133 lines
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133 lines
17 KiB
XML
<document id="37B354C95886E2E9B1DC60BC4C75BC60" ID-CLB-Dataset="90145" ID-DOI="10.5252/geodiversitas2022v44a22" ID-GBIF-Dataset="ff4dff07-723c-467d-84e1-c0c009173d67" ID-ISSN="1638-9395" ID-Zenodo-Dep="6795405" ID-ZooBank="urn:lsid:zoobank.org:pub:B8864E92-2EC0-4E0A-97F1-BFAE94DC24D8" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656955284745" checkinUser="felipe" docAuthor="Muizon, Christian de & Ladevèze, Sandrine" docDate="2022" docId="03D2AB06FFCD5134FBAE2592DA05E288" docLanguage="en" docName="Geodiversitas.44.22.609-643.pdf.imf" docOrigin="Geodiversitas 44 (22)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pucadelphyoidea Muizon & Ladevèze 2022, n. superfam." docType="treatment" docVersion="9" lastPageNumber="612" masterDocId="FFEBD37EFFC85131F82D2627DF27E426" masterDocTitle="New material of Incadelphys antiquus (Pucadelphyda, Metatheria, Mammalia) from the early Palaeocene of Bolivia reveals phylogenetic affinities with enigmatic North and South American metatherians" masterLastPageNumber="643" masterPageNumber="609" pageNumber="612" updateTime="1699356357151" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="21C236157FAD926A5601E03A6D90AD0D">
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<mods:title id="539F1364245F6900623E503254A89214">New material of Incadelphys antiquus (Pucadelphyda, Metatheria, Mammalia) from the early Palaeocene of Bolivia reveals phylogenetic affinities with enigmatic North and South American metatherians</mods:title>
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<mods:namePart id="C7DA90E17D28D8712ECF9AF8F7D6F865">Muizon, Christian de</mods:namePart>
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<mods:name id="78FB94EF03BAD1112771C3E92EABCBEF" type="personal">
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<mods:namePart id="CE77BC58F1BB0AFF7718932B78BB13B6">Ladevèze, Sandrine</mods:namePart>
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<mods:typeOfResource id="106626A7323C1E6532FB00FF3FF683FA">text</mods:typeOfResource>
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<mods:title id="2F9C4D568E2FB6FC555382E58504CB95">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="4A70227167232CA445FEB843E6FFBA7D">
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<mods:date id="38EC85FE6C01371047C1BF1464BCDF13">2022</mods:date>
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<mods:number id="9232AA03D441CEBB01C0D794C8AB3667">2022-06-30</mods:number>
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<mods:number id="EACE27DAB8F4070DCF0C0816BB4CBB89">44</mods:number>
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<mods:identifier id="6C6DFC1F5B161E7B98ADDB0222871DAB" type="DOI">10.5252/geodiversitas2022v44a22</mods:identifier>
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<treatment id="03D2AB06FFCD5134FBAE2592DA05E288" ID-DOI="http://doi.org/10.5281/zenodo.6802428" ID-GBIF-Taxon="196398373" ID-Zenodo-Dep="6802428" LSID="urn:lsid:plazi:treatment:03D2AB06FFCD5134FBAE2592DA05E288" httpUri="http://treatment.plazi.org/id/03D2AB06FFCD5134FBAE2592DA05E288" lastPageNumber="612" pageId="5" pageNumber="612">
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<subSubSection id="C361499BFFCD5134FBAE2592DA7EE7E9" box="[899,1369,949,976]" pageId="5" pageNumber="612" type="nomenclature">
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<paragraph id="8BC41A10FFCD5134FBAE2592DA7EE7E9" blockId="5.[899,1369,949,976]" box="[899,1369,949,976]" pageId="5" pageNumber="612">
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<heading id="D08CAD7CFFCD5134FBAE2592DA7EE7E9" box="[899,1369,949,976]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="612" reason="7">
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Superfamily
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<taxonomicName id="4C7B6193FFCD5134FC252591DBF7E7F6" authority="Muizon & Ladevèze, 2022" authorityName="Muizon & Ladevèze" authorityYear="2022" box="[1032,1232,950,976]" kingdom="Animalia" pageId="5" pageNumber="612" rank="superFamily" status="superfam. nov." superFamily="Pucadelphyoidea">PUCADELPHYOIDEA</taxonomicName>
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<taxonomicNameLabel id="A23C7B79FFCD5134FCF82591DA7EE7E9" box="[1237,1369,949,976]" pageId="5" pageNumber="612" rank="superFamily">n. superfam.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C361499BFFCD5134FB0025D1DC4BE036" box="[813,876,1014,1040]" pageId="5" pageNumber="612" type="multiple">
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<paragraph id="8BC41A10FFCD5134FB0025D1DC4BE036" blockId="5.[812,1457,1014,1710]" box="[813,876,1014,1040]" pageId="5" pageNumber="612">NOTE</paragraph>
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</subSubSection>
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<subSubSection id="C361499BFFCD5134FB002232DAB9E0E9" pageId="5" pageNumber="612" type="synonymic_list">
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<paragraph id="8BC41A10FFCD5134FB002232DAB9E0E9" blockId="5.[812,1457,1014,1710]" pageId="5" pageNumber="612">
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The superfamily
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<taxonomicName id="4C7B6193FFCD5134FBCD2232DBB1E009" authority="Muizon & Ladevèze, 2022" authorityName="Muizon & Ladevèze" authorityYear="2022" box="[992,1174,1045,1071]" kingdom="Animalia" pageId="5" pageNumber="612" rank="superFamily" status="superfam. nov." superFamily="Pucadelphyoidea">Pucadelphyoidea</taxonomicName>
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<taxonomicNameLabel id="A23C7B79FFCD5134FCB62231DA04E009" box="[1179,1315,1045,1072]" pageId="5" pageNumber="612" rank="superFamily">n. superfam.</taxonomicNameLabel>
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includes two clades, the family
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<taxonomicName id="4C7B6193FFCD5134FBD52212DB82E069" authorityName=", Muizon" authorityYear="1998" box="[1016,1189,1077,1103]" class="Mammalia" family="Pucadelphyidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="612" phylum="Chordata" rank="family">Pucadelphyidae</taxonomicName>
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and an unnamed clade defined below. The
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<taxonomicName id="4C7B6193FFCD5134FC212272DB9EE049" authorityName=", Muizon" authorityYear="1998" box="[1036,1209,1109,1135]" class="Mammalia" family="Pucadelphyidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="612" phylum="Chordata" rank="family">Pucadelphyidae</taxonomicName>
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include the following genera:
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<taxonomicName id="4C7B6193FFCD5134FBAE2252DA09E0A9" authority="Marshall & Muizon, 1988" authorityName="Marshall & Muizon" authorityYear="1988" box="[899,1326,1141,1167]" class="Mammalia" family="Didelphidae" genus="Pucadelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FBAE2252DCD8E0A9" box="[899,1023,1141,1167]" italics="true" pageId="5" pageNumber="612">Pucadelphys</emphasis>
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<bibRefCitation id="EFEA67E1FFCD5134FC252252DA09E0A9" author="MARSHALL L. G. & MUIZON C. DE" box="[1032,1326,1141,1167]" pageId="5" pageNumber="612" pagination="23 - 55" refId="ref22591" refString="MARSHALL L. G. & MUIZON C. DE 1988. - The dawn of the age of mammals in South America. National Geographic Research 4 (1): 23 - 55." type="journal article" year="1988">Marshall & Muizon, 1988</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName id="4C7B6193FFCD5134FD102252DBA3E089" authority="Marshall & Muizon, 1988" authorityName="Marshall & Muizon" authorityYear="1988" class="Mammalia" family="Didelphidae" genus="Andinodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FD102252DC7FE089" italics="true" pageId="5" pageNumber="612">Andinodelphys</emphasis>
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<bibRefCitation id="EFEA67E1FFCD5134FB4D22B2DBA3E089" author="MARSHALL L. G. & MUIZON C. DE" box="[864,1156,1173,1199]" pageId="5" pageNumber="612" pagination="23 - 55" refId="ref22591" refString="MARSHALL L. G. & MUIZON C. DE 1988. - The dawn of the age of mammals in South America. National Geographic Research 4 (1): 23 - 55." type="journal article" year="1988">Marshall & Muizon, 1988</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName id="4C7B6193FFCD5134FCBF22B2DCECE0E9" authority="Marshall & Muizon, 1988" authorityName="Marshall & Muizon" authorityYear="1988" class="Mammalia" family="Didelphidae" genus="Mizquedelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FCBF22B2DA0CE089" box="[1170,1323,1173,1199]" italics="true" pageId="5" pageNumber="612">Mizquedelphys</emphasis>
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<bibRefCitation id="EFEA67E1FFCD5134FD1F22B2DCECE0E9" author="MARSHALL L. G. & MUIZON C. DE" pageId="5" pageNumber="612" pagination="23 - 55" refId="ref22591" refString="MARSHALL L. G. & MUIZON C. DE 1988. - The dawn of the age of mammals in South America. National Geographic Research 4 (1): 23 - 55." type="journal article" year="1988">Marshall & Muizon, 1988</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C7B6193FFCD5134FBF82292DABEE0E9" authority="Marshall & Muizon, 1984" authorityName="Marshall & Muizon" authorityYear="1984" box="[981,1433,1205,1231]" class="Mammalia" family="Didelphidae" genus="Itaboraidelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FBF82292DB56E0E9" box="[981,1137,1205,1231]" italics="true" pageId="5" pageNumber="612">Itaboraidelphys</emphasis>
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<bibRefCitation id="EFEA67E1FFCD5134FC552292DABEE0E9" author="MARSHALL L. G. & MUIZON C. DE" box="[1144,1433,1205,1231]" pageId="5" pageNumber="612" pagination="1297 - 1300" refId="ref22522" refString="MARSHALL L. G. & MUIZON C. DE 1984. - Un nouveau Marsupial didelphide (Itaboraidelphys camposi nov. gen. nov. sp.) du Paleocene moyen (Itaboraien) de Sao Jose de Itaborai (Bresil). Comptes Rendus hebdomadaires des Seances de l'Academie des Sciences, Paris, ser. II, 299 (18): 1297 - 1300." type="journal article" year="1984">Marshall & Muizon, 1984</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C361499BFFCD5134FB6E22F2DA05E288" pageId="5" pageNumber="612" type="discussion">
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<paragraph id="8BC41A10FFCD5134FB6E22F2DA05E288" blockId="5.[812,1457,1014,1710]" pageId="5" pageNumber="612">
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The unnamed clade includes, in addition to
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<taxonomicName id="4C7B6193FFCD5134FD1922F2DB78E129" authority="Marshall & Muizon, 1988" authorityName="Marshall & Muizon" authorityYear="1988" class="Mammalia" family="Didelphidae" genus="Incadelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FD1922F2DA88E0C9" box="[1332,1455,1237,1263]" italics="true" pageId="5" pageNumber="612">Incadelphys</emphasis>
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<bibRefCitation id="EFEA67E1FFCD5134FB0022D3DB78E129" author="MARSHALL L. G. & MUIZON C. DE" box="[813,1119,1268,1295]" pageId="5" pageNumber="612" pagination="23 - 55" refId="ref22591" refString="MARSHALL L. G. & MUIZON C. DE 1988. - The dawn of the age of mammals in South America. National Geographic Research 4 (1): 23 - 55." type="journal article" year="1988">Marshall & Muizon, 1988</bibRefCitation>
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</taxonomicName>
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the following genera:
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<taxonomicName id="4C7B6193FFCD5134FD4522D2DBEBE109" authority="Cifelli & Johanson, 1994" authorityName="Cifelli & Johanson" authorityYear="1994" class="Mammalia" family="Asiatheriidae" genus="Aenigmadelphys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FD4522D2DCBBE108" italics="true" pageId="5" pageNumber="612">Aenigmadelphys</emphasis>
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<bibRefCitation id="EFEA67E1FFCD5134FB8B2333DBEBE109" author="CIFELLI R. L. & JOHANSON Z." box="[934,1228,1300,1327]" pageId="5" pageNumber="612" pagination="292 - 295" refId="ref20372" refString="CIFELLI R. L. & JOHANSON Z. 1994. - New mammal from the upper Cretaceous of Utah. Journal of Vertebrate Paleontology 14 (2): 292 - 295. https: // doi. org / 10.1073 / pnas. 90.20.9413" type="journal article" year="1994">Cifelli & Johanson, 1994</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C7B6193FFCD5134FCF32332DCE6E168" authority="Paula Couto, 1962" authorityName=", Paula Couto" authorityYear="1962" class="Mammalia" family="Didelphidae" genus="Marmosopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FCF32332DA4EE108" box="[1246,1385,1301,1326]" italics="true" pageId="5" pageNumber="612">Marmosopsis</emphasis>
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Paula Couto, 1962
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</taxonomicName>
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, and
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<taxonomicName id="4C7B6193FFCD5134FC2A2313DAA1E168" authority="Muizon & Cifelli, 2001" authorityName="Muizon & Cifelli" authorityYear="2001" box="[1031,1414,1332,1358]" class="Mammalia" genus="Szalinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FC2A2313DB45E168" box="[1031,1122,1332,1358]" italics="true" pageId="5" pageNumber="612">Szalinia</emphasis>
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<bibRefCitation id="EFEA67E1FFCD5134FC412313DAA1E168" author="MUIZON C. DE & CIFELLI R. L." box="[1132,1414,1332,1358]" pageId="5" pageNumber="612" pagination="87 - 97" refId="ref22894" refString="MUIZON C. DE & CIFELLI R. L. 2001. - A new basal " didelphoid " (Marsupialia, Mammalia) from the early Paleocene of Tiupampa (Bolivia). Journal of Vertebrate paleontology 21 (1): 87 - 97. https: // doi. org / brt 8 zq" type="journal article" year="2001">Muizon & Cifelli, 2001</bibRefCitation>
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</taxonomicName>
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. As discussed in the phylogeny section below, we favor the topology resulting from the analysis with implied weighting of characters, which retrieved the four genera in a clade. However, we are reluctant to formally name this clade because we consider that
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<taxonomicName id="4C7B6193FFCD5134FCB323F3DA13E1C8" authorityName="Marshall & de Muizon" authorityYear="1988" box="[1182,1332,1492,1518]" class="Mammalia" family="Macropididae" genus="Jaskhadelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FCB323F3DA13E1C8" box="[1182,1332,1492,1518]" italics="true" pageId="5" pageNumber="612">Jaskhadelphys</emphasis>
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</taxonomicName>
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should be included in the taxon list (which we did not) but with substantial new and more complete specimens than the single maxillary fragment with M2-M3 known so far. If
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<taxonomicName id="4C7B6193FFCD5134FB022074DCE6E24B" authorityName="Marshall & de Muizon" authorityYear="1988" box="[815,961,1619,1645]" class="Mammalia" family="Macropididae" genus="Jaskhadelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="5" pageNumber="612" phylum="Chordata" rank="genus">
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<emphasis id="B90FC602FFCD5134FB022074DCE6E24B" box="[815,961,1619,1645]" italics="true" pageId="5" pageNumber="612">Jaskhadelphys</emphasis>
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</taxonomicName>
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was to be included in this clade (what is so far highly uncertain), then the resulting clade should be the family
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<taxonomicName id="4C7B6193FFCD5134FB8520B4DA3AE288" authority="(Muizon 1992)" baseAuthorityName="Muizon" baseAuthorityYear="1992" box="[936,1309,1683,1710]" class="Mammalia" family="Jaskhadelphyidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="612" phylum="Chordata" rank="family">
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Jaskhadelphyidae (
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<bibRefCitation id="EFEA67E1FFCD5134FC5420B4DA35E28B" author="MUIZON C. DE" box="[1145,1298,1683,1709]" pageId="5" pageNumber="612" pagination="575 - 624" refId="ref22721" refString="MUIZON C. DE 1992. - La fauna de mamiferos de Tiupampa (Paleoceno inferior, Formacion Santa Lucia), Bolivia, in SUAREZ RIGLOS M. (ed.), Fosiles y facies de Bolivia. Vol. I. Vertebrados. Revista Tecnica de Yacimientos Petroliferos y Fiscales de Bolivia 12 (3 - 4): 575 - 624." type="journal article" year="1992">Muizon 1992</bibRefCitation>
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)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |