName>
|
||
(Charentese amber, uppermost Albian, Lower Cretaceous,
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||
<collectingCountry id="F3507631D148513808DB560735032A23" box="[762,843,784,810]" name="France" pageId="7" pageNumber="418">France</collectingCountry>
|
||
);
|
||
<taxonomicName id="4C474D22D1485138097E5607335C2A23" ID-CoL="9HM3S" authority="Bramuzzo & Nel, 2017: 590" authorityName="Bramuzzo & Nel" authorityPageNumber="590" authorityYear="2017" box="[863,1300,783,810]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="erikai">
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||
<emphasis id="B933EAB3D1485138097E5607353E2A20" box="[863,886,784,809]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D148513809A75618358F2A20" box="[902,967,783,809]" italics="true" pageId="7" pageNumber="418">erikai</emphasis>
|
||
<bibRefCitation id="EFD64B50D148513809F15607335C2A23" author="Bramuzzo, S. & Nel, A." box="[976,1300,783,810]" pageId="7" pageNumber="418" pagination="590 - 594" refId="ref13466" refString="Bramuzzo, S. & Nel, A. (2017) Youngest representative of the extinct genus Microphorites in the Eocene amber of France (Diptera: Dolichopodidae: Microphorinae). Zootaxa, 4231 (4), 590 - 594. https: // doi. org / 10.11646 / zootaxa. 4231.4.11" type="journal article" year="2017">Bramuzzo & Nel, 2017: 590</bibRefCitation>
|
||
</taxonomicName>
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||
(lowermost Eocene, Oise amber,
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||
<collectingCountry id="F3507631D14851380BA45623379E2A47" box="[389,470,820,846]" name="France" pageId="7" pageNumber="418">France</collectingCountry>
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||
);
|
||
<taxonomicName id="4C474D22D14851380BC8562335742A44" ID-CoL="8CVGQ" authority="Hennig, 1971: 16" authorityName="Hennig" authorityPageNumber="16" authorityYear="1971" box="[489,828,819,846]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D14851380BC8562334482A44" box="[489,512,820,845]" italics="true" pageId="7" pageNumber="418">M</emphasis>
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||
.
|
||
<emphasis id="B933EAB3D1485138082E5623343B2A44" box="[527,627,820,845]" italics="true" pageId="7" pageNumber="418">extinctus</emphasis>
|
||
<bibRefCitation id="EFD64B50D14851380858562435742A44" author="Hennig, W." box="[633,828,819,845]" pageId="7" pageNumber="418" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig, 1971: 16</bibRefCitation>
|
||
</taxonomicName>
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||
(Lebanese amber, Lower Cretaceous, Barremian, Jezzine,
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||
<collectingCountry id="F3507631D14851380AF4564037742A7B" box="[213,316,855,882]" name="Lebanon" pageId="7" pageNumber="418">Lebanon</collectingCountry>
|
||
);
|
||
<taxonomicName id="4C474D22D14851380B71564F356E2A78" authority="Perrichot & Engel, 2014: 31" authorityName="Perrichot & Engel" authorityPageNumber="31" authorityYear="2014" box="[336,806,855,882]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="magaliae">
|
||
<emphasis id="B933EAB3D14851380B71564F372F2A78" box="[336,359,856,881]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380B57564037962A78" box="[374,478,855,881]" italics="true" pageId="7" pageNumber="418">magaliae</emphasis>
|
||
<bibRefCitation id="EFD64B50D14851380BC75640356E2A78" author="Perrichot, V. & Engel, M. S." box="[486,806,855,881]" pageId="7" pageNumber="418" pagination="30 - 33" refId="ref15852" refString="Perrichot, V. & Engel, M. S. (2014) Youngest occurrence of the genus Microphorites (Diptera: Dolichopodidae): a new species in Late Cretaceous Vendean amber. Paleontological Contributions, 10 G, 30 - 33. https: // doi. org / 10.17161 / PC. 1808.15987" type="journal article" year="2014">Perrichot & Engel, 2014: 31</bibRefCitation>
|
||
</taxonomicName>
|
||
(Vendean amber, Upper Cretaceous,
|
||
<collectingCountry id="F3507631D14851380EED564F33552A7B" box="[1228,1309,856,882]" name="France" pageId="7" pageNumber="418">France</collectingCountry>
|
||
);
|
||
<taxonomicName id="4C474D22D14851380F13564F340E2A9F" ID-CoL="9HM3T" authority="Tkoc, Nel & Prokop, 2016: 185" authorityName="Tkoc, Nel & Prokop" authorityPageNumber="185" authorityYear="2016" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B933EAB3D14851380F13564F33012A78" box="[1330,1353,856,881]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380F79564F36992A9C" italics="true" pageId="7" pageNumber="418">moravicus</emphasis>
|
||
<bibRefCitation id="EFD64B50D14851380AFB566C340E2A9F" author="Tkoc, M. & Nel, A. & Prokop, J." box="[218,582,891,918]" pageId="7" pageNumber="418" pagination="181 - 193" refId="ref16568" refString="Tkoc, M., Nel, A. & Prokop, J. (2016) Discovery of a new species of the Cretaceous genus Microphorites Hennig, 1971 (Diptera: Dolichopodidae s. lat.) in Paleogene amber from eastern Moravia (Czech Republic). Insect Systematics & Evolution, 47, 181 - 193. https: // doi. org / 10.1163 / 1876312 X- 47022139" type="journal article" year="2016">Tkoč, Nel & Prokop, 2016: 185</bibRefCitation>
|
||
</taxonomicName>
|
||
(Studlov amber, Cretaceous (Cenomanian)/lower Paleogene, Moravia);
|
||
<taxonomicName id="4C474D22D14851380F5F566B34172AB3" ID-CoL="8CVGR" authority="Grimaldi & Cumming, 1999: 59" authorityName="Grimaldi & Cumming" authorityPageNumber="59" authorityYear="1999" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="oculeus">
|
||
<emphasis id="B933EAB3D14851380F5F566B33DD2A9C" box="[1406,1429,892,917]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380AB6568836A52AB0" box="[151,237,927,953]" italics="true" pageId="7" pageNumber="418">oculeus</emphasis>
|
||
<bibRefCitation id="EFD64B50D14851380AD5568834172AB3" author="Grimaldi, D. A. & Cumming, J. M." box="[244,607,927,954]" pageId="7" pageNumber="418" pagination="1 - 124" refId="ref14337" refString="Grimaldi, D. A. & Cumming, J. M. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 1 - 124." type="journal article" year="1999">Grimaldi & Cumming, 1999: 59</bibRefCitation>
|
||
</taxonomicName>
|
||
(Lebanese amber, Lower Cretaceous);
|
||
<taxonomicName id="4C474D22D14851380E3B56B733372AB0" ID-CoL="9HM3V" authority="Ngo-Muller & Nel" authorityName="Ngo-Muller & Neal" authorityYear="2020" box="[1050,1407,927,954]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="pouilloni">
|
||
<emphasis id="B933EAB3D14851380E3B56B732792AB0" box="[1050,1073,928,953]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380E61568832ED2AB0" box="[1088,1189,927,953]" italics="true" pageId="7" pageNumber="418">pouilloni</emphasis>
|
||
Ngô-Muller & Nel
|
||
</taxonomicName>
|
||
<emphasis id="B933EAB3D14851380FA756B733D42AB0" box="[1414,1436,928,953]" italics="true" pageId="7" pageNumber="418">in</emphasis>
|
||
<bibRefCitation id="EFD64B50D14851380AB656D437F62AD4" author="Ngo-Muller, V. & Garrouste, R. & Pouillon, J. - M. & Nel, A." box="[151,446,963,990]" pageId="7" pageNumber="418" pagination="1 - 4" refId="ref15786" refString="Ngo-Muller, V., Garrouste, R., Pouillon, J. - M. & Nel, A. (2020) A new species of the long-legged fly genus Microphorites in Burmese amber (Dolichopodidae: Microphorinae). Cretaceous Research, 107, 1 - 4. https: // doi. org / 10.1016 / j. cretres. 2019.104284" type="journal article" year="2020">
|
||
Ngô-Muller
|
||
<emphasis id="B933EAB3D14851380B0556D3371D2AD4" box="[292,341,963,989]" italics="true" pageId="7" pageNumber="418">et al</emphasis>
|
||
., 2020: 2
|
||
</bibRefCitation>
|
||
(lowermost Cenomanian, Burmese amber,
|
||
<collectingCountry id="F3507631D1485138098256D3325D2AD7" box="[931,1045,964,990]" name="Myanmar" pageId="7" pageNumber="418">Myanmar</collectingCountry>
|
||
);
|
||
<taxonomicName id="4C474D22D14851380E0956D336B32D0B" ID-CoL="8CVGS" authority="Grimaldi & Cumming, 1999: 57" authorityName="Grimaldi & Cumming" authorityPageNumber="57" authorityYear="1999" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B933EAB3D14851380E0956D332772AD4" box="[1064,1087,964,989]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380E6C56D432DE2AD4" box="[1101,1174,963,989]" italics="true" pageId="7" pageNumber="418">similis</emphasis>
|
||
<bibRefCitation id="EFD64B50D14851380EBC56D436B32D0B" author="Grimaldi, D. A. & Cumming, J. M." pageId="7" pageNumber="418" pagination="1 - 124" refId="ref14337" refString="Grimaldi, D. A. & Cumming, J. M. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 1 - 124." type="journal article" year="1999">Grimaldi & Cumming, 1999: 57</bibRefCitation>
|
||
</taxonomicName>
|
||
(Lebanese amber, Lower Cretaceous);
|
||
<taxonomicName id="4C474D22D1485138089B56FF35F32D08" ID-CoL="8CVGT" authority="Penalver" authorityName="Penalver" authorityYear="2008" box="[698,955,999,1026]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="B933EAB3D1485138089B56FF34992D08" box="[698,721,1000,1025]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D148513808C056F035192D08" box="[737,849,999,1025]" italics="true" pageId="7" pageNumber="418">utrillensis</emphasis>
|
||
Peñalver
|
||
</taxonomicName>
|
||
<emphasis id="B933EAB3D148513809E256FF35912D08" box="[963,985,1000,1025]" italics="true" pageId="7" pageNumber="418">in</emphasis>
|
||
<bibRefCitation id="EFD64B50D148513809C056F032912D08" author="Arillo, A. & Penalver, E. & Delclos, X." box="[993,1241,999,1026]" pageId="7" pageNumber="418" pagination="29 - 40" refId="ref13350" refString="Arillo, A., Penalver, E. & Delclos, X. (2008) Microphorites (Diptera: Dolichopodidae) from the Lower Cretaceous amber of San Just (Spain), and the co-occurrence of two ceratopogonid species in Spanish amber deposits. Zootaxa, 1920 (1), 29 - 40. https: // doi. org / 10.11646 / zootaxa. 1920.1.2" type="journal article" year="2008">
|
||
Arillo
|
||
<emphasis id="B933EAB3D14851380E0D56FF32172D08" box="[1068,1119,999,1025]" italics="true" pageId="7" pageNumber="418">et al</emphasis>
|
||
., 2008: 31
|
||
</bibRefCitation>
|
||
(San Just amber, upper Albian, Lower Cretaceous,
|
||
<collectingCountry id="F3507631D14851380830511C341E2D2F" box="[529,598,1035,1062]" name="Spain" pageId="7" pageNumber="418">Spain</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D652AD14851360AE6513832E82AC0" lastPageId="9" lastPageNumber="420" pageId="7" pageNumber="418" type="discussion">
|
||
<paragraph id="8BF836A1D14851380AE6513832922DF4" blockId="7.[151,1437,747,2034]" pageId="7" pageNumber="418">
|
||
<emphasis id="B933EAB3D14851380AE6513837732D40" bold="true" box="[199,315,1071,1097]" pageId="7" pageNumber="418">Remarks.</emphasis>
|
||
<bibRefCitation id="EFD64B50D14851380B65513837A22D40" author="Hennig, W." box="[324,490,1071,1097]" pageId="7" pageNumber="418" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971)</bibRefCitation>
|
||
established this genus to include the single species
|
||
<taxonomicName id="4C474D22D14851380E63512732862D40" authorityName="Hennig" authorityYear="1971" box="[1090,1230,1072,1098]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D14851380E63512732112D40" box="[1090,1113,1072,1097]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380E4B512732862D40" box="[1130,1230,1072,1097]" italics="true" pageId="7" pageNumber="418">extinctus</emphasis>
|
||
</taxonomicName>
|
||
described by him from Lebanese amber (Lower Cretaceous, Barremian, Jezzine,
|
||
<collectingCountry id="F3507631D14851380972514435FD2D64" box="[851,949,1107,1133]" name="Lebanon" pageId="7" pageNumber="418">Lebanon</collectingCountry>
|
||
(129.4–125 Ma)). Currently,
|
||
<taxonomicName id="4C474D22D14851380EDC514433D42D64" authorityName="Hennig" authorityYear="1971" box="[1277,1436,1107,1133]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14851380EDC514433D42D64" box="[1277,1436,1107,1133]" italics="true" pageId="7" pageNumber="418">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
unites nine named species, which were found from different deposits dated over a very broad geological temporal range from Lower Cretaceous to lowermost Eocene: Lower Cretaceous—five species; Upper Cretaceous—three (?two); Eocene—one (?two). In terms of current geography, eight species are known from the Palaearctic Region (
|
||
<collectingCountry id="F3507631D14851380ABE51F4371D2DF4" box="[159,341,1251,1277]" name="Czech Republic" pageId="7" pageNumber="418">Czech Republic</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507631D14851380B4051F337F82DF7" box="[353,432,1252,1278]" name="France" pageId="7" pageNumber="418">France</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507631D14851380B9A51F434682DF4" box="[443,544,1251,1277]" name="Lebanon" pageId="7" pageNumber="418">Lebanon</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3507631D1485138080A51F434242DF7" box="[555,620,1251,1278]" name="Spain" pageId="7" pageNumber="418">Spain</collectingCountry>
|
||
) and one species from the Oriental Region (
|
||
<collectingCountry id="F3507631D14851380E7A51F432872DF4" box="[1115,1231,1251,1277]" name="Myanmar" pageId="7" pageNumber="418">Myanmar</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14851380AE6501035992CB8" blockId="7.[151,1437,747,2034]" pageId="7" pageNumber="418">
|
||
<bibRefCitation id="EFD64B50D14851380AE6501034472C28" author="Grimaldi, D. A. & Cumming, J. M." box="[199,527,1287,1313]" pageId="7" pageNumber="418" pagination="1 - 124" refId="ref14337" refString="Grimaldi, D. A. & Cumming, J. M. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 1 - 124." type="journal article" year="1999">Grimaldi & Cumming (1999)</bibRefCitation>
|
||
described from Lebanese amber the next two species of
|
||
<taxonomicName id="4C474D22D14851380EA3501033692C28" authorityName="Hennig" authorityYear="1971" box="[1154,1313,1287,1313]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14851380EA3501033692C28" box="[1154,1313,1287,1313]" italics="true" pageId="7" pageNumber="418">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C474D22D14851380F0E501F33D42C28" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[1327,1436,1287,1314]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B933EAB3D14851380F0E501F330E2C28" box="[1327,1350,1288,1313]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380F72501033D42C28" box="[1363,1436,1287,1313]" italics="true" pageId="7" pageNumber="418">similis</emphasis>
|
||
</taxonomicName>
|
||
(only female) and
|
||
<taxonomicName id="4C474D22D14851380B43503B37942C4C" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[354,476,1323,1350]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="oculeus">
|
||
<emphasis id="B933EAB3D14851380B43503B37312C4C" box="[354,377,1324,1349]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380BA7503C37942C4C" box="[390,476,1323,1349]" italics="true" pageId="7" pageNumber="418">oculeus</emphasis>
|
||
</taxonomicName>
|
||
(only male). Two other Lower Cretaceous species of
|
||
<taxonomicName id="4C474D22D14851380E0B503C32812C4C" authorityName="Hennig" authorityYear="1971" box="[1066,1225,1323,1349]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14851380E0B503C32812C4C" box="[1066,1225,1323,1349]" italics="true" pageId="7" pageNumber="418">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
are much younger. They were described from Charentese amber of
|
||
<collectingCountry id="F3507631D1485138088D504734B02C63" box="[684,760,1360,1386]" name="France" pageId="7" pageNumber="418">France</collectingCountry>
|
||
(Albian, 105.3–99.6 Ma) (
|
||
<bibRefCitation id="EFD64B50D14851380E00505832802C60" author="Nel, A. & Perrichot, V. & Daugeron, C. & Neraudeau, D." box="[1057,1224,1359,1386]" pageId="7" pageNumber="418" pagination="23 - 29" refId="ref15681" refString="Nel, A., Perrichot, V., Daugeron, C. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France, New Series, 40, 23 - 29. https: // doi. org / 10.1080 / 00379271.2004.10697401" type="journal article" year="2004">
|
||
Nel
|
||
<emphasis id="B933EAB3D14851380E73504732CA2C60" box="[1106,1154,1359,1385]" italics="true" pageId="7" pageNumber="418">et al</emphasis>
|
||
. 2004
|
||
</bibRefCitation>
|
||
) and from San Just amber of
|
||
<collectingCountry id="F3507631D14851380B275064370E2C84" box="[262,326,1395,1421]" name="Spain" pageId="7" pageNumber="418">Spain</collectingCountry>
|
||
(Albian, 112–105.3 Ma) (
|
||
<bibRefCitation id="EFD64B50D14851380851506435752C84" author="Arillo, A. & Penalver, E. & Delclos, X." box="[624,829,1395,1422]" pageId="7" pageNumber="418" pagination="29 - 40" refId="ref13350" refString="Arillo, A., Penalver, E. & Delclos, X. (2008) Microphorites (Diptera: Dolichopodidae) from the Lower Cretaceous amber of San Just (Spain), and the co-occurrence of two ceratopogonid species in Spanish amber deposits. Zootaxa, 1920 (1), 29 - 40. https: // doi. org / 10.11646 / zootaxa. 1920.1.2" type="journal article" year="2008">
|
||
Arillo
|
||
<emphasis id="B933EAB3D1485138089C506334B82C84" box="[701,752,1395,1421]" italics="true" pageId="7" pageNumber="418">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
):
|
||
<taxonomicName id="4C474D22D14851380970506333D42C84" authority="Nel, Perrishot, Daugeron & Neraudeau" authorityName="Nel, Perrishot, Daugeron & Neraudeau" box="[849,1436,1395,1422]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="B933EAB3D14851380970506335902C84" box="[849,984,1395,1421]" italics="true" pageId="7" pageNumber="418">M. deploegi</emphasis>
|
||
Nel, Perrishot, Daugeron & Néraudeau
|
||
</taxonomicName>
|
||
(male and female) and
|
||
<taxonomicName id="4C474D22D14851380BB8508F34D02CB8" authority="Penalver" authorityName="Penalver" authorityYear="2008" box="[409,664,1431,1457]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="B933EAB3D14851380BB8508F34672CB8" box="[409,559,1431,1457]" italics="true" pageId="7" pageNumber="418">M. utrillensis</emphasis>
|
||
Peñalver
|
||
</taxonomicName>
|
||
(only female), respectively.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14851380AE650AC34642FFC" blockId="7.[151,1437,747,2034]" pageId="7" pageNumber="418">
|
||
The Upper Cretaceous fauna of
|
||
<taxonomicName id="4C474D22D1485138081850AC34902CDC" authorityName="Hennig" authorityYear="1971" box="[569,728,1467,1493]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D1485138081850AC34902CDC" box="[569,728,1467,1493]" italics="true" pageId="7" pageNumber="418">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
is known at least after two species. The first species was described from Vendean amber of
|
||
<collectingCountry id="F3507631D1485138082A50F7341F2CF3" box="[523,599,1504,1530]" name="France" pageId="7" pageNumber="418">France</collectingCountry>
|
||
(Cenomanian to Turonian, 99.6–89.8 Ma) (
|
||
<bibRefCitation id="EFD64B50D14851380E6E50C833202CF0" author="Perrichot, V. & Engel, M. S." box="[1103,1384,1503,1529]" pageId="7" pageNumber="418" pagination="30 - 33" refId="ref15852" refString="Perrichot, V. & Engel, M. S. (2014) Youngest occurrence of the genus Microphorites (Diptera: Dolichopodidae): a new species in Late Cretaceous Vendean amber. Paleontological Contributions, 10 G, 30 - 33. https: // doi. org / 10.17161 / PC. 1808.15987" type="journal article" year="2014">Perrichot & Engel 2014</bibRefCitation>
|
||
):
|
||
<taxonomicName id="4C474D22D14851380F5F50F737982F14" authority="Perrichot & Engel" authorityName="Perrichot & Engel" authorityYear="2014" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="magaliae">
|
||
<emphasis id="B933EAB3D14851380F5F50F736B72F14" italics="true" pageId="7" pageNumber="418">M. magaliae</emphasis>
|
||
Perrichot & Engel
|
||
</taxonomicName>
|
||
(only male). The second species was found in Burmese amber of
|
||
<collectingCountry id="F3507631D14851380E8B5313335F2F17" box="[1194,1303,1540,1566]" name="Myanmar" pageId="7" pageNumber="418">Myanmar</collectingCountry>
|
||
(lowermost Cenomanian, ~99 Ma):
|
||
<taxonomicName id="4C474D22D14851380B85533F355D2F48" authority="Ngo-Muller & Neal" authorityName="Ngo-Muller & Neal" authorityYear="2020" box="[420,789,1575,1602]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="pouilloni">
|
||
<emphasis id="B933EAB3D14851380B85533F37F32F48" box="[420,443,1576,1601]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380BE8533034662F48" box="[457,558,1575,1601]" italics="true" pageId="7" pageNumber="418">pouilloni</emphasis>
|
||
Ngô-Muller & Neal
|
||
</taxonomicName>
|
||
(only male).
|
||
<bibRefCitation id="EFD64B50D1485138098C533032D82F48" author="Jouault, C. & Ngo-Muller, V. & Zhang, Q. & Nel, A." box="[941,1168,1575,1602]" pageId="7" pageNumber="418" pagination="204 - 211" refId="ref14465" refString="Jouault, C., Ngo-Muller, V., Zhang, Q. & Nel, A. (2020) New empidoid flies (Diptera: Atelestidae; Dolichopodidae) from mid- Cretaceous Burmese amber. Palaeoentomology, 3 (2), 204 - 211. https: // doi. org / 10.11646 / palaeoentomology. 3.2.10" type="journal article" year="2020">
|
||
Jouault
|
||
<emphasis id="B933EAB3D14851380E27533F327F2F48" box="[1030,1079,1575,1601]" italics="true" pageId="7" pageNumber="418">et al</emphasis>
|
||
. (2020)
|
||
</bibRefCitation>
|
||
published a description of the female of
|
||
<taxonomicName id="4C474D22D14851380B73535B37942F6C" authorityName="Ngo-Muller & Neal" authorityYear="2020" box="[338,476,1611,1638]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="pouilloni">
|
||
<emphasis id="B933EAB3D14851380B73535B37212F6C" box="[338,361,1612,1637]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380B56535C37942F6C" box="[375,476,1611,1637]" italics="true" pageId="7" pageNumber="418">pouilloni</emphasis>
|
||
</taxonomicName>
|
||
. However, the female described by these authors is not conspecific with
|
||
<taxonomicName id="4C474D22D14851380F30535B33D42F6C" authorityName="Ngo-Muller & Neal" authorityYear="2020" box="[1297,1436,1611,1638]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="pouilloni">
|
||
<emphasis id="B933EAB3D14851380F30535B33602F6C" box="[1297,1320,1612,1637]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380F16535C33D42F6C" box="[1335,1436,1611,1637]" italics="true" pageId="7" pageNumber="418">pouilloni</emphasis>
|
||
</taxonomicName>
|
||
because of the differences with the male are beyond the range of sexual dimorphic variability in microphorines.
|
||
<taxonomicName id="4C474D22D14851380F55536737DB2FA4" authorityName="Tkoc, Nel & Prokop" authorityYear="2016" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B933EAB3D14851380F55536737DB2FA4" italics="true" pageId="7" pageNumber="418">Microphorites moravicus</emphasis>
|
||
</taxonomicName>
|
||
described after a female from Študlov amber of
|
||
<collectingCountry id="F3507631D148513809955384322E2FA4" box="[948,1126,1683,1709]" name="Czech Republic" pageId="7" pageNumber="418">Czech Republic</collectingCountry>
|
||
was dated from the Eocene (Lutetian, 47.8 Ma), although, the Upper Cretaceous age (upper Cenomanian, 91 Ma) cannot be precluded (
|
||
<bibRefCitation id="EFD64B50D14851380F4053A0375A2FFC" author="Tkoc, M. & Nel, A. & Prokop, J." pageId="7" pageNumber="418" pagination="181 - 193" refId="ref16568" refString="Tkoc, M., Nel, A. & Prokop, J. (2016) Discovery of a new species of the Cretaceous genus Microphorites Hennig, 1971 (Diptera: Dolichopodidae s. lat.) in Paleogene amber from eastern Moravia (Czech Republic). Insect Systematics & Evolution, 47, 181 - 193. https: // doi. org / 10.1163 / 1876312 X- 47022139" type="journal article" year="2016">
|
||
Tkoč
|
||
<emphasis id="B933EAB3D14851380AB653CB36802FFC" box="[151,200,1755,1781]" italics="true" pageId="7" pageNumber="418">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD64B50D14851380B3C53CB34682FFC" author="Bramuzzo, S. & Nel, A." box="[285,544,1755,1782]" pageId="7" pageNumber="418" pagination="590 - 594" refId="ref13466" refString="Bramuzzo, S. & Nel, A. (2017) Youngest representative of the extinct genus Microphorites in the Eocene amber of France (Diptera: Dolichopodidae: Microphorinae). Zootaxa, 4231 (4), 590 - 594. https: // doi. org / 10.11646 / zootaxa. 4231.4.11" type="journal article" year="2017">Bramuzzo & Nel 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14851380AE653E835C22E6B" blockId="7.[151,1437,747,2034]" pageId="7" pageNumber="418">
|
||
Finally, a species described from Oise amber of
|
||
<collectingCountry id="F3507631D148513808C5521735782E13" box="[740,816,1792,1818]" name="France" pageId="7" pageNumber="418">France</collectingCountry>
|
||
(lowermost Eocene, 53 Ma) would support that
|
||
<taxonomicName id="4C474D22D14851380F71521736BB2E34" authorityName="Hennig" authorityYear="1971" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14851380F71521736BB2E34" italics="true" pageId="7" pageNumber="418">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
really occurred in the Eocene (
|
||
<bibRefCitation id="EFD64B50D14851380877523435162E34" author="Bramuzzo, S. & Nel, A." box="[598,862,1827,1853]" pageId="7" pageNumber="418" pagination="590 - 594" refId="ref13466" refString="Bramuzzo, S. & Nel, A. (2017) Youngest representative of the extinct genus Microphorites in the Eocene amber of France (Diptera: Dolichopodidae: Microphorinae). Zootaxa, 4231 (4), 590 - 594. https: // doi. org / 10.11646 / zootaxa. 4231.4.11" type="journal article" year="2017">Bramuzzo & Nel 2017</bibRefCitation>
|
||
):
|
||
<taxonomicName id="4C474D22D14851380953523332EA2E34" authority="Bramuzzo & Nel" authorityName="Bramuzzo & Nel" authorityYear="2017" box="[882,1186,1827,1854]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="erikai">
|
||
<emphasis id="B933EAB3D14851380953523335912E34" box="[882,985,1827,1853]" italics="true" pageId="7" pageNumber="418">M. erikai</emphasis>
|
||
Bramuzzo & Nel
|
||
</taxonomicName>
|
||
(only female). Therefore, this species could be the youngest representative of the genus.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14851370AE6527C3219286F" blockId="7.[151,1437,747,2034]" lastBlockId="8.[151,1437,151,2020]" lastPageId="8" lastPageNumber="419" pageId="7" pageNumber="418">
|
||
<bibRefCitation id="EFD64B50D14851380AE6527C37242E8C" author="Hennig, W." box="[199,364,1899,1925]" pageId="7" pageNumber="418" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971)</bibRefCitation>
|
||
indicated the main features distinguishing
|
||
<taxonomicName id="4C474D22D14851380979527C35BF2E8C" authorityName="Hennig" authorityYear="1971" box="[856,1015,1899,1925]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14851380979527C35BF2E8C" box="[856,1015,1899,1925]" italics="true" pageId="7" pageNumber="418">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C474D22D14851380E1F527C32FE2E8C" box="[1086,1206,1899,1925]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14851380E1F527C32FE2E8C" box="[1086,1206,1899,1925]" italics="true" pageId="7" pageNumber="418">Microphor</emphasis>
|
||
</taxonomicName>
|
||
(anal angle of wing strongly rounded) as well as from
|
||
<taxonomicName id="4C474D22D14851380834529834FE2EA0" authorityName="Becker" authorityYear="1909" box="[533,694,1935,1961]" class="Insecta" family="Dolichopodidae" genus="Microphorella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14851380834529834FE2EA0" box="[533,694,1935,1961]" italics="true" pageId="7" pageNumber="418">Microphorella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D148513808CD529835DC2EA0" box="[748,916,1935,1961]" class="Insecta" family="Dolichopodidae" genus="Parathalassius" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D148513808CD529835DC2EA0" box="[748,916,1935,1961]" italics="true" pageId="7" pageNumber="418">Parathalassius</emphasis>
|
||
</taxonomicName>
|
||
(antennae situated approximately in middle of anterior eye margin, arista two-segmented, basal crossvein of wing complete, eye lack hairs). Figure 21 (
|
||
<bibRefCitation id="EFD64B50D14851380F6052A436B32EF8" author="Hennig, W." pageId="7" pageNumber="418" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig, 1971: 17</bibRefCitation>
|
||
) shows that his expression “the anal angle of the wing is strongly rounded” means that in
|
||
<taxonomicName id="4C474D22D14851380ECB52CF333C2EF8" authorityName="Hennig" authorityYear="1971" box="[1258,1396,2008,2034]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D14851380ECB52CF33492EF8" box="[1258,1281,2008,2033]" italics="true" pageId="7" pageNumber="418">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14851380F3152CF333C2EF8" box="[1296,1396,2008,2033]" italics="true" pageId="7" pageNumber="418">extinctus</emphasis>
|
||
</taxonomicName>
|
||
the anal lobe is weakly developed and the anal angle is obtuse (not right-angled as
|
||
<taxonomicName id="4C474D22D14751370E3B558032DA29B8" box="[1050,1170,151,177]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370E3B558032DA29B8" box="[1050,1170,151,177]" italics="true" pageId="8" pageNumber="419">Microphor</emphasis>
|
||
</taxonomicName>
|
||
). A review of available illustrations of the wing of other species assigned to
|
||
<taxonomicName id="4C474D22D147513708CF55AC35C529DC" authorityName="Hennig" authorityYear="1971" box="[750,909,187,213]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D147513708CF55AC35C529DC" box="[750,909,187,213]" italics="true" pageId="8" pageNumber="419">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
shows that the degree of curvature of the anal lobe is variable (anal lobe margin invisible in
|
||
<taxonomicName id="4C474D22D147513708B655C8354E29F0" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[663,774,223,249]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B933EAB3D147513708B655C834E629F1" box="[663,686,223,248]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137089C55C8354E29F0" box="[701,774,223,249]" italics="true" pageId="8" pageNumber="419">similis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D1475137093255C835C629F0" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[787,910,223,249]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="oculeus">
|
||
<emphasis id="B933EAB3D1475137093255C8356229F1" box="[787,810,223,248]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137091955C835C629F0" box="[824,910,223,249]" italics="true" pageId="8" pageNumber="419">oculeus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D147513709E555C8326229F0" authorityName="Bramuzzo & Nel" authorityYear="2017" box="[964,1066,223,249]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="erikai">
|
||
<emphasis id="B933EAB3D147513709E555C8359329F1" box="[964,987,223,248]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D147513709C855C8326229F0" box="[1001,1066,223,249]" italics="true" pageId="8" pageNumber="419">erikai</emphasis>
|
||
</taxonomicName>
|
||
). It is especially clear in the case of
|
||
<taxonomicName id="4C474D22D14751370A965414378E2814" authorityName="Perrichot & Engel" authorityYear="2014" box="[183,454,259,285]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="magaliae">
|
||
<emphasis id="B933EAB3D14751370A965414378E2814" box="[183,454,259,285]" italics="true" pageId="8" pageNumber="419">Microphorites magaliae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFD64B50D14751370BF4541434A12814" author="Perrichot, V. & Engel, M. S." box="[469,745,259,285]" pageId="8" pageNumber="419" pagination="30 - 33" refId="ref15852" refString="Perrichot, V. & Engel, M. S. (2014) Youngest occurrence of the genus Microphorites (Diptera: Dolichopodidae): a new species in Late Cretaceous Vendean amber. Paleontological Contributions, 10 G, 30 - 33. https: // doi. org / 10.17161 / PC. 1808.15987" type="journal article" year="2014">Perrichot & Engel 2014</bibRefCitation>
|
||
: fig. G2;
|
||
<bibRefCitation id="EFD64B50D147513709745414325C2817" author="Tkoc, M. & Nel, A. & Prokop, J." box="[853,1044,259,286]" pageId="8" pageNumber="419" pagination="181 - 193" refId="ref16568" refString="Tkoc, M., Nel, A. & Prokop, J. (2016) Discovery of a new species of the Cretaceous genus Microphorites Hennig, 1971 (Diptera: Dolichopodidae s. lat.) in Paleogene amber from eastern Moravia (Czech Republic). Insect Systematics & Evolution, 47, 181 - 193. https: // doi. org / 10.1163 / 1876312 X- 47022139" type="journal article" year="2016">
|
||
Tkoč
|
||
<emphasis id="B933EAB3D147513709B7541435812814" box="[918,969,259,285]" italics="true" pageId="8" pageNumber="419">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
). In this species, the wing shape is the same as in the females of some
|
||
<taxonomicName id="4C474D22D14751370809543034B12848" box="[552,761,295,321]" pageId="8" pageNumber="419">
|
||
<emphasis id="B933EAB3D14751370809543034E82848" box="[552,672,295,321]" italics="true" pageId="8" pageNumber="419">Microphor</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and in both sexes of many species of
|
||
<taxonomicName id="4C474D22D14751370E87543033722848" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[1190,1338,295,321]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="419" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14751370E87543033722848" box="[1190,1338,295,321]" italics="true" pageId="8" pageNumber="419">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
; i.e., the anal lobe is strongly developed (
|
||
<bibRefCitation id="EFD64B50D14751370823545C3564286C" author="Brooks, S. E. & Cumming, J. M." box="[514,812,331,357]" pageId="8" pageNumber="419" pagination="1 - 141" refId="ref13522" refString="Brooks, S. E. & Cumming, J. M. (2022) Revision of the Nearctic Microphorinae (Diptera: Dolichopodidae sensu lato). Zootaxa, 5095 (1), 1 - 141. https: // doi. org / 10.11646 / zootaxa. 5095.1.1" type="journal article" year="2022">Brooks & Cumming 2022</bibRefCitation>
|
||
; Shamshev, unpubl. data).
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14751370AE6547A35A12BEA" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="419">
|
||
<taxonomicName id="4C474D22D14751370AE6547A379B288E" authorityName="Hennig" authorityYear="1971" box="[199,467,365,391]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D14751370AE6547A379B288E" box="[199,467,365,391]" italics="true" pageId="8" pageNumber="419">Microphorites extinctus</emphasis>
|
||
</taxonomicName>
|
||
(as well as species of
|
||
<taxonomicName id="4C474D22D147513708F7547A3506288E" box="[726,846,365,391]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D147513708F7547A3506288E" box="[726,846,365,391]" italics="true" pageId="8" pageNumber="419">Microphor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D147513709A9547A3254288E" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[904,1052,365,391]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="419" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D147513709A9547A3254288E" box="[904,1052,365,391]" italics="true" pageId="8" pageNumber="419">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
) differs from all species assigned later to
|
||
<taxonomicName id="4C474D22D14751370ACF548737C528A3" authorityName="Hennig" authorityYear="1971" box="[238,397,400,426]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370ACF548737C528A3" box="[238,397,400,426]" italics="true" pageId="8" pageNumber="419">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
primarily by the complete (circumambient) costal vein (
|
||
<emphasis id="B933EAB3D14751370E2A5486321B28A3" box="[1035,1107,401,426]" italics="true" pageId="8" pageNumber="419">versus</emphasis>
|
||
incomplete, i.e., ending at or just beyond R
|
||
<subScript id="17C334E4D14751370B1454D2370728DA" attach="left" box="[309,335,453,467]" fontSize="6" pageId="8" pageNumber="419">4+5</subScript>
|
||
, or at M
|
||
<subScript id="17C334E4D14751370B9054D237F128DA" attach="left" box="[433,441,453,467]" fontSize="6" pageId="8" pageNumber="419">1</subScript>
|
||
, leaving posterior margin of wing membranous). The costal termination is an important diagnostic character in higher level
|
||
<taxonomicName id="4C474D22D1475137080654C234FA28E6" authorityName="Latreille" authorityYear="1804" box="[551,690,469,495]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="426" phylum="Arthropoda" rank="superFamily" superFamily="Empidoidea">Empidoidea</taxonomicName>
|
||
. Phylogenetically, it is probable undisputable that the circumambient costa is the plesiomorphic condition (e.g.,
|
||
<bibRefCitation id="EFD64B50D1475137088754EF324B2B1A" author="Sinclair, B. J. & Cumming, J. M." box="[678,1027,504,531]" pageId="8" pageNumber="419" pagination="1 - 172" refId="ref16254" refString="Sinclair, B. J. & Cumming, J. M. (2006) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa, 1180 (1), 1 - 172. https: // doi. org / 10.11646 / zootaxa. 1180.1.1" type="journal article" year="2006">Sinclair & Cumming 2006: 35</bibRefCitation>
|
||
, character 50). However, the reduction of the costa evolved independently in several groups of
|
||
<taxonomicName id="4C474D22D1475137091D570C358F2B3C" authorityName="Latreille" authorityYear="1804" box="[828,967,539,565]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="426" phylum="Arthropoda" rank="superFamily" superFamily="Empidoidea">Empidoidea</taxonomicName>
|
||
. This character has some “disadvantages” because, in some empidoids, the costa does not end abruptly (when incomplete) or it is weakened beyond meeting point with R
|
||
<subScript id="17C334E4D14751370B05576537762B89" attach="left" box="[292,318,626,640]" fontSize="6" pageId="8" pageNumber="419">4+5</subScript>
|
||
or M
|
||
<subScript id="17C334E4D14751370B5C576537CD2B89" attach="left" box="[381,389,626,640]" fontSize="6" pageId="8" pageNumber="419">1</subScript>
|
||
; i.e. along posterior margin of the wing (when complete) (see
|
||
<bibRefCitation id="EFD64B50D14751370E695777333C2B73" author="Sinclair, B. J. & Cumming, J. M." box="[1096,1396,608,634]" pageId="8" pageNumber="419" pagination="1 - 172" refId="ref16254" refString="Sinclair, B. J. & Cumming, J. M. (2006) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa, 1180 (1), 1 - 172. https: // doi. org / 10.11646 / zootaxa. 1180.1.1" type="journal article" year="2006">Sinclair & Cumming 2006</bibRefCitation>
|
||
for some examples). Within the
|
||
<taxonomicName id="4C474D22D14751370BFE579434CD2B94" authorityName="Collin" authorityYear="1960" box="[479,645,643,669]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="subFamily" subFamily="Microphorinae">Microphorinae</taxonomicName>
|
||
lineage, an incomplete costa was described in
|
||
<taxonomicName id="4C474D22D14751370E83579433612B94" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[1186,1321,643,669]" class="Insecta" family="Dolichopodidae" genus="Avenaphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370E83579433612B94" box="[1186,1321,643,669]" italics="true" pageId="8" pageNumber="419">Avenaphora</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14751370F45579437622BB6" class="Insecta" family="Dolichopodidae" genus="Pristinmicrophor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370F45579437622BB6" italics="true" pageId="8" pageNumber="419">Pristinmicrophor</emphasis>
|
||
</taxonomicName>
|
||
, within the
|
||
<taxonomicName id="4C474D22D14751370B9957B234242BB6" authorityName="Chvála" authorityYear="1981" box="[440,620,677,703]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="subFamily" subFamily="Parathalassiinae">Parathalassiinae</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C474D22D147513708B157B2350B2BB6" authorityName="Latreille" authorityYear="1809" box="[656,835,677,703]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
|
||
<emphasis id="B933EAB3D1475137096C57B135952BB6" box="[845,989,678,703]" italics="true" pageId="8" pageNumber="419">
|
||
<taxonomicName id="4C474D22D1475137096C57B135C32BB6" box="[845,907,678,703]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
lineage—in extinct
|
||
<taxonomicName id="4C474D22D14751370EE957B233DD2BB6" authorityName="Negrobov" authorityYear="1978" box="[1224,1429,677,703]" class="Insecta" family="Empididae" genus="Cretomicrophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370EE957B233DD2BB6" box="[1224,1429,677,703]" italics="true" pageId="8" pageNumber="419">Cretomicrophorus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D14751370AB657DF37082BEB" authorityName="Negrobov" authorityYear="1978" box="[151,320,712,738]" class="Insecta" family="Dolichopodidae" genus="Archichrysotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370AB657DF37082BEB" box="[151,320,712,738]" italics="true" pageId="8" pageNumber="419">Archichrysotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14751370B5757DE37922BEB" authorityName="Negrobov" authorityYear="1978" box="[374,474,713,738]" class="Insecta" family="Empididae" genus="Retinitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370B5757DE37922BEB" box="[374,474,713,738]" italics="true" pageId="8" pageNumber="419">Retinitus</emphasis>
|
||
</taxonomicName>
|
||
, as well as in all
|
||
<taxonomicName id="4C474D22D147513708BA57DF35062BEB" authorityName="Latreille" authorityYear="1809" box="[667,846,712,738]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
|
||
<emphasis id="B933EAB3D1475137097457DE35AA2BEB" box="[853,994,713,738]" italics="true" pageId="8" pageNumber="419">
|
||
<taxonomicName id="4C474D22D1475137097457DE35DB2BEB" box="[853,915,713,738]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14751370AE657FC35772D34" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="419">
|
||
In addition,
|
||
<bibRefCitation id="EFD64B50D14751370B6F57FC37B92A0C" author="Hennig, W." box="[334,497,747,773]" pageId="8" pageNumber="419" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971)</bibRefCitation>
|
||
noted that the ommatidia in
|
||
<taxonomicName id="4C474D22D1475137090F57FC35FF2A0D" authorityName="Hennig" authorityYear="1971" box="[814,951,747,773]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D1475137090F57FC350D2A0D" box="[814,837,747,772]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137097257FC35FF2A0D" box="[851,951,747,772]" italics="true" pageId="8" pageNumber="419">extinctus</emphasis>
|
||
</taxonomicName>
|
||
are distinctly larger on the upper half of the eye and the mesonotum is fairly uniformly covered with fine hairs (besides macrosetae). Both features are unique for microphorines, but their phylogenetic value is unclear. The first character is applicable only to females because the male of
|
||
<taxonomicName id="4C474D22D14751370B3C564437EF2A65" authorityName="Hennig" authorityYear="1971" box="[285,423,851,877]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D14751370B3C5644377C2A65" box="[285,308,851,876]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370B62564437EF2A65" box="[323,423,851,876]" italics="true" pageId="8" pageNumber="419">extinctus</emphasis>
|
||
</taxonomicName>
|
||
remains unknown. An eye with enlarged upper ommatidia is a curious feature for females of microphorines, as
|
||
<bibRefCitation id="EFD64B50D14751370BA3566234042A99" author="Chvala, M." box="[386,588,885,912]" pageId="8" pageNumber="419" pagination="1 - 279" refId="ref13639" refString="Chvala, M. (1983) The Empidoidea (Diptera) of Fennoscandia and Denmark. II. General Part. The families Hybotidae, Atelestidae and Microphoridae. Fauna Entomologica Scandinavica, 12, 1 - 279." type="journal article" year="1983">Chvála (1983: 57)</bibRefCitation>
|
||
has already noted. In many species of these flies, the enlarged upper ommatidia are usually present in males (when compound eyes are holoptic), whereas in females they are uniformly small. Undifferentiated ommatidia (i.e., equally small) were described in males with holoptic eyes of all species assigned to
|
||
<taxonomicName id="4C474D22D14751370A9556CA371B2AFE" authorityName="Hennig" authorityYear="1971" box="[180,339,989,1015]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370A9556CA371B2AFE" box="[180,339,989,1015]" italics="true" pageId="8" pageNumber="419">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
(Grimaldi & Cumming 19991;
|
||
<bibRefCitation id="EFD64B50D1475137089856CA352B2AFE" author="Nel, A. & Perrichot, V. & Daugeron, C. & Neraudeau, D." box="[697,867,989,1016]" pageId="8" pageNumber="419" pagination="23 - 29" refId="ref15681" refString="Nel, A., Perrichot, V., Daugeron, C. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France, New Series, 40, 23 - 29. https: // doi. org / 10.1080 / 00379271.2004.10697401" type="journal article" year="2004">
|
||
Nel
|
||
<emphasis id="B933EAB3D147513708C956C935512AFE" box="[744,793,989,1015]" italics="true" pageId="8" pageNumber="419">et al</emphasis>
|
||
. 2004
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD64B50D1475137094F56CA32352AFE" author="Perrichot, V. & Engel, M. S." box="[878,1149,989,1015]" pageId="8" pageNumber="419" pagination="30 - 33" refId="ref15852" refString="Perrichot, V. & Engel, M. S. (2014) Youngest occurrence of the genus Microphorites (Diptera: Dolichopodidae): a new species in Late Cretaceous Vendean amber. Paleontological Contributions, 10 G, 30 - 33. https: // doi. org / 10.17161 / PC. 1808.15987" type="journal article" year="2014">Perrichot & Engel 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD64B50D14751370EA956CA33D82AFE" author="Ngo-Muller, V. & Garrouste, R. & Pouillon, J. - M. & Nel, A." box="[1160,1424,989,1016]" pageId="8" pageNumber="419" pagination="1 - 4" refId="ref15786" refString="Ngo-Muller, V., Garrouste, R., Pouillon, J. - M. & Nel, A. (2020) A new species of the long-legged fly genus Microphorites in Burmese amber (Dolichopodidae: Microphorinae). Cretaceous Research, 107, 1 - 4. https: // doi. org / 10.1016 / j. cretres. 2019.104284" type="journal article" year="2020">
|
||
Ngô-Muller
|
||
<emphasis id="B933EAB3D14751370F3456C9330E2AFE" box="[1301,1350,989,1015]" italics="true" pageId="8" pageNumber="419">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
), which is known in some recent species of
|
||
<taxonomicName id="4C474D22D1475137084F5117354A2D13" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[622,770,1024,1050]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="419" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D1475137084F5117354A2D13" box="[622,770,1024,1050]" italics="true" pageId="8" pageNumber="419">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
as well (
|
||
<bibRefCitation id="EFD64B50D14751370944511735B12D13" author="Chvala, M." box="[869,1017,1024,1050]" pageId="8" pageNumber="419" pagination="133 - 155" refId="ref13708" refString="Chvala, M. (1987) Revision of Palearctic Microphoridae (Diptera) 2. Schistostoma Beck. Acta Entomologica Bohemoslovaca, 84, 133 - 155." type="journal article" year="1987">Chvála 1987</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFD64B50D14751370E2D5117333A2D12" author="Sinclair, B. J. & Cumming, J. M." box="[1036,1394,1024,1051]" pageId="8" pageNumber="419" pagination="1 - 172" refId="ref16254" refString="Sinclair, B. J. & Cumming, J. M. (2006) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa, 1180 (1), 1 - 172. https: // doi. org / 10.11646 / zootaxa. 1180.1.1" type="journal article" year="2006">Sinclair & Cumming (2006: 60)</bibRefCitation>
|
||
did not include the facet size character in their cladistic analysis.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14751370AE65151334E2C1A" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="419">
|
||
In the remaining species assigned later to
|
||
<taxonomicName id="4C474D22D147513708B8515235702D56" authorityName="Hennig" authorityYear="1971" box="[665,824,1093,1119]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D147513708B8515235702D56" box="[665,824,1093,1119]" italics="true" pageId="8" pageNumber="419">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
, the costa is incomplete, as it follows from the original descriptions (a helpful summary, with some other characters, was provided by
|
||
<bibRefCitation id="EFD64B50D14751370E3D517E32B12D8D" author="Tkoc, M. & Nel, A. & Prokop, J." box="[1052,1273,1129,1156]" pageId="8" pageNumber="419" pagination="181 - 193" refId="ref16568" refString="Tkoc, M., Nel, A. & Prokop, J. (2016) Discovery of a new species of the Cretaceous genus Microphorites Hennig, 1971 (Diptera: Dolichopodidae s. lat.) in Paleogene amber from eastern Moravia (Czech Republic). Insect Systematics & Evolution, 47, 181 - 193. https: // doi. org / 10.1163 / 1876312 X- 47022139" type="journal article" year="2016">
|
||
Tkoč
|
||
<emphasis id="B933EAB3D14751370E7F517D32D92D8A" box="[1118,1169,1129,1155]" italics="true" pageId="8" pageNumber="419">et al</emphasis>
|
||
. (2016))
|
||
</bibRefCitation>
|
||
. However, the authors described somewhat different conditions of costal termination: costa ending at M
|
||
<subScript id="17C334E4D14751370EB751B732D62DA7" attach="left" box="[1174,1182,1184,1198]" fontSize="6" pageId="8" pageNumber="419">1</subScript>
|
||
—
|
||
<taxonomicName id="4C474D22D14751370E9B5199331A2DAE" authorityName="Penalver" authorityYear="2008" box="[1210,1362,1165,1192]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="B933EAB3D14751370E9B519932992DAE" box="[1210,1233,1166,1191]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370EC3519A331A2DAE" box="[1250,1362,1165,1191]" italics="true" pageId="8" pageNumber="419">utrillensis</emphasis>
|
||
</taxonomicName>
|
||
; costa ending between R
|
||
<subScript id="17C334E4D14751370B4051D337332DDB" attach="left" box="[353,379,1220,1234]" fontSize="6" pageId="8" pageNumber="419">4+5</subScript>
|
||
and M
|
||
<subScript id="17C334E4D14751370BE851D337992DDB" attach="left" box="[457,465,1220,1234]" fontSize="6" pageId="8" pageNumber="419">1</subScript>
|
||
—
|
||
<taxonomicName id="4C474D22D14751370BCF51A5343B2DC2" box="[494,627,1201,1228]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="B933EAB3D14751370BCF51A5344D2DC2" box="[494,517,1202,1227]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137083251A6343B2DC2" box="[531,627,1201,1227]" italics="true" pageId="8" pageNumber="419">deploegi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D147513708A051A535432DC2" authorityName="Ngo-Muller & Neal" authorityYear="2020" box="[641,779,1201,1228]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="pouilloni">
|
||
<emphasis id="B933EAB3D147513708A051A534D02DC2" box="[641,664,1202,1227]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137088751A635432DC2" box="[678,779,1201,1227]" italics="true" pageId="8" pageNumber="419">pouilloni</emphasis>
|
||
</taxonomicName>
|
||
; costa ending at R
|
||
<subScript id="17C334E4D147513709FA51D335BD2DDB" attach="left" box="[987,1013,1220,1234]" fontSize="6" pageId="8" pageNumber="419">4+5</subScript>
|
||
(or just beyond)—
|
||
<taxonomicName id="4C474D22D14751370EE851A5337F2DC2" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[1225,1335,1201,1228]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B933EAB3D14751370EE851A532A82DC2" box="[1225,1248,1202,1227]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370ECF51A6337F2DC2" box="[1262,1335,1201,1227]" italics="true" pageId="8" pageNumber="419">similis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D14751370F6451A5368C2DE6" authorityName="Grimaldi & Cumming" authorityYear="1999" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="oculeus">
|
||
<emphasis id="B933EAB3D14751370F6451A533142DC2" box="[1349,1372,1202,1227]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370F4B51A5368C2DE6" italics="true" pageId="8" pageNumber="419">oculeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D14751370AF551C1372B2DE6" authorityName="Perrichot & Engel" authorityYear="2014" box="[212,355,1237,1264]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="magaliae">
|
||
<emphasis id="B933EAB3D14751370AF551C136A32DE6" box="[212,235,1238,1263]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370ADA51C2372B2DE6" box="[251,355,1237,1263]" italics="true" pageId="8" pageNumber="419">magaliae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D14751370B5351C134462DE6" authorityName="Tkoc, Nel & Prokop" authorityYear="2016" box="[370,526,1238,1264]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B933EAB3D14751370B5351C137C12DE6" box="[370,393,1238,1263]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370BB851C134462DE6" box="[409,526,1238,1263]" italics="true" pageId="8" pageNumber="419">moravicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D1475137083C51C134CD2DE6" authorityName="Bramuzzo & Nel" authorityYear="2017" box="[541,645,1237,1264]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="erikai">
|
||
<emphasis id="B933EAB3D1475137083C51C1347C2DE6" box="[541,564,1238,1263]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137086551C234CD2DE6" box="[580,645,1237,1263]" italics="true" pageId="8" pageNumber="419">erikai</emphasis>
|
||
</taxonomicName>
|
||
. As far as we are aware, there are no similar examples of such broad variations in the costal termination among genera of empidoids possessing an incomplete costal vein.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14751370AE6500A32772CCE" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="419">
|
||
Although the costal vein extends around the entire wing margin in species of
|
||
<taxonomicName id="4C474D22D14751370E11500A32E02C3E" box="[1072,1192,1309,1335]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370E11500A32E02C3E" box="[1072,1192,1309,1335]" italics="true" pageId="8" pageNumber="419">Microphor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14751370EC1500A333C2C3E" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[1248,1396,1309,1335]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="419" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14751370EC1500A333C2C3E" box="[1248,1396,1309,1335]" italics="true" pageId="8" pageNumber="419">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
, its thickness is somewhat variable posteriorly. In
|
||
<taxonomicName id="4C474D22D147513708BE5056355F2C52" box="[671,791,1345,1371]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D147513708BE5056355F2C52" box="[671,791,1345,1371]" italics="true" pageId="8" pageNumber="419">Microphor</emphasis>
|
||
</taxonomicName>
|
||
, the costa becomes much thinner just beyond the apex of M
|
||
<subScript id="17C334E4D14751370A91506F36F02C8F" attach="left" box="[176,184,1400,1414]" fontSize="6" pageId="8" pageNumber="419">1</subScript>
|
||
(including extinct
|
||
<taxonomicName id="4C474D22D14751370BB25071345B2C76" baseAuthorityName="Meunier" baseAuthorityYear="1908" box="[403,531,1382,1408]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="rusticus">
|
||
<emphasis id="B933EAB3D14751370BB2507137E22C76" box="[403,426,1382,1407]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370B9B5071345B2C76" box="[442,531,1382,1407]" italics="true" pageId="8" pageNumber="419">rusticus</emphasis>
|
||
</taxonomicName>
|
||
). In
|
||
<taxonomicName id="4C474D22D1475137086C507234A92C76" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[589,737,1381,1407]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="419" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D1475137086C507234A92C76" box="[589,737,1381,1407]" italics="true" pageId="8" pageNumber="419">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
(including fossil species), the costa becomes thinner just beyond R
|
||
<subScript id="17C334E4D14751370AC8508B374B2CA3" attach="left" box="[233,259,1436,1450]" fontSize="6" pageId="8" pageNumber="419">4+5</subScript>
|
||
or, in some extant species, of nearly uniform thickness. It is somewhat curious but these features greatly resemble the distribution of costal termination in
|
||
<taxonomicName id="4C474D22D147513708E150BA35172CCE" authorityName="Hennig" authorityYear="1971" box="[704,863,1453,1479]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D147513708E150BA35172CCE" box="[704,863,1453,1479]" italics="true" pageId="8" pageNumber="419">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
summarised above.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14751370AE650C537CC2FCD" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="419">
|
||
A cladistic analysis of
|
||
<taxonomicName id="4C474D22D14751370BF250C6343A2CE2" authorityName="Hennig" authorityYear="1971" box="[467,626,1489,1515]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370BF250C6343A2CE2" box="[467,626,1489,1515]" italics="true" pageId="8" pageNumber="419">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
is beyond the scope of our paper. Nevertheless, very provisionally, a complex of species including
|
||
<taxonomicName id="4C474D22D14751370BC250E134202F06" box="[483,616,1525,1552]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="B933EAB3D14751370BC250E137B22F06" box="[483,506,1526,1551]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137082950E234202F06" box="[520,616,1525,1551]" italics="true" pageId="8" pageNumber="419">deploegi</emphasis>
|
||
</taxonomicName>
|
||
(Lower Cretaceous, uppermost Albian),
|
||
<taxonomicName id="4C474D22D14751370E1150E132F42F06" authorityName="Perrichot & Engel" authorityYear="2014" box="[1072,1212,1525,1552]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="magaliae">
|
||
<emphasis id="B933EAB3D14751370E1150E1320F2F06" box="[1072,1095,1526,1551]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370E7550E232F42F06" box="[1108,1212,1525,1551]" italics="true" pageId="8" pageNumber="419">magaliae</emphasis>
|
||
</taxonomicName>
|
||
(Upper Cretaceous, Cenomanian to Turonian) and
|
||
<taxonomicName id="4C474D22D14751370BD1530D34C32F3A" authorityName="Tkoc, Nel & Prokop" authorityYear="2016" box="[496,651,1562,1588]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B933EAB3D14751370BD1530D344F2F3A" box="[496,519,1562,1587]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370837530D34C32F3A" box="[534,651,1562,1587]" italics="true" pageId="8" pageNumber="419">moravicus</emphasis>
|
||
</taxonomicName>
|
||
(?Upper Cretaceous/Lower Paleogene) may be supported by the synapomorphic setation of the scutellum. These species share one pair of scutellar setae
|
||
<emphasis id="B933EAB3D14751370E1A532932CB2F5E" box="[1083,1155,1598,1623]" italics="true" pageId="8" pageNumber="419">versus</emphasis>
|
||
two pairs that are present in the oldest species of
|
||
<taxonomicName id="4C474D22D14751370B875376340D2F72" authorityName="Hennig" authorityYear="1971" box="[422,581,1633,1659]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370B875376340D2F72" box="[422,581,1633,1659]" italics="true" pageId="8" pageNumber="419">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName id="4C474D22D147513708E65375351A2F72" authorityName="Hennig" authorityYear="1971" box="[711,850,1634,1660]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D147513708E6537534962F72" box="[711,734,1634,1659]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D147513708CF5375351A2F72" box="[750,850,1634,1659]" italics="true" pageId="8" pageNumber="419">extinctus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D14751370940537535992F72" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[865,977,1633,1660]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B933EAB3D14751370940537535302F72" box="[865,888,1634,1659]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D147513709A9537635992F72" box="[904,977,1633,1659]" italics="true" pageId="8" pageNumber="419">similis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14751370E2A537532EA2F72" authorityName="Penalver" authorityYear="2008" box="[1035,1186,1633,1660]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="B933EAB3D14751370E2A5375326A2F72" box="[1035,1058,1634,1659]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370E13537632EA2F72" box="[1074,1186,1633,1659]" italics="true" pageId="8" pageNumber="419">utrillensis</emphasis>
|
||
</taxonomicName>
|
||
(unknown in
|
||
<taxonomicName id="4C474D22D14751370F625375368C2F96" authorityName="Grimaldi & Cumming" authorityYear="1999" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="oculeus">
|
||
<emphasis id="B933EAB3D14751370F62537533122F72" box="[1347,1370,1634,1659]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370F4B5375368C2F96" italics="true" pageId="8" pageNumber="419">oculeus</emphasis>
|
||
</taxonomicName>
|
||
)), as well as in potential outgroups. One pair of scutellar setae is present in some extant species of
|
||
<taxonomicName id="4C474D22D14751370F29539233D42F96" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[1288,1436,1669,1695]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="419" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14751370F29539233D42F96" box="[1288,1436,1669,1695]" italics="true" pageId="8" pageNumber="419">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
(two pairs in fossils).
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14751370AE653DA35A02EEA" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="419">
|
||
The postpedicel of
|
||
<taxonomicName id="4C474D22D14751370BBE53DA34762FEE" authorityName="Hennig" authorityYear="1971" box="[415,574,1741,1767]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370BBE53DA34762FEE" box="[415,574,1741,1767]" italics="true" pageId="8" pageNumber="419">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
varies considerably in shape and length. Although it is somewhat difficult to formalise it seems that the postpedicel of
|
||
<taxonomicName id="4C474D22D1475137084253E534992E02" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[611,721,1777,1804]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B933EAB3D1475137084253E534322E02" box="[611,634,1778,1803]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D147513708A953E634992E02" box="[648,721,1777,1803]" italics="true" pageId="8" pageNumber="419">similis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D1475137092753E535D22E02" authorityName="Penalver" authorityYear="2008" box="[774,922,1777,1804]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="B933EAB3D1475137092753E535552E02" box="[774,797,1778,1803]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137090B53E635D22E02" box="[810,922,1777,1803]" italics="true" pageId="8" pageNumber="419">utrillensis</emphasis>
|
||
</taxonomicName>
|
||
(as well as
|
||
<taxonomicName id="4C474D22D14751370E3A53E532EC2E02" authorityName="Hennig" authorityYear="1971" box="[1051,1188,1778,1804]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D14751370E3A53E5327A2E02" box="[1051,1074,1778,1803]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370E6153E532EC2E02" box="[1088,1188,1778,1803]" italics="true" pageId="8" pageNumber="419">extinctus</emphasis>
|
||
</taxonomicName>
|
||
) refers more or less to the plesiomorphic condition suggested by
|
||
<bibRefCitation id="EFD64B50D14751370852520235502E26" author="Chvala, M." box="[627,792,1813,1839]" pageId="8" pageNumber="419" pagination="1 - 279" refId="ref13639" refString="Chvala, M. (1983) The Empidoidea (Diptera) of Fennoscandia and Denmark. II. General Part. The families Hybotidae, Atelestidae and Microphoridae. Fauna Entomologica Scandinavica, 12, 1 - 279." type="journal article" year="1983">Chvála (1983)</bibRefCitation>
|
||
: moderately long conical postpedicel and at most equally long stylus. However, it should be noted that the postpedicel of microphorines is often not quite conical and bilaterally symmetrical, which can lead to differences in the shape depending on angle of view (e.g.,
|
||
<bibRefCitation id="EFD64B50D14751370E96524A332E2E71" author="Hennig, W." box="[1207,1382,1885,1912]" pageId="8" pageNumber="419" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig 1971: 5</bibRefCitation>
|
||
, figs 2, 3). In
|
||
<taxonomicName id="4C474D22D14751370ADA529537302E92" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[251,376,1921,1948]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="oculeus">
|
||
<emphasis id="B933EAB3D14751370ADA5295375A2E92" box="[251,274,1922,1947]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370B03529637302E92" box="[290,376,1921,1947]" italics="true" pageId="8" pageNumber="419">oculeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D14751370BA6529534452E92" box="[391,525,1921,1948]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="B933EAB3D14751370BA6529537D62E92" box="[391,414,1922,1947]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370B8C529634452E92" box="[429,525,1921,1947]" italics="true" pageId="8" pageNumber="419">deploegi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D1475137083C529534E42E92" authorityName="Perrichot & Engel" authorityYear="2014" box="[541,684,1921,1948]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="magaliae">
|
||
<emphasis id="B933EAB3D1475137083C5295347C2E92" box="[541,564,1922,1947]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14751370865529634E42E92" box="[580,684,1921,1947]" italics="true" pageId="8" pageNumber="419">magaliae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D147513708C4529535C92E92" authorityName="Tkoc, Nel & Prokop" authorityYear="2016" box="[741,897,1922,1948]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B933EAB3D147513708C4529534B42E92" box="[741,764,1922,1947]" italics="true" pageId="8" pageNumber="419">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1475137092D529535C92E92" box="[780,897,1922,1947]" italics="true" pageId="8" pageNumber="419">moravicus</emphasis>
|
||
</taxonomicName>
|
||
, the postpedicel is abruptly tapered (broadened at base and constricted distally). Similar transformations of the shape of the postpedicel are present in species of
|
||
<taxonomicName id="4C474D22D14751370AB652DE37472EEA" box="[151,271,1993,2019]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14751370AB652DE37472EEA" box="[151,271,1993,2019]" italics="true" pageId="8" pageNumber="419">Microphor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14751370B6752DE37922EEA" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[326,474,1993,2019]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="419" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14751370B6752DE37922EEA" box="[326,474,1993,2019]" italics="true" pageId="8" pageNumber="419">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
, as well as in some groups of
|
||
<taxonomicName id="4C474D22D1475137090D52DE35AC2EEA" authorityName="Chvála" authorityYear="1981" box="[812,996,1993,2019]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="subFamily" subFamily="Parathalassiinae">Parathalassiinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14651360AE6558032C02814" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="420">
|
||
It seems that
|
||
<taxonomicName id="4C474D22D14651360B415580342429B8" authorityName="Ngo-Muller & Neal" authorityYear="2020" box="[352,620,151,177]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="pouilloni">
|
||
<emphasis id="B933EAB3D14651360B415580342429B8" box="[352,620,151,177]" italics="true" pageId="9" pageNumber="420">Microphorites pouilloni</emphasis>
|
||
</taxonomicName>
|
||
from Burmese amber is placed somewhat aside among other species of the genus. Although
|
||
<taxonomicName id="4C474D22D14651360BA355AC344529DC" authorityName="Ngo-Muller & Neal" authorityYear="2020" box="[386,525,187,213]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="pouilloni">
|
||
<emphasis id="B933EAB3D14651360BA355AC37D129DD" box="[386,409,187,212]" italics="true" pageId="9" pageNumber="420">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14651360B8955AC344529DC" box="[424,525,187,213]" italics="true" pageId="9" pageNumber="420">pouilloni</emphasis>
|
||
</taxonomicName>
|
||
retains two pairs of scutellar setae, this species possesses a longer list of unique and potentially apomorphic features, including onion-shaped postpedicel, long basal section of M
|
||
<subScript id="17C334E4D14651360EF855E632A929F6" attach="left" box="[1241,1249,241,255]" fontSize="6" pageId="9" pageNumber="420">2</subScript>
|
||
(= m-cu), which is nearly as long as crossvein dm-m (= dm-cu), and some others (
|
||
<bibRefCitation id="EFD64B50D14651360950541432342814" author="Ngo-Muller, V. & Garrouste, R. & Pouillon, J. - M. & Nel, A." box="[881,1148,259,285]" pageId="9" pageNumber="420" pagination="1 - 4" refId="ref15786" refString="Ngo-Muller, V., Garrouste, R., Pouillon, J. - M. & Nel, A. (2020) A new species of the long-legged fly genus Microphorites in Burmese amber (Dolichopodidae: Microphorinae). Cretaceous Research, 107, 1 - 4. https: // doi. org / 10.1016 / j. cretres. 2019.104284" type="journal article" year="2020">
|
||
Ngô-Muller
|
||
<emphasis id="B933EAB3D14651360E21541432792814" box="[1024,1073,259,285]" italics="true" pageId="9" pageNumber="420">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14651360AE65430347A28A4" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="420">
|
||
Two species assigned to
|
||
<taxonomicName id="4C474D22D14651360BFB543034312848" authorityName="Hennig" authorityYear="1971" box="[474,633,295,321]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360BFB543034312848" box="[474,633,295,321]" italics="true" pageId="9" pageNumber="420">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C474D22D146513608A85430356B2849" authorityName="Tkoc, Nel & Prokop" authorityYear="2016" box="[649,803,295,321]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B933EAB3D146513608A8543034E82849" box="[649,672,295,320]" italics="true" pageId="9" pageNumber="420">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1465136088F5430356B2849" box="[686,803,295,320]" italics="true" pageId="9" pageNumber="420">moravicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14651360979543035F62848" authorityName="Bramuzzo & Nel" authorityYear="2017" box="[856,958,295,321]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="erikai">
|
||
<emphasis id="B933EAB3D14651360979543035272849" box="[856,879,295,320]" italics="true" pageId="9" pageNumber="420">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D1465136095C543035F62848" box="[893,958,295,321]" italics="true" pageId="9" pageNumber="420">erikai</emphasis>
|
||
</taxonomicName>
|
||
) share the wing with a brownish pterostigma.Amongst microphorines the pterostigma is present in
|
||
<taxonomicName id="4C474D22D14651360932545C35C3286C" box="[787,907,331,357]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360932545C35C3286C" box="[787,907,331,357]" italics="true" pageId="9" pageNumber="420">Microphor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C474D22D146513609B6545C3272286C" authorityName="Meunier" authorityYear="1908" box="[919,1082,331,357]" class="Insecta" family="Empididae" genus="Meghyperiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D146513609B6545C3272286C" box="[919,1082,331,357]" italics="true" pageId="9" pageNumber="420">Meghyperiella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14651360E4F545C3367286C" box="[1134,1327,331,357]" class="Insecta" family="Dolichopodidae" genus="Pristinmicrophor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360E4F545C3367286C" box="[1134,1327,331,357]" italics="true" pageId="9" pageNumber="420">Pristinmicrophor</emphasis>
|
||
</taxonomicName>
|
||
; in
|
||
<taxonomicName id="4C474D22D14651360F79545C36B82881" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="420" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14651360F79545C36B82881" italics="true" pageId="9" pageNumber="420">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
the pterostigma is usually present, but sometimes weakly so, or absent (in fossil species). The pterostigma is absent in
|
||
<taxonomicName id="4C474D22D14651360B3D548437EB28A4" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[284,419,403,429]" class="Insecta" family="Dolichopodidae" genus="Avenaphora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360B3D548437EB28A4" box="[284,419,403,429]" italics="true" pageId="9" pageNumber="420">Avenaphora</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14651360BFB5484346328A4" authorityName="Kaddumi" authorityYear="2005" box="[474,555,403,429]" class="Insecta" family="Chironomidae" genus="Curvus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360BFB5484346328A4" box="[474,555,403,429]" italics="true" pageId="9" pageNumber="420">Curvus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14651360AE654A035D42B68" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="420">
|
||
<taxonomicName id="4C474D22D14651360AE654A037AB28D9" authorityName="Tkoc, Nel & Prokop" authorityYear="2016" box="[199,483,439,465]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B933EAB3D14651360AE654A037AB28D9" box="[199,483,439,465]" italics="true" pageId="9" pageNumber="420">Microphorites moravicus</emphasis>
|
||
</taxonomicName>
|
||
is a distinct species with rather clear affinities to some Cretaceous species of
|
||
<taxonomicName id="4C474D22D14651360F7154A036BB28FC" authorityName="Hennig" authorityYear="1971" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360F7154A036BB28FC" italics="true" pageId="9" pageNumber="420">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
, as have been mentioned above.
|
||
<taxonomicName id="4C474D22D1465136084F54CC351F28FC" authorityName="Bramuzzo & Nel" authorityYear="2017" box="[622,855,475,501]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="erikai">
|
||
<emphasis id="B933EAB3D1465136084F54CC351F28FC" box="[622,855,475,501]" italics="true" pageId="9" pageNumber="420">Microphorites erikai</emphasis>
|
||
</taxonomicName>
|
||
resembles more an extant species of
|
||
<taxonomicName id="4C474D22D14651360F2554CC333428FC" box="[1284,1404,475,501]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360F2554CC333428FC" box="[1284,1404,475,501]" italics="true" pageId="9" pageNumber="420">Microphor</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C474D22D14651360AB654E837632B10" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[151,299,511,537]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="420" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14651360AB654E837632B10" box="[151,299,511,537]" italics="true" pageId="9" pageNumber="420">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
. It possesses even three pairs of scutellar setae that are usually present in
|
||
<taxonomicName id="4C474D22D14651360E4654E832972B10" box="[1127,1247,511,537]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360E4654E832972B10" box="[1127,1247,511,537]" italics="true" pageId="9" pageNumber="420">Microphor</emphasis>
|
||
</taxonomicName>
|
||
and, sometimes, in
|
||
<taxonomicName id="4C474D22D14651360A95573437002B34" baseAuthorityName="Brooks & Cumming" baseAuthorityYear="2022" box="[180,328,547,573]" class="Lecanoromycetes" family="Graphidaceae" genus="Schistostoma" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="420" phylum="Ascomycota" rank="genus">
|
||
<emphasis id="B933EAB3D14651360A95573437002B34" box="[180,328,547,573]" italics="true" pageId="9" pageNumber="420">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
. There are no characters suggesting closer relationships of
|
||
<taxonomicName id="4C474D22D146513609C0573432342B35" authorityName="Tkoc, Nel & Prokop" authorityYear="2016" box="[993,1148,547,573]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B933EAB3D146513609C0573435B02B35" box="[993,1016,547,572]" italics="true" pageId="9" pageNumber="420">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14651360E26573432342B35" box="[1031,1148,547,572]" italics="true" pageId="9" pageNumber="420">moravicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C474D22D14651360E93573433512B34" authorityName="Bramuzzo & Nel" authorityYear="2017" box="[1202,1305,547,573]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="erikai">
|
||
<emphasis id="B933EAB3D14651360E93573432812B35" box="[1202,1225,547,572]" italics="true" pageId="9" pageNumber="420">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14651360EF9573433512B34" box="[1240,1305,547,573]" italics="true" pageId="9" pageNumber="420">erikai</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, in these species the pterostigma has probably evolved independently.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14651360AE6577C35A12BF8" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="420">
|
||
To conclude, an assemblage of the species known nowadays as the genus
|
||
<taxonomicName id="4C474D22D14651360E39577C32FF2B8C" authorityName="Hennig" authorityYear="1971" box="[1048,1207,619,645]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360E39577C32FF2B8C" box="[1048,1207,619,645]" italics="true" pageId="9" pageNumber="420">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
cannot be precisely defined. The species assigned to
|
||
<taxonomicName id="4C474D22D14651360828579834E02BA0" authorityName="Hennig" authorityYear="1971" box="[521,680,655,681]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360828579834E02BA0" box="[521,680,655,681]" italics="true" pageId="9" pageNumber="420">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
can be separated into two groups. The first group includes only
|
||
<taxonomicName id="4C474D22D14651360F5F579836B32BC5" authorityName="Hennig" authorityYear="1971" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="B933EAB3D14651360F5F579833DD2BA1" box="[1406,1429,655,680]" italics="true" pageId="9" pageNumber="420">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14651360AB657A436B32BC5" box="[151,251,691,716]" italics="true" pageId="9" pageNumber="420">extinctus</emphasis>
|
||
</taxonomicName>
|
||
(the
|
||
<typeStatus id="54FC8803D14651360B1657A4372F2BC4" box="[311,359,691,717]" pageId="9" pageNumber="420">type</typeStatus>
|
||
species) and, therefore, it is the genus
|
||
<taxonomicName id="4C474D22D1465136090757A432432BC5" box="[806,1035,691,717]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="sensu">
|
||
<emphasis id="B933EAB3D1465136090757A432432BC5" box="[806,1035,691,717]" italics="true" pageId="9" pageNumber="420">Microphorites sensu</emphasis>
|
||
</taxonomicName>
|
||
Hennig’s original concept (
|
||
<bibRefCitation id="EFD64B50D14651360F6957A4369B2BF8" author="Hennig, W." pageId="9" pageNumber="420" pagination="1 - 28" refId="ref14414" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphorinae ") aus der unteren Kreide und aus dem baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig 1971</bibRefCitation>
|
||
). However, the question about monophyly of this genus remains open.
|
||
</paragraph>
|
||
<paragraph id="8BF836A1D14651360AE657EC32E82AC0" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="420">
|
||
The second group (
|
||
<taxonomicName id="4C474D22D14651360B8257EC345A2A1C" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[419,530,763,789]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B933EAB3D14651360B8257EC37F22A1D" box="[419,442,763,788]" italics="true" pageId="9" pageNumber="420">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14651360BE857EC345A2A1C" box="[457,530,763,789]" italics="true" pageId="9" pageNumber="420">similis</emphasis>
|
||
</taxonomicName>
|
||
species group) includes all other species assigned later to
|
||
<taxonomicName id="4C474D22D14651360E8957EC330F2A1C" authorityName="Hennig" authorityYear="1971" box="[1192,1351,763,789]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360E8957EC330F2A1C" box="[1192,1351,763,789]" italics="true" pageId="9" pageNumber="420">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
. Provisionally, this group could be supported by the apomorphic state of the costal vein (incomplete) and would deserve a generic level status. However, a curiously broad variation of the costal termination has been described. Since it is unclear if this pattern has phylogenetic significance, a misinterpretation of the costal termination by the authors, as one of the possible explanations, cannot be precluded. Therefore, we retained provisionally the
|
||
<taxonomicName id="4C474D22D14651360EC4569C331C2AAC" authorityName="Grimaldi & Cumming" authorityYear="1999" box="[1253,1364,907,933]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B933EAB3D14651360EC4569C32B42AAD" box="[1253,1276,907,932]" italics="true" pageId="9" pageNumber="420">M</emphasis>
|
||
.
|
||
<emphasis id="B933EAB3D14651360F2A569C331C2AAC" box="[1291,1364,907,933]" italics="true" pageId="9" pageNumber="420">similis</emphasis>
|
||
</taxonomicName>
|
||
group within
|
||
<taxonomicName id="4C474D22D14651360AC756B837CD2AC0" authorityName="Hennig" authorityYear="1971" box="[230,389,943,969]" class="Insecta" family="Dolichopodidae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EAB3D14651360AC756B837CD2AC0" box="[230,389,943,969]" italics="true" pageId="9" pageNumber="420">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
as an assemblage of the species with questionable systematic position.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |