treatments-xml/data/03/EE/87/03EE8796FF99B505E3E1B7ECFD5452A0.xml
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<document id="FC3A53579DFC7CCFE7B06DBF233F8C6B" ID-CLB-Dataset="7585" ID-DOI="10.11646/zootaxa.4942.4.3" ID-GBIF-Dataset="ddf1c8ab-ddf2-476d-a319-63046f6d99fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="4612451" ID-ZooBank="38464B00-0721-4C27-98F9-599D862E4CB1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615992940915" checkinUser="plazi" docAuthor="Strümpher, Werner P. &amp; Stals, Riaan" docDate="2021" docId="03EE8796FF99B505E3E1B7ECFD5452A0" docLanguage="en" docName="zootaxa.4942.4.3.pdf" docOrigin="Zootaxa 4942 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phoberus cyrtus" docType="treatment" docVersion="7" lastPageNumber="549" masterDocId="FFD7FFEEFF9BB503E376B44EFF905649" masterDocTitle="Proper cleaning of keratin beetles reveals a cryptic species: Phoberus fumarius (Haaf, 1953) from southern Africa is reinstated as valid (Coleoptera: Trogidae)" masterLastPageNumber="557" masterPageNumber="543" pageNumber="545" updateTime="1698933609828" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="16176BE7842F7D30C272B2AA0ADA7C37">Proper cleaning of keratin beetles reveals a cryptic species: Phoberus fumarius (Haaf, 1953) from southern Africa is reinstated as valid (Coleoptera: Trogidae)</mods:title>
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<mods:name id="8A75EBCD792A62EC21EAE649588FC218" type="personal">
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<mods:namePart id="237DC091B7A58FED28AE78775CE78B3A">Strümpher, Werner P.</mods:namePart>
<mods:affiliation id="5ADBBD32747975C7E4ADD3E7FA4987FC">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa.</mods:affiliation>
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<mods:namePart id="2A8A3E32FB20D7E285FEDB7E23164404">Stals, Riaan</mods:namePart>
<mods:affiliation id="034F095F9E3C3C869BB00915B542BDC1">South African National Collection of Insects, Agricultural Research Council, Private Bag X 134, Queenswood 0121, Pretoria, South Africa.</mods:affiliation>
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<mods:title id="595541719B8F65EC147FB4ECB6DFAB5E">Zootaxa</mods:title>
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<mods:date id="D613043E55B8FD5EE9434510B9597226">2021</mods:date>
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<mods:number id="099E8884667EA2592E35C452F43F8889">2021-03-16</mods:number>
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<mods:identifier id="23DECDDFB3E52920727867F16B40F413" type="DOI">10.11646/zootaxa.4942.4.3</mods:identifier>
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<treatment id="03EE8796FF99B505E3E1B7ECFD5452A0" ID-DOI="http://doi.org/10.5281/zenodo.4644585" ID-GBIF-Taxon="180777882" ID-Zenodo-Dep="4644585" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0" httpUri="http://treatment.plazi.org/id/03EE8796FF99B505E3E1B7ECFD5452A0" lastPageId="6" lastPageNumber="549" pageId="2" pageNumber="545">
<subSubSection id="C35D650BFF99B501E3E1B7ECFE6D55F5" box="[151,509,930,957]" pageId="2" pageNumber="545" type="nomenclature">
<paragraph id="8BF83680FF99B501E3E1B7ECFE6D55F5" blockId="2.[151,509,930,993]" box="[151,509,930,957]" pageId="2" pageNumber="545">
<heading id="D0B081ECFF99B501E3E1B7ECFE6D55F5" bold="true" box="[151,509,930,957]" fontSize="11" level="1" pageId="2" pageNumber="545" reason="1">
<taxonomicName id="4C474D03FF99B501E3E1B7ECFE6D55F5" ID-CoL="4GJWC" authority="(Haaf, 1953)" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[151,509,930,957]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF99B501E3E1B7ECFE6D55F5" bold="true" box="[151,509,930,957]" pageId="2" pageNumber="545">
<emphasis id="B933EA92FF99B501E3E1B7ECFECB55F4" bold="true" box="[151,347,930,957]" italics="true" pageId="2" pageNumber="545">Phoberus cyrtus</emphasis>
(
<bibRefCitation id="EFD64B71FF99B501E21DB7ECFE6555F5" author="Haaf, E." box="[363,501,930,956]" pageId="2" pageNumber="545" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF99B501E3E1B789FEF055A8" box="[151,352,966,993]" pageId="2" pageNumber="545" type="description">
<paragraph id="8BF83680FF99B501E3E1B789FEF055A8" blockId="2.[151,509,930,993]" box="[151,352,966,993]" pageId="2" pageNumber="545">
(
<figureCitation id="137C2A05FF99B501E3E9B789FE9455A8" box="[159,260,967,993]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 47" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="5.[151,264,1943,1967]" captionTargetBox-0="[216,1385,195,1920]" captionTargetBox-1="[187,1416,192,1903]" captionTargetId-0="figure-31@4.[185,1403,181,1920]" captionTargetId-1="figure-17@5.[170,1416,181,1920]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 13. Holotype of Trox cyrtus Haaf, 1953 (MNHN). 1, habitus, dorsal view; 2, aedeagus, dorsal view; 3, holotype labels. Photographs by Antoine Mantilleri, copyright MNHN." captionText-1="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi-0="http://doi.org/10.5281/zenodo.4612453" figureDoi-1="http://doi.org/10.5281/zenodo.4612457" httpUri-0="https://zenodo.org/record/4612453/files/figure.png" httpUri-1="https://zenodo.org/record/4612457/files/figure.png" pageId="2" pageNumber="545">Figs 17</figureCitation>
,
<figureCitation id="137C2A05FF99B501E266B788FEBE55A8" box="[272,302,966,993]" captionStart="FIGURES 1516" captionStartId="10.[151,264,1442,1466]" captionTargetBox="[171,1416,181,1419]" captionTargetId="figure-31@10.[170,1416,181,1419]" captionTargetPageId="10" captionText="FIGURES 1516. Pronotal disc, dorsal view. 15, Phoberus cyrtus (Haaf, 1953); 16, Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612465" httpUri="https://zenodo.org/record/4612465/files/figure.png" pageId="2" pageNumber="545">15</figureCitation>
,
<figureCitation id="137C2A05FF99B501E24CB789FEC755A8" box="[314,343,967,993]" captionStart="FIGURE 21" captionStartId="12.[151,250,1140,1164]" captionTargetBox="[151,1434,181,1117]" captionTargetId="figure-31@12.[151,1435,181,1117]" captionTargetPageId="12" captionText="FIGURE 21. Known distribution of Phoberus cyrtus (Haaf, 1953) and Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612475" httpUri="https://zenodo.org/record/4612475/files/figure.png" pageId="2" pageNumber="545">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF99B501E3E1B049FBD15233" pageId="2" pageNumber="545" type="reference_group">
<paragraph id="8BF83680FF99B501E3E1B049FD085257" blockId="2.[151,1361,1030,1146]" box="[151,664,1030,1054]" pageId="2" pageNumber="545">
<treatmentCitationGroup id="AB5711AEFF99B501E3E1B049FD085257" box="[151,664,1030,1054]" pageId="2" pageNumber="545">
<treatmentCitation id="0AE61091FF99B501E3E1B049FE235257" author="Haaf, E." box="[151,435,1030,1054]" page="333" pageId="2" pageNumber="545" year="1953">
<taxonomicName id="4C474D03FF99B501E3E1B049FE235257" ID-CoL="59DJJ" authority="Haaf, 1953: 333" authorityName="Haaf" authorityPageNumber="333" authorityYear="1953" box="[151,435,1030,1054]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF99B501E3E1B049FE995257" box="[151,265,1031,1054]" italics="true" pageId="2" pageNumber="545">Trox cyrtus</emphasis>
<bibRefCitation id="EFD64B71FF99B501E279B048FE235257" author="Haaf, E." box="[271,435,1030,1054]" pageId="2" pageNumber="545" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953: 333</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF83680FF99B501E3E1B06BFAC15275" blockId="2.[151,1361,1030,1146]" box="[151,1361,1060,1085]" pageId="2" pageNumber="545">
<treatmentCitationGroup id="AB5711AEFF99B501E3E1B06BFAC15275" box="[151,1361,1060,1085]" pageId="2" pageNumber="545">
<treatmentCitation id="0AE61091FF99B501E3E1B06BFD855275" author="Scholtz, C. H." box="[151,533,1060,1085]" page="66" pageId="2" pageNumber="545" year="1980">
<taxonomicName id="4C474D03FF99B501E3E1B06BFD855275" ID-CoL="59DJJ" authority=": Scholtz 1980 a: 66" authorityName="Scholtz" authorityPageNumber="66" authorityYear="1980" box="[151,533,1060,1085]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="cyrtus" subGenus="Trox">
<emphasis id="B933EA92FF99B501E3E1B06BFF555275" box="[151,197,1061,1084]" italics="true" pageId="2" pageNumber="545">Trox</emphasis>
(
<emphasis id="B933EA92FF99B501E3A5B06BFE915275" box="[211,257,1061,1084]" italics="true" pageId="2" pageNumber="545">Trox</emphasis>
)
<emphasis id="B933EA92FF99B501E266B06BFEDE5275" box="[272,334,1061,1084]" italics="true" pageId="2" pageNumber="545">cyrtus</emphasis>
:
<bibRefCitation id="EFD64B71FF99B501E22DB06AFD855275" author="Scholtz, C. H." box="[347,533,1060,1085]" pageId="2" pageNumber="545" pagination="1 - 104" refId="ref8073" refString="Scholtz, C. H. (1980 a) Monograph of the genus Trox F. (Coleoptera: Trogidae) of subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz 1980a: 66</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 68, 87, 97 (redescription, illustrations, distribution);
<treatmentCitation id="0AE61091FF99B501E745B06AFB425274" author="Scholtz, C. H." box="[1075,1234,1060,1085]" page="3" pageId="2" pageNumber="545" year="1982">
<bibRefCitation id="EFD64B71FF99B501E745B06AFB425274" author="Scholtz, C. H." box="[1075,1234,1060,1085]" pageId="2" pageNumber="545" pagination="1 - 27" refId="ref8146" refString="Scholtz, C. H. (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries, Republic of South Africa, 54, 1 - 27." type="journal article" year="1982">Scholtz 1982: 3</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF83680FF99B501E3E1B00AFD055212" blockId="2.[151,1361,1030,1146]" box="[151,661,1091,1115]" pageId="2" pageNumber="545">
<treatmentCitationGroup id="AB5711AEFF99B501E3E1B00AFD055212" box="[151,661,1091,1115]" pageId="2" pageNumber="545">
<treatmentCitation id="0AE61091FF99B501E3E1B00AFD8D5212" author="Zidek, J." box="[151,541,1091,1115]" page="9" pageId="2" pageNumber="545" year="2013">
<taxonomicName id="4C474D03FF99B501E3E1B00AFD8D5212" ID-CoL="59DJJ" authority=": Zidek 2013: 9" authorityName="Zidek" authorityPageNumber="9" authorityYear="2013" box="[151,541,1091,1115]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="cyrtus" subGenus="Phoberus">
<emphasis id="B933EA92FF99B501E3E1B00AFF555212" box="[151,197,1092,1115]" italics="true" pageId="2" pageNumber="545">Trox</emphasis>
(
<emphasis id="B933EA92FF99B501E3A5B00DFEA45212" box="[211,308,1091,1115]" italics="true" pageId="2" pageNumber="545">Phoberus</emphasis>
)
<emphasis id="B933EA92FF99B501E235B00AFE115212" box="[323,385,1092,1115]" italics="true" pageId="2" pageNumber="545">cyrtus</emphasis>
:
<bibRefCitation id="EFD64B71FF99B501E2F9B00DFD8D5212" author="Zidek, J." box="[399,541,1091,1115]" pageId="2" pageNumber="545" pagination="1 - 38" refId="ref8671" refString="Zidek, J. (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi, 314, 1 - 38." type="journal article" year="2013">Zidek 2013: 9</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF83680FF99B501E3E1B02CFBD15233" blockId="2.[151,1361,1030,1146]" box="[151,1089,1122,1146]" pageId="2" pageNumber="545">
<treatmentCitationGroup id="AB5711AEFF99B501E3E1B02CFBD15233" box="[151,1089,1122,1146]" pageId="2" pageNumber="545">
<treatmentCitation id="0AE61091FF99B501E3E1B02CFDCA5233" author="Strumpher, W. P. &amp; Villet, M. H. &amp; Sole, C. L. &amp; Scholtz, C. H." box="[151,602,1122,1146]" page="78" pageId="2" pageNumber="545" year="2016">
<taxonomicName id="4C474D03FF99B501E3E1B02CFDCA5233" ID-CoL="4GJWC" authority=": Strumpher et al. 2016 b: 78" authorityName="Strumpher" authorityPageNumber="78" authorityYear="2016" box="[151,602,1122,1146]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF99B501E3E1B02CFEAD5233" box="[151,317,1122,1146]" italics="true" pageId="2" pageNumber="545">Phoberus cyrtus</emphasis>
:
<bibRefCitation id="EFD64B71FF99B501E23CB02CFDCA5233" author="Strumpher, W. P. &amp; Villet, M. H. &amp; Sole, C. L. &amp; Scholtz, C. H." box="[330,602,1122,1146]" pageId="2" pageNumber="545" pagination="53 - 82" refId="ref8543" refString="Strumpher, W. P., Villet, M. H., Sole, C. L. &amp; Scholtz, C. H. (2016 b) Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics &amp; Evolution, 47 (1), 53 - 82. https: // doi. org / 10.1163 / 1876312 X- 46052133" type="journal article" year="2016">
Strümpher
<emphasis id="B933EA92FF99B501E2CDB02DFE7E5233" box="[443,494,1122,1146]" italics="true" pageId="2" pageNumber="545">et al.</emphasis>
2016b: 78
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(new combination);
<treatmentCitation id="0AE61091FF99B501E058B02CFC585233" author="Zidek, J." box="[814,968,1122,1146]" page="99" pageId="2" pageNumber="545" year="2017">
<bibRefCitation id="EFD64B71FF99B501E058B02CFC585233" author="Zidek, J." box="[814,968,1122,1146]" pageId="2" pageNumber="545" pagination="93 - 127" refId="ref8699" refString="Zidek, J. (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana, Series A, 25 (1), 93 - 127." type="journal article" year="2017">Zídek 2017: 99</bibRefCitation>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF99B500E3E1B0EDFC5A572C" lastPageId="3" lastPageNumber="546" pageId="2" pageNumber="545" type="materials_examined">
<paragraph id="8BF83680FF99B501E3E1B0EDFED052AB" blockId="2.[151,1437,1186,2041]" pageId="2" pageNumber="545">
<materialsCitation id="3B2F3CDDFF99B501E3E1B0EDFED052AB" ID-GBIF-Occurrence="3062300334" collectorName="Vallee du Haute Orange" country="Lesotho" location="Districts" pageId="2" pageNumber="545" specimenCount="1" stateProvince="Mohale's Hoek" typeStatus="holotype">
<emphasis id="B933EA92FF99B501E3E1B0EDFEA652F5" bold="true" box="[151,310,1186,1213]" pageId="2" pageNumber="545">
<typeStatus id="54FC8822FF99B501E3E1B0EDFF4252F4" box="[151,210,1187,1213]" pageId="2" pageNumber="545">Type</typeStatus>
locality.
</emphasis>
<collectorName id="26B25356FF99B501E24BB0EDFDDA52F4" box="[317,586,1187,1213]" pageId="2" pageNumber="545">Vallée du Haute Orange</collectorName>
[Valley of high Orange River] (border of
<collectingRegion id="4983F862FF99B501E753B0EDFB4152F4" box="[1061,1233,1187,1213]" country="Lesotho" name="Mohale's Hoek" pageId="2" pageNumber="545">Mohales Hoek</collectingRegion>
and
<collectingRegion id="4983F862FF99B501E67FB0EDFAF552F4" box="[1289,1381,1187,1213]" country="Lesotho" name="Quthing" pageId="2" pageNumber="545">Quthing</collectingRegion>
<location id="8E98605BFF99B501E61AB0EDFF4252A8" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF99B501E61AB0EDFF4252A8" country="Lesotho" name="Districts" pageId="2" pageNumber="545" stateProvince="Mohale's Hoek">Districts</location>
,
<collectingCountry id="F3507610FF99B501E3ABB089FEA952AB" box="[221,313,1223,1250]" name="Lesotho" pageId="2" pageNumber="545">Lesotho</collectingCountry>
)
</materialsCitation>
</paragraph>
<paragraph id="8BF83680FF99B501E3B1B0A5FA8B5024" blockId="2.[151,1437,1186,2041]" pageId="2" pageNumber="545">
<emphasis id="B933EA92FF99B501E3B1B0A5FE67534D" bold="true" box="[199,503,1258,1285]" pageId="2" pageNumber="545">Type material examined.</emphasis>
All type specimens were examined through detailed photographs.
<materialsCitation id="3B2F3CDDFF99B501E670B0A5FB4453DC" ID-GBIF-Occurrence="3062300332" collectionCode="MNHN" collectorName="Rectangle &amp; Rectangular" country="Lesotho" determinerName="Dr. E." elevation="1500" latitude="-30.283333" location="Rectangle" longLatPrecision="1222" longitude="27.766666" municipality="Rectangle" pageId="2" pageNumber="545" specimenCode="EC10025" specimenCount="2" typeStatus="holotype">
<emphasis id="B933EA92FF99B501E670B0A5FAEC534A" bold="true" box="[1286,1404,1259,1285]" pageId="2" pageNumber="545">
<typeStatus id="54FC8822FF99B501E670B0A5FAEC534A" box="[1286,1404,1259,1285]" pageId="2" pageNumber="545" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
Ƌ (
<collectionCode id="ED56AE45FF99B501E3E9B141FF695360" box="[159,249,1295,1321]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
, aedeagus extracted) (
<figureCitation id="137C2A05FF99B501E287B141FDC75360" box="[497,599,1295,1321]" captionStart="FIGURES 13" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[216,1385,195,1920]" captionTargetId="figure-31@4.[185,1403,181,1920]" captionTargetPageId="4" captionText="FIGURES 13. Holotype of Trox cyrtus Haaf, 1953 (MNHN). 1, habitus, dorsal view; 2, aedeagus, dorsal view; 3, holotype labels. Photographs by Antoine Mantilleri, copyright MNHN." figureDoi="http://doi.org/10.5281/zenodo.4612453" httpUri="https://zenodo.org/record/4612453/files/figure.png" pageId="2" pageNumber="545">Figs 13</figureCitation>
):
<location id="8E98605BFF99B501E11AB141FD4F5360" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF99B501E11AB141FD4F5360" box="[620,735,1295,1321]" country="Lesotho" latitude="-30.283333" longLatPrecision="1222" longitude="27.766666" municipality="Rectangle" name="Rectangle" pageId="2" pageNumber="545">Rectangle</location>
, faded blue cardstock: “MUSEUM PARIS | Basoutoland, Lessouto [
<collectingCountry id="F3507610FF99B501E395B17CFED35304" box="[227,323,1330,1357]" name="Lesotho" pageId="2" pageNumber="545">Lesotho</collectingCountry>
] | Vallée du
<superScript id="7C329BC8FF99B501E2A5B17DFE7C5308" attach="left" box="[467,492,1331,1357]" fontSize="6" pageId="2" pageNumber="545">Ht</superScript>
Orange [Valley of high Orange River,
<geoCoordinate id="EE735047FF99B501E0DEB17AFB915306" box="[936,1025,1332,1359]" degrees="30" direction="south" minutes="17" orientation="latitude" pageId="2" pageNumber="545" precision="925" value="-30.283333">30°17S</geoCoordinate>
<geoCoordinate id="EE735047FF99B501E77CB17AFBF95306" box="[1034,1129,1332,1359]" degrees="27" direction="east" minutes="46" orientation="longitude" pageId="2" pageNumber="545" precision="925" value="27.766666">27°46E</geoCoordinate>
],
<quantity id="4CBF9B65FF99B501E70BB17CFB475304" box="[1149,1239,1330,1357]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="545" unit="m" value="1500.0">
<elevation id="006AD1B3FF99B501E70BB17CFB475304" box="[1149,1239,1330,1357]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="545" unit="m" value="1500.0">1500 m</elevation>
</quantity>
. | (legit R. Ellenberger) | E. HAUG 1906” ||
<collectingMunicipality id="6B9CACFAFF99B501E2A3B119FDD85338" box="[469,584,1367,1393]" pageId="2" pageNumber="545">Rectangle</collectingMunicipality>
, red cardstock: “
<typeStatus id="54FC8822FF99B501E072B119FC0E5338" box="[772,926,1367,1393]" pageId="2" pageNumber="545" type="holotype">HOLOTYPE</typeStatus>
” ||
<collectorName id="26B25356FF99B501E0B4B119FBA55338" box="[962,1077,1367,1393]" pageId="2" pageNumber="545">Rectangle</collectorName>
, minimally foxing white: “
<taxonomicName id="4C474D03FF99B501E612B119FA0C5338" authorityName="Fabricius" authorityYear="1775" box="[1380,1436,1367,1393]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="genus">Trox</taxonomicName>
|
<taxonomicName id="4C474D03FF99B501E3D0B135FEB953DC" authority="Haaf" authorityName="Haaf" box="[166,297,1403,1429]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="cyrtus">cyrtus Haaf</taxonomicName>
| det.
<determinerName id="7E24DA44FF99B501E21FB135FE3253DC" box="[361,418,1403,1429]" pageId="2" pageNumber="545">Dr.E.</determinerName>
<bibRefCitation id="EFD64B71FF99B501E2D7B135FD8F53DC" author="Haaf, E." box="[417,543,1402,1429]" pageId="2" pageNumber="545" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf 1953</bibRefCitation>
” ||
<collectorName id="26B25356FF99B501E131B135FD5E53DC" box="[583,718,1403,1429]" pageId="2" pageNumber="545">Rectangular</collectorName>
curatorial label, white: “
<collectionCode id="ED56AE45FF99B501E09CB135FBD353DC" box="[1002,1091,1403,1429]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
|
<specimenCode id="DBE19EFBFF99B501E72CB134FB5A53DC" box="[1114,1226,1402,1429]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">EC10025</specimenCode>
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF99B501E796B135FCD25394" ID-GBIF-Occurrence="3062300309" collectionCode="MNHN" collectorName="Foxy &amp; White" country="Lesotho" location="Blue" municipality="Rectangle" pageId="2" pageNumber="545" specimenCode="EC10027, EC10029" specimenCount="2" typeStatus="paratype">
<emphasis id="B933EA92FF99B501E796B135FACC53DA" bold="true" box="[1248,1372,1403,1429]" pageId="2" pageNumber="545">
<typeStatus id="54FC8822FF99B501E796B135FACC53DA" box="[1248,1372,1403,1429]" pageId="2" pageNumber="545" type="paratype">PARATYPES</typeStatus>
</emphasis>
2ƋƋ (
<collectionCode id="ED56AE45FF99B501E3E9B1D1FF6C53F0" box="[159,252,1439,1465]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
):
<location id="8E98605BFF99B501E278B1D1FED353F0" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF99B501E278B1D1FED353F0" box="[270,323,1439,1465]" country="Lesotho" municipality="Rectangle" name="Blue" pageId="2" pageNumber="545">Blue</location>
label identical ||
<collectingMunicipality id="6B9CACFAFF99B501E289B1D1FDE253F0" box="[511,626,1439,1465]" pageId="2" pageNumber="545">Rectangle</collectingMunicipality>
, red cardstock: “
<typeStatus id="54FC8822FF99B501E05CB1D1FC2C53F0" box="[810,956,1439,1465]" pageId="2" pageNumber="545" type="paratype">PARATYPE</typeStatus>
” ||
<collectorName id="26B25356FF99B501E0ABB1D1FB8753F0" box="[989,1047,1439,1465]" pageId="2" pageNumber="545">Foxy</collectorName>
white label identical ||
<collectorName id="26B25356FF99B501E66EB1D1FACC53F0" box="[1304,1372,1439,1465]" pageId="2" pageNumber="545">White</collectorName>
curatorial labels: “
<collectionCode id="ED56AE45FF99B501E243B18DFE1E5394" box="[309,398,1475,1501]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
|
<specimenCode id="DBE19EFBFF99B501E2D5B18DFD9F5394" box="[419,527,1475,1501]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">EC10027</specimenCode>
” and “
<collectionCode id="ED56AE45FF99B501E12DB18DFD245394" box="[603,692,1475,1501]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
|
<specimenCode id="DBE19EFBFF99B501E1BFB18DFCA85394" box="[713,824,1475,1501]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">EC10029</specimenCode>
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF99B501E038B18DFF3E5048" ID-GBIF-Occurrence="3062300301" collectionCode="MNHN" collectorName="Labels" country="Lesotho" pageId="2" pageNumber="545" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="B933EA92FF99B501E038B18DFC595392" bold="true" box="[846,969,1475,1501]" pageId="2" pageNumber="545">
<typeStatus id="54FC8822FF99B501E038B18DFC595392" box="[846,969,1475,1501]" pageId="2" pageNumber="545" type="paratype">PARATYPES</typeStatus>
</emphasis>
<specimenCount id="9D41FD09FF99B501E0A7B18DFB995396" box="[977,1033,1475,1503]" pageId="2" pageNumber="545" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED56AE45FF99B501E76EB18DFBE55394" box="[1048,1141,1475,1501]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
):
<collectorName id="26B25356FF99B501E7FEB18DFB435394" box="[1160,1235,1475,1501]" pageId="2" pageNumber="545">Labels</collectorName>
identical to those of
</materialsCitation>
<materialsCitation id="3B2F3CDDFF99B501E3A3B1A9FDC85000" ID-GBIF-Occurrence="3062300336" collectionCode="MNHN" country="Lesotho" pageId="2" pageNumber="545" specimenCode="EC10028, EC10026" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54FC8822FF99B501E3A3B1A9FED35048" box="[213,323,1511,1537]" pageId="2" pageNumber="545" type="paratype">paratypes</typeStatus>
, but each specimen with an additional foxy white square: “
<specimenCount id="9D41FD09FF99B501E098B1A9FB905048" box="[1006,1024,1511,1537]" pageId="2" pageNumber="545" type="female"></specimenCount>
” || “
<collectionCode id="ED56AE45FF99B501E743B1A9FB1E5048" box="[1077,1166,1511,1537]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
|
<specimenCode id="DBE19EFBFF99B501E7D3B1A9FA845048" box="[1189,1300,1511,1537]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">EC10028</specimenCode>
”; “
<collectionCode id="ED56AE45FF99B501E643B1A9FA1E5048" box="[1333,1422,1511,1537]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
|
<specimenCode id="DBE19EFBFF99B501E3E1B245FE94506C" box="[151,260,1547,1573]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">EC10026</specimenCode>
” with red label: “
<typeStatus id="54FC8822FF99B501E2A6B245FDFA506C" box="[464,618,1547,1573]" pageId="2" pageNumber="545" type="allotype">ALLOTYPE</typeStatus>
”; “
<collectionCode id="ED56AE45FF99B501E1FCB245FD73506C" box="[650,739,1547,1573]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
| 10030” with different determination label, white rectangle: “
<typeStatus id="54FC8822FF99B501E3D6B261FEBF5000" box="[160,303,1583,1609]" pageId="2" pageNumber="545" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="4C474D03FF99B501E236B261FEE25001" authorityName="Fabricius" authorityYear="1775" box="[320,370,1583,1608]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="genus">
<emphasis id="B933EA92FF99B501E236B261FEE25001" box="[320,370,1583,1608]" italics="true" pageId="2" pageNumber="545">Trox</emphasis>
</taxonomicName>
|
<taxonomicName id="4C474D03FF99B501E2F5B261FDDD5000" authority="Haaf, 1953" authorityName="Haaf" authorityYear="1953" box="[387,589,1582,1609]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF99B501E2F5B261FE575001" box="[387,455,1583,1608]" italics="true" pageId="2" pageNumber="545">cyrtus</emphasis>
<bibRefCitation id="EFD64B71FF99B501E2BBB261FDDD5000" author="Haaf, E." box="[461,589,1582,1609]" pageId="2" pageNumber="545" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF99B501E117B261FA885024" ID-GBIF-Occurrence="3062300335" collectionCode="MNHB" country="Lesotho" determinerName="Dr. E." location="Blue" municipality="Rectangle" pageId="2" pageNumber="545" specimenCount="2" typeStatus="paratype">
<emphasis id="B933EA92FF99B501E117B261FD42500E" bold="true" box="[609,722,1583,1609]" pageId="2" pageNumber="545">
<typeStatus id="54FC8822FF99B501E117B261FD42500E" box="[609,722,1583,1609]" pageId="2" pageNumber="545" type="paratype">PARATYPE</typeStatus>
</emphasis>
1Ƌ (
<collectionCode id="ED56AE45FF99B501E07EB261FCF35000" box="[776,867,1583,1609]" pageId="2" pageNumber="545">MNHB</collectionCode>
):
<location id="8E98605BFF99B501E002B261FC395000" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF99B501E002B261FC395000" box="[884,937,1583,1609]" country="Lesotho" municipality="Rectangle" name="Blue" pageId="2" pageNumber="545">Blue</location>
label identical ||
<collectingMunicipality id="6B9CACFAFF99B501E714B261FB455000" box="[1122,1237,1583,1609]" pageId="2" pageNumber="545">Rectangle</collectingMunicipality>
, white, preprinted red text interspersed with black handwriting: “
<typeStatus id="54FC8822FF99B501E1EBB21DFCAE5024" box="[669,830,1619,1645]" pageId="2" pageNumber="545" type="paratype">PARATYPUS</typeStatus>
|
<taxonomicName id="4C474D03FF99B501E027B21DFC165024" authorityName="Fabricius" authorityYear="1775" box="[849,902,1619,1645]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="545" phylum="Arthropoda" rank="genus">Trox</taxonomicName>
| cyrtus
<taxonomicNameLabel id="A20057E9FF99B501E093B21DFB8A5024" box="[997,1050,1619,1645]" pageId="2" pageNumber="545" rank="species">sp.n.</taxonomicNameLabel>
| det.
<determinerName id="7E24DA44FF99B501E720B21DFB1E5024" box="[1110,1166,1619,1645]" pageId="2" pageNumber="545">Dr.E.</determinerName>
<bibRefCitation id="EFD64B71FF99B501E7F8B21DFA9D5024" author="Haaf, E." box="[1166,1293,1618,1645]" pageId="2" pageNumber="545" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf 1953</bibRefCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF83680FF99B501E3B1B239FC6D5090" blockId="2.[151,1437,1186,2041]" pageId="2" pageNumber="545">
<bibRefCitation id="EFD64B71FF99B501E3B1B239FEC050D8" author="Haaf, E." box="[199,336,1654,1681]" pageId="2" pageNumber="545" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf (1953)</bibRefCitation>
described this species from six males and four females, all with the same label data. We studied four males and three females of the type series, including the
<typeStatus id="54FC8822FF99B501E049B2D5FC3550FC" box="[831,933,1691,1717]" pageId="2" pageNumber="545" type="holotype">holotype</typeStatus>
.
<bibRefCitation id="EFD64B71FF99B501E0D8B2D5FBA550FC" author="Haaf, E." box="[942,1077,1690,1717]" pageId="2" pageNumber="545" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf (1953)</bibRefCitation>
indicated that all ten type specimens would be lodged in the
<collectionCode id="ED56AE45FF99B501E297B2F1FDA95090" box="[481,569,1727,1753]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="545">MNHN</collectionCode>
, but we located one male in the MNHB.
</paragraph>
<paragraph id="8BF83680FF99B500E3B1B2ACFA0D56F8" blockId="2.[151,1437,1186,2041]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="546" pageId="2" pageNumber="545">
<emphasis id="B933EA92FF99B501E3B1B2ACFDAD50B5" bold="true" box="[199,573,1762,1788]" pageId="2" pageNumber="545">Additional material examined.</emphasis>
<materialsCitation id="3B2F3CDDFF99B501E13EB2ACFD4D5168" ID-GBIF-Occurrence="3062300318" collectingDate="1971-02-15" collectingDateMax="1971-02-25" collectingDateMin="1971-02-15" collectionCode="TMSA" collectorName="Snyman &amp; Jones" country="South Africa" latitude="-30.7" location="Aliwal North" longLatPrecision="1220" longitude="26.7" pageId="2" pageNumber="545" specimenCount="1" stateProvince="Eastern Cape">
<emphasis id="B933EA92FF99B501E13EB2ACFBC650B5" bold="true" box="[584,1110,1762,1789]" pageId="2" pageNumber="545">
<collectingCountry id="F3507610FF99B501E13EB2ACFCB350B5" box="[584,803,1762,1788]" name="South Africa" pageId="2" pageNumber="545">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="4983F862FF99B501E042B2ADFBC150B5" box="[820,1105,1762,1789]" country="South Africa" name="Eastern Cape" pageId="2" pageNumber="545">Eastern Cape Province</collectingRegion>
.
</emphasis>
1♁
<location id="8E98605BFF99B501E7F9B2ADFABA50B4" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF99B501E7F9B2ADFABA50B4" box="[1167,1322,1763,1789]" country="South Africa" latitude="-30.7" longLatPrecision="1220" longitude="26.7" name="Aliwal North" pageId="2" pageNumber="545" stateProvince="Eastern Cape">Aliwal North</location>
, [
<geoCoordinate id="EE735047FF99B501E637B2AAFA0C50B6" box="[1345,1436,1764,1791]" degrees="30" direction="south" minutes="42" orientation="latitude" pageId="2" pageNumber="545" precision="925" value="-30.7">30°42S</geoCoordinate>
<geoCoordinate id="EE735047FF99B501E3E1B346FF67516A" box="[151,247,1800,1827]" degrees="26" direction="east" minutes="42" orientation="longitude" pageId="2" pageNumber="545" precision="925" value="26.7">26°42E</geoCoordinate>
],
<date id="FFF91040FF99B501E27FB348FE375168" box="[265,423,1798,1825]" pageId="2" pageNumber="545" value="1971-02-15" valueMax="1971-02-25" valueMin="1971-02-15">
<collectingDate id="EFBDE9A8FF99B501E27FB348FE375168" box="[265,423,1798,1825]" pageId="2" pageNumber="545" value="1971-02-15" valueMax="1971-02-25" valueMin="1971-02-15">1525.ii.1971</collectingDate>
</date>
,
<collectorName id="26B25356FF99B501E2C5B349FD9F5168" box="[435,527,1799,1825]" pageId="2" pageNumber="545">Snyman</collectorName>
&amp;
<collectorName id="26B25356FF99B501E142B349FDE25168" box="[564,626,1799,1825]" pageId="2" pageNumber="545">Jones</collectorName>
(
<collectionCode id="ED56AE45FF99B501E1F7B349FD455168" box="[641,725,1799,1825]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="2" pageNumber="545">TMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF99B501E19EB349FA08510C" ID-GBIF-Occurrence="3062300310" collectingDate="1976-01-19" collectingDateMax="1976-01-22" collectingDateMin="1976-01-19" collectionCode="BMSA" collectorName="GW Ferreira &amp; S van Ee" country="South Africa" latitude="-29.116667" location="Ab" longLatPrecision="1228" longitude="27.366667" municipality="Nova" pageId="2" pageNumber="545" specimenCount="1" stateProvince="Free State">
<emphasis id="B933EA92FF99B501E19EB349FC4A5169" bold="true" box="[744,986,1798,1825]" pageId="2" pageNumber="545">
<collectingRegion id="4983F862FF99B501E19EB349FC445169" box="[744,980,1798,1825]" country="South Africa" name="Free State" pageId="2" pageNumber="545">Free State Province</collectingRegion>
.
</emphasis>
13♁
<collectingMunicipality id="6B9CACFAFF99B501E76FB349FBC65168" box="[1049,1110,1799,1825]" pageId="2" pageNumber="545">Nova</collectingMunicipality>
667 [farm], near
<collectorName id="26B25356FF99B501E66AB349FF27510C" pageId="2" pageNumber="545">Ladybrand, SE</collectorName>
2927
<location id="8E98605BFF99B501E386B365FE83510C" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF99B501E386B365FE83510C" box="[240,275,1835,1861]" country="South Africa" latitude="-29.116667" longLatPrecision="1228" longitude="27.366667" municipality="Nova" name="Ab" pageId="2" pageNumber="545" stateProvince="Free State">Ab</location>
[
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<geoCoordinate id="EE735047FF99B501E2F7B362FE70510E" box="[385,480,1836,1863]" degrees="27" direction="east" minutes="22" orientation="longitude" pageId="2" pageNumber="545" precision="925" value="27.366667">27°22E</geoCoordinate>
],
<date id="FFF91040FF99B501E287B365FD18510C" box="[497,648,1835,1861]" pageId="2" pageNumber="545" value="1976-01-19" valueMax="1976-01-22" valueMin="1976-01-19">
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</date>
, GW
<collectorName id="26B25356FF99B501E1B1B365FCB3510C" box="[711,803,1835,1861]" pageId="2" pageNumber="545">Ferreira</collectorName>
,
<collectorName id="26B25356FF99B501E05AB365FC1E510C" box="[812,910,1835,1861]" pageId="2" pageNumber="545">S van Ee</collectorName>
(
<collectionCode id="ED56AE45FF99B501E0EAB365FC61510C" box="[924,1009,1835,1861]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
:
<collectionCode id="ED56AE45FF99B501E08DB365FBDC510C" box="[1019,1100,1835,1861]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
(C)-2103221044) (
<figureCitation id="137C2A05FF99B501E65CB365FA00510C" box="[1322,1424,1835,1861]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="2" pageNumber="545">Figs 47</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF99B501E3E1B31EFCF451C4" ID-GBIF-Occurrence="3062300348" collectingDate="1977-02-01" collectingDateMax="1977-02-12" collectingDateMin="1977-02-01" collectionCode="BMSA" collectorName="Ferreira &amp; Strydom" country="South Africa" latitude="-29.116667" location="Ab" longLatPrecision="1228" longitude="27.366667" municipality="Nova" pageId="2" pageNumber="545" specimenCount="3" specimenCount-female="3" stateProvince="Free State">
3♁,
<specimenCount id="9D41FD09FF99B501E3BDB301FF7D5120" box="[203,237,1871,1897]" pageId="2" pageNumber="545" type="female">3♀</specimenCount>
<emphasis id="B933EA92FF99B501E381B301FEBD5120" box="[247,301,1871,1897]" italics="true" pageId="2" pageNumber="545">idem</emphasis>
but
<date id="FFF91040FF99B501E213B301FD925120" box="[357,514,1871,1897]" pageId="2" pageNumber="545" value="1977-02-01" valueMax="1977-02-12" valueMin="1977-02-01">
<collectingDate id="EFBDE9A8FF99B501E213B301FD925120" box="[357,514,1871,1897]" pageId="2" pageNumber="545" value="1977-02-01" valueMax="1977-02-12" valueMin="1977-02-01">0112.ii.1977</collectingDate>
</date>
, GW &amp; MC
<collectorName id="26B25356FF99B501E1EBB301FD695120" box="[669,761,1871,1897]" pageId="2" pageNumber="545">Ferreira</collectorName>
, A
<collectorName id="26B25356FF99B501E057B301FC115120" box="[801,897,1871,1897]" pageId="2" pageNumber="545">Strydom</collectorName>
(
<collectionCode id="ED56AE45FF99B501E0E5B301FC785120" box="[915,1000,1871,1897]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
:
<collectionCode id="ED56AE45FF99B501E083B300FBD75120" box="[1013,1095,1870,1897]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
(C)-2104521046;
<collectionCode id="ED56AE45FF99B501E668B301FAE25120" box="[1310,1394,1871,1897]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
(C)- 21975;
<collectionCode id="ED56AE45FF99B501E399B33DFED151C4" box="[239,321,1907,1933]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
(C)-21978;
<collectionCode id="ED56AE45FF99B501E2B5B33DFD8651C4" box="[451,534,1907,1933]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
(C)-22031;
<collectionCode id="ED56AE45FF99B501E1E1B33DFD7A51C4" box="[663,746,1907,1933]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
(C)-22293)
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF99B501E002B33AFB5E51F8" ID-GBIF-Occurrence="3062300305" collectingDate="1976-11-08" collectingDateMax="1976-11-18" collectingDateMin="1976-11-08" collectionCode="BMSA" collectorName="Museum" country="South Africa" county="Kerkenberg" latitude="-28.50389" location="Ac" longLatPrecision="20" longitude="29.108334" municipality="Harrismith" pageId="2" pageNumber="545" specimenCount="4" specimenCount-female="4" stateProvince="Free State">
4♁,
<specimenCount id="9D41FD09FF99B501E0DFB33DFC5C51C4" box="[937,972,1907,1933]" pageId="2" pageNumber="545" type="female">4♀</specimenCount>
<collectingCounty id="62994E0CFF99B501E0A1B33DFBCB51C4" box="[983,1115,1907,1933]" pageId="2" pageNumber="545">Kerkenberg</collectingCounty>
, near
<collectingMunicipality id="6B9CACFAFF99B501E7D0B33DFA8F51C4" box="[1190,1311,1907,1933]" pageId="2" pageNumber="545">Harrismith</collectingMunicipality>
, 2829
<location id="8E98605BFF99B501E611B33DFA1751C4" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF99B501E611B33DFA1751C4" box="[1383,1415,1907,1933]" country="South Africa" county="Kerkenberg" latitude="-28.50389" longLatPrecision="20" longitude="29.108334" municipality="Harrismith" name="Ac" pageId="2" pageNumber="545" stateProvince="Free State">Ac</location>
/d [
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<geoCoordinate id="EE735047FF99B501E246B3D6FE2851FA" box="[304,440,1944,1971]" degrees="29" direction="east" minutes="06" orientation="longitude" pageId="2" pageNumber="545" precision="15" seconds="30" value="29.108334">29°0630″E</geoCoordinate>
],
<date id="FFF91040FF99B501E2A7B3D9FDE551F8" box="[465,629,1943,1969]" pageId="2" pageNumber="545" value="1976-11-08" valueMax="1976-11-18" valueMin="1976-11-08">
<collectingDate id="EFBDE9A8FF99B501E2A7B3D9FDE551F8" box="[465,629,1943,1969]" pageId="2" pageNumber="545" value="1976-11-08" valueMax="1976-11-18" valueMin="1976-11-08">0818.xi.1976</collectingDate>
</date>
,
<collectorName id="26B25356FF99B501E1FFB3D9FD7B51F8" box="[649,747,1943,1969]" pageId="2" pageNumber="545">Museum</collectorName>
staff (
<collectionCode id="ED56AE45FF99B501E034B3D9FC0751F8" box="[834,919,1943,1969]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
:
<collectionCode id="ED56AE45FF99B501E0DFB3D9FC6B51F8" box="[937,1019,1943,1969]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="2" pageNumber="545">BMSA</collectionCode>
(C)-2102021027)
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF99B501E796B3D9FD3A51B0" ID-GBIF-Occurrence="3062300314" collectingDate="1981-11-13" collectingDateMax="1981-11-14" collectingDateMin="1981-11-13" collectionCode="SANC" collectorName="Tonder &amp; Kok" country="South Africa" latitude="-29.116667" location="Mdedelelo Forestry Area" longLatPrecision="1228" longitude="29.433332" pageId="2" pageNumber="545" specimenCount="1" stateProvince="KwaZulu-Natal">
<emphasis id="B933EA92FF99B501E796B3D9FE98519D" bold="true" pageId="2" pageNumber="545">
<collectingRegion id="4983F862FF99B501E796B3D9FE92519D" country="South Africa" name="KwaZulu-Natal" pageId="2" pageNumber="545">KwaZulu-Natal Province</collectingRegion>
.
</emphasis>
1♁ [uKhahlamba-Drakensberg Park World Heritage Site],
<location id="8E98605BFF99B501E0DBB3F5FB5C519C" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF99B501E0DBB3F5FB5C519C" box="[941,1228,1979,2005]" country="South Africa" latitude="-29.116667" longLatPrecision="1228" longitude="29.433332" name="Mdedelelo Forestry Area" pageId="2" pageNumber="545" stateProvince="KwaZulu-Natal">Mdedelelo Forestry Area</location>
,
<geoCoordinate id="EE735047FF99B501E7AFB3F2FAA2519E" box="[1241,1330,1980,2007]" degrees="29" direction="south" minutes="07" orientation="latitude" pageId="2" pageNumber="545" precision="925" value="-29.116667">29°07S</geoCoordinate>
<geoCoordinate id="EE735047FF99B501E64DB3F2FA07519E" box="[1339,1431,1980,2007]" degrees="29" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="545" precision="925" value="29.433332">29°26E</geoCoordinate>
,
<date id="FFF91040FF99B501E3E1B391FEAC51B0" box="[151,316,2015,2041]" pageId="2" pageNumber="545" value="1981-11-13" valueMax="1981-11-14" valueMin="1981-11-13">
<collectingDate id="EFBDE9A8FF99B501E3E1B391FEAC51B0" box="[151,316,2015,2041]" pageId="2" pageNumber="545" value="1981-11-13" valueMax="1981-11-14" valueMin="1981-11-13">1314.xi.1981</collectingDate>
</date>
, SJ van
<collectorName id="26B25356FF99B501E2ECB391FE7B51B0" box="[410,491,2015,2041]" pageId="2" pageNumber="545">Tonder</collectorName>
, C
<collectorName id="26B25356FF99B501E167B391FDD151B0" box="[529,577,2015,2041]" pageId="2" pageNumber="545">Kok</collectorName>
(
<collectionCode id="ED56AE45FF99B501E127B391FD3251B0" box="[593,674,2015,2041]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="545" type="Museum">SANC</collectionCode>
)
</materialsCitation>
.
<specimenCount id="9D41FD09FF99B501E1C3B391FD4851B0" box="[693,728,2015,2041]" pageId="2" pageNumber="545" type="female">1♀</specimenCount>
[uKhahlamba-Drakensberg Park World Heritage Site], Giants
<materialsCitation id="3B2F3CDDFF98B500E3E1B4D9FA0956F8" ID-GBIF-Occurrence="3062300316" box="[151,1433,151,179]" collectingDate="1994-04-24" collectionCode="TMSA" collectorName="Harrison &amp; R Stals" elevation="2300" latitude="-29.25" location="Castle Game Reserve" longLatPrecision="1228" longitude="29.433332" municipality="Bannerman Hut" pageId="3" pageNumber="546" specimenCount="1">
<location id="8E98605BFF98B500E3E1B4D9FE1B56F8" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF98B500E3E1B4D9FE1B56F8" box="[151,395,151,177]" latitude="-29.25" longLatPrecision="1228" longitude="29.433332" municipality="Bannerman Hut" name="Castle Game Reserve" pageId="3" pageNumber="546">Castle Game Reserve</location>
, at
<collectingMunicipality id="6B9CACFAFF98B500E2C7B4D9FDF556F8" box="[433,613,151,177]" pageId="3" pageNumber="546">Bannerman Hut</collectingMunicipality>
,
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,
<quantity id="4CBF9B65FF98B500E04EB4D9FC0656F8" box="[824,918,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="546" unit="m" value="2300.0">
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</quantity>
,
<date id="FFF91040FF98B500E0D4B4D9FB8B56F8" box="[930,1051,151,177]" pageId="3" pageNumber="546" value="1994-04-24">
<collectingDate id="EFBDE9A8FF98B500E0D4B4D9FB8B56F8" box="[930,1051,151,177]" pageId="3" pageNumber="546" value="1994-04-24">24.iv.1994</collectingDate>
</date>
, J duG
<collectorName id="26B25356FF98B500E718B4D9FB4356F8" box="[1134,1235,151,177]" pageId="3" pageNumber="546">Harrison</collectorName>
,
<collectorName id="26B25356FF98B500E7A8B4D9FABE56F8" box="[1246,1326,151,177]" pageId="3" pageNumber="546">R Stals</collectorName>
(
<collectionCode id="ED56AE45FF98B500E64BB4D9FA0156F8" box="[1341,1425,151,177]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="546">TMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF83680FF98B500E3E1B4F5FC5A572C" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="546">
<materialsCitation id="3B2F3CDDFF98B500E3E1B4F5FD1956B0" ID-GBIF-Occurrence="3062300320" collectingDate="1994-01-28" collectionCode="TMSA" collectorName="S Endrody-Younga" latitude="-24.983334" location="Skukuza Rest Camp" longLatPrecision="1248" longitude="31.6" municipality="Kruger National Park" pageId="3" pageNumber="546" specimenCount="1" stateProvince="Mpumalanga Province">
<emphasis id="B933EA92FF98B500E3E1B4F5FE21569D" bold="true" box="[151,433,186,213]" pageId="3" pageNumber="546">
<collectingRegion id="4983F862FF98B500E3E1B4F5FE3C569D" box="[151,428,186,213]" inferredBySuffix="true" pageId="3" pageNumber="546">Mpumalanga Province</collectingRegion>
.
</emphasis>
1♁
<collectingMunicipality id="6B9CACFAFF98B500E290B4F5FD4E569C" box="[486,734,187,213]" pageId="3" pageNumber="546">Kruger National Park</collectingMunicipality>
,
<location id="8E98605BFF98B500E19DB4F5FC46569C" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF98B500E19DB4F5FC46569C" box="[747,982,187,213]" latitude="-24.983334" longLatPrecision="1248" longitude="31.6" municipality="Kruger National Park" name="Skukuza Rest Camp" pageId="3" pageNumber="546" stateProvince="Mpumalanga Province">Skukuza Rest Camp</location>
,
<geoCoordinate id="EE735047FF98B500E095B4F2FBAC569E" box="[995,1084,188,215]" degrees="24" direction="south" minutes="59" orientation="latitude" pageId="3" pageNumber="546" precision="925" value="-24.983334">24°59S</geoCoordinate>
<geoCoordinate id="EE735047FF98B500E733B4F2FB32569E" box="[1093,1186,188,215]" degrees="31" direction="east" minutes="36" orientation="longitude" pageId="3" pageNumber="546" precision="925" value="31.6">31°36E</geoCoordinate>
,
<date id="FFF91040FF98B500E7D9B4F5FA8B569C" box="[1199,1307,187,213]" pageId="3" pageNumber="546" value="1994-01-28">
<collectingDate id="EFBDE9A8FF98B500E7D9B4F5FA8B569C" box="[1199,1307,187,213]" pageId="3" pageNumber="546" value="1994-01-28">28.i.1994</collectingDate>
</date>
, E-Y:2952,
<collectingMethod id="52064E97FF98B500E3E1B491FEA456B0" box="[151,308,223,249]" pageId="3" pageNumber="546">UV light trap</collectingMethod>
,
<collectorName id="26B25356FF98B500E234B491FD8B56B0" box="[322,539,223,249]" pageId="3" pageNumber="546">S Endrödy-Younga</collectorName>
(
<collectionCode id="ED56AE45FF98B500E15BB491FD1156B0" box="[557,641,223,249]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="546">TMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF98B500E1E0B4AEFEE75754" ID-GBIF-Occurrence="3062300338" collectingDate="1970-10" collectionCode="TMSA" collectorName="L Schulze" latitude="-26.5" location="Ermelo district" longLatPrecision="1241" longitude="29.116667" municipality="Winkelhaak" pageId="3" pageNumber="546" specimenCount="1" stateProvince="Mpumalanga Province">
1♁
<collectingMunicipality id="6B9CACFAFF98B500E1B4B491FCD856B0" box="[706,840,223,249]" pageId="3" pageNumber="546">Winkelhaak</collectingMunicipality>
[mine],
<location id="8E98605BFF98B500E0DBB491FBCA56B0" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF98B500E0DBB491FBCA56B0" box="[941,1114,223,249]" latitude="-26.5" longLatPrecision="1241" longitude="29.116667" municipality="Winkelhaak" name="Ermelo district" pageId="3" pageNumber="546" stateProvince="Mpumalanga Province">Ermelo district</location>
, [
<geoCoordinate id="EE735047FF98B500E719B4AEFB5A56B2" box="[1135,1226,224,251]" degrees="26" direction="south" minutes="30" orientation="latitude" pageId="3" pageNumber="546" precision="925" value="-26.5">26°30S</geoCoordinate>
<geoCoordinate id="EE735047FF98B500E7A2B4AEFAA356B2" box="[1236,1331,224,251]" degrees="29" direction="east" minutes="07" orientation="longitude" pageId="3" pageNumber="546" precision="925" value="29.116667">29°07E</geoCoordinate>
],
<date id="FFF91040FF98B500E63EB491FA0856B0" box="[1352,1432,223,249]" pageId="3" pageNumber="546" value="1970-10">
<collectingDate id="EFBDE9A8FF98B500E63EB491FA0856B0" box="[1352,1432,223,249]" pageId="3" pageNumber="546" value="1970-10">x.1970</collectingDate>
</date>
,
<collectorName id="26B25356FF98B500E3E1B54DFE9A5754" box="[151,266,259,285]" pageId="3" pageNumber="546">L Schulze</collectorName>
(
<collectionCode id="ED56AE45FF98B500E26DB54DFEFF5754" box="[283,367,259,285]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="546">TMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2F3CDDFF98B500E2F2B54DFA9D5709" ID-GBIF-Occurrence="3062300340" collectingDate="1956-10-28" collectingDateMax="1961-08-13" collectingDateMin="1956-10-28" collectionCode="NHMB" collectorName="L Schulze" country="Botswana" latitude="-25.65" location="Rustenburg" longLatPrecision="1245" longitude="27.233334" municipality="Winkelhaak" pageId="3" pageNumber="546" specimenCount="1" stateProvince="North West">
<emphasis id="B933EA92FF98B500E2F2B54DFD1A5755" bold="true" box="[388,650,258,285]" pageId="3" pageNumber="546">
<collectingRegion id="4983F862FF98B500E2F2B54DFD155755" box="[388,645,258,285]" country="Botswana" name="North West" pageId="3" pageNumber="546">North West Province</collectingRegion>
.
</emphasis>
2♁, 7 unsexed:
<location id="8E98605BFF98B500E03AB54DFC405754" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF98B500E03AB54DFC405754" box="[844,976,259,285]" country="Botswana" latitude="-25.65" longLatPrecision="1245" longitude="27.233334" name="Rustenburg" pageId="3" pageNumber="546" stateProvince="North West">Rustenburg</location>
, [
<geoCoordinate id="EE735047FF98B500E093B54AFBD05756" box="[997,1088,260,287]" degrees="25" direction="south" minutes="39" orientation="latitude" pageId="3" pageNumber="546" precision="925" value="-25.65">25°39S</geoCoordinate>
<geoCoordinate id="EE735047FF98B500E73CB54AFB385756" box="[1098,1192,260,287]" degrees="27" direction="east" minutes="14" orientation="longitude" pageId="3" pageNumber="546" precision="925" value="27.233334">27°14E</geoCoordinate>
],
<date id="FFF91040FF98B500E7C8B54CFABE5754" box="[1214,1326,258,285]" pageId="3" pageNumber="546" value="1956-10-28">
<collectingDate id="EFBDE9A8FF98B500E7C8B54CFABE5754" box="[1214,1326,258,285]" pageId="3" pageNumber="546" value="1956-10-28">28.x.1956</collectingDate>
</date>
(
<collectionCode id="ED56AE45FF98B500E636B54DFA085754" box="[1344,1432,259,285]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="546">NHMB</collectionCode>
, through photographs). 7 unsexed:
<emphasis id="B933EA92FF98B500E167B569FDD75708" box="[529,583,295,321]" italics="true" pageId="3" pageNumber="546">idem</emphasis>
but
<date id="FFF91040FF98B500E101B569FD6E5708" box="[631,766,295,321]" pageId="3" pageNumber="546" value="1961-08-13">
<collectingDate id="EFBDE9A8FF98B500E101B569FD6E5708" box="[631,766,295,321]" pageId="3" pageNumber="546" value="1961-08-13">13.viii.1961</collectingDate>
</date>
(
<collectionCode id="ED56AE45FF98B500E07AB569FCF45708" box="[780,868,295,321]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="546">NHMB</collectionCode>
, through photographs).
<collectingCountry id="F3507610FF98B500E71DB568FA9D5709" box="[1131,1293,294,320]" name="Botswana" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E71DB568FA9D5709" bold="true" box="[1131,1293,294,320]" pageId="3" pageNumber="546">BOTSWANA</emphasis>
</collectingCountry>
</materialsCitation>
<emphasis id="B933EA92FF98B500E67BB568FA825709" bold="true" box="[1293,1298,294,320]" pageId="3" pageNumber="546">.</emphasis>
<materialsCitation id="3B2F3CDDFF98B500E66EB568FC56572C" ID-GBIF-Occurrence="3062300342" collectingDate="1963-09-02" collectionCode="TMSA" collectorName="JA Nagle" country="Botswana" latitude="-25.033333" location="Otse" longLatPrecision="925" longitude="25.73889" pageId="3" pageNumber="546" specimenCount="1" stateProvince="South East">
<emphasis id="B933EA92FF98B500E66EB568FF69572D" bold="true" pageId="3" pageNumber="546">
<collectingRegion id="4983F862FF98B500E66EB568FF63572D" country="Botswana" name="South East" pageId="3" pageNumber="546">South-East District</collectingRegion>
.
</emphasis>
1♁
<location id="8E98605BFF98B500E25FB505FECF572C" LSID="urn:lsid:plazi:treatment:03EE8796FF99B505E3E1B7ECFD5452A0:8E98605BFF98B500E25FB505FECF572C" box="[297,351,331,357]" country="Botswana" latitude="-25.033333" longLatPrecision="925" longitude="25.73889" name="Otse" pageId="3" pageNumber="546" stateProvince="South East">Otse</location>
, [
<geoCoordinate id="EE735047FF98B500E204B502FE5D572E" box="[370,461,332,359]" degrees="25" direction="south" minutes="02" orientation="latitude" pageId="3" pageNumber="546" precision="925" value="-25.033333">25°02S</geoCoordinate>
<geoCoordinate id="EE735047FF98B500E2A2B502FDCC572E" box="[468,604,332,359]" degrees="25" direction="east" minutes="44" orientation="longitude" pageId="3" pageNumber="546" precision="15" seconds="20" value="25.73889">25°4420″E</geoCoordinate>
],
<date id="FFF91040FF98B500E11BB505FD78572C" box="[621,744,331,357]" pageId="3" pageNumber="546" value="1963-09-02">
<collectingDate id="EFBDE9A8FF98B500E11BB505FD78572C" box="[621,744,331,357]" pageId="3" pageNumber="546" value="1963-09-02">02.ix.1963</collectingDate>
</date>
,
<collectorName id="26B25356FF98B500E185B505FCCB572C" box="[755,859,331,357]" pageId="3" pageNumber="546">JA Nagle</collectorName>
(
<collectionCode id="ED56AE45FF98B500E01FB505FC2D572C" box="[873,957,331,357]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="546">TMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF98B500E3B1B520FC915194" pageId="3" pageNumber="546" type="description">
<paragraph id="8BF83680FF98B500E3B1B520FC7957C0" blockId="3.[151,1437,151,2013]" box="[199,1001,366,393]" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E3B1B520FEE557C1" bold="true" box="[199,373,366,392]" pageId="3" pageNumber="546">Redescription.</emphasis>
<emphasis id="B933EA92FF98B500E20AB521FE3957C0" box="[380,425,367,393]" italics="true" pageId="3" pageNumber="546">Size</emphasis>
: Length:
<quantity id="4CBF9B65FF98B500E160B521FD1457C0" box="[534,644,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="3" pageNumber="546" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">913 mm</quantity>
. Width across humeri:
<quantity id="4CBF9B65FF98B500E0F3B520FC7557C0" box="[901,997,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="546" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
.
</paragraph>
<paragraph id="8BF83680FF98B500E3B1B5DDFAC157E4" blockId="3.[151,1437,151,2013]" box="[199,1361,402,429]" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E3B1B5DDFE8657E4" box="[199,278,403,429]" italics="true" pageId="3" pageNumber="546">Colour</emphasis>
(
<figureCitation id="137C2A05FF98B500E253B5DDFEFC57E4" box="[293,364,403,429]" captionStart="FIGURES 13" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[216,1385,195,1920]" captionTargetId="figure-31@4.[185,1403,181,1920]" captionTargetPageId="4" captionText="FIGURES 13. Holotype of Trox cyrtus Haaf, 1953 (MNHN). 1, habitus, dorsal view; 2, aedeagus, dorsal view; 3, holotype labels. Photographs by Antoine Mantilleri, copyright MNHN." figureDoi="http://doi.org/10.5281/zenodo.4612453" httpUri="https://zenodo.org/record/4612453/files/figure.png" pageId="3" pageNumber="546">Figs 1</figureCitation>
,
<figureCitation id="137C2A05FF98B500E20CB5DCFE3957E4" box="[378,425,402,429]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="3" pageNumber="546">45</figureCitation>
): Matt black, with dull reflection. Body setae yellow unless stated otherwise below.
</paragraph>
<paragraph id="8BF83680FF98B500E3B1B5F9FDCD5484" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E3B1B5F9FE945798" box="[199,260,439,465]" italics="true" pageId="3" pageNumber="546">Head</emphasis>
(
<figureCitation id="137C2A05FF98B500E263B5F9FE105798" box="[277,384,438,465]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="3" pageNumber="546">Figs 45</figureCitation>
): Surface sculpted with discrete punctures, surfaces between punctures with dull reflection. Clypeus triangular, straight, apically deflexed and rounded; anterior clypeal margin entire, bare; clypeogenal and frontoclypeal sutures distinctly raised; frontoclypeal suture complete, with a low clypeogenal tubercle on either side, tubercles punctate. Frons raised, with paired, arched, setose frontoclypeal carinae delimiting two semicircular depressions, separated by a distinct, setose median carina. Genal angles obtuse, wide, recurved; genal margin with straight setae. Eyes large and bulbous, barely visible in dorsal view. Antennal scape rounded, longer than wide, punctate, with long light brown setae; pedicel attached apically; antennal club dark brown, velutinous. Setae around mouthparts dense, yellow to rust-brown.
</paragraph>
<paragraph id="8BF83680FF98B500E3B1B699FD315580" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E3B1B699FEA654B9" box="[199,310,727,752]" italics="true" pageId="3" pageNumber="546">Pronotum</emphasis>
(
<figureCitation id="137C2A05FF98B500E231B699FE3E54B8" box="[327,430,726,753]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="3" pageNumber="546">Figs 45</figureCitation>
,
<figureCitation id="137C2A05FF98B500E2CAB698FE4B54B8" box="[444,475,726,753]" captionStart="FIGURES 1516" captionStartId="10.[151,264,1442,1466]" captionTargetBox="[171,1416,181,1419]" captionTargetId="figure-31@10.[170,1416,181,1419]" captionTargetPageId="10" captionText="FIGURES 1516. Pronotal disc, dorsal view. 15, Phoberus cyrtus (Haaf, 1953); 16, Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612465" httpUri="https://zenodo.org/record/4612465/files/figure.png" pageId="3" pageNumber="546">15</figureCitation>
): Slightly narrower than elytra. Surface with discrete, round punctures, surfaces between punctures dull to shiny. Lateral margins broad and flat, somewhat explanate, entire to irregularly, coarsely dentate, strongly attenuate anteriad, with fringes of long, straight setae. Apex with row of long, recurved, evenly spaced setae. Base with dense fringe of straight setae. Median discal area only slightly raised, broad, narrowing towards base to form a distinct triangular disc; dorsal outline evenly arched in anterior view; median discal depression shallow; median basal tubercles distinct; lateral basal tubercles fused to those in front of them to form a low, broad carina; all tubercles and ridges with setal tufts.
</paragraph>
<paragraph id="8BF83680FF98B500E3B1B79DFCDC5258" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="546">
<taxonomicName id="4C474D03FF98B500E3B1B79DFEA555A4" box="[199,309,979,1005]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="CoL" kingdom="Fungi" order="Corynexochida" pageId="3" pageNumber="546" phylum="Arthropoda" rank="genus">
<emphasis id="B933EA92FF98B500E3B1B79DFEA555A4" box="[199,309,979,1005]" italics="true" pageId="3" pageNumber="546">Scutellum</emphasis>
</taxonomicName>
(
<figureCitation id="137C2A05FF98B500E233B79DFE3E55A4" box="[325,430,978,1005]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="3" pageNumber="546">Figs 45</figureCitation>
): Longer than wide; surface smooth; apicolaterally rounded, lateral margins straight; mediobasally depressed, depression weakly sculpted to punctate.
</paragraph>
<paragraph id="8BF83680FF98B500E3B1B055FC965064" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E3B1B055FE9D527C" box="[199,269,1051,1077]" italics="true" pageId="3" pageNumber="546">Elytra</emphasis>
(
<figureCitation id="137C2A05FF98B500E26DB055FEF1527C" box="[283,353,1051,1077]" captionStart="FIGURES 13" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[216,1385,195,1920]" captionTargetId="figure-31@4.[185,1403,181,1920]" captionTargetPageId="4" captionText="FIGURES 13. Holotype of Trox cyrtus Haaf, 1953 (MNHN). 1, habitus, dorsal view; 2, aedeagus, dorsal view; 3, holotype labels. Photographs by Antoine Mantilleri, copyright MNHN." figureDoi="http://doi.org/10.5281/zenodo.4612453" httpUri="https://zenodo.org/record/4612453/files/figure.png" pageId="3" pageNumber="546">Figs 1</figureCitation>
,
<figureCitation id="137C2A05FF98B500E218B054FE0C527C" box="[366,412,1050,1077]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="3" pageNumber="546">45</figureCitation>
): Elongate, length 2.72.8 pronotum length, lateral margins subparallel, sides narrow, lateral margin finely dentate and bearing long setae; widest at c. 0.6 length; profile slightly convex, attaining maximum height approximately in the middle, strongly declivous posteriorly. Humeral calli prominent, with short setae. Sutural margin raised, surface smooth, with low (sometimes flat) round to oval tubercles, widely separated and irregularly distributed along margin, roughly of similar size for 0.75 elytral length, thereafter decreasing in size until barely discernible; these tubercles smooth, dull shiny, each with a posterior setal tuft. Elytral costae distinct, evennumbered costae more prominent than odd-numbered. Even-numbered costae with raised oval tubercles widely and fairly evenly spaced along elytral length; these tubercles smooth, shiny, each with a posterior setal tuft. Apical callosity present on fourth costal interval at top of elytral declivity. Elytral costae 2 and 4 with 58 tubercles for 0.75 elytral length (up to apical callosity); odd-numbered costae marked by small round to oval, shiny tubercles, each with one or two short setae, these tubercles regularly spaced, roughly of equal size for 0.75 elytral length, thereafter decreasing in size until barely discernible. Dorsally costae 14 separated from intercostae as slightly elevated bands; laterally costae 59 barely discernible as bands. Intercostae with very shallow, undulating depressions separated by transverse ridges; dorsally, margins of intercostae 14 marked by weak narrow longitudinal ridges; intercostal carinae laterally vestigial, barely discernible or lacking.
</paragraph>
<paragraph id="8BF83680FF98B500E3B1B279FEEB5160" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E3B1B279FF6C5019" box="[199,252,1591,1616]" italics="true" pageId="3" pageNumber="546">Legs</emphasis>
(
<figureCitation id="137C2A05FF98B500E27DB279FEE25018" box="[267,370,1590,1617]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="3" pageNumber="546">Figs 45</figureCitation>
): Surfaces rugose and coarsely punctate. Protibia dorsally keeled, with small apical process and row of short, stout transverse setae; outer margin with 2 distinct median teeth and 35 smaller basal teeth; inner margin with long, dense setae; protibial apex divided into two spines, distinct in males, females with spines partly fused; protibial spur as long as, or longer than, third tarsomere, flattened laterally, apex pointed and recurved. Mesotibia and metatibia with outer margins dentate and with long, dense yellow setae; with 2 apical spurs on inner margin, spurs as long as, or slightly longer than, second tarsomere. Tarsomeres ventrally with sparse setae. Tarsal claws simple, equal.
</paragraph>
<paragraph id="8BF83680FF98B500E3B1B37DFDBA5104" blockId="3.[151,1437,151,2013]" box="[199,554,1843,1869]" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E3B1B37DFE365105" box="[199,422,1843,1869]" italics="true" pageId="3" pageNumber="546">Metathoracic wings</emphasis>
: Complete.
</paragraph>
<paragraph id="8BF83680FF98B500E3B1B319FC915194" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="546">
<emphasis id="B933EA92FF98B500E3B1B319FEFB5138" box="[199,363,1879,1905]" italics="true" pageId="3" pageNumber="546">Male genitalia</emphasis>
(
<figureCitation id="137C2A05FF98B500E20CB319FE525138" box="[378,450,1879,1905]" captionStart="FIGURES 13" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[216,1385,195,1920]" captionTargetId="figure-31@4.[185,1403,181,1920]" captionTargetPageId="4" captionText="FIGURES 13. Holotype of Trox cyrtus Haaf, 1953 (MNHN). 1, habitus, dorsal view; 2, aedeagus, dorsal view; 3, holotype labels. Photographs by Antoine Mantilleri, copyright MNHN." figureDoi="http://doi.org/10.5281/zenodo.4612453" httpUri="https://zenodo.org/record/4612453/files/figure.png" pageId="3" pageNumber="546">Figs 2</figureCitation>
,
<figureCitation id="137C2A05FF98B500E2B8B319FE4F513B" box="[462,479,1879,1906]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="3" pageNumber="546">6</figureCitation>
): Symmetrical, slender; phallobase arched, slightly longer than parameres. Parameres long, extending beyond median lobe, apices pointed, long and decurved. Median lobe divided into two symmetrical parts, these narrow, subparallel, in dorsal view concave upward; apices acutely rounded; two bladelike projections and hooks visible between parameres and median lobe.
</paragraph>
</subSubSection>
<caption id="DF386608FF9FB507E3E1B3D9FC98519A" ID-DOI="http://doi.org/10.5281/zenodo.4612453" ID-Zenodo-Dep="4612453" httpUri="https://zenodo.org/record/4612453/files/figure.png" pageId="4" pageNumber="547" startId="4.[151,264,1943,1967]" targetBox="[216,1385,195,1920]" targetPageId="4">
<paragraph id="8BF83680FF9FB507E3E1B3D9FC98519A" blockId="4.[151,1437,1943,2004]" pageId="4" pageNumber="547">
<emphasis id="B933EA92FF9FB507E3E1B3D9FEAC51E6" bold="true" box="[151,316,1943,1967]" pageId="4" pageNumber="547">FIGURES 13.</emphasis>
Holotype of
<taxonomicName id="4C474D03FF9FB507E2B0B3D6FD2351E6" authority="Haaf, 1953" authorityName="Haaf" authorityYear="1953" box="[454,691,1943,1968]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="547" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9FB507E2B0B3D6FDA951E6" box="[454,569,1944,1967]" italics="true" pageId="4" pageNumber="547">Trox cyrtus</emphasis>
<bibRefCitation id="EFD64B71FF9FB507E137B3D6FD2351E6" author="Haaf, E." box="[577,691,1943,1968]" pageId="4" pageNumber="547" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
</taxonomicName>
(MNHN). 1, habitus, dorsal view; 2, aedeagus, dorsal view; 3, holotype labels. Photographs by Antoine Mantilleri, copyright MNHN.
</paragraph>
</caption>
<caption id="DF386608FF9EB506E3E1B3D9FCF6519D" ID-DOI="http://doi.org/10.5281/zenodo.4612457" ID-Zenodo-Dep="4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="5" pageNumber="548" startId="5.[151,264,1943,1967]" targetBox="[187,1416,192,1903]" targetPageId="5">
<paragraph id="8BF83680FF9EB506E3E1B3D9FCF6519D" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="548">
<emphasis id="B933EA92FF9EB506E3E1B3D9FEAA51E6" bold="true" box="[151,314,1943,1967]" pageId="5" pageNumber="548">FIGURES 47.</emphasis>
<taxonomicName id="4C474D03FF9EB506E249B3D6FDFC51E6" authority="(Haaf, 1953)" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[319,620,1943,1968]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="548" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9EB506E249B3D6FE7351E6" box="[319,483,1944,1968]" italics="true" pageId="5" pageNumber="548">Phoberus cyrtus</emphasis>
(
<bibRefCitation id="EFD64B71FF9EB506E286B3D6FDF451E6" author="Haaf, E." box="[496,612,1943,1968]" pageId="5" pageNumber="548" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
)
</taxonomicName>
, specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels.
</paragraph>
</caption>
<subSubSection id="C35D650BFF9DB505E3B1B4D8FD5452A0" pageId="6" pageNumber="549" type="diagnosis">
<paragraph id="8BF83680FF9DB505E3B1B4D8FAD057BC" blockId="6.[151,1437,150,1257]" pageId="6" pageNumber="549">
<emphasis id="B933EA92FF9DB505E3B1B4D8FED256F9" bold="true" box="[199,322,150,176]" pageId="6" pageNumber="549">Diagnosis.</emphasis>
<taxonomicName id="4C474D03FF9DB505E23FB4D9FE6F56F9" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[329,511,151,177]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E23FB4D9FE6F56F9" box="[329,511,151,177]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C474D03FF9DB505E141B4D9FC9D56F8" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[567,781,151,177]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E141B4D9FC9D56F8" box="[567,781,151,177]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
</taxonomicName>
are very similar, but the former can be distinguished from the latter in having a less coarsely punctate pronotum (
<figureCitation id="137C2A05FF9DB505E077B4F5FC14569C" box="[769,900,186,213]" captionStart="FIGURES 1516" captionStartId="10.[151,264,1442,1466]" captionTargetBox="[171,1416,181,1419]" captionTargetId="figure-31@10.[170,1416,181,1419]" captionTargetPageId="10" captionText="FIGURES 1516. Pronotal disc, dorsal view. 15, Phoberus cyrtus (Haaf, 1953); 16, Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612465" httpUri="https://zenodo.org/record/4612465/files/figure.png" pageId="6" pageNumber="549">Figs 1516</figureCitation>
). Additionally, the arrangement of the tubercles on elytral costae 2 and 4 differs between the species:
<taxonomicName id="4C474D03FF9DB505E197B491FC0656B1" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[737,918,223,249]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E197B491FC0656B1" box="[737,918,223,249]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
has these tubercles widely separated, with 58 tubercles between the elytral base and the apical callosity, whereas these tubercles in
<taxonomicName id="4C474D03FF9DB505E72DB54DFAA25754" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[1115,1330,259,285]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E72DB54DFAA25754" box="[1115,1330,259,285]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
</taxonomicName>
are close together, with 812 tubercles between the elytral base and the apical callosity. The male genitalia of the two species are distinctive with the most important differences found on the median lobes.
<taxonomicName id="4C474D03FF9DB505E084B505FB36572D" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[1010,1190,331,357]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E084B505FB36572D" box="[1010,1190,331,357]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
have the lobes narrow and their apices sharper or angular, whereas in
<taxonomicName id="4C474D03FF9DB505E1EBB521FCE157C0" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[669,881,367,393]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E1EBB521FCE157C0" box="[669,881,367,393]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
</taxonomicName>
the lobes are broad and their apices more rounded (as described in the text and as shown in
<figureCitation id="137C2A05FF9DB505E12AB5DDFD3157E4" box="[604,673,403,429]" captionStart="FIGURES 13" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[216,1385,195,1920]" captionTargetId="figure-31@4.[185,1403,181,1920]" captionTargetPageId="4" captionText="FIGURES 13. Holotype of Trox cyrtus Haaf, 1953 (MNHN). 1, habitus, dorsal view; 2, aedeagus, dorsal view; 3, holotype labels. Photographs by Antoine Mantilleri, copyright MNHN." figureDoi="http://doi.org/10.5281/zenodo.4612453" httpUri="https://zenodo.org/record/4612453/files/figure.png" pageId="6" pageNumber="549">Figs 2</figureCitation>
and
<figureCitation id="137C2A05FF9DB505E1A0B5DDFD7457E4" box="[726,740,403,429]" captionStart="FIGURES 47" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[187,1416,192,1903]" captionTargetId="figure-17@5.[170,1416,181,1920]" captionTargetPageId="5" captionText="FIGURES 47. Phoberus cyrtus (Haaf, 1953), specimen with obscuring layer of dirt removed (BMSA). 4, habitus, dorsal view; 5, habitus, lateral view; 6, aedeagus, dorsal and lateral views; 7, labels." figureDoi="http://doi.org/10.5281/zenodo.4612457" httpUri="https://zenodo.org/record/4612457/files/figure.png" pageId="6" pageNumber="549">6</figureCitation>
versus
<figureCitation id="137C2A05FF9DB505E04EB5DDFCEC57E4" box="[824,892,403,429]" captionStart="FIGURES 810" captionStartId="8.[151,264,1932,1956]" captionTargetBox="[182,1402,193,1906]" captionTargetId="figure-31@8.[167,1419,181,1908]" captionTargetPageId="8" captionText="FIGURES 810. Holotype of Trox fumarius Haaf, 1953 (NHMB): 8, habitus, dorsal view; 9, aedeagus, dorsal view; 10, holotype labels. Photographs by Christoph Germann, copyright NHMB." figureDoi="http://doi.org/10.5281/zenodo.4612459" httpUri="https://zenodo.org/record/4612459/files/figure.png" pageId="6" pageNumber="549">Figs 9</figureCitation>
and
<figureCitation id="137C2A05FF9DB505E0C4B5DDFC4057E4" box="[946,976,403,429]" captionStart="FIGURES 1114" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 1114. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="6" pageNumber="549">13</figureCitation>
). Available information strongly suggests that the two species are allopatric and existent only in different, non-overlapping biomes,
<taxonomicName id="4C474D03FF9DB505E71FB5F9FA8D5799" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[1129,1309,439,465]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E71FB5F9FA8D5799" box="[1129,1309,439,465]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
in the summer rainfall area of southern Africa and
<taxonomicName id="4C474D03FF9DB505E123B595FCB957BC" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[597,809,475,501]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E123B595FCB957BC" box="[597,809,475,501]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
</taxonomicName>
in the winter rainfall area (see text and
<figureCitation id="137C2A05FF9DB505E794B595FAA357BC" box="[1250,1331,475,501]" captionStart="FIGURE 21" captionStartId="12.[151,250,1140,1164]" captionTargetBox="[151,1434,181,1117]" captionTargetId="figure-31@12.[151,1435,181,1117]" captionTargetPageId="12" captionText="FIGURE 21. Known distribution of Phoberus cyrtus (Haaf, 1953) and Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612475" httpUri="https://zenodo.org/record/4612475/files/figure.png" pageId="6" pageNumber="549">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BF83680FF9DB505E3B1B5B1FADB5570" blockId="6.[151,1437,150,1257]" pageId="6" pageNumber="549">
<taxonomicName id="4C474D03FF9DB505E3B1B5B1FDCE5450" authority="(Thunberg, 1787)" baseAuthorityName="Thunberg" baseAuthorityYear="1787" box="[199,606,511,537]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B933EA92FF9DB505E3B1B5B1FE045450" box="[199,404,511,537]" italics="true" pageId="6" pageNumber="549">Phoberus sulcatus</emphasis>
(Thunberg, 1787)
</taxonomicName>
(
<figureCitation id="137C2A05FF9DB505E11DB5B1FD7A5450" box="[619,746,511,537]" captionStart="FIGURES 1720" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[185,1404,195,1919]" captionTargetId="figure-17@11.[166,1422,181,1919]" captionTargetPageId="11" captionText="FIGURES 1720. Phoberus sulcatus (Thunberg, 1787), cleaned specimen (BMSA). 17, habitus, dorsal view; 18, aedeagus, dorsal view; 19, pronotal disc, with red arrow indicating distinct median pronotal depression; 20, specimen labels." figureDoi="http://doi.org/10.5281/zenodo.4612471" httpUri="https://zenodo.org/record/4612471/files/figure.png" pageId="6" pageNumber="549">Figs 1720</figureCitation>
) is widespread and common throughout
<collectingCountry id="F3507610FF9DB505E7DAB5B1FAAA5450" box="[1196,1338,511,537]" name="South Africa" pageId="6" pageNumber="549">South Africa</collectingCountry>
(
<bibRefCitation id="EFD64B71FF9DB505E631B5B1FF705474" author="Scholtz, C. H." pageId="6" pageNumber="549" pagination="1 - 104" refId="ref8073" refString="Scholtz, C. H. (1980 a) Monograph of the genus Trox F. (Coleoptera: Trogidae) of subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz 1980a</bibRefCitation>
). Both
<taxonomicName id="4C474D03FF9DB505E244B66DFE795475" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[306,489,547,573]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E244B66DFE795475" box="[306,489,547,573]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C474D03FF9DB505E157B66DFD685474" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[545,760,547,573]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E157B66DFD685474" box="[545,760,547,573]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
</taxonomicName>
have an external appearance similar to that species, and the three may be confused. In collections they have certainly been confused. Of the two species treated in this work,
<taxonomicName id="4C474D03FF9DB505E3E1B625FEFE54CC" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[151,366,619,645]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E3E1B625FEFE54CC" box="[151,366,619,645]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
</taxonomicName>
is more likely to be confused in this way.
<taxonomicName id="4C474D03FF9DB505E031B625FB8554CC" baseAuthorityName="Thunberg" baseAuthorityYear="1787" box="[839,1045,619,645]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B933EA92FF9DB505E031B625FB8554CC" box="[839,1045,619,645]" italics="true" pageId="6" pageNumber="549">Phoberus sulcatus</emphasis>
</taxonomicName>
differs from both the other species in bearing a distinct median pronotal depression (
<figureCitation id="137C2A05FF9DB505E1B5B6C1FC9E54E0" box="[707,782,655,681]" captionStart="FIGURES 1720" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[185,1404,195,1919]" captionTargetId="figure-17@11.[166,1422,181,1919]" captionTargetPageId="11" captionText="FIGURES 1720. Phoberus sulcatus (Thunberg, 1787), cleaned specimen (BMSA). 17, habitus, dorsal view; 18, aedeagus, dorsal view; 19, pronotal disc, with red arrow indicating distinct median pronotal depression; 20, specimen labels." figureDoi="http://doi.org/10.5281/zenodo.4612471" httpUri="https://zenodo.org/record/4612471/files/figure.png" pageId="6" pageNumber="549">Fig 19</figureCitation>
) and having the pronotal and elytral carinae and tubercles more pronounced and higher (
<figureCitation id="137C2A05FF9DB505E29DB6FDFDFE5484" box="[491,622,691,717]" captionStart="FIGURES 1720" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[185,1404,195,1919]" captionTargetId="figure-17@11.[166,1422,181,1919]" captionTargetPageId="11" captionText="FIGURES 1720. Phoberus sulcatus (Thunberg, 1787), cleaned specimen (BMSA). 17, habitus, dorsal view; 18, aedeagus, dorsal view; 19, pronotal disc, with red arrow indicating distinct median pronotal depression; 20, specimen labels." figureDoi="http://doi.org/10.5281/zenodo.4612471" httpUri="https://zenodo.org/record/4612471/files/figure.png" pageId="6" pageNumber="549">Figs 17, 19</figureCitation>
). The male genitalia of
<taxonomicName id="4C474D03FF9DB505E00DB6FDFBDA5484" baseAuthorityName="Thunberg" baseAuthorityYear="1787" box="[891,1098,691,717]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B933EA92FF9DB505E00DB6FDFBDA5484" box="[891,1098,691,717]" italics="true" pageId="6" pageNumber="549">Phoberus sulcatus</emphasis>
</taxonomicName>
are indubitably different with the median lobe of this species divided into two broadly curved lobes (
<figureCitation id="137C2A05FF9DB505E0D4B699FC7E54B8" box="[930,1006,727,753]" captionStart="FIGURES 1720" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[185,1404,195,1919]" captionTargetId="figure-17@11.[166,1422,181,1919]" captionTargetPageId="11" captionText="FIGURES 1720. Phoberus sulcatus (Thunberg, 1787), cleaned specimen (BMSA). 17, habitus, dorsal view; 18, aedeagus, dorsal view; 19, pronotal disc, with red arrow indicating distinct median pronotal depression; 20, specimen labels." figureDoi="http://doi.org/10.5281/zenodo.4612471" httpUri="https://zenodo.org/record/4612471/files/figure.png" pageId="6" pageNumber="549">Fig 18</figureCitation>
). We found that
<taxonomicName id="4C474D03FF9DB505E7D4B699FAE054B8" baseAuthorityName="Thunberg" baseAuthorityYear="1787" box="[1186,1392,727,753]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B933EA92FF9DB505E7D4B699FAE054B8" box="[1186,1392,727,753]" italics="true" pageId="6" pageNumber="549">Phoberus sulcatus</emphasis>
</taxonomicName>
can readily be distinguished from these other two species by the larger shadows cast under oblique lighting by its more prominent paramedian pronotal carinae; specimens should not be too soiled for this comparison to succeed.
</paragraph>
<paragraph id="8BF83680FF9DB505E3B1B70CFE9D55EC" blockId="6.[151,1437,150,1257]" pageId="6" pageNumber="549">
<emphasis id="B933EA92FF9DB505E3B1B70CFE605515" bold="true" box="[199,496,834,860]" pageId="6" pageNumber="549">Geographic distribution.</emphasis>
<taxonomicName id="4C474D03FF9DB505E28FB70DFD215515" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[505,689,835,861]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E28FB70DFD215515" box="[505,689,835,861]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F3507610FF9DB505E01EB70DFC6A5514" box="[872,1018,835,861]" name="South Africa" pageId="6" pageNumber="549">South Africa</collectingCountry>
,
<collectingCountry id="F3507610FF9DB505E771B70DFBF15514" box="[1031,1121,835,861]" name="Lesotho" pageId="6" pageNumber="549">Lesotho</collectingCountry>
and
<collectingCountry id="F3507610FF9DB505E7EDB70DFA995514" box="[1179,1289,835,861]" name="Botswana" pageId="6" pageNumber="549">Botswana</collectingCountry>
(
<figureCitation id="137C2A05FF9DB505E66CB70DFAFF5514" box="[1306,1391,835,861]" captionStart="FIGURE 21" captionStartId="12.[151,250,1140,1164]" captionTargetBox="[151,1434,181,1117]" captionTargetId="figure-31@12.[151,1435,181,1117]" captionTargetPageId="12" captionText="FIGURE 21. Known distribution of Phoberus cyrtus (Haaf, 1953) and Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612475" httpUri="https://zenodo.org/record/4612475/files/figure.png" pageId="6" pageNumber="549">Fig. 21</figureCitation>
). In
<collectingCountry id="F3507610FF9DB505E3E1B729FEB755C8" box="[151,295,871,897]" name="South Africa" pageId="6" pageNumber="549">South Africa</collectingCountry>
the species has been recorded from the
<collectingRegion id="4983F862FF9DB505E19BB729FC1555C8" box="[749,901,871,897]" country="South Africa" name="Eastern Cape" pageId="6" pageNumber="549">Eastern Cape</collectingRegion>
,
<collectingRegion id="4983F862FF9DB505E0E4B729FBD555C8" box="[914,1093,871,897]" country="South Africa" name="KwaZulu-Natal" pageId="6" pageNumber="549">KwaZulu-Natal</collectingRegion>
,
<collectingRegion id="4983F862FF9DB505E726B729FB7455C8" box="[1104,1252,871,897]" country="South Africa" name="Mpumalanga" pageId="6" pageNumber="549">Mpumalanga</collectingRegion>
and
<collectingRegion id="4983F862FF9DB505E66AB729FA0C55C8" box="[1308,1436,871,897]" country="South Africa" name="North West" pageId="6" pageNumber="549">North West</collectingRegion>
Provinces.
</paragraph>
<paragraph id="8BF83680FF9DB505E3B1B7E1FD5452A0" blockId="6.[151,1437,150,1257]" pageId="6" pageNumber="549">
<emphasis id="B933EA92FF9DB505E3B1B7E1FD985581" bold="true" box="[199,520,942,969]" pageId="6" pageNumber="549">Broad habitat associations.</emphasis>
As far as it is known,
<taxonomicName id="4C474D03FF9DB505E18CB7E1FC205581" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[762,944,943,969]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E18CB7E1FC205581" box="[762,944,943,969]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
is restricted to the summer rainfall region of southern Africa, in contrast to its putative sister species
<taxonomicName id="4C474D03FF9DB505E071B79DFC4D55A4" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[775,989,979,1005]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E071B79DFC4D55A4" box="[775,989,979,1005]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
</taxonomicName>
, which is confined to the winter rainfall region.
<taxonomicName id="4C474D03FF9DB505E39BB7B9FE355259" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[237,421,1015,1041]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E39BB7B9FE355259" box="[237,421,1015,1041]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
is known from two biomes. In the south of its known distribution, the species lives in the Grassland Biome, where it is present in the Mesic Highveld Grassland, Drakensberg Grassland and Dry Highveld Grassland Bioregions (biomes and bioregions following
<bibRefCitation id="EFD64B71FF9DB505E07CB071FBAD5210" author="Mucina, L. &amp; Rutherford, M. C." box="[778,1085,1087,1113]" pageId="6" pageNumber="549" pagination="1 - 807" refId="ref7875" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. South African National Biodiversity Institute, Pretoria, Strelitzia, 19, i - viii + 1 - 807." type="journal article" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
;
<bibRefCitation id="EFD64B71FF9DB505E730B071FABC5210" author="Dayaram, A. &amp; Harris, L. R. &amp; Grobler, B. A. &amp; Van der Merwe, S. &amp; Rebelo, A. G. &amp; Powrie, L. W. &amp; Vlok, J. H. J. &amp; Desmet, P. G. &amp; Qabaqaba, M. &amp; Hlahane, K. M. &amp; Skowno, A. L." box="[1094,1324,1087,1113]" pageId="6" pageNumber="549" pagination="1 - 11" refId="ref7440" refString="Dayaram, A., Harris, L. R., Grobler, B. A., Van der Merwe, S., Rebelo, A. G., Powrie, L. W., Vlok, J. H. J., Desmet, P. G., Qabaqaba, M., Hlahane, K. M. &amp; Skowno, A. L. (2019) Vegetation map of South Africa, Lesotho and Swaziland 2018: a description of changes since 2006. Bothalia, 49 (1), a 2452, 1 - 11. https: // doi. org / 10.4102 / abc. v 49 i 1.2452" type="journal article" year="2019">
Dayaram
<emphasis id="B933EA92FF9DB505E7C4B071FB795210" box="[1202,1257,1087,1113]" italics="true" pageId="6" pageNumber="549">et al.</emphasis>
2019
</bibRefCitation>
). Towards the north and west of its range, the species occurs in the Savanna Biome (Central Bushveld and Lowveld Bioregions). Whereas the grassland habitats are cool and mesic, the savanna habitats are warm to hot and experience extended dry periods. This discordant distribution among two sets of habitats with significantly different bioregional climate regimes may indicate a eurytopic species.
</paragraph>
</subSubSection>
</treatment>
</document>