136 lines
18 KiB
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136 lines
18 KiB
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<document id="D601BC751872588C89B4B392A8EE3A69" ID-DOI="10.5281/zenodo.186035" ID-GBIF-Dataset="8469ec5c-856b-4223-80d5-5dbe44ec5cd2" ID-ISSN="1175-5326" ID-Zenodo-Dep="186035" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460049915467" checkinUser="plazi" docAuthor="Kammerer, Christian F. & Angielczyk, Kenneth D." docDate="2009" docId="03EE87A896100F1EDBAFFBE3FC5AF9F8" docLanguage="en" docName="zt02018p024.pdf" docOrigin="Zootaxa 2018" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oudenodontidae Cope 1871" docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="FFD7FFD0961B0F15DB38FFF3F90FFF93" masterDocTitle="A proposed higher taxonomy of anomodont therapsids" masterLastPageNumber="24" masterPageNumber="1" pageNumber="12" updateTime="1698234516456" updateUser="plazi">
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<mods:mods id="2E0F5727064766B193178A5E02F23321" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A8689A2D2C5147924370A5D803593178">
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<mods:title id="4A9FAAC4C4B72D34DFB5358B2D789A11">A proposed higher taxonomy of anomodont therapsids</mods:title>
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<mods:name id="4A4DA794683500368F467445507FBBDE" type="personal">
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<mods:namePart id="8673F2C7D1F91283EFC42D5878165759">Kammerer, Christian F.</mods:namePart>
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<mods:namePart id="8C8889D34929D4EFE4A93C8559A5BF5D">Angielczyk, Kenneth D.</mods:namePart>
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<mods:typeOfResource id="42645F9CEA0A40C71B87F9AFB0CE303E">text</mods:typeOfResource>
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<mods:title id="0DF108CEA353D75E3F35294048367737">Zootaxa</mods:title>
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<mods:part id="9E7AEC3626A8417978DC07C23B23FBA2">
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<mods:date id="C47E19B258DDDC191B083AB6E3F4FC95">2009</mods:date>
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<mods:detail id="BD5700241ED2D66DB437D117A9F2E304" type="volume">
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<mods:number id="D17119B08AB1E018FA32E73513381706">2018</mods:number>
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<mods:identifier id="9B243ABF51E7EFFE361B4D4C50CF4D41" type="DOI">10.5281/zenodo.186035</mods:identifier>
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<mods:identifier id="C98EA981F5808E0A570440C45851E3BD" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B6B928EF87CC3CD5DCF07E0926D8750D" type="Zenodo-Dep">186035</mods:identifier>
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<treatment id="03EE87A896100F1EDBAFFBE3FC5AF9F8" ID-DOI="http://doi.org/10.5281/zenodo.6222280" ID-GBIF-Taxon="124874312" ID-Zenodo-Dep="6222280" LSID="urn:lsid:plazi:treatment:03EE87A896100F1EDBAFFBE3FC5AF9F8" httpUri="http://treatment.plazi.org/id/03EE87A896100F1EDBAFFBE3FC5AF9F8" lastPageNumber="12" pageId="11" pageNumber="12">
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<subSubSection id="C35D653596100F1EDBAFFBE3FA8BFB08" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8BF836BE96100F1EDBAFFBE3F8F7FBB9" blockId="11.[151,504,1040,1066]" box="[151,504,1040,1066]" pageId="11" pageNumber="12">
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<heading id="D0B081D296100F1EDBAFFBE3F8F7FBB9" bold="true" box="[151,504,1040,1066]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
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<taxonomicName id="4C474D3D96100F1EDBAFFBE3F8F7FBB9" authority="Cope, 1871" authorityName="Cope" authorityYear="1871" box="[151,504,1040,1066]" class="Reptilia" family="Oudenodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="family">
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<emphasis id="B933EAAC96100F1EDBAFFBE3F8F7FBB9" bold="true" box="[151,504,1040,1066]" pageId="11" pageNumber="12">
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Oudenodontidae
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<bibRefCitation id="EFD64B4F96100F1EDA56FBE3F8F7FBB9" author="Cope" box="[366,504,1040,1066]" pageId="11" pageNumber="17" refString="Cope, E. D. (1871) On the homologies of some of the cranial bones of the Reptilia, and on the systematic arrangement of the class. Proceedings of the American Association for the Advancement of Science, 19, 194 - 247." type="journal article" year="1871">Cope, 1871</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BF836BE96100F1EDBAFFBA9FA8BFB08" blockId="11.[151,1438,1114,1643]" pageId="11" pageNumber="12">
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<emphasis id="B933EAAC96100F1EDBAFFBA9F810FBE7" bold="true" box="[151,287,1114,1140]" pageId="11" pageNumber="12">Definition:</emphasis>
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All taxa more closely related to
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<taxonomicName id="4C474D3D96100F1ED99EFBA8FD17FBE7" authority="Owen, 1860" authorityName="Owen" authorityYear="1860" box="[678,1048,1114,1140]" class="Reptilia" family="Cryptodontidae" genus="Oudenodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="bainii">
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<emphasis id="B933EAAC96100F1ED99EFBA8FA72FBE7" box="[678,893,1115,1140]" italics="true" pageId="11" pageNumber="12">Oudenodon bainii</emphasis>
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<bibRefCitation id="EFD64B4F96100F1ED8BDFBA9FD17FBE7" author="Owen" box="[901,1048,1114,1140]" pageId="11" pageNumber="19" refString="Owen, R. (1860 a) On some reptilian fossils from South Africa. Quarterly Journal of the Geological Society of London, 16, 49 - 63." type="journal article" year="1860" yearSuffix="a">Owen, 1860</bibRefCitation>
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</taxonomicName>
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a, than
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<taxonomicName id="4C474D3D96100F1EDF55FBA8F8D6FB08" authority="(Owen, 1876) Seeley, 1898" authorityName="(Owen, 1876) Seeley" authorityYear="1898" class="Reptilia" family="Oudenodontidae" genus="Rhachiocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="magnus">
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<emphasis id="B933EAAC96100F1EDF55FBA8FC93FBE7" box="[1133,1436,1115,1140]" italics="true" pageId="11" pageNumber="12">Rhachiocephalus magnus</emphasis>
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(
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<bibRefCitation id="EFD64B4F96100F1EDBA7FB72F83EFB08" author="Owen" box="[159,305,1153,1179]" pageId="11" pageNumber="19" refString="Owen, R. (1876) Descriptive and Illustrated Catalogue of the Fossil Reptilia of South Africa in the Collection of the British Museum. Taylor and Francis, London, 88 pp." type="book" year="1876">Owen, 1876</bibRefCitation>
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)
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<bibRefCitation id="EFD64B4F96100F1EDA79FB72F8D6FB08" author="Seeley" box="[321,473,1153,1179]" pageId="11" pageNumber="20" refString="Seeley, H. G. (1898) On Oudenodon (Aulacocephalus) pithecops from the Dicynodon Beds of East London, Cape Colony. Geological Magazine, 4, 107 - 110." type="journal article" year="1898">Seeley, 1898</bibRefCitation>
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</taxonomicName>
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, or
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<taxonomicName id="4C474D3D96100F1ED93CFB72FA8FFB08" authority="Newton, 1893" authorityName="Newton" authorityYear="1893" box="[516,896,1153,1179]" class="Reptilia" family="Dicynodontidae" genus="Geikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="elginensis">
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<emphasis id="B933EAAC96100F1ED93CFB72FBC0FB09" box="[516,719,1153,1178]" italics="true" pageId="11" pageNumber="12">Geikia elginensis</emphasis>
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<bibRefCitation id="EFD64B4F96100F1ED9EFFB72FA8FFB08" author="Newton" box="[727,896,1153,1179]" pageId="11" pageNumber="19" refString="Newton, E. T. (1893) On some new reptiles from the Elgin Sandstone. Philosophical Transactions of the Royal Society of London B, 184, 431 - 503." type="journal article" year="1893">Newton, 1893</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D653596100F1EDBFEFB54FC5AF9F8" pageId="11" pageNumber="12" type="description">
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<paragraph id="8BF836BE96100F1EDBFEFB54F88DFA64" blockId="11.[151,1438,1114,1643]" pageId="11" pageNumber="12">
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<emphasis id="B933EAAC96100F1EDBFEFB54F8E1FB52" bold="true" box="[198,494,1191,1217]" pageId="11" pageNumber="12">Taxonomic comments:</emphasis>
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Recent research strongly supports a sister-group relationship between
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<taxonomicName id="4C474D3D96100F1EDBAFFB3CF8C7FB7B" authority="Owen, 1860" authorityName="Owen" authorityYear="1860" box="[151,456,1230,1256]" class="Reptilia" family="Cryptodontidae" genus="Oudenodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
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<emphasis id="B933EAAC96100F1EDBAFFB3CF829FB7B" box="[151,294,1231,1256]" italics="true" pageId="11" pageNumber="12">Oudenodon</emphasis>
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<bibRefCitation id="EFD64B4F96100F1EDA16FB3DF8C7FB7B" author="Owen" box="[302,456,1230,1256]" pageId="11" pageNumber="19" refString="Owen, R. (1860 a) On some reptilian fossils from South Africa. Quarterly Journal of the Geological Society of London, 16, 49 - 63." type="journal article" year="1860" yearSuffix="a">Owen, 1860</bibRefCitation>
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</taxonomicName>
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a, and
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<taxonomicName id="4C474D3D96100F1ED920FB3CFA65FB7B" authority="Seeley, 1889" authorityName="Seeley" authorityYear="1889" box="[536,874,1230,1256]" class="Reptilia" family="Oudenodontidae" genus="Tropidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
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<emphasis id="B933EAAC96100F1ED920FB3CFBCDFB7B" box="[536,706,1231,1256]" italics="true" pageId="11" pageNumber="12">Tropidostoma</emphasis>
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<bibRefCitation id="EFD64B4F96100F1ED9F3FB3DFA65FB7B" author="Seeley" box="[715,874,1230,1256]" pageId="11" pageNumber="20" refString="Seeley, H. G. (1889) Researches on the structure, organisation, and classification of the fossil Reptilia. - VI. On the anomodont Reptilia and their allies. Philosophical Transactions of the Royal Society of London B, 180, 215 - 296." type="journal article" year="1889">Seeley, 1889</bibRefCitation>
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</taxonomicName>
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(traditionally placed in separate family-level groups: Oudenodontinae van
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<bibRefCitation id="EFD64B4F96100F1EDACBFB06FB94FA9C" author="Hoepen" box="[499,667,1269,1295]" pageId="11" pageNumber="20" refString="van Hoepen, E. C. N. (1934) Oor die indeling van die Dicynodontidae na aanleiding van nuew vorme. Paleontologiese Navorsing van die Nasionale Museum, 2, 67 - 101." type="journal article" year="1934">Hoepen, 1934</bibRefCitation>
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, Oudenodontinae
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<bibRefCitation id="EFD64B4F96100F1ED84AFB06FD61FA9C" author="Cluver" box="[882,1134,1269,1295]" pageId="11" pageNumber="17" refString="Cluver, M. A. & King, G. M. (1983) A reassessment of the relationships of Permian Dicynodontia (Reptilia, Therapsida) and a new classification of dicynodonts. Annals of the South African Museum, 91, 195 - 273." type="journal article" year="1983">Cluver & King, 1983</bibRefCitation>
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[
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<emphasis id="B933EAAC96100F1EDFB8FB06FDAFFA9D" box="[1152,1184,1269,1294]" italics="true" pageId="11" pageNumber="12">sic</emphasis>
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], and Oudenodontini
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<bibRefCitation id="EFD64B4F96100F1EDBAFFAEFF821FAA5" author="King" box="[151,302,1308,1334]" pageId="11" pageNumber="18" refString="King, G. M. (1988) Anomodontia. Handbuch der Palaoherpetologie 17 C. Gustav Fischer Verlag, Stuttgart, 174 pp." type="book" year="1988">King, 1988</bibRefCitation>
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, versus Tropidostominae [
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<emphasis id="B933EAAC96100F1ED9B4FAEFFBBEFAA6" box="[652,689,1308,1333]" italics="true" pageId="11" pageNumber="12">sic</emphasis>
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, properly Tropidostomatinae]
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<bibRefCitation id="EFD64B4F96100F1EDF7DFAEFFC6FFAA5" author="Cluver" box="[1093,1376,1308,1334]" pageId="11" pageNumber="17" refString="Cluver, M. A. & King, G. M. (1983) A reassessment of the relationships of Permian Dicynodontia (Reptilia, Therapsida) and a new classification of dicynodonts. Annals of the South African Museum, 91, 195 - 273." type="journal article" year="1983">Cluver & King, 1983</bibRefCitation>
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, and Tropidostomini [
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<emphasis id="B933EAAC96100F1EDA65FAB0F872FACF" box="[349,381,1347,1372]" italics="true" pageId="11" pageNumber="12">sic</emphasis>
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]
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<bibRefCitation id="EFD64B4F96100F1EDAB7FAB1FB19FACF" author="King" box="[399,534,1346,1372]" pageId="11" pageNumber="18" refString="King, G. M. (1988) Anomodontia. Handbuch der Palaoherpetologie 17 C. Gustav Fischer Verlag, Stuttgart, 174 pp." type="book" year="1988">King, 1988</bibRefCitation>
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), to the extent that the generic distinction between these taxa has been called into question (
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<bibRefCitation id="EFD64B4F96100F1EDA7BFA9AFB4EFA10" author="Cluver" box="[323,577,1385,1411]" pageId="11" pageNumber="17" refString="Cluver, M. A. & King, G. M. (1983) A reassessment of the relationships of Permian Dicynodontia (Reptilia, Therapsida) and a new classification of dicynodonts. Annals of the South African Museum, 91, 195 - 273." type="journal article" year="1983">Cluver & King 1983</bibRefCitation>
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;
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<bibRefCitation id="EFD64B4F96100F1ED976FA9AFA9FFA10" author="Botha" box="[590,912,1385,1411]" pageId="11" pageNumber="16" refString="Botha, J. & Angielczyk, K. D. (2007) An integrative approach to distinguishing the Late Permian dicynodont species Oudenodon bainii and Tropidostoma microtrema (Therapsida: Anomodontia). Palaeontology, 50, 1175 - 1209." type="journal article" year="2007">Botha & Angielczyk 2007</bibRefCitation>
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). Here, we unite these taxa and the Russian genus
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<emphasis id="B933EAAC96100F1EDBDCFA63F8B9FA3A" box="[228,438,1424,1449]" italics="true" pageId="11" pageNumber="12">Australobarbarus</emphasis>
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<bibRefCitation id="EFD64B4F96100F1EDA87FA63FB6FFA39" author="Kurkin" box="[447,608,1424,1450]" pageId="11" pageNumber="18" refString="Kurkin, A. A. (2000) New dicynodonts from the Upper Permian of the Vyatka Basin. Paleontological Journal, 34, S 203 - S 210." type="journal article" year="2000">Kurkin, 2000</bibRefCitation>
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, (following
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<bibRefCitation id="EFD64B4F96100F1ED9CAFA63FD46FA39" author="Angielczyk" box="[754,1097,1424,1450]" pageId="11" pageNumber="16" refString="Angielczyk, K. D. & Kurkin, A. A. (2003 a) Phylogenetic analysis of Russian Permian dicynodonts (Therapsida: Anomodontia): implications for Permian biostratigraphy and Pangaean biogeography. Zoological Journal of the Linnean Society, 139, 157 - 212." type="journal article" year="2003" yearSuffix="a">Angielczyk & Kurkin 2003a</bibRefCitation>
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,
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<bibRefCitation id="EFD64B4F96100F1EDF6EFA63FDAEFA39" author="Angielczyk" box="[1110,1185,1424,1450]" pageId="11" pageNumber="16" refString="Angielczyk, K. D. & Kurkin, A. A. (2003 b) Has the utility of Dicynodon for Upper Permian terrestrial biostratigraphy been overstated? Geology, 31, 363 - 366." type="journal article" year="2003" yearSuffix="b">2003b</bibRefCitation>
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) in
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<taxonomicName id="4C474D3D96100F1EDFE9FA63FC98FA39" box="[1233,1431,1424,1450]" class="Reptilia" family="Oudenodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Oudenodontidae</taxonomicName>
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, based on the earliest available name for this group (
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<taxonomicName id="4C474D3D96100F1ED82AFA45FD7AFA43" authority="Cope, 1871" authorityName="Cope" authorityYear="1871" box="[786,1141,1462,1488]" class="Reptilia" family="Oudenodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="family">
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Oudenodontidae
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<bibRefCitation id="EFD64B4F96100F1ED8DDFA45FD7AFA43" author="Cope" box="[997,1141,1462,1488]" pageId="11" pageNumber="17" refString="Cope, E. D. (1871) On the homologies of some of the cranial bones of the Reptilia, and on the systematic arrangement of the class. Proceedings of the American Association for the Advancement of Science, 19, 194 - 247." type="journal article" year="1871">Cope, 1871</bibRefCitation>
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</taxonomicName>
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, validated under Article 12.2.4 of the Code).
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</paragraph>
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<paragraph id="8BF836BE96100F1EDBFEF9F0FC5AF9F8" blockId="11.[151,1438,1114,1643]" pageId="11" pageNumber="12">
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<emphasis id="B933EAAC96100F1EDBFEF9F0F8A0F98E" bold="true" box="[198,431,1539,1565]" pageId="11" pageNumber="12">Revised diagnosis:</emphasis>
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The analyses of Angielczyk (2007),
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<bibRefCitation id="EFD64B4F96100F1ED857F9F7FD3FF98D" author="Frobisch" box="[879,1072,1540,1566]" pageId="11" pageNumber="17" refString="Frobisch, J. (2007) The cranial anatomy of Kombuisia frerensis Hotton (Synapsida, Dicynodontia) and a new phylogeny of anomodont therapsids. Zoological Journal of the Linnean Society, 150, 117 - 144." type="journal article" year="2007">Fröbisch (2007)</bibRefCitation>
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, and
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<bibRefCitation id="EFD64B4F96100F1EDF4AF9F7FC93F98D" author="Frobisch" box="[1138,1436,1540,1566]" pageId="11" pageNumber="17" refString="Frobisch, J. & Reisz, R. R. (2008) A new species of Emydops (Synapsida, Anomodontia) and a discussion of dental variability and pathology in dicynodonts. Journal of Vertebrate Paleontology, 28, 770 - 787." type="journal article" year="2008">Fröbisch & Reisz (2008)</bibRefCitation>
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identified two autapomorphies diagnostic of
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<taxonomicName id="4C474D3D96100F1ED99DF9D9FA63F9D7" box="[677,876,1578,1604]" class="Reptilia" family="Oudenodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Oudenodontidae</taxonomicName>
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: (1) interpterygoid vacuity long and reaches the level of the palatal exposure of the palatines; (2) postparietal contributes to the intertemporal skull roof.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |