266 lines
31 KiB
XML
266 lines
31 KiB
XML
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<mods:title id="0458AF58AE4F17687E43DA9E27442231">From the dark side of paradise: a new natural replication of cave planthopper evolution from Hawaiian lava tubes (Hemiptera: Fulgoromorpha: Cixiidae)</mods:title>
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<mods:namePart id="534650CF450EE8123A4062A2AA92EF8B">Hoch, Hannelore</mods:namePart>
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<mods:affiliation id="4BB48D2650119B4D5CC164200043614D">Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversitv Science, Humboldt-Universität zu Berlin, Invalidenstrasse 43, 10115 Berlin, Germanv</mods:affiliation>
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<mods:namePart id="B481192EB1651B8446F9251A933AA715">Porter, Megan L.</mods:namePart>
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<mods:affiliation id="D7BAEE2892023C577B7CD9E2E37F8C2B">School of Life Sciences, Universitv of Hawaiʻi at Mānoa, Honolulu, HI 96822, USA</mods:affiliation>
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<mods:namePart id="B6ABCDFF1CE9F500E0FA87AE03A7C8BE">Slav, Christv M.</mods:namePart>
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<mods:affiliation id="70AE7DB5CC18A8F8F572887C04ED7163">Ŋe Sustainabilitv Consortium, Julie Ann Wriglev Global Futures Laboratorv, Arizona State Universitv, Tempe, AZ 85281, USA</mods:affiliation>
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<mods:namePart id="D74CEBEF4DB42835F1C209D14F53C9E2">Slav, Michael E.</mods:namePart>
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<mods:affiliation id="1C77199291E54161D6022B203C69B484">Arkansas Field Office, Ŋe Nature Conservancv, Liưle Rock, AR 72201, USA</mods:affiliation>
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<mods:namePart id="5D2B96DD4C687844DD1EECC8DA39F4FB">Steck, Mireille</mods:namePart>
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<mods:affiliation id="F0CD935A1B9E46531622B9C149FA9B5A">School of Life Sciences, Universitv of Hawaiʻi at Mānoa, Honolulu, HI 96822, USA</mods:affiliation>
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<mods:namePart id="A677474EB62C0879312CBE1A0B106CAE">Chong, Rebecca A.</mods:namePart>
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<mods:affiliation id="D045C1C1DFA2306B06072B376C6BC8D8">School of Life Sciences, Universitv of Hawaiʻi at Mānoa, Honolulu, HI 96822, USA</mods:affiliation>
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<mods:date id="3F535006A37F03C282F92129E06ECAA8">2024</mods:date>
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<treatment id="03E3EA7C3150BF011C74FE5DFA75F865" LSID="urn:lsid:plazi:treatment:03E3EA7C3150BF011C74FE5DFA75F865" httpUri="http://treatment.plazi.org/id/03E3EA7C3150BF011C74FE5DFA75F865" lastPageNumber="5" pageId="4" pageNumber="5">
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<paragraph id="8BF55B6A3150BF011C74FE5DFD80FDDA" blockId="4.[312,594,494,521]" box="[312,594,494,521]" pageId="4" pageNumber="5">
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<heading id="D0BDEC063150BF011C74FE5DFD80FDDA" bold="true" box="[312,594,494,521]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
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<emphasis id="B93E87783150BF011C74FE5DFD80FDDA" bold="true" box="[312,594,494,521]" pageId="4" pageNumber="5">
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<taxonomicName id="4C4A20E93150BF011C74FE5DFE33FDDA" ID-CoL="54QG" ID-ENA="2831207" authority="Kirkaldy" authorityName="Kirkaldy" authorityYear="1902" box="[312,481,494,521]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E87783150BF011C74FE5DFE52FDDB" bold="true" box="[312,384,494,521]" italics="true" pageId="4" pageNumber="5">Iolania</emphasis>
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Kirkaldy
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19 0 2: 118
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</emphasis>
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<subSubSection id="C35008E13150BF011DCDFD93FDBAFDE5" box="[129,616,543,567]" pageId="4" pageNumber="5" type="type_taxon">
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<paragraph id="8BF55B6A3150BF011DCDFD93FDBAFDE5" blockId="4.[129,616,543,567]" box="[129,616,543,567]" pageId="4" pageNumber="5">
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<emphasis id="B93E87783150BF011DCDFD93FF2EFDE5" box="[129,252,544,567]" italics="true" pageId="4" pageNumber="5">Type species:</emphasis>
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<taxonomicName id="4C4A20E93150BF011C4DFDACFDB1FDE5" authority="Kirkaldv 1902: 119" authorityName="Kirkaldv" authorityPageNumber="119" authorityYear="1902" box="[257,611,543,567]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="perkinsi">
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<emphasis id="B93E87783150BF011C4DFDACFE45FDE5" box="[257,407,543,567]" italics="true" pageId="4" pageNumber="5">Iolania perkinsi</emphasis>
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<bibRefCitation id="EFDB269B3150BF011CD0FDACFDB1FDE5" author="Kirkaldv GW" box="[412,611,543,567]" pageId="4" pageNumber="5" pagination="93 - 174" refId="ref9471" refString="Kirkaldv GW. Hemiptera. In: Sharp D (ed.), Fauna Hawaiiensis, Vol. III, 2. Cambridge: Cambridge Universitv Press, 1902; 93 - 174, pls 4, 5." type="journal article" year="1902">Kirkaldv 1902: 119</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35008E13150BF011C22FDE4FA6DFB7B" pageId="4" pageNumber="5" type="diagnosis">
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<paragraph id="8BF55B6A3150BF011C22FDE4FDCEFDA3" blockId="4.[129,778,599,1194]" box="[366,540,599,625]" pageId="4" pageNumber="5">
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||
<emphasis id="B93E87783150BF011C22FDE4FDCEFDA3" bold="true" box="[366,540,599,625]" pageId="4" pageNumber="5">Genus diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BF55B6A3150BF011DCDFDCEFD1EFBDF" blockId="4.[129,778,599,1194]" pageId="4" pageNumber="5">
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||
Hoch (2006) provided a detailed diagnosis of the genus
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||
<taxonomicName id="4C4A20E93150BF011DCDFD2EFF14FD67" authorityName="Kirkaldy" authorityYear="1902" box="[129,198,669,693]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B93E87783150BF011DCDFD2EFF14FD67" box="[129,198,669,693]" italics="true" pageId="4" pageNumber="5">Iolania</emphasis>
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</taxonomicName>
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, which then contained exclusivelv epigean species. The cavernicolous species
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<taxonomicName id="4C4A20E93150BF011C24FD0EFDD0FD06" box="[360,514,700,724]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ssankanstonei">
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<emphasis id="B93E87783150BF011C24FD0EFDD0FD06" box="[360,514,700,724]" italics="true" pageId="4" pageNumber="5">I. frankanstonei</emphasis>
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</taxonomicName>
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from
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<collectingRegion id="498E95883150BF011F0AFD0FFD5CFD06" box="[582,654,700,724]" country="United States of America" name="Hawaii" pageId="4" pageNumber="5">Hawaii</collectingRegion>
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Island (described below) features characters pertaining to its external morphologv that have evolved in the course of adaptation bv the species to its subterranean habitat (troglomorphies). These include the reduction of compound eves, tegmina and hindwings, and bodilv pigmentation. As has been shown in other cavernicolous planthopper taxa (
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<bibRefCitation id="EFDB269B3150BF011CA8FCCBFD2BFC42" author="Hoch H & Howarth FG" box="[484,761,888,912]" pageId="4" pageNumber="5" pagination="377 - 402" refId="ref8822" refString="Hoch H, Howarth FG. Six new cavernicolous cixiid planthoppers in the genus Solonaima from Australia (Homoptera: Fulgoroidea). Systematic Entomology 1989; 14: 377 - 402. hups: // doi. org / 10.1111 / j. 1365 - 3113.1989. tb 00291. x" type="journal article" year="1989">Hoch and Howarth 1989</bibRefCitation>
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), these reductions of single traits can influence the configuration of other traits, such as head width, sculpturing (e.g. carination), and general bodv shape. It therefore seems appropriate to provide a separate diagnosis for cavernicolous
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<taxonomicName id="4C4A20E93150BF011F7FFC46FD1EFBDF" box="[563,716,1013,1037]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B93E87783150BF011F7FFC46FDAAFBDF" box="[563,632,1013,1037]" italics="true" pageId="4" pageNumber="5">Iolania</emphasis>
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species.
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</taxonomicName>
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</paragraph>
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<paragraph id="8BF55B6A3150BF011DD0FBA7FA8EFF15" blockId="4.[129,778,599,1194]" lastBlockId="4.[825,1474,144,199]" pageId="4" pageNumber="5">
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All species of the genus
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<taxonomicName id="4C4A20E93150BF011CC0FBA7FE03FBFE" authorityName="Kirkaldy" authorityYear="1902" box="[396,465,1044,1068]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E87783150BF011CC0FBA7FE03FBFE" box="[396,465,1044,1068]" italics="true" pageId="4" pageNumber="5">Iolania</emphasis>
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</taxonomicName>
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are easilv distinguishable from the other Hawaiian cixiid genus,
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<taxonomicName id="4C4A20E93150BF011CA4FB87FDE0FB9E" authorityName="Stal" authorityYear="1862" box="[488,562,1076,1100]" class="Insecta" family="Cixiidae" genus="Oliarus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E87783150BF011CA4FB87FDE0FB9E" box="[488,562,1076,1100]" italics="true" pageId="4" pageNumber="5">Oliarus</emphasis>
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</taxonomicName>
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, bv their tricarinate pronotum (vs. pentecarinate in
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<taxonomicName id="4C4A20E93150BF011C91FBE0FDF5FBB9" authorityName="Stal" authorityYear="1862" box="[477,551,1107,1131]" class="Insecta" family="Cixiidae" genus="Oliarus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B93E87783150BF011C91FBE0FDF5FBB9" box="[477,551,1107,1131]" italics="true" pageId="4" pageNumber="5">Oliarus</emphasis>
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</taxonomicName>
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) and from all other known Cixiini bv the morphological configuration of the male copulatorv organ, the aedeagus (shaft bilaterallv svmmetrical, smooth; flagellum rigid, short, not movable against shaft; phallotreme near base of flagellum, exposed dorsallv).
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</paragraph>
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<paragraph id="8BF55B6A3150BF011E2BFF5BFB60FEF2" blockId="4.[871,1428,231,288]" pageId="4" pageNumber="5">
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<emphasis id="B93E87783150BF011E2BFF5BFB60FEF2" italics="true" pageId="4" pageNumber="5">
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Diagnosis of epigean species of
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<taxonomicName id="4C4A20E93150BF0119CDFF54FB19FED3" authorityName="Kirkaldy" authorityYear="1902" box="[1153,1227,231,257]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Iolania</taxonomicName>
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(see also Hoch 2006: 303–304)
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</emphasis>
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</paragraph>
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<paragraph id="8BF55B6A3150BF011E75FE9EFAD6FD29" blockId="4.[825,1475,301,763]" pageId="4" pageNumber="5">
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Total length (tip of head to distal margin of tegmina): males,
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<quantity id="4CB2F68F3150BF0118D9FE9EFC54FEB1" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.3" metricValueMin="4.8" pageId="4" pageNumber="5" unit="mm" value="5.55" valueMax="6.3" valueMin="4.8">4.8– 6.3 mm</quantity>
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; females,
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<quantity id="4CB2F68F3150BF011EA7FEFFFBB6FEB1" box="[1003,1124,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.1" metricValueMin="6.2" pageId="4" pageNumber="5" unit="mm" value="6.65" valueMax="7.1" valueMin="6.2">6.2–7.1 mm</quantity>
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. Moderatelv large cixiids of robust appearance (
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<figureCitation id="137147EF3150BF011EF2FED8FBF6FE51" box="[958,1060,363,387]" captionStart="Figure 3" captionStartId="3.[113,178,1836,1860]" captionTargetBox="[121,1451,145,1807]" captionTargetId="figure-6@3.[120,1452,144,1808]" captionTargetPageId="3" captionText="Figure 3. Images of epigean (Iolania perkinsi) and cavernicolous (Iolania Ƒankanstonei) Iolania species, illustrating external morphological features.A, B, Iolania perkinsi Kirkaldv habitus (A) and head and thorax (B), dorsal aspect. C, D, Iolania ssankanstonei habitus (C) and head and thorax (D), dorsal aspect. Specimen from OHS entrance (images bv B. Schurian)." figureDoi="http://doi.org/10.5281/zenodo.14423203" httpUri="https://zenodo.org/record/14423203/files/figure.png" pageId="4" pageNumber="5">Fig. 3A, B</figureCitation>
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); tegmina ~2.6× longer than maximallv wide, more or less shallowlv tectiform, surpassing tip of abdomen with approximatelv one-third of total length. Costal veins of tegmina nearlv parallel in dorsal aspect. Tegmina and hindwings translucent to hvaline. Tegmina with colour pattern variable within populations, ranging from hvaline with little recognizable patterning to those with distinct dark brown spots or stripes, especiallv at the tegmen base and in the distal third. Granules on veins usuallv brownish. Bodv coloration inconspicuous; intraspecific variation ranging from vellowish brown to darker brown. Coloration of bodv and tegmina usuallv slightlv darker in females. Vertex short, with lateral margins diverging slightlv posteriorlv. Hind tibiae laterallv with three minute spines.
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</paragraph>
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<paragraph id="8BF55B6A3150BF011E02FC81FAC0FCB8" blockId="4.[825,1475,817,1193]" pageId="4" pageNumber="5">
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<emphasis id="B93E87783150BF011E02FC81FAC0FCB8" italics="true" pageId="4" pageNumber="5">
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Diagnosis of cavernicolous species of
|
||
<taxonomicName id="4C4A20E93150BF0119D1FC82FB35FC99" authorityName="Kirkaldy" authorityYear="1902" box="[1181,1255,817,843]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Iolania</taxonomicName>
|
||
(
|
||
<figureCitation id="137147EF3150BF0119BAFC81FA88FC99" box="[1270,1370,817,843]" captionStart="Figure 3" captionStartId="3.[113,178,1836,1860]" captionTargetBox="[121,1451,145,1807]" captionTargetId="figure-6@3.[120,1452,144,1808]" captionTargetPageId="3" captionText="Figure 3. Images of epigean (Iolania perkinsi) and cavernicolous (Iolania Ƒankanstonei) Iolania species, illustrating external morphological features.A, B, Iolania perkinsi Kirkaldv habitus (A) and head and thorax (B), dorsal aspect. C, D, Iolania ssankanstonei habitus (C) and head and thorax (D), dorsal aspect. Specimen from OHS entrance (images bv B. Schurian)." figureDoi="http://doi.org/10.5281/zenodo.14423203" httpUri="https://zenodo.org/record/14423203/files/figure.png" pageId="4" pageNumber="5">Figs 3C, D</figureCitation>
|
||
,
|
||
<figureCitation id="137147EF3150BF01182FFC82FAA2FC99" box="[1379,1392,817,843]" captionStart="Figure 4" captionStartId="5.[113,178,1954,1978]" captionTargetBox="[359,1212,820,1927]" captionTargetId="figure-313@5.[356,1216,817,1928]" captionTargetPageId="5" captionText="Figure 4. Iolania Ƒankanstonei, male, wing morphologv. A, right tegmen. B, right hind wing.Scale bar:0.5 mm. Male from CKM entrance." figureDoi="http://doi.org/10.5281/zenodo.14423205" httpUri="https://zenodo.org/record/14423205/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
|
||
,
|
||
<figureCitation id="137147EF3150BF011835FC82FA54FC99" box="[1401,1414,817,843]" captionStart="Figure 5" captionStartId="7.[113,178,1768,1792]" captionTargetBox="[115,1456,146,1738]" captionTargetId="figure-6@7.[112,1459,144,1740]" captionTargetPageId="7" captionText="Figure 5. Iolania Ƒankanstonei, male genitalia.A, genital segment, less lateral aspect. B, same, caudal aspect. C, anal segment, right lateral aspect.D, same, caudal aspect. E, less gonostvle, maximal aspect.F, aedeagus, dorsal aspect.G, same, less lateral aspect. Scale bars: 0.1 mm. Male from CKM entrance, same as in Figure 4." figureDoi="http://doi.org/10.5281/zenodo.14423207" httpUri="https://zenodo.org/record/14423207/files/figure.png" pageId="4" pageNumber="5">5</figureCitation>
|
||
), as represented by
|
||
<taxonomicName id="4C4A20E93150BF01193DFCE2FAC0FCB8" box="[1137,1298,848,874]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ssankanstonei">I. frankanstonei</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF55B6A3150BF011E75FCC4FA6DFB7B" blockId="4.[825,1475,817,1193]" pageId="4" pageNumber="5">
|
||
Total length: males,
|
||
<quantity id="4CB2F68F3150BF01194BFCC5FB52FC5C" box="[1031,1152,886,910]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="4" pageNumber="5" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
|
||
; females,
|
||
<quantity id="4CB2F68F3150BF0119A9FCC4FA8CFC5C" box="[1253,1374,887,911]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="4" pageNumber="5" unit="mm" value="4.5" valueMax="4.6" valueMin="4.4">4.4–4.6 mm</quantity>
|
||
. Tegmina 2× longer than maximallv wide, shallowlv tectiform, not surpassing tip of anal segment.Costal veins of tegmina in dorsal aspect convex, together creating an oval outline (
|
||
<figureCitation id="137147EF3150BF0119A7FC66FAE7FC3F" box="[1259,1333,981,1005]" captionStart="Figure 4" captionStartId="5.[113,178,1954,1978]" captionTargetBox="[359,1212,820,1927]" captionTargetId="figure-313@5.[356,1216,817,1928]" captionTargetPageId="5" captionText="Figure 4. Iolania Ƒankanstonei, male, wing morphologv. A, right tegmen. B, right hind wing.Scale bar:0.5 mm. Male from CKM entrance." figureDoi="http://doi.org/10.5281/zenodo.14423205" httpUri="https://zenodo.org/record/14423205/files/figure.png" pageId="4" pageNumber="5">Fig. 4A</figureCitation>
|
||
). Vertex with lateral margins stronglv diverging posteriorlv and with posterior margin shallowlv concave; compound eves small, pronotum in dorsal aspect mediallv narrow and laterallv expanded. Compound eves with light red pigmentation, coloration of bodv vellowish to light brown, tegmina with darker spots and stripes, variable among specimens studied. Hind tibiae laterallv with three minute spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35008E13150BF011F58FB5FFA75F865" pageId="4" pageNumber="5" type="key">
|
||
<paragraph id="8BF55B6A3150BF011F58FB5FFA75FA51" blockId="4.[155,1449,1259,1975]" pageId="4" pageNumber="5">
|
||
Kev to
|
||
<taxonomicName id="4C4A20E93150BF011F2CFB58FD28FAD7" box="[608,762,1259,1285]" pageId="4" pageNumber="5">
|
||
<emphasis id="B93E87783150BF011F2CFB58FD7AFAD7" box="[608,680,1259,1285]" italics="true" pageId="4" pageNumber="5">Iolania</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(modified from Hoch 2006)
|
||
<key id="FFDB0C5B3150BF011DD7FA9EFA75F865" pageId="4" pageNumber="5">
|
||
<keyStep id="30BE43CF3150BF011DD7FA9EFA75F865" pageId="4" pageNumber="5">
|
||
<keyLead id="30BBF85F3150BF011DD7FA9EFA75FA51" pageId="4" pageNumber="5">
|
||
(1) Tegmina longer than twice their maximal width, well surpassing tip of abdomen; compound eves large and darklv pigmented; vertex with lateral margins slightlv diverging posteriorlv and with posterior margin deeplv concave; pronotum in dorsal aspect narrow throughout; bodv size of males
|
||
<quantity id="4CB2F68F3150BF011FA8FAD8FC8DFA51" box="[740,863,1387,1411]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.3" metricValueMin="4.8" pageId="4" pageNumber="5" unit="mm" value="5.55" valueMax="6.3" valueMin="4.8">4.8–6.3 mm</quantity>
|
||
, females
|
||
<quantity id="4CB2F68F3150BF011EF7FAD8FBE4FA51" box="[955,1078,1387,1411]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.1" metricValueMin="6.2" pageId="4" pageNumber="5" unit="mm" value="6.65" valueMax="7.1" valueMin="6.2">6.2–7.1 mm</quantity>
|
||
............................................................. 2
|
||
</keyLead>
|
||
<paragraph id="8BF55B6A3150BF011DD7FA38FA75F9F2" blockId="4.[155,1449,1259,1975]" pageId="4" pageNumber="5">
|
||
<keyLead id="30BBF85F3150BF011DD7FA38FA75F9F2" pageId="4" pageNumber="5">
|
||
- Tegmina twice as long as maximallv wide, not surpassing tip of anal segment; compound eves small and reddish; vertex with lateral margins stronglv diverging posteriorlv and with posterior margin shallowlv concave; pronotum in dorsal aspect mediallv narrow and laterallv expanded; bodv size of males
|
||
<quantity id="4CB2F68F3150BF011E69FA7AFC70FA33" box="[805,930,1481,1505]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="4" pageNumber="5" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
|
||
, females
|
||
<quantity id="4CB2F68F3150BF011EB2FA7AFBABFA33" box="[1022,1145,1481,1505]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="4" pageNumber="5" unit="mm" value="4.5" valueMax="4.6" valueMin="4.4">4.4–4.6 mm</quantity>
|
||
.................................................... .....................................................................................................................................................................
|
||
<emphasis id="B93E87783150BF01193CFA54FA7AF9D3" bold="true" box="[1136,1448,1511,1537]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C4A20E93150BF01193CFA54FA80F9D2" authority="Fankanstonei" authorityName="Fankanstonei. B" authorityYear="2024" box="[1136,1362,1511,1536]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="sp. nov.">
|
||
<emphasis id="B93E87783150BF01193CFA54FA80F9D2" bold="true" box="[1136,1362,1511,1536]" italics="true" pageId="4" pageNumber="5">Iolania frankanstonei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20D3A033150BF01181AFA5AFA7AF9D3" box="[1366,1448,1513,1537]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(2) Male genitalia with aedeagus svmmetrical; anal segment without spinose processes ................................................................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF55B6A3150BF011DD7F99BFA75F9AC" blockId="4.[155,1449,1259,1975]" pageId="4" pageNumber="5">
|
||
<keyLead id="30BBF85F3150BF011DD7F99BFA75F9AC" pageId="4" pageNumber="5">
|
||
- Aedeagus with asvmmetrical paired spines; anal segment caudallv bent ventrad, with two distinct ventrolateral spinose processes ........................................................................................................................................................
|
||
<taxonomicName id="4C4A20E93150BF01192AF9F4FA75F98C" authority="Hoch, 2006" authorityName="Hoch" authorityYear="2006" box="[1126,1447,1606,1631]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kraussohana">
|
||
<emphasis id="B93E87783150BF01192AF9F4FAFBF98D" box="[1126,1321,1607,1631]" italics="true" pageId="4" pageNumber="5">Iolania kraussohana</emphasis>
|
||
Hoch, 2006
|
||
</taxonomicName>
|
||
(3) Shaft of aedeagus in lateral aspect curved dorsad; flagellum without sculpturing; gonostvli not ventrallv ridged .................. 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF55B6A3150BF011DD7F935FA75F90E" blockId="4.[155,1449,1259,1975]" pageId="4" pageNumber="5">
|
||
<keyLead id="30BBF85F3150BF011DD7F935FA75F90E" pageId="4" pageNumber="5">
|
||
- Shaft of aedeagus straight; flagellum sculptured with numerous distinct tuberculous protuberances; gonostvli ventrallv ridged .............................................................................................................................................................................................................. 5 (4) Paired spines of aedeagus foliatelv dilated ...................................................................................................................
|
||
<taxonomicName id="4C4A20E93150BF0119B3F977FA75F90E" authorityName="Giffard" authorityYear="1925" box="[1279,1447,1732,1756]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mauiensis">
|
||
<emphasis id="B93E87783150BF0119B3F977FA75F90E" box="[1279,1447,1732,1756]" italics="true" pageId="4" pageNumber="5">Iolania mauiensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF55B6A3150BF011DD7F957FA75F8C8" blockId="4.[155,1449,1259,1975]" pageId="4" pageNumber="5">
|
||
<keyLead id="30BBF85F3150BF011DD7F957FA75F8C8" pageId="4" pageNumber="5">
|
||
- Paired spines of aedeagus not dilated, arm-like, curved dorsad at midlength ....................................................
|
||
<taxonomicName id="4C4A20E93150BF0119BFF950FA75F929" authorityName="Giffard" authorityYear="1925" box="[1267,1447,1763,1787]" class="Insecta" family="Cixiidae" genus="Iolanaia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lanaiensis">
|
||
<emphasis id="B93E87783150BF0119BFF950FA75F929" box="[1267,1447,1763,1787]" italics="true" pageId="4" pageNumber="5">Iolanaia lanaiensis</emphasis>
|
||
</taxonomicName>
|
||
(5) Gonostvli with caudal margin triangular; tips of aedeagal spines directed straight basad or mediobasad ................................. 6
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF55B6A3150BF011DD7F891FA7AF8AB" blockId="4.[155,1449,1259,1975]" pageId="4" pageNumber="5">
|
||
<keyLead id="30BBF85F3150BF011DD7F891FA7AF8AB" pageId="4" pageNumber="5">
|
||
- Gonostvli with caudal margin rounded; tips of aedeagal spines directed dorsad ..................................................
|
||
<taxonomicName id="4C4A20E93150BF01185DF891FA75F8E8" authorityName="Kirkaldv" authorityYear="1902" box="[1297,1447,1826,1850]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="perkinsi">
|
||
<emphasis id="B93E87783150BF01185DF891FA75F8E8" box="[1297,1447,1826,1850]" italics="true" pageId="4" pageNumber="5">Iolania perkinsi</emphasis>
|
||
</taxonomicName>
|
||
(6) Male anal segment distallv with produced lateroventral corners; shaft of aedeagus in lateral aspect with ventral margin distinctlv narrowing in distal half; tips of aedeagal spines curved mediad .................................................................
|
||
<taxonomicName id="4C4A20E93150BF01184CF8D2FA7AF8AB" authorityName="Giffard" authorityYear="1925" box="[1280,1448,1889,1913]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="oahuensis">
|
||
<emphasis id="B93E87783150BF01184CF8D2FA7AF8AB" box="[1280,1448,1889,1913]" italics="true" pageId="4" pageNumber="5">Iolania oahuensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF55B6A3150BF011DD7F833FA75F865" blockId="4.[155,1449,1259,1975]" pageId="4" pageNumber="5">
|
||
<keyLead id="30BBF85F3150BF011DD7F833FA75F865" pageId="4" pageNumber="5">
|
||
- Male anal segment in caudal aspect ventrallv rounded; shaft of aedeagus in lateral aspect not narrowing towards apex; tips of aedeagal spines directed straight basad .....................................................................................................................
|
||
<taxonomicName id="4C4A20E93150BF0119A3F82CFA75F865" authorityName="Giffard" authorityYear="1925" box="[1263,1447,1951,1975]" class="Insecta" family="Cixiidae" genus="Iolania" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="koolauensis">
|
||
<emphasis id="B93E87783150BF0119A3F82CFA75F865" box="[1263,1447,1951,1975]" italics="true" pageId="4" pageNumber="5">Iolania koolauensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |