treatments-xml/data/03/D4/6C/03D46C492E75FFDAFF54FA12FB71F88E.xml

<document id="D9444B31B0B2FE1E8908015EB1F9659C" ID-CLB-Dataset="58288" ID-DOI="10.11646/zootaxa.5141.2.1" ID-GBIF-Dataset="ddfca40f-e77f-4fc3-aeb9-ca5d91ce5572" ID-ISSN="1175-5326" ID-Zenodo-Dep="6581580" ID-ZooBank="DDD8181B-416B-4B8E-972D-5B62F6EE5399" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653496954074" checkinUser="plazi" docAuthor="Malonza, Patrick K. &amp; Bauer, Aaron M." docDate="2022" docId="03D46C492E75FFDAFF54FA12FB71F88E" docLanguage="en" docName="zootaxa.5141.2.1.pdf" docOrigin="Zootaxa 5141 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ancylodactylus chyuluensis Malonza &amp; Bauer 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="131" masterDocId="FFED14312E69FFC4FFC3FF90FF9CFFFF" masterDocTitle="Resurrection of the African gecko genus Ancylodactylus Müller, 1907 (Squamata: Gekkonidae) and description of six new species from Kenya" masterLastPageNumber="139" masterPageNumber="101" pageNumber="129" updateTime="1699338080367" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B85B62D80418109729B571EAF50B5C36">Resurrection of the African gecko genus Ancylodactylus Müller, 1907 (Squamata: Gekkonidae) and description of six new species from Kenya</mods:title>
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<mods:namePart id="134CDA741C9DAF670A6ADC10A5B8ADC6">Malonza, Patrick K.</mods:namePart>
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<mods:affiliation id="9317F2EF0BACFE3B5F58CBCBC054018B">National Museums of Kenya, Herpetology Section, Museum Hill Road, P. O. Box 40658 - 00100, Nairobi, Kenya malonzapk @ gmail. com; https: // orcid. org / 0000 - 0003 - 1426 - 2397</mods:affiliation>
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<mods:namePart id="C625BFE2E488730F0377B2CF45C084BF">Bauer, Aaron M.</mods:namePart>
<mods:affiliation id="EF291F87FC4ABE082B3C70C7D269264B">Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA.</mods:affiliation>
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<mods:date id="F12548DBC38741F6D56E640A94D56761">2022</mods:date>
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<treatment id="03D46C492E75FFDAFF54FA12FB71F88E" ID-DOI="http://doi.org/10.5281/zenodo.6587160" ID-GBIF-Taxon="195657748" ID-Zenodo-Dep="6587160" LSID="urn:lsid:plazi:treatment:03D46C492E75FFDAFF54FA12FB71F88E" httpUri="http://treatment.plazi.org/id/03D46C492E75FFDAFF54FA12FB71F88E" lastPageId="30" lastPageNumber="131" pageId="28" pageNumber="129">
<subSubSection id="C3678ED42E75FFD8FF54FA12FD86FA3E" pageId="28" pageNumber="129" type="nomenclature">
<paragraph id="8BC2DD5F2E75FFD8FF54FA12FDD7FA62" blockId="28.[151,587,1410,1509]" box="[151,587,1410,1437]" pageId="28" pageNumber="129">
<heading id="D08A6A332E75FFD8FF54FA12FDD7FA62" bold="true" box="[151,587,1410,1437]" fontSize="11" level="1" pageId="28" pageNumber="129" reason="1">
<emphasis id="B909014D2E75FFD8FF54FA12FDD7FA62" bold="true" box="[151,587,1410,1437]" pageId="28" pageNumber="129">
<taxonomicName id="4C7DA6DC2E75FFD8FF54FA12FE7AFA62" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[151,486,1410,1437]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="129" phylum="Chordata" rank="species" species="chyuluensis" status="sp. nov.">
<emphasis id="B909014D2E75FFD8FF54FA12FE7AFA62" bold="true" box="[151,486,1410,1437]" italics="true" pageId="28" pageNumber="129">Ancylodactylus chyuluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23ABC362E75FFD8FE2EFA13FDD7FA62" box="[493,587,1411,1437]" pageId="28" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC2DD5F2E75FFD8FF54FA37FD86FA3E" blockId="28.[151,587,1410,1509]" box="[151,538,1447,1473]" pageId="28" pageNumber="129">Chyulu Hills Pygmy Forest Gecko</paragraph>
</subSubSection>
<subSubSection id="C3678ED42E75FFD8FF54FA5BFE8AFA1A" box="[151,278,1483,1509]" pageId="28" pageNumber="129" type="description">
<paragraph id="8BC2DD5F2E75FFD8FF54FA5BFE8AFA1A" blockId="28.[151,587,1410,1509]" box="[151,278,1483,1509]" pageId="28" pageNumber="129">
(
<figureCitation id="1346C1DA2E75FFD8FF5CFA5BFE91FA1A" box="[159,269,1483,1509]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="28" pageNumber="129">Figure 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3678ED42E75FFD8FF54F983FA85F9AE" pageId="28" pageNumber="129" type="materials_examined">
<paragraph id="8BC2DD5F2E75FFD8FF54F983FA85F9AE" blockId="28.[151,1437,1554,2013]" pageId="28" pageNumber="129">
<materialsCitation id="3B15D7022E75FFD8FF54F983FA85F9AE" ID-GBIF-Occurrence="3785192319" collectingDate="2016-09-21" collectorName="Patrick K. Malonza &amp; Felista K. Kilunda &amp; Justus Ochong." country="Kenya" elevation="1200" latitude="-2.56017" location="Chyulu Hills National Park" longLatPrecision="1" longitude="37.88877" pageId="28" pageNumber="129" specimenCount="1" stateProvince="Makueni" typeStatus="holotype">
<emphasis id="B909014D2E75FFD8FF54F983FE97F9D2" bold="true" box="[151,267,1555,1581]" pageId="28" pageNumber="129">
<typeStatus id="54C663FD2E75FFD8FF54F983FE9AF9D2" box="[151,262,1555,1581]" pageId="28" pageNumber="129" type="holotype">Holotype</typeStatus>
.
</emphasis>
NMK-L3869, adult male, 
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, 
<collectingRegion id="49B913BD2E75FFD8FCB2F983FBB2F9D2" box="[881,1070,1555,1581]" country="Kenya" name="Makueni" pageId="28" pageNumber="129">Makueni County</collectingRegion>
, 
<collectingCountry id="F36A9DCF2E75FFD8FBFBF983FB1DF9D2" box="[1080,1153,1555,1581]" name="Kenya" pageId="28" pageNumber="129">Kenya</collectingCountry>
(
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, 
<geoCoordinate id="EE49BB982E75FFD8FAEBF983FF37F9AE" degrees="37.88877" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="1" value="37.88877">37.88877° E</geoCoordinate>
; 
<quantity id="4C8570BA2E75FFD8FF74F9A7FE93F9AE" box="[183,271,1591,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="129" unit="m" value="1200.0">
<elevation id="00503A6C2E75FFD8FF74F9A7FE93F9AE" box="[183,271,1591,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="129" unit="m" value="1200.0">1200 m</elevation>
</quantity>
), collected 
<date id="FFC3FB9F2E75FFD8FE4CF9A7FDF5F9AE" box="[399,617,1591,1617]" pageId="28" pageNumber="129" value="2016-09-21">
<collectingDate id="EF8702772E75FFD8FE4CF9A7FDF5F9AE" box="[399,617,1591,1617]" pageId="28" pageNumber="129" value="2016-09-21">September 21 2016</collectingDate>
</date>
by 
<collectorName id="2688B8892E75FFD8FD50F9A7FCEDF9AE" box="[659,881,1591,1617]" pageId="28" pageNumber="129">Patrick K. Malonza</collectorName>
, 
<collectorName id="2688B8892E75FFD8FCBFF9A7FBD5F9AE" box="[892,1097,1591,1617]" pageId="28" pageNumber="129">Felista K. Kilunda</collectorName>
&amp; 
<collectorName id="2688B8892E75FFD8FBAEF9A7FA85F9AE" box="[1133,1305,1591,1617]" pageId="28" pageNumber="129">Justus Ochong.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3678ED42E75FFD9FF04F9CAFCDAFC12" lastPageId="29" lastPageNumber="130" pageId="28" pageNumber="129" type="diagnosis">
<paragraph id="8BC2DD5F2E75FFD8FF04F9CAFD19F86A" blockId="28.[151,1437,1554,2013]" pageId="28" pageNumber="129">
<emphasis id="B909014D2E75FFD8FF04F9CAFEDEF98B" bold="true" box="[199,322,1626,1652]" pageId="28" pageNumber="129">Diagnosis.</emphasis>
An extremely small-sized 
<taxonomicName id="4C7DA6DC2E75FFD8FDAFF9CBFC8BF98A" authorityName="Muller" authorityYear="1907" box="[620,791,1627,1653]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="129" phylum="Chordata" rank="genus">
<emphasis id="B909014D2E75FFD8FDAFF9CBFC8BF98A" box="[620,791,1627,1653]" italics="true" pageId="28" pageNumber="129">Ancylodactylus</emphasis>
</taxonomicName>
with a SVL of 
<quantity id="4C8570BA2E75FFD8FC7CF9CBFB92F98A" box="[959,1038,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="28" pageNumber="129" unit="mm" value="28.0">28 mm</quantity>
in the unique 
<typeStatus id="54C663FD2E75FFD8FB69F9CBFB46F98A" box="[1194,1242,1627,1653]" pageId="28" pageNumber="129">type</typeStatus>
. Dorsal scalation mostly homogeneous with a single pair of rows of very small tubercles along the flanks; other scales minute and granular. Limbs and digits long, with enlarged basal lamellae under and proximal to penultimate interphalangeal joint (3 under digit IV). Length of intact original tail slightly longer than SVL. Tail dorsum distal to the pygal portion of the tail atuberculate; median subcaudal scales in a single row of large, but not transversely widened scales. Male precloacal pores in a single continuous row of 8. Dorsal pattern of pale fleurs-de-lis and chevrons as well as pale and dark spots on spots on a gray-brown to olive background; tail with pairs of dark brown and ashy chevrons or crossbands. Ventral coloration grayish with darker markings on the edges of scales yielding a mottled pattern; throat and chin mottled with a bluish center.
</paragraph>
<paragraph id="8BC2DD5F2E75FFD9FF04F80EFCD6FD0E" blockId="28.[151,1437,1554,2013]" lastBlockId="29.[151,1437,151,2013]" lastPageId="29" lastPageNumber="130" pageId="28" pageNumber="129">
<emphasis id="B909014D2E75FFD8FF04F80EFDAFF847" bold="true" box="[199,563,1950,1976]" pageId="28" pageNumber="129">Comparisons with Congeners.</emphasis>
<taxonomicName id="4C7DA6DC2E75FFD8FDFFF80FFCEFF846" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[572,883,1951,1977]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="129" phylum="Chordata" rank="species" species="chyuluensis" status="sp. nov.">
<emphasis id="B909014D2E75FFD8FDFFF80FFCEFF846" box="[572,883,1951,1977]" italics="true" pageId="28" pageNumber="129">Ancylodactylus chyuluensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E75FFD8FCB8F80FFC4BF846" bold="true" box="[891,983,1951,1977]" pageId="28" pageNumber="129">
<taxonomicNameLabel id="A23ABC362E75FFD8FCB8F80FFC4BF846" box="[891,983,1951,1977]" pageId="28" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
may be distinguished from 
<taxonomicName id="4C7DA6DC2E75FFD8FADAF80FFF2DF822" authorityName="Muller" authorityYear="1907" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="129" phylum="Chordata" rank="species" species="spinicollis">
<emphasis id="B909014D2E75FFD8FADAF80FFF2DF822" italics="true" pageId="28" pageNumber="129">A. spinicollis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7DA6DC2E75FFD8FF2AF853FE17F822" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[233,395,1987,2013]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="129" phylum="Chordata" rank="species" species="petrodroma">
<emphasis id="B909014D2E75FFD8FF2AF853FE17F822" box="[233,395,1987,2013]" italics="true" pageId="28" pageNumber="129">A. petrodroma</emphasis>
</taxonomicName>
in lacking an enlarged preaxial metatarsal scale and from these two species plus 
<taxonomicName id="4C7DA6DC2E75FFD8FAE2F853FA00F822" authorityName="Bauer, Chirio, Ineich &amp; LeBreton" authorityYear="2006" box="[1313,1436,1987,2013]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="129" phylum="Chordata" rank="species" species="alantika">
<emphasis id="B909014D2E75FFD8FAE2F853FA00F822" box="[1313,1436,1987,2013]" italics="true" pageId="28" pageNumber="129">A. alantika</emphasis>
</taxonomicName>
in having a series of flattened, rectangular lamellae subtending the second and third phalanges of the pedal digits, rather than single enlarged, rounded scale (plaque of 
<bibRefCitation id="EFECA0AE2E74FFD9FD34FF2BFCE0FF29" author="Perret, J. - L." box="[759,892,187,214]" pageId="29" pageNumber="130" pagination="457 - 505" refId="ref27183" refString="Perret, J. - L. (1986) Revision des especes africaines du genre Cnemaspis Strauch, sous-genre Ancylodactylus Muller (Lacertilia, Gekkonidae), avec la description de quatre especes Nouvelles. Revue Suisse de Zoologie, 93, 457 - 505. https: // doi. org / 10.5962 / bhl. part. 79705" type="journal article" year="1986">Perret 1986</bibRefCitation>
) at the penultimate joint of each digit. It is distinguished from 
<taxonomicName id="4C7DA6DC2E74FFD9FE91FF4FFE7EFF07" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[338,482,223,249]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="uzungwae">
<emphasis id="B909014D2E74FFD9FE91FF4FFE7EFF07" box="[338,482,223,249]" italics="true" pageId="29" pageNumber="130">A. uzungwae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FE32FF4FFD56FF07" authorityName="Loveridge" authorityYear="1947" box="[497,714,223,249]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="quattuorseriatus">
<emphasis id="B909014D2E74FFD9FE32FF4FFD56FF07" box="[497,714,223,249]" italics="true" pageId="29" pageNumber="130">A. quattuorseriatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FD1AFF4FFC1CFF06" baseAuthorityName="Schmidt" baseAuthorityYear="1919" box="[729,896,223,249]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="dickersonae">
<emphasis id="B909014D2E74FFD9FD1AFF4FFC1CFF06" box="[729,896,223,249]" italics="true" pageId="29" pageNumber="130">A. dickersonae</emphasis>
</taxonomicName>
(but see 
<bibRefCitation id="EFECA0AE2E74FFD9FC2AFF4FFB23FF06" author="Spawls, S. &amp; Howell, K. &amp; Hinkel, H. &amp; Menegon, M." box="[1001,1215,223,249]" pageId="29" pageNumber="130" refId="ref27759" refString="Spawls, S., Howell, K. Hinkel, H. &amp; Menegon, M. (2018) Field Guide to East African Reptiles. 2 nd Edition. Bloomsbury Publishing, London, 624 pp." type="book" year="2018">
Spawls 
<emphasis id="B909014D2E74FFD9FB81FF4FFBE7FF06" box="[1090,1147,223,249]" italics="true" pageId="29" pageNumber="130">et al.</emphasis>
2018
</bibRefCitation>
), 
<taxonomicName id="4C7DA6DC2E74FFD9FB10FF4FFAE9FF06" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[1235,1397,223,249]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="petrodroma">
<emphasis id="B909014D2E74FFD9FB10FF4FFAE9FF06" box="[1235,1397,223,249]" italics="true" pageId="29" pageNumber="130">A. petrodroma</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FA47FF4FFE82FEE2" baseAuthorityName="Angel" baseAuthorityYear="1943" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="B909014D2E74FFD9FA47FF4FFE82FEE2" italics="true" pageId="29" pageNumber="130">A. occidentalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FEEDFE93FE5DFEE2" authorityName="Muller" authorityYear="1907" box="[302,449,259,285]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="spinicollis">
<emphasis id="B909014D2E74FFD9FEEDFE93FE5DFEE2" box="[302,449,259,285]" italics="true" pageId="29" pageNumber="130">A. spinicollis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FE12FE93FDD1FEE2" authorityName="Bauer, Chirio, Ineich &amp; LeBreton" authorityYear="2006" box="[465,589,259,285]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="alantika">
<emphasis id="B909014D2E74FFD9FE12FE93FDD1FEE2" box="[465,589,259,285]" italics="true" pageId="29" pageNumber="130">A. alantika</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FD9FFE93FD7BFEE2" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[604,743,259,285]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="kituiensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FD9FFE93FD7BFEE2" box="[604,743,259,285]" italics="true" pageId="29" pageNumber="130">A. kituiensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FD2CFE93FCD5FEE2" bold="true" box="[751,841,259,285]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FD2CFE93FCD5FEE2" box="[751,841,259,285]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and 
<taxonomicName id="4C7DA6DC2E74FFD9FC4AFE93FC62FEE2" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[905,1022,259,285]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="spawlsi" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FC4AFE93FC62FEE2" box="[905,1022,259,285]" italics="true" pageId="29" pageNumber="130">A. spawlsi</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FBC5FE93FBFDFEE2" bold="true" box="[1030,1121,259,285]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FBC5FE93FBFDFEE2" box="[1030,1121,259,285]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by lacking tubercles on the post-pygal (autotomic) portion of the tail dorsum. It has only two tubercle rows on the trunk, one on each flank, a feature that differentiates it form all congeners except some 
<taxonomicName id="4C7DA6DC2E74FFD9FC8AFEDBFC6DFE9A" baseAuthorityName="Schmidt" baseAuthorityYear="1919" box="[841,1009,331,357]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="dickersonae">
<emphasis id="B909014D2E74FFD9FC8AFEDBFC6DFE9A" box="[841,1009,331,357]" italics="true" pageId="29" pageNumber="130">A. dickersonae</emphasis>
</taxonomicName>
(0–6 rows fide 
<bibRefCitation id="EFECA0AE2E74FFD9FB69FEDBFAACFE9A" author="Perret, J. - L." box="[1194,1328,331,357]" pageId="29" pageNumber="130" pagination="457 - 505" refId="ref27183" refString="Perret, J. - L. (1986) Revision des especes africaines du genre Cnemaspis Strauch, sous-genre Ancylodactylus Muller (Lacertilia, Gekkonidae), avec la description de quatre especes Nouvelles. Revue Suisse de Zoologie, 93, 457 - 505. https: // doi. org / 10.5962 / bhl. part. 79705" type="journal article" year="1986">Perret 1986</bibRefCitation>
; 0–4 fide 
<bibRefCitation id="EFECA0AE2E74FFD9FF54FEFFFEFBFE76" author="Spawls, S. &amp; Howell, K. &amp; Hinkel, H. &amp; Menegon, M." box="[151,359,367,393]" pageId="29" pageNumber="130" refId="ref27759" refString="Spawls, S., Howell, K. Hinkel, H. &amp; Menegon, M. (2018) Field Guide to East African Reptiles. 2 nd Edition. Bloomsbury Publishing, London, 624 pp." type="book" year="2018">
Spawls 
<emphasis id="B909014D2E74FFD9FF2CFEFFFEBCFE76" box="[239,288,367,393]" italics="true" pageId="29" pageNumber="130">et al</emphasis>
. 2018
</bibRefCitation>
) and 
<taxonomicName id="4C7DA6DC2E74FFD9FE67FEFFFDC1FE76" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[420,605,367,393]" class="Reptilia" family="Lacertidae" genus="Adolfus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="mathewsensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FE67FEFFFDC1FE76" box="[420,605,367,393]" italics="true" pageId="29" pageNumber="130">A. mathewsensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FDA0FEFFFD21FE76" bold="true" box="[611,701,367,393]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FDA0FEFFFD21FE76" box="[611,701,367,393]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(0–2 rows), and 
<taxonomicName id="4C7DA6DC2E74FFD9FCB9FEFFFC70FE76" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[890,1004,367,393]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="spawlsi" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FCB9FEFFFC70FE76" box="[890,1004,367,393]" italics="true" pageId="29" pageNumber="130">A. spawlsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23ABC362E74FFD9FC30FEFFFBD2FE76" box="[1011,1102,367,393]" pageId="29" pageNumber="130" rank="species">
<emphasis id="B909014D2E74FFD9FC30FEFFFBDBFE76" bold="true" box="[1011,1095,367,393]" pageId="29" pageNumber="130">sp. nov</emphasis>
.
</taxonomicNameLabel>
(2 rows). It possesses a lower number of precloacal pores (8) than 
<taxonomicName id="4C7DA6DC2E74FFD9FDF7FE03FD22FE52" box="[564,702,403,429]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="africanus">
<emphasis id="B909014D2E74FFD9FDF7FE03FD22FE52" box="[564,702,403,429]" italics="true" pageId="29" pageNumber="130">A. africanus</emphasis>
</taxonomicName>
(9–12), 
<taxonomicName id="4C7DA6DC2E74FFD9FCDCFE03FC3FFE52" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[799,931,403,429]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="barbouri">
<emphasis id="B909014D2E74FFD9FCDCFE03FC3FFE52" box="[799,931,403,429]" italics="true" pageId="29" pageNumber="130">A. barbouri</emphasis>
</taxonomicName>
(14), 
<taxonomicName id="4C7DA6DC2E74FFD9FC2BFE03FBF8FE52" authorityName="Bauer, Chirio, Ineich &amp; LeBreton" authorityYear="2006" box="[1000,1124,403,429]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="alantika">
<emphasis id="B909014D2E74FFD9FC2BFE03FBF8FE52" box="[1000,1124,403,429]" italics="true" pageId="29" pageNumber="130">A. alantika</emphasis>
</taxonomicName>
(11), and 
<taxonomicName id="4C7DA6DC2E74FFD9FB1BFE03FAA9FE53" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[1240,1333,403,429]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="gigas">
<emphasis id="B909014D2E74FFD9FB1BFE03FAA9FE53" box="[1240,1333,403,429]" italics="true" pageId="29" pageNumber="130">A. gigas</emphasis>
</taxonomicName>
(15–16), but has a greater number than 
<taxonomicName id="4C7DA6DC2E74FFD9FE21FE27FD06FE2E" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[482,666,439,465]" class="Reptilia" family="Lacertidae" genus="Adolfus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="mathewsensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FE21FE27FD06FE2E" box="[482,666,439,465]" italics="true" pageId="29" pageNumber="130">A. mathewsensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FD63FE27FD64FE2E" bold="true" box="[672,760,439,465]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FD63FE27FD64FE2E" box="[672,760,439,465]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(6–7), 
<taxonomicName id="4C7DA6DC2E74FFD9FC84FE27FC79FE2E" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[839,997,439,465]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="laikipiensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FC84FE27FC79FE2E" box="[839,997,439,465]" italics="true" pageId="29" pageNumber="130">A. laikipiensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FC29FE27FBDEFE2E" bold="true" box="[1002,1090,439,465]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FC29FE27FBDEFE2E" box="[1002,1090,439,465]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(7), and 
<taxonomicName id="4C7DA6DC2E74FFD9FB60FE27FA89FE2E" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[1187,1301,439,465]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="spawlsi" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FB60FE27FA89FE2E" box="[1187,1301,439,465]" italics="true" pageId="29" pageNumber="130">A. spawlsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23ABC362E74FFD9FAD8FE27FAEEFE2D" box="[1307,1394,439,466]" pageId="29" pageNumber="130" rank="species">
<emphasis id="B909014D2E74FFD9FAD8FE27FAF2FE2E" bold="true" box="[1307,1390,439,465]" pageId="29" pageNumber="130">sp. nov</emphasis>
.
</taxonomicNameLabel>
(6). It may also be distinguished in having a single median series of enlarged, but not transversely widened subcaudals in contrast to 
<taxonomicName id="4C7DA6DC2E74FFD9FEF7FE6FFE5BFDE6" authorityName="Muller" authorityYear="1907" box="[308,455,511,537]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="spinicollis">
<emphasis id="B909014D2E74FFD9FEF7FE6FFE5BFDE6" box="[308,455,511,537]" italics="true" pageId="29" pageNumber="130">A. spinicollis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FE15FE6FFDE4FDE6" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[470,632,511,537]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="petrodroma">
<emphasis id="B909014D2E74FFD9FE15FE6FFDE4FDE6" box="[470,632,511,537]" italics="true" pageId="29" pageNumber="130">A. petrodroma</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7DA6DC2E74FFD9FD7BFE6FFCC3FDE6" baseAuthorityName="Angel" baseAuthorityYear="1943" box="[696,863,511,537]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="B909014D2E74FFD9FD7BFE6FFCC3FDE6" box="[696,863,511,537]" italics="true" pageId="29" pageNumber="130">A. occidentalis</emphasis>
</taxonomicName>
(irregular subcaudals), 
<taxonomicName id="4C7DA6DC2E74FFD9FBAFFE6FFA9EFDE6" baseAuthorityName="Loveridge" baseAuthorityYear="1936" box="[1132,1282,511,537]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="elgonensis">
<emphasis id="B909014D2E74FFD9FBAFFE6FFA9EFDE6" box="[1132,1282,511,537]" italics="true" pageId="29" pageNumber="130">A. elgonensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FAD2FE6FFA0AFDE6" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[1297,1430,511,537]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="barbouri">
<emphasis id="B909014D2E74FFD9FAD2FE6FFA0AFDE6" box="[1297,1430,511,537]" italics="true" pageId="29" pageNumber="130">A. barbouri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FF54FDB3FEBAFDC3" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[151,294,547,573]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="uzungwae">
<emphasis id="B909014D2E74FFD9FF54FDB3FEBAFDC3" box="[151,294,547,573]" italics="true" pageId="29" pageNumber="130">A. uzungwae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FEF0FDB3FE57FDC2" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[307,459,547,573]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="kenyaensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FEF0FDB3FED8FDC2" box="[307,324,547,573]" italics="true" pageId="29" pageNumber="130">A</emphasis>
. 
<emphasis id="B909014D2E74FFD9FE91FDB3FE57FDC2" box="[338,459,547,573]" italics="true" pageId="29" pageNumber="130">kenyaensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FE11FDB3FDB7FDC2" bold="true" box="[466,555,547,573]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FE11FDB3FDB7FDC2" box="[466,555,547,573]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and 
<taxonomicName id="4C7DA6DC2E74FFD9FDABFDB3FD6CFDC2" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[616,752,547,573]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="kituiensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FDABFDB3FD6CFDC2" box="[616,752,547,573]" italics="true" pageId="29" pageNumber="130">A. kituiensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FD34FDB3FCCCFDC2" bold="true" box="[759,848,547,573]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FD34FDB3FCCCFDC2" box="[759,848,547,573]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(alternating single and paired scales), and 
<taxonomicName id="4C7DA6DC2E74FFD9FAEFFDB3FA0AFDC2" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1324,1430,547,573]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="dilepis">
<emphasis id="B909014D2E74FFD9FAEFFDB3FA0AFDC2" box="[1324,1430,547,573]" italics="true" pageId="29" pageNumber="130">A. dilepis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FF54FDD7FF68FD9F" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[151,244,583,609]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="gigas">
<emphasis id="B909014D2E74FFD9FF54FDD7FF68FD9F" box="[151,244,583,609]" italics="true" pageId="29" pageNumber="130">A. gigas</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FEC1FDD7FE17FD9E" box="[258,395,583,609]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="africanus">
<emphasis id="B909014D2E74FFD9FEC1FDD7FE17FD9E" box="[258,395,583,609]" italics="true" pageId="29" pageNumber="130">A. africanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FE59FDD7FDEEFD9F" authorityName="Loveridge" authorityYear="1947" box="[410,626,583,609]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="quattuorseriatus">
<emphasis id="B909014D2E74FFD9FE59FDD7FE37FD9E" box="[410,427,583,609]" italics="true" pageId="29" pageNumber="130">A</emphasis>
. 
<emphasis id="B909014D2E74FFD9FE7AFDD7FDEEFD9F" box="[441,626,583,608]" italics="true" pageId="29" pageNumber="130">quattuorseriatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FD42FDD7FCBBFD9E" baseAuthorityName="Schmidt" baseAuthorityYear="1919" box="[641,807,583,609]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="dickersonae">
<emphasis id="B909014D2E74FFD9FD42FDD7FCBBFD9E" box="[641,807,583,609]" italics="true" pageId="29" pageNumber="130">A. dickersonae</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7DA6DC2E74FFD9FCA5FDD7FC7EFD9E" baseAuthorityName="Mertens" baseAuthorityYear="1937" box="[870,994,583,609]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="koehleri">
<emphasis id="B909014D2E74FFD9FCA5FDD7FC7EFD9E" box="[870,994,583,609]" italics="true" pageId="29" pageNumber="130">A. koehleri</emphasis>
</taxonomicName>
, (single row of median subcaudals, but transversely widened or not uniform throughout). In lacking any yellow or orange ventral coloration 
<taxonomicName id="4C7DA6DC2E74FFD9FB38FDFBFA01FD7A" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[1275,1437,619,645]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="chyuluensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FB38FDFBFA01FD7A" box="[1275,1437,619,645]" italics="true" pageId="29" pageNumber="130">A. chyuluensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FF54FD1FFF6CFD56" bold="true" box="[151,240,655,681]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FF54FD1FFF6CFD56" box="[151,240,655,681]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
may be distinguished from 
<taxonomicName id="4C7DA6DC2E74FFD9FDE7FD1FFD30FD56" box="[548,684,655,681]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="africanus">
<emphasis id="B909014D2E74FFD9FDE7FD1FFD30FD56" box="[548,684,655,681]" italics="true" pageId="29" pageNumber="130">A. africanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FD7AFD1FFCD1FD56" baseAuthorityName="Loveridge" baseAuthorityYear="1936" box="[697,845,655,681]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="elgonensis">
<emphasis id="B909014D2E74FFD9FD7AFD1FFCD1FD56" box="[697,845,655,681]" italics="true" pageId="29" pageNumber="130">A. elgonensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FC99FD1FFC48FD56" baseAuthorityName="Mertens" baseAuthorityYear="1937" box="[858,980,655,681]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="koehleri">
<emphasis id="B909014D2E74FFD9FC99FD1FFC48FD56" box="[858,980,655,681]" italics="true" pageId="29" pageNumber="130">A. koehleri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FC22FD1FFBD6FD56" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[993,1098,655,681]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="dilepis">
<emphasis id="B909014D2E74FFD9FC22FD1FFBD6FD56" box="[993,1098,655,681]" italics="true" pageId="29" pageNumber="130">A. dilepis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FB94FD1FFB74FD56" authorityName="Muller" authorityYear="1907" box="[1111,1256,655,681]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="spinicollis">
<emphasis id="B909014D2E74FFD9FB94FD1FFB74FD56" box="[1111,1256,655,681]" italics="true" pageId="29" pageNumber="130">A. spinicollis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FB36FD1FFA09FD56" baseAuthorityName="Perret" baseAuthorityYear="1986" box="[1269,1429,655,681]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="petrodroma">
<emphasis id="B909014D2E74FFD9FB36FD1FFA09FD56" box="[1269,1429,655,681]" italics="true" pageId="29" pageNumber="130">A. petrodroma</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FF54FD23FEA3FD32" baseAuthorityName="Angel" baseAuthorityYear="1943" box="[151,319,691,717]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="occidentalis">
<emphasis id="B909014D2E74FFD9FF54FD23FEA3FD32" box="[151,319,691,717]" italics="true" pageId="29" pageNumber="130">A. occidentalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FE8CFD23FE50FD32" authorityName="Bauer, Chirio, Ineich &amp; LeBreton" authorityYear="2006" box="[335,460,691,717]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="alantika">
<emphasis id="B909014D2E74FFD9FE8CFD23FE50FD32" box="[335,460,691,717]" italics="true" pageId="29" pageNumber="130">A. alantika</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FE1FFD23FD0BFD32" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[476,663,691,717]" class="Reptilia" family="Lacertidae" genus="Adolfus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="mathewsensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FE1FFD23FD0BFD32" box="[476,663,691,717]" italics="true" pageId="29" pageNumber="130">A. mathewsensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23ABC362E74FFD9FD5CFD23FD67FD32" box="[671,763,691,717]" pageId="29" pageNumber="130" rank="species">
<emphasis id="B909014D2E74FFD9FD5CFD23FD6AFD32" bold="true" box="[671,758,691,717]" pageId="29" pageNumber="130">sp. nov</emphasis>
.
</taxonomicNameLabel>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FCCFFD23FC31FD32" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[780,941,691,717]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="laikipiensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FCCFFD23FC31FD32" box="[780,941,691,717]" italics="true" pageId="29" pageNumber="130">A. laikipiensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FC76FD23FB8DFD32" bold="true" box="[949,1041,691,717]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FC76FD23FB8DFD32" box="[949,1041,691,717]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and 
<taxonomicName id="4C7DA6DC2E74FFD9FB91FD23FB5BFD32" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[1106,1223,691,717]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="spawlsi" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FB91FD23FB5BFD32" box="[1106,1223,691,717]" italics="true" pageId="29" pageNumber="130">A. spawlsi</emphasis>
</taxonomicName>
, 
<emphasis id="B909014D2E74FFD9FB14FD23FAAEFD32" bold="true" box="[1239,1330,691,717]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FB14FD23FAAEFD32" box="[1239,1330,691,717]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
This last feature also distinguishes the new species from 
<taxonomicName id="4C7DA6DC2E74FFD9FD6EFD47FCDEFD0E" baseAuthorityName="Loveridge" baseAuthorityYear="1936" box="[685,834,727,753]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="elgonensis">
<emphasis id="B909014D2E74FFD9FD6EFD47FCDEFD0E" box="[685,834,727,753]" italics="true" pageId="29" pageNumber="130">A. elgonensis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph id="8BC2DD5F2E74FFD9FF04FD6BFCDAFC12" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="130">
With the only known specimen having a SVL of 
<quantity id="4C8570BA2E74FFD9FCC7FD6BFCC4FCEA" box="[772,856,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="29" pageNumber="130" unit="mm" value="28.0">28 mm</quantity>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FCA5FD6BFB02FCEA" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[870,1182,763,789]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="chyuluensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FCA5FD6BFB02FCEA" box="[870,1182,763,789]" italics="true" pageId="29" pageNumber="130">Ancylodactylus chyuluensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FB64FD6BFA98FCEA" bold="true" box="[1191,1284,763,789]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FB64FD6BFA98FCEA" box="[1191,1284,763,789]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the smallest known 
<taxonomicName id="4C7DA6DC2E74FFD9FED4FC8FFE5EFCC6" authorityName="Muller" authorityYear="1907" box="[279,450,799,825]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="genus">
<emphasis id="B909014D2E74FFD9FED4FC8FFE5EFCC6" box="[279,450,799,825]" italics="true" pageId="29" pageNumber="130">Ancylodactylus</emphasis>
</taxonomicName>
. Although diminutive (&lt;
<quantity id="4C8570BA2E74FFD9FD2EFC8FFCA3FCC6" box="[749,831,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="130" unit="mm" value="30.0">30 mm</quantity>
SVL) species of South Asian 
<taxonomicName id="4C7DA6DC2E74FFD9FB60FC8FFA82FCC6" authorityName="Strauch" authorityYear="1887" box="[1187,1310,799,825]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="genus">
<emphasis id="B909014D2E74FFD9FB60FC8FFA82FCC6" box="[1187,1310,799,825]" italics="true" pageId="29" pageNumber="130">Cnemaspis</emphasis>
</taxonomicName>
have been identified (
<bibRefCitation id="EFECA0AE2E74FFD9FED1FCD3FE3CFCA2" author="Rosler, H." box="[274,416,835,861]" pageId="29" pageNumber="130" pagination="1 - 94" refId="ref27356" refString="Rosler, H. (2016) Bemerkungen uber einige Gckoarten der Zoologischen Staatsammlung Munchen. 2 Teil: Asien, mit Anmerkungen zur Arten der Gattungen Cnemaspis Strauch, 1887 s. lat., Cyrtopodion Fitzinger, 1843 und Hemidactylus Oken, 1817. Gekkota, Supplement No. 3, 1 - 94." type="book chapter" year="2016">Rösler 2016</bibRefCitation>
; 
<bibRefCitation id="EFECA0AE2E74FFD9FE6FFCD3FD08FCA2" author="Batuwita, S. &amp; Agarwal, I. &amp; Bauer, A. M." box="[428,660,835,861]" pageId="29" pageNumber="130" pagination="223 - 234" refId="ref23590" refString="Batuwita, S., Agarwal, I. &amp; Bauer, A. M. (2019) Description of a new diminutive, rupicolous species of day-gecko (Squamata: Gekkonidae: Cnemaspis) from southern Sri Lanka. Zootaxa, 4565 (2), 223 - 234. https: // doi. org / 10.11646 / zootaxa. 4565.2.6" type="journal article" year="2019">
Batuwita 
<emphasis id="B909014D2E74FFD9FDDBFCD3FDCDFCA2" box="[536,593,835,861]" italics="true" pageId="29" pageNumber="130">et al.</emphasis>
2019
</bibRefCitation>
; 
<bibRefCitation id="EFECA0AE2E74FFD9FD63FCD3FCEBFCA2" author="Sayyed, A. &amp; Cyriac, V. P. &amp; Pardeshi, A. &amp; Sulakhe, S." box="[672,887,835,861]" pageId="29" pageNumber="130" pagination="25 - 38" refId="ref27542" refString="Sayyed, A., Cyriac, V. P., Pardeshi, A. &amp; Sulakhe, S. (2021) Dwarfs of the fortress: a new cryptic species of dwarf gecko of the genus Cnemaspis Strauch, 1887 (Squamata, Gekkonidae) from Rajgad Fort in the northern Western Ghats of Maharashtra, India. Evolutionary Systematics, 5, 25 - 38. https: // doi. org / 10.3897 / evolsyst. 5.62929" type="journal article" year="2021">
Sayyed 
<emphasis id="B909014D2E74FFD9FD39FCD3FCB0FCA2" box="[762,812,835,861]" italics="true" pageId="29" pageNumber="130">et al</emphasis>
. 2021
</bibRefCitation>
), 
<taxonomicName id="4C7DA6DC2E74FFD9FC48FCD3FC62FCA2" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[907,1022,835,861]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="spawlsi" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FC48FCD3FC62FCA2" box="[907,1022,835,861]" italics="true" pageId="29" pageNumber="130">A. spawlsi</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FBC5FCD3FBFCFCA2" bold="true" box="[1030,1120,835,861]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FBC5FCD3FBFCFCA2" box="[1030,1120,835,861]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C7DA6DC2E74FFD9FB5BFCD3FAA7FCA2" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[1176,1339,835,861]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="chyuluensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FB5BFCD3FAA7FCA2" box="[1176,1339,835,861]" italics="true" pageId="29" pageNumber="130">A. chyuluensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FA81FCD3FA00FCA2" bold="true" box="[1346,1436,835,861]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FA81FCD3FA00FCA2" box="[1346,1436,835,861]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
represent the first such tiny forms for 
<taxonomicName id="4C7DA6DC2E74FFD9FD89FCF7FD69FC7E" authorityName="Muller" authorityYear="1907" box="[586,757,871,897]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="genus">
<emphasis id="B909014D2E74FFD9FD89FCF7FD69FC7E" box="[586,757,871,897]" italics="true" pageId="29" pageNumber="130">Ancylodactylus</emphasis>
</taxonomicName>
. Among Kenyan congeners both of these species are easily separated on size from 
<taxonomicName id="4C7DA6DC2E74FFD9FE56FC1BFD81FC5A" box="[405,541,907,933]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="africanus">
<emphasis id="B909014D2E74FFD9FE56FC1BFD81FC5A" box="[405,541,907,933]" italics="true" pageId="29" pageNumber="130">A. africanus</emphasis>
</taxonomicName>
(maximum 
<quantity id="4C8570BA2E74FFD9FD60FC1AFD6DFC5A" box="[675,753,906,933]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="29" pageNumber="130" unit="mm" value="54.0">54 mm</quantity>
SVL), 
<taxonomicName id="4C7DA6DC2E74FFD9FC82FC1BFC48FC5A" baseAuthorityName="Loveridge" baseAuthorityYear="1936" box="[833,980,907,933]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="elgonensis">
<emphasis id="B909014D2E74FFD9FC82FC1BFC48FC5A" box="[833,980,907,933]" italics="true" pageId="29" pageNumber="130">A. elgonensis</emphasis>
</taxonomicName>
(maximum SVL 
<quantity id="4C8570BA2E74FFD9FB57FC1BFB7AFC5A" box="[1172,1254,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="29" pageNumber="130" unit="mm" value="61.0">61 mm</quantity>
), 
<taxonomicName id="4C7DA6DC2E74FFD9FB34FC1BFA00FC5A" baseAuthorityName="Schmidt" baseAuthorityYear="1919" box="[1271,1436,907,933]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="dickersonae">
<emphasis id="B909014D2E74FFD9FB34FC1BFA00FC5A" box="[1271,1436,907,933]" italics="true" pageId="29" pageNumber="130">A. dickersonae</emphasis>
</taxonomicName>
(maximum SVL 
<quantity id="4C8570BA2E74FFD9FE9BFC3FFE30FC36" box="[344,428,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="29" pageNumber="130" unit="mm" value="41.0">41 mm</quantity>
), 
<taxonomicName id="4C7DA6DC2E74FFD9FE02FC3FFDC7FC36" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[449,603,943,969]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="kenyaensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FE02FC3FFDC7FC36" box="[449,603,943,969]" italics="true" pageId="29" pageNumber="130">A. kenyaensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FDA7FC3FFD5CFC36" bold="true" box="[612,704,943,969]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FDA7FC3FFD5CFC36" box="[612,704,943,969]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(maximum SVL 
<quantity id="4C8570BA2E74FFD9FC4AFC3EFC40FC36" box="[905,988,942,969]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="29" pageNumber="130" unit="mm" value="65.0">65 mm</quantity>
, 
<taxonomicName id="4C7DA6DC2E74FFD9FC2AFC3FFBCFFC36" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[1001,1107,943,969]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="kituiensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FC2AFC3FFBCFFC36" box="[1001,1107,943,969]" italics="true" pageId="29" pageNumber="130">kituiensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FB9FFC3FFB2BFC36" bold="true" box="[1116,1207,943,969]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FB9FFC3FFB2BFC36" box="[1116,1207,943,969]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(maximum SVL 
<quantity id="4C8570BA2E74FFD9FA43FC3EFF5BFC12" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="130" unit="mm" value="50.0">50 mm</quantity>
), and 
<taxonomicName id="4C7DA6DC2E74FFD9FECAFC43FE5FFC12" authority="Malonza &amp; Bauer, 2022" authorityName="Malonza &amp; Bauer" authorityYear="2022" box="[265,451,979,1005]" class="Reptilia" family="Lacertidae" genus="Adolfus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="130" phylum="Chordata" rank="species" species="mathewsensis" status="sp. nov.">
<emphasis id="B909014D2E74FFD9FECAFC43FE5FFC12" box="[265,451,979,1005]" italics="true" pageId="29" pageNumber="130">A. mathewsensis</emphasis>
</taxonomicName>
<emphasis id="B909014D2E74FFD9FE09FC43FDBFFC12" bold="true" box="[458,547,979,1005]" pageId="29" pageNumber="130">
<taxonomicNameLabel id="A23ABC362E74FFD9FE09FC43FDBFFC12" box="[458,547,979,1005]" pageId="29" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(maximum SVL 
<quantity id="4C8570BA2E74FFD9FD24FC43FCA6FC12" box="[743,826,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="130" unit="mm" value="40.0">40 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C3678ED42E74FFDAFF04FC66FB2DFB16" lastPageId="30" lastPageNumber="131" pageId="29" pageNumber="130" type="description">
<paragraph id="8BC2DD5F2E74FFD9FF04FC66FBBBFB16" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="130">
<emphasis id="B909014D2E74FFD9FF04FC66FE78FBEE" bold="true" box="[199,484,1014,1041]" pageId="29" pageNumber="130">
Description of 
<typeStatus id="54C663FD2E74FFD9FEB5FC67FE43FBEE" box="[374,479,1015,1041]" pageId="29" pageNumber="130" type="holotype">holotype</typeStatus>
.
</emphasis>
Specimen generally in good condition. Body dorsoventrally flattened, tail entire (
<figureCitation id="1346C1DA2E74FFD9FAB6FC67FF2AFBCA" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="29" pageNumber="130">Fig 13</figureCitation>
). Measurements: SVL = 27.2; TAL = 27.1; TRL = 12.5; HL = 8.7; HW = 5.7; OD = 1.3; SE = 3.3. Head elongate (HL/SVL = 0.32), moderately wide (HW/HL = 0.65), depressed, and distinct from the neck, loreal region flattened, canthus rostralis slightly pointed. Scales on snout and loreal region domed, larger than scales of interorbital region and crown. Eyes very small (OD/HL = 0.15), ear opening slit-like; two large internasals, 4 infralabials and 5 supralabials. Mental scale subtriangular; 3 postmentals, the outer pair in contact with the first infralabials and mental, central postmental hexagonal and smaller than its neighbors; 5 post-post mentals.
</paragraph>
<paragraph id="8BC2DD5F2E74FFD9FF04FB63FD9EFACE" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="130">Mental scale pentagonal and divided, 3 postmentals the outer larger and in contact with the infralabials and mental, and 5 post-post mentals.</paragraph>
<paragraph id="8BC2DD5F2E74FFD9FF04FAABFAD0F98A" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="130">
Dorsal pholidosis mostly homogenous, body covered by minute granular scales (
<figureCitation id="1346C1DA2E74FFD9FB89FAABFB70FAAA" box="[1098,1260,1339,1365]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="29" pageNumber="130">Fig. 13A, 13D</figureCitation>
) 13. Very small (~ twice size of granular scales), somewhat elongate tubercles present in a sparse row from the neck, across the flanks to the tail base (
<figureCitation id="1346C1DA2E74FFD9FE66FA13FD8DFA62" box="[421,529,1411,1437]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="29" pageNumber="130">Fig. 13A</figureCitation>
). Ventral scales much larger than dorsals, smooth, imbricate, slightly larger in precloacal and femoral regions than on chest and belly, smallest in gular region; scales on lateral aspect of neck granular. Ventrolateral folds feebly developed. Fore-and hind limbs relatively long, slender, covered by granular to slightly enlarged sub-imbricating scales, the latter chiefly on the preaxial surfaces. All digits moderately long and slender, strongly clawed; penultimate phalanx of all digits curved, arising angularly from distal portion of wider basal toe pad; three wide basal lamellae, the distalmost much larger than the more proximal, and 9 narrower distal lamellae under digit IV of pes (
<figureCitation id="1346C1DA2E74FFD9FE36F9CBFDC6F98A" box="[501,602,1627,1653]" captionStart="FIGURE 11" captionStartId="26.[151,250,1857,1881]" captionTargetBox="[151,1435,181,1833]" captionTargetId="figure-28@26.[151,1435,181,1833]" captionTargetPageId="26" captionText="FIGURE 11. Ancylodactylus spawlsi sp. nov. A) Dorsal view of holotype, NMK-L3470. Scale bar = 10 mm. B) Precloacal region of holotype showing series of 6 precloacal pores. C) Ventral view of right hind foot of paratype, NMK-L4007/1. D) Dorsum of tail of holotype illustrating rows of tubercles near the posterior margins of each tail segment." figureDoi="http://doi.org/10.5281/zenodo.6581608" httpUri="https://zenodo.org/record/6581608/files/figure.png" pageId="29" pageNumber="130">Fig. 11C</figureCitation>
). Enlarged basal lamellae on digits of pes: I:1, II:2, III:4, IV:3, V:1.
</paragraph>
<paragraph id="8BC2DD5F2E74FFD9FF04F9EFFB12F8D6" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="130">
Male precloacal pores in a single continuous row of 8 (
<figureCitation id="1346C1DA2E74FFD9FCF6F9EFFC07F966" box="[821,923,1663,1689]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="29" pageNumber="130">Fig. 13E</figureCitation>
). Hemipenial bulge pronounced, a single pair of small postcloacal spurs (
<figureCitation id="1346C1DA2E74FFD9FE06F933FDB0F942" box="[453,556,1699,1725]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="29" pageNumber="130">Fig. 13A</figureCitation>
). Tail slightly depressed in cross section; segmentation not pronounced. Length of tail 99% of SVL. Scales of tail dorsum granular, squarish to rectangular or rounded, larger than trunk granules; lacking tubercles in post-pygal portion of tail (
<figureCitation id="1346C1DA2E74FFD9FD63F97BFCD7F8FA" box="[672,843,1771,1797]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="29" pageNumber="130">Figs. 13C, 13F</figureCitation>
). Subcaudal scales larger than dorsals; enlarged midventral subcaudal scales in a single row, but not greatly broadened transversely (
<figureCitation id="1346C1DA2E74FFD9FBD8F89FFB1EF8D6" box="[1051,1154,1807,1833]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="29" pageNumber="130">Fig. 13B</figureCitation>
).
</paragraph>
<paragraph id="8BC2DD5F2E74FFDAFF12F8A2FD80FF2A" blockId="29.[151,1437,151,2013]" lastBlockId="30.[151,1437,151,1905]" lastPageId="30" lastPageNumber="131" pageId="29" pageNumber="130">
<emphasis id="B909014D2E74FFD9FF12F8A2FDB5F8B3" bold="true" box="[209,553,1842,1868]" pageId="29" pageNumber="130">Coloration (in preservative).</emphasis>
Dorsum dark grayish-brown with a series of light gray blotches or chevrons along the back, separated from one another by narrow dark transverse bands (
<figureCitation id="1346C1DA2E74FFD9FC3CF8C7FBF9F88E" box="[1023,1125,1879,1905]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="29" pageNumber="130">Fig. 13C</figureCitation>
). Dark brown spots parallel to chevrons and also forming a lower row along the flanks, though weakly differentiated. Crown of the head dark grayish-brown, mostly uniform except for a light grayish marking on the occiput and scattered darker patches, e.g. on the snout. Labial scales with alternating light and dark markings. Limbs grayish-brown with darker mottling or spotting; dark bands on all digits. Tail dorsum slightly lighter than trunk with thin transverse dark brown bands. Venter whitish with dark stippling.
</paragraph>
<paragraph id="8BC2DD5F2E77FFDAFF04FF4EFB2DFB16" blockId="30.[151,1437,151,1905]" pageId="30" pageNumber="131">
<emphasis id="B909014D2E77FFDAFF04FF4EFE2DFF07" bold="true" box="[199,433,222,248]" pageId="30" pageNumber="131">Coloration (in life).</emphasis>
Based on photographs of freshly euthanized 
<typeStatus id="54C663FD2E77FFDAFC7BFF4FFB86FF06" box="[952,1050,223,249]" pageId="30" pageNumber="131" type="holotype">holotype</typeStatus>
(see 
<figureCitation id="1346C1DA2E77FFDAFB9AFF4FFA90FF06" box="[1113,1292,223,249]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="30" pageNumber="131">Figs. 13A, 13B</figureCitation>
). During the day the dorsal color is mainly shades of light to dark gray-brown to olive, although like its congeners, body color was changeable based on substrate, environmental conditions and physiological state. Dorsum bearing a series of eight partly overlapping blue-greyish fleurs-de-lis from the nape to the sacrum. An additional smaller peach-colored marking (cream to orangish) present on the occiput. Anterior margins of fleurs-de-lis with dark brown anterior borders, expanding laterally to form pairs of blotches lateral to the anterior apices of the fleurs-de-lis (
<figureCitation id="1346C1DA2E77FFDAFB24FE03FAC6FE52" box="[1255,1370,403,429]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="30" pageNumber="131">Figs. 13A</figureCitation>
). Lateral surfaces with a series of approximately 12 small, roundish, blue-grayish spots extending from the neck, across the shoulder and down the flanks to the sacrum; largest spots between the limb insertions. The sparse flank tubercles are bright white and are each enclosed within a pale spot, although not all spots contain tubercles. A parallel series of smaller, less conspicuous pale spots runs between the fleurs-de-lis and larger spots. Much of the flank area not occupied by pale markings supports irregular, almost blackish spots of different sizes. The head grayish with a dark brown line passing from the snout and through the eye and another extending posteroventrally from the corner of the mouth. Broken transverse blackish bands pass over the crown and between the orbits and there are whitish granules speckled over the lower margins of the head. Posterior to the angle of the jaws there is a single pale peach-colored marking (
<figureCitation id="1346C1DA2E77FFDAFEC5FD47FEEDFD0E" box="[262,369,727,753]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="30" pageNumber="131">Fig. 13A</figureCitation>
). Limbs predominantly gray with blackish markings forming irregular banding on the limbs. Thighs with whitish banding alternating with gray and black. Alternating bands of blackish, gray, and ashy white on the digits. The small postcloacal spur is bright white. Tail dorsum mostly pale grayish-olive and bearing a series of 12 dark brown chevrons or cross bands from tail base to tail tip. Proximal dark bands are thin and are followed by a more diffuse ashy border. These markings become less regular near the middle of the tail and both the dark (now blackish) and ashy marks become thicker, excluding the grayish-olive base color from the tail tip, which terminates in a dark brown tip. Individual dark brown or ashy scales are scattered on the spaces between the bands and larger spots alternate along the ventrolateral margin of the tail (
<figureCitation id="1346C1DA2E77FFDAFCDFFC43FC1AFC12" box="[796,902,979,1005]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="30" pageNumber="131">Fig. 13A</figureCitation>
). Ventral color of trunk dark overall (
<figureCitation id="1346C1DA2E77FFDAFAECFC43FA08FC12" box="[1327,1428,979,1005]" captionStart="FIGURE 13" captionStartId="31.[151,250,1900,1924]" captionTargetBox="[217,1370,181,1876]" captionTargetId="figure-17@31.[217,1370,181,1876]" captionTargetPageId="31" captionText="FIGURE 13. Holotype of Ancylodactylus chyuluensis sp. nov., NMK-L3869.A) Dorsal view. B) Ventral view showing the bluish throat and extensive dark pigmentation on most of the ventral scales. C) Dorsal view of preserved specimen. Scale bar = 10 mm. D) Trunk dorsum at midbody showing mostly granular surface. E) Precloacal region showing single row of 8 pores (arrows mark the lateralmost pores). F) Tail dorsum showing atubercular surface. Photos A–B © P.K. Malonza." figureDoi="http://doi.org/10.5281/zenodo.6581612" httpUri="https://zenodo.org/record/6581612/files/figure.png" pageId="30" pageNumber="131">Fig. 13B</figureCitation>
) with most individual ventral scales including scattered blackish pigment granules. Distal margins of most scales from neck to groin appear almost pale yellowish as do some of the lateralmost of the ventral scales. Undersides of limbs similarly dark, darker beneath thighs where dark pigment is present at the periphery of all scales. Palms and soles gray with scattered darker punctations. Throat and chin dull whitish to bluish (centrally) and heavily marked by purplish spotting anteriorly and complex reticulations posteriorly that are confluent with the dark trunk markings. Subcaudal surface of the tail pale greenish-white with most scales having their distal and lateral margins darkly pigmented. Pigmentation denser towards tail tip, where some scales are almost entirely black.
</paragraph>
</subSubSection>
<subSubSection id="C3678ED42E77FFDAFF04FB63FB71F88E" pageId="30" pageNumber="131" type="etymology">
<paragraph id="8BC2DD5F2E77FFDAFF04FB63FAE5FAF2" blockId="30.[151,1437,151,1905]" box="[199,1401,1267,1293]" pageId="30" pageNumber="131">
<emphasis id="B909014D2E77FFDAFF04FB63FED2FAF2" bold="true" box="[199,334,1267,1293]" pageId="30" pageNumber="131">Etymology.</emphasis>
Named for the Chyulu Hills, the 
<typeStatus id="54C663FD2E77FFDAFD06FB63FD69FAF2" box="[709,757,1267,1293]" pageId="30" pageNumber="131">type</typeStatus>
locality where the single known specimen was collected.
</paragraph>
<paragraph id="8BC2DD5F2E77FFDAFF04FA87FE18F9AE" blockId="30.[151,1437,151,1905]" pageId="30" pageNumber="131">
<emphasis id="B909014D2E77FFDAFF04FA87FE16FACF" bold="true" box="[199,394,1302,1329]" pageId="30" pageNumber="131">Natural History.</emphasis>
This specimen was found at approximately 8h00 in a pitfall trap in an X-shaped drift fence array in a dry forest patch growing within volcanic rocks with fallen dry tree branches/logs in the vicinity. We suspect that the species is arboreal but can descend to utilize fallen logs and trunk bases as well as the crevices in volcanic rocks. It is likely to be a diurnal species and we assume it had entered the trap on the morning of capture or during late afternoon of the previous day, as it was found resting during the cold morning. However, we have no evidence to rule out the possibility of it being nocturnal. Two-day searches had been done in the area before the trapping with the only commonly recorded gecko species at the site being 
<taxonomicName id="4C7DA6DC2E77FFDAFC8DFA7FFE3CF9D2" authority="Malonza, Bauer, Granthon, Williams &amp; Wojnowski" authorityName="Malonza, Bauer, Granthon, Williams &amp; Wojnowski" authorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="131" phylum="Chordata" rank="species" species="tsavoensis">
<emphasis id="B909014D2E77FFDAFC8DFA7FFBC3F9F7" box="[846,1119,1519,1545]" italics="true" pageId="30" pageNumber="131">Lygodactylus tsavoensis</emphasis>
Malonza, Bauer, Granthon, Williams &amp; Wojnowski
</taxonomicName>
on trees and fallen logs/branches. The sampling was carried out during the peak of the dry season in September.
</paragraph>
<paragraph id="8BC2DD5F2E77FFDAFF04F9CAFB71F88E" blockId="30.[151,1437,151,1905]" pageId="30" pageNumber="131">
<emphasis id="B909014D2E77FFDAFF04F9CAFE60F98B" bold="true" box="[199,508,1626,1653]" pageId="30" pageNumber="131">Habitat and Distribution.</emphasis>
This is a mid- to high altitude montane species. In the Chyulu Hills there is dry forest on volcanic rocks from elevations of around 
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that gives way to largely open grasslands from about 
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. The grassland continues up to the hilltops’ maximum elevation of around 
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with hilltop patches of moist forests surrounded by grasslands from elevations of about 
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and above. We suspect the species to be present in these forest patches on tree trunk crevices and cracks, like its congeners. The Chyulu Hills are prone to perennial wildfires. The area where the specimen was collected had been burned in previous years and fires occurred during the sampling period. This is a major conservation threat for this species because the dry trees increase the fire intensity, permitting the fires to burn for long periods, even underground within the volcanic rocks.
</paragraph>
</subSubSection>
</treatment>
</document>