302 lines
31 KiB
XML
302 lines
31 KiB
XML
<document id="295515B24017E6762FA8D280F32EB8B3" ID-DOI="10.1080/002229301750384301" ID-ISSN="1464-5262" ID-Zenodo-Dep="5277273" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630023468274" checkinUser="felipe" docAuthor="Gofas, Serge" docDate="2001" docId="03CA537AF927FFAD5E00956A06FFFAA1" docLanguage="en" docName="JNatHist.35.9.1277-1369.pdf" docOrigin="Journal of Natural History 35 (9)" docSource="http://www.tandfonline.com/doi/abs/10.1080/002229301750384301" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Cochlostoma bicostulatum Gofas 1989" docType="treatment" docVersion="4" lastPageNumber="1319" masterDocId="FFF32B02F90DFF815F5F91120439FFDB" masterDocTitle="The systematics of Pyrenean and Cantabrian Cochlostoma (Gastropoda, Cyclophoroidea) revisited" masterLastPageNumber="1369" masterPageNumber="1277" pageNumber="1317" updateTime="1699137454917" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5F48C6F47CB49020CD3008E12A1DD879">The systematics of Pyrenean and Cantabrian Cochlostoma (Gastropoda, Cyclophoroidea) revisited</mods:title>
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<mods:name id="7A4C4200E8CF703C93C3CA9E52D87DD5" type="personal">
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<mods:namePart id="4917B4BA4233A2F55C218FC6A6F1811F">Gofas, Serge</mods:namePart>
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<mods:typeOfResource id="27EF85289B58615D429246DC5B2BB644">text</mods:typeOfResource>
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<mods:title id="B95DDA1B8B876986498F3567815CE500">Journal of Natural History</mods:title>
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<mods:part id="58C53571B6FDB7F10AE5FC5713CCCCAE">
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<mods:date id="D4703610E1405C2FB3823188F365B333">2001</mods:date>
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<mods:number id="6AC68C91789CB60A6E330F7BAFA5241A">2010-12-03</mods:number>
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<mods:number id="4C7B9C6579B9C347C9D7D4901B68259C">35</mods:number>
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<mods:number id="EE841B7CF178454E15D1BCFC4280EEC7">9</mods:number>
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<mods:start id="E0A62FDBBDD505B6DF9C630FC2143BFD">1277</mods:start>
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<mods:end id="30B76B7DBC383B1329D2DF97F6A48744">1369</mods:end>
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<mods:identifier id="38CAEA95737C00BCA3A2A3EF12C17DA4" type="DOI">10.1080/002229301750384301</mods:identifier>
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<mods:identifier id="7C2C4AB07520562455C35E6688AF0EDF" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03CA537AF927FFAD5E00956A06FFFAA1" ID-DOI="http://doi.org/10.5281/zenodo.5279394" ID-Zenodo-Dep="5279394" LSID="urn:lsid:plazi:treatment:03CA537AF927FFAD5E00956A06FFFAA1" httpUri="http://treatment.plazi.org/id/03CA537AF927FFAD5E00956A06FFFAA1" lastPageId="44" lastPageNumber="1319" pageId="42" pageNumber="1317">
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<subSubSection id="C379B1E7F927FFAB5E00956A06A8FB6B" pageId="42" pageNumber="1317" type="nomenclature">
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<paragraph id="8BDCE26CF927FFAB5E00956A072FFB4B" blockId="42.[351,790,1144,1168]" box="[351,790,1144,1168]" pageId="42" pageNumber="1317">
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<heading id="D0945500F927FFAB5E00956A072FFB4B" box="[351,790,1144,1168]" centered="true" fontSize="10" level="2" pageId="42" pageNumber="1317" reason="2">
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<taxonomicName id="4C6399EFF927FFAB5E00956A072FFB4B" authority="Gofas, 1989" authorityName="Gofas" authorityYear="1989" box="[351,790,1144,1168]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="42" pageNumber="1317" phylum="Mollusca" rank="species" species="bicostulatum">
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<emphasis id="B9173E7EF927FFAB5E00956A06BCFB4B" bold="true" box="[351,645,1144,1168]" italics="true" pageId="42" pageNumber="1317">Cochlostoma bicostulatum</emphasis>
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<bibRefCitation id="EFF29F9DF927FFAB5DD2956A072FFB4B" author="GOFAS, S." box="[653,790,1144,1168]" pageId="42" pageNumber="1317" pagination="43" refId="ref24637" refString="GOFAS, S., 1989, DescripcioAn preliminar de dos nuevos Cochlostoma (Gastropoda, Prosobranchia) del CantaAbrico, PublicacEoAes ocasionais da Sociedade Portuguesa de Malacologia, 14, 43 ± 48." type="journal article" year="1989">Gofas, 1989</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BDCE26CF927FFAB5EBA958A06A8FB6B" blockId="42.[93,1049,1176,1391]" box="[485,657,1176,1200]" pageId="42" pageNumber="1317">(®gures 31±34)</paragraph>
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</subSubSection>
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||
<subSubSection id="C379B1E7F927FFAB5F0295A8062AFADE" pageId="42" pageNumber="1317" type="reference_group">
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<paragraph id="8BDCE26CF927FFAB5F0295A8003CFB31" blockId="42.[93,1049,1176,1391]" pageId="42" pageNumber="1317">
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<treatmentCitationGroup id="AB73C542F927FFAB5F0295A8003CFB31" pageId="42" pageNumber="1317">
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<treatmentCitation id="0AC2C47DF927FFAB5F0295A8061CFB0B" author="GOFAS, S." box="[93,549,1210,1232]" page="45" pageId="42" pageNumber="1317" year="1989">
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<taxonomicName id="4C6399EFF927FFAB5F0295A8061CFB0B" authority="Gofas, 1989: 45" authorityName="Gofas" authorityPageNumber="45" authorityYear="1989" box="[93,549,1210,1232]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="42" pageNumber="1317" phylum="Mollusca" rank="species" species="bicostulatum">
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<emphasis id="B9173E7EF927FFAB5F0295A80553FB0B" box="[93,362,1210,1232]" italics="true" pageId="42" pageNumber="1317">Cochlostoma bicostulatum</emphasis>
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<bibRefCitation id="EFF29F9DF927FFAB5E2795A8061CFB0B" author="GOFAS, S." box="[376,549,1210,1232]" pageId="42" pageNumber="1317" pagination="43" refId="ref24637" refString="GOFAS, S., 1989, DescripcioAn preliminar de dos nuevos Cochlostoma (Gastropoda, Prosobranchia) del CantaAbrico, PublicacEoAes ocasionais da Sociedade Portuguesa de Malacologia, 14, 43 ± 48." type="journal article" year="1989">Gofas, 1989: 45</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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±46, ®gure 6±9. Type locality: Seldesuto, near Matienzo (Santander)ÐHOLOTYPE MNHN.
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<materialsCitation id="3B0BE831F927FFAB5D0D95C707C6FB31" box="[594,1023,1236,1258]" collectionCode="MNHN, MNCN, MZB, UPV" isTailEnumeration="true" pageId="42" pageNumber="1317" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="54D85CCEF927FFAB5D0D95C70687FB31" box="[594,702,1236,1258]" pageId="42" pageNumber="1317" type="paratype">PARATYPES</typeStatus>
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<collectionCode id="ED727AA9F927FFAB5D9995C70720FB31" box="[710,793,1237,1258]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="42" pageNumber="1317">MNHN</collectionCode>
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,
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<collectionCode id="ED727AA9F927FFAB5C7895C6074EFB31" box="[807,887,1236,1258]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="42" pageNumber="1317" type="Museum">MNCN</collectionCode>
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,
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<collectionCode id="ED727AA9F927FFAB5CD995C70784FB31" box="[902,957,1237,1258]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="42" pageNumber="1317" type="Museum">MZB</collectionCode>
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,
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<collectionCode id="ED727AA9F927FFAB5C9395C707C6FB31" box="[972,1023,1237,1258]" pageId="42" pageNumber="1317">UPV</collectionCode>
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</materialsCitation>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BDCE26CF927FFAB5F0295FD062AFADE" blockId="42.[93,1049,1176,1391]" box="[93,531,1263,1285]" pageId="42" pageNumber="1317">Manuscript names in Museum collections:</paragraph>
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</subSubSection>
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||
<subSubSection id="C379B1E7F927FFAD5F02941806FFFAA1" lastPageId="44" lastPageNumber="1319" pageId="42" pageNumber="1317" type="description">
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<paragraph id="8BDCE26CF927FFAB5F0294180577FAE1" blockId="42.[93,1049,1176,1391]" pageId="42" pageNumber="1317">
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<taxonomicName id="4C6399EFF927FFAB5F02941805ECFAC4" authority="Aguiar-Amat" authorityName="Aguiar-Amat" box="[93,469,1289,1311]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="42" pageNumber="1317" phylum="Mollusca" rank="species" species="bicostulatum">
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<emphasis id="B9173E7EF927FFAB5F0294180578FAC4" box="[93,321,1289,1311]" italics="true" pageId="42" pageNumber="1317">Pomatias bicostulatum</emphasis>
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Aguiar-Amat
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</taxonomicName>
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`des®ladero de Beyos,
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<collectingRegion id="49A72C8EF927FFAB5D94941B0718FAC4" box="[715,801,1289,1311]" country="Spain" name="Asturias" pageId="42" pageNumber="1317">Asturias</collectingRegion>
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, D. Quatrecasas leg. et det.’ MZB 80.0047.
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</paragraph>
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<paragraph id="8BDCE26CF927FFAB5F02942C07B8FA8F" blockId="42.[93,1049,1176,1391]" box="[93,897,1342,1364]" pageId="42" pageNumber="1317">
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<taxonomicName id="4C6399EFF927FFAB5F02942C063FFA8F" authority="Ortiz de Zarate" authorityName="Ortiz de Zarate" box="[93,518,1342,1364]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="42" pageNumber="1317" phylum="Mollusca" rank="species" species="azpeitianum">
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<emphasis id="B9173E7EF927FFAB5F02942C0566FA8F" box="[93,351,1342,1364]" italics="true" pageId="42" pageNumber="1317">Cochlostoma azpeitianum</emphasis>
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Ortiz de Zarate
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</taxonomicName>
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in coll. Altimira (®de
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<bibRefCitation id="EFF29F9DF927FFAB5DAF942D074BFA8F" author="RAVEN, J. G. M." box="[752,882,1342,1364]" pageId="42" pageNumber="1317" pagination="17" refId="ref25752" refString="RAVEN, J. G. M., 1990, A revision of Obscurella Clessin, 1889 (Gastropoda Prosobranchia: Cyclophoridae, Basteria, 54, 17 ± 62." type="journal article" year="1990">Raven, 1990</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BDCE26CF927FFAB5F02944B0602FAB4" blockId="42.[93,1049,1176,1391]" box="[93,571,1369,1391]" pageId="42" pageNumber="1317">
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<taxonomicName id="4C6399EFF927FFAB5F02944B0595FAB4" authority="Raven" authorityName="Raven" box="[93,428,1369,1391]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="42" pageNumber="1317" phylum="Mollusca" rank="species" species="cantabricum">
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<emphasis id="B9173E7EF927FFAB5F02944B055BFAB4" box="[93,354,1369,1391]" italics="true" pageId="42" pageNumber="1317">Cochlostoma cantabricum</emphasis>
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Raven
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</taxonomicName>
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in coll. UPV.
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</paragraph>
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<paragraph id="8BDCE26CF927FFAB5F02949A04E2FA44" blockId="42.[93,1049,1416,1662]" box="[93,219,1416,1439]" pageId="42" pageNumber="1317">
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<emphasis id="B9173E7EF927FFAB5F02949A04E2FA44" box="[93,219,1416,1439]" italics="true" pageId="42" pageNumber="1317">Description</emphasis>
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</paragraph>
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<paragraph id="8BDCE26CF927FFAB5FD994B505BFFA25" blockId="42.[93,1049,1416,1662]" pageId="42" pageNumber="1317">
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Shell up to 12.5
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<emphasis id="B9173E7EF927FFAB5E6394B5056DFA64" bold="true" box="[316,340,1447,1471]" pageId="42" pageNumber="1317">Ö</emphasis>
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<quantity id="4C9B4F89F927FFAB5E0B94B505A8FA64" box="[340,401,1447,1471]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="1317" unit="mm" value="5.0">5 mm</quantity>
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with 7.5 to 8.5 quite rounded whorls. Early whorls with a peripheral angle which is slightly apparent next to it; body whorl de®nitely constricted, well rounded.
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</paragraph>
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<paragraph id="8BDCE26CF927FFAB5FD997140773F9A6" blockId="42.[93,1049,1416,1662]" pageId="42" pageNumber="1317">Sculpture of weak wrinkles on the ®rst embryonic half whorl, then grading to crowded (25±30/mm) ribs on the second whorl. Spire whorls with alternating very strong ribs (9±11 ribs/mm) with 1± 4 ®ner ones; ribs de®nitely thickened at their termination along the suture, so as to give it a crenulated aspect.</paragraph>
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<caption id="DF1CB2E4F926FFAA5F02917B0788FF6F" pageId="43" pageNumber="1318" startId="43.[93,150,105,127]">
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<paragraph id="8BDCE26CF926FFAA5F02917B0788FF6F" blockId="43.[93,1049,105,180]" pageId="43" pageNumber="1318">
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Table 4. Observed genotype frequencies for the LAP, ODH and IDH loci in
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<taxonomicName id="4C6399EFF926FFAA5C21917B07D3FFA4" box="[894,1002,105,127]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="1318" phylum="Mollusca" rank="species" species="hidalgoi">
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<emphasis id="B9173E7EF926FFAA5C21917B07D3FFA4" box="[894,1002,105,127]" italics="true" pageId="43" pageNumber="1318">C. hidalgoi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6399EFF926FFAA5FFA91910517FF42" box="[165,302,131,153]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="1318" phylum="Mollusca" rank="species" species="asturicum">
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<emphasis id="B9173E7EF926FFAA5FFA91910517FF42" box="[165,302,131,153]" italics="true" pageId="43" pageNumber="1318">C. asturicum</emphasis>
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</taxonomicName>
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from Covadonga, compared to expected frequencies under Hardy- Weinberg equilibrium when considered separately (left) and pooled (right).
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</paragraph>
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</caption>
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<paragraph id="8BDCE26CF926FFAA5F9491C1054CFB50" pageId="43" pageNumber="1318">
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<table id="F96310CCF926007E5F0291C107C2FB50" box="[93,1019,211,1163]" gridcols="6" gridrows="11" pageId="43" pageNumber="1318">
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||
<tr id="3553E02EF926007E5F0291C107C2FF32" box="[93,1019,211,233]" gridrow="0" pageId="43" pageNumber="1318">
|
||
<th id="76828952F926007E5F0291C10637FF32" box="[93,526,211,233]" colspan="3" colspanRight="2" gridcol="0" gridrow="0" pageId="43" pageNumber="1318">
|
||
<taxonomicName id="4C6399EFF926FFAA5F9491C10584FF32" authority=", Covadonga" box="[203,445,211,233]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="1318" phylum="Mollusca" rank="species" species="hidalgoi">
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||
<emphasis id="B9173E7EF926FFAA5F9491C10500FF32" box="[203,313,211,233]" italics="true" pageId="43" pageNumber="1318">C. hidalgoi</emphasis>
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||
, Covadonga
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||
</taxonomicName>
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||
</th>
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||
<th id="76828952F926007E5D3C91C107C2FF32" box="[611,1019,211,233]" colspan="3" colspanRight="2" gridcol="3" gridrow="0" pageId="43" pageNumber="1318">
|
||
<taxonomicName id="4C6399EFF926FFAA5DDB91C106CBFF32" box="[644,754,211,233]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="1318" phylum="Mollusca" rank="species" species="hidalgoi">
|
||
<emphasis id="B9173E7EF926FFAA5DDB91C106CBFF32" box="[644,754,211,233]" italics="true" pageId="43" pageNumber="1318">C. hidalgoi</emphasis>
|
||
</taxonomicName>
|
||
1
|
||
<taxonomicName id="4C6399EFF926FFAA5C5491C107B5FF32" box="[779,908,211,233]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="1318" phylum="Mollusca" rank="species" species="asturicum">
|
||
<emphasis id="B9173E7EF926FFAA5C5491C107B5FF32" box="[779,908,211,233]" italics="true" pageId="43" pageNumber="1318">C. asturicum</emphasis>
|
||
</taxonomicName>
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||
, pooled
|
||
</th>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F02901A07C2FEE3" box="[93,1019,264,312]" gridrow="1" pageId="43" pageNumber="1318">
|
||
<th id="76828952F926007E5F02901A04F2FEE3" box="[93,203,264,312]" gridcol="0" gridrow="1" pageId="43" pageNumber="1318">LAP Category</th>
|
||
<td id="76828952F926007E5E4C901A054EFEE3" box="[275,375,264,312]" gridcol="1" gridrow="1" pageId="43" pageNumber="1318">observed frequency</td>
|
||
<td id="76828952F926007E5EF5901A0637FEE3" box="[426,526,264,312]" gridcol="2" gridrow="1" pageId="43" pageNumber="1318">expected frequency</td>
|
||
<td id="76828952F926007E5D3C901A06E8FEE3" box="[611,721,264,312]" gridcol="3" gridrow="1" pageId="43" pageNumber="1318">LAP Category</td>
|
||
<td id="76828952F926007E5C5F901A075DFEE3" box="[768,868,264,312]" gridcol="4" gridrow="1" pageId="43" pageNumber="1318">observed frequency</td>
|
||
<td id="76828952F926007E5CC8901A07C2FEE3" box="[919,1019,264,312]" gridcol="5" gridrow="1" pageId="43" pageNumber="1318">expected frequency</td>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F02904507C2FE64" box="[93,1019,343,447]" gridrow="2" pageId="43" pageNumber="1318">
|
||
<th id="76828952F926007E5F02904504F2FE64" box="[93,203,343,447]" gridcol="0" gridrow="2" pageId="43" pageNumber="1318">B-B B-C C-C x2 523.633</th>
|
||
<td id="76828952F926007E5E4C9045054EFE64" box="[275,375,343,447]" gridcol="1" gridrow="2" pageId="43" pageNumber="1318">27 0 4 df5 1</td>
|
||
<td id="76828952F926007E5EF590450637FE64" box="[426,526,343,447]" gridcol="2" gridrow="2" pageId="43" pageNumber="1318">
|
||
23.516 6.968 0.516
|
||
<emphasis id="B9173E7EF926FFAA5E9D90BA05EBFE66" box="[450,466,424,445]" italics="true" pageId="43" pageNumber="1318">P</emphasis>
|
||
50
|
||
</td>
|
||
<td id="76828952F926007E5D3C904506E8FE64" box="[611,721,343,447]" gridcol="3" gridrow="2" pageId="43" pageNumber="1318">B-B B-C C-C x2 538.979</td>
|
||
<td id="76828952F926007E5C5F9045075DFE64" box="[768,868,343,447]" gridcol="4" gridrow="2" pageId="43" pageNumber="1318">27 1 18 df5 1</td>
|
||
<td id="76828952F926007E5CC8904507C2FE64" box="[919,1019,343,447]" gridcol="5" gridrow="2" pageId="43" pageNumber="1318">
|
||
16.440 22.120 7.440
|
||
<emphasis id="B9173E7EF926FFAA5CF090BA0786FE66" box="[943,959,424,445]" italics="true" pageId="43" pageNumber="1318">P</emphasis>
|
||
50
|
||
</td>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F0290DD07C2FD94" box="[93,1019,463,591]" gridrow="3" pageId="43" pageNumber="1318">
|
||
<th id="76828952F926007E5F0290DD04F2FD94" box="[93,203,463,591]" gridcol="0" gridrow="3" pageId="43" pageNumber="1318">ODH Category D-D D-E D-G</th>
|
||
<td id="76828952F926007E5E4C90DD054EFD94" box="[275,375,463,591]" gridcol="1" gridrow="3" pageId="43" pageNumber="1318">observed frequency 27 3 1</td>
|
||
<td id="76828952F926007E5EF590DD0637FD94" box="[426,526,463,591]" gridcol="2" gridrow="3" pageId="43" pageNumber="1318">expected frequency 27.129 2.806 0.935</td>
|
||
<td id="76828952F926007E5D3C90DD06E8FD94" box="[611,721,463,591]" gridcol="3" gridrow="3" pageId="43" pageNumber="1318">ODH Category D-D D-E D-G</td>
|
||
<td id="76828952F926007E5C5F90DD075DFD94" box="[768,868,463,591]" gridcol="4" gridrow="3" pageId="43" pageNumber="1318">observed frequency 41 3 2</td>
|
||
<td id="76828952F926007E5CC890DD07C2FD94" box="[919,1019,463,591]" gridcol="5" gridrow="3" pageId="43" pageNumber="1318">expected frequency 41.136 2.837 1.891</td>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F02934607C2FD60" box="[93,1019,596,699]" gridrow="4" pageId="43" pageNumber="1318">
|
||
<th id="76828952F926007E5F02934604F2FD60" box="[93,203,596,699]" gridcol="0" gridrow="4" pageId="43" pageNumber="1318">E-E E-G G-G x2 50</th>
|
||
<td id="76828952F926007E5E4C9346054EFD60" box="[275,375,596,699]" gridcol="1" gridrow="4" pageId="43" pageNumber="1318">0 0 0 df5 3</td>
|
||
<td id="76828952F926007E5EF593460637FD60" box="[426,526,596,699]" gridcol="2" gridrow="4" pageId="43" pageNumber="1318">
|
||
0.073 0.048 0.008
|
||
<emphasis id="B9173E7EF926FFAA5E9D93B705EBFD61" box="[450,466,677,698]" italics="true" pageId="43" pageNumber="1318">P</emphasis>
|
||
51
|
||
</td>
|
||
<td id="76828952F926007E5D3C934606E8FD60" box="[611,721,596,699]" gridcol="3" gridrow="4" pageId="43" pageNumber="1318">E-E E-G G-G x2 50</td>
|
||
<td id="76828952F926007E5C5F9346075DFD60" box="[768,868,596,699]" gridcol="4" gridrow="4" pageId="43" pageNumber="1318">0 0 0 df5 3</td>
|
||
<td id="76828952F926007E5CC8934607C2FD60" box="[919,1019,596,699]" gridcol="5" gridrow="4" pageId="43" pageNumber="1318">
|
||
0.049 0.065 0.022
|
||
<emphasis id="B9173E7EF926FFAA5CF093B70786FD61" box="[943,959,677,698]" italics="true" pageId="43" pageNumber="1318">P</emphasis>
|
||
51
|
||
</td>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F0293DE07C2FCEA" box="[93,1019,716,817]" gridrow="5" pageId="43" pageNumber="1318">
|
||
<th id="76828952F926007E5F0293DE04F2FCEA" box="[93,203,716,817]" gridcol="0" gridrow="5" pageId="43" pageNumber="1318">IDH Category A-A A-B</th>
|
||
<td id="76828952F926007E5E4C93DE054EFCEA" box="[275,375,716,817]" gridcol="1" gridrow="5" pageId="43" pageNumber="1318">observed frequency 4 0</td>
|
||
<td id="76828952F926007E5EF593DE0637FCEA" box="[426,526,716,817]" gridcol="2" gridrow="5" pageId="43" pageNumber="1318">expected frequency 0.516 6.968</td>
|
||
<td id="76828952F926007E5D3C93DE06E8FCEA" box="[611,721,716,817]" gridcol="3" gridrow="5" pageId="43" pageNumber="1318">IDH Category A-A A-B</td>
|
||
<td id="76828952F926007E5C5F93DE075DFCEA" box="[768,868,716,817]" gridcol="4" gridrow="5" pageId="43" pageNumber="1318">observed frequency 19 0</td>
|
||
<td id="76828952F926007E5CC893DE07C2FCEA" box="[919,1019,716,817]" gridcol="5" gridrow="5" pageId="43" pageNumber="1318">expected frequency 7.848 22.304</td>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F02922407C2FCB3" box="[93,1019,822,872]" gridrow="6" pageId="43" pageNumber="1318">
|
||
<th id="76828952F926007E5F02922404F2FCB3" box="[93,203,822,872]" gridcol="0" gridrow="6" pageId="43" pageNumber="1318">B-B x2 523.633</th>
|
||
<td id="76828952F926007E5E4C9224054EFCB3" box="[275,375,822,872]" gridcol="1" gridrow="6" pageId="43" pageNumber="1318">27 df5 1</td>
|
||
<td id="76828952F926007E5EF592240637FCB3" box="[426,526,822,872]" gridcol="2" gridrow="6" pageId="43" pageNumber="1318">
|
||
23.516
|
||
<emphasis id="B9173E7EF926FFAA5E9D924305EBFCBD" box="[450,466,849,870]" italics="true" pageId="43" pageNumber="1318">P</emphasis>
|
||
50
|
||
</td>
|
||
<td id="76828952F926007E5D3C922406E8FCB3" box="[611,721,822,872]" gridcol="3" gridrow="6" pageId="43" pageNumber="1318">B-B x2 542.934</td>
|
||
<td id="76828952F926007E5C5F9224075DFCB3" box="[768,868,822,872]" gridcol="4" gridrow="6" pageId="43" pageNumber="1318">27 df5 1</td>
|
||
<td id="76828952F926007E5CC8922407C2FCB3" box="[919,1019,822,872]" gridcol="5" gridrow="6" pageId="43" pageNumber="1318">
|
||
15.848
|
||
<emphasis id="B9173E7EF926FFAA5CF092430786FCBD" box="[943,959,849,870]" italics="true" pageId="43" pageNumber="1318">P</emphasis>
|
||
50
|
||
</td>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F02929407C2FC47" box="[93,1019,902,924]" gridrow="7" pageId="43" pageNumber="1318" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76828952F926007E5F0292940637FC47" box="[93,526,902,924]" colspan="3" colspanRight="2" gridcol="0" gridrow="7" pageId="43" pageNumber="1318">
|
||
<taxonomicName id="4C6399EFF926FFAA5F9D929405FFFC47" authority=", Covadonga" box="[194,454,902,924]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="1318" phylum="Mollusca" rank="species" species="asturicum">
|
||
<emphasis id="B9173E7EF926FFAA5F9D9294057AFC40" box="[194,323,902,924]" italics="true" pageId="43" pageNumber="1318">C. asturicum</emphasis>
|
||
, Covadonga
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F0292A907C2FC37" box="[93,1019,955,1004]" gridrow="8" pageId="43" pageNumber="1318" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76828952F926007E5F0292A904F2FC37" box="[93,203,955,1004]" gridcol="0" gridrow="8" pageId="43" pageNumber="1318">LAP Category</th>
|
||
<td id="76828952F926007E5E4C92A9054EFC37" box="[275,375,955,1004]" gridcol="1" gridrow="8" pageId="43" pageNumber="1318">observed frequency</td>
|
||
<td id="76828952F926007E5EF592A90637FC37" box="[426,526,955,1004]" gridcol="2" gridrow="8" pageId="43" pageNumber="1318">expected frequency</td>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F02951907C2FBAA" box="[93,1019,1035,1137]" gridrow="9" pageId="43" pageNumber="1318" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76828952F926007E5F02951904F2FBAA" box="[93,203,1035,1137]" gridcol="0" gridrow="9" pageId="43" pageNumber="1318">B-B B-C C-C x2 50</th>
|
||
<td id="76828952F926007E5E4C9519054EFBAA" box="[275,375,1035,1137]" gridcol="1" gridrow="9" pageId="43" pageNumber="1318">0 1 14 df5 1</td>
|
||
<td id="76828952F926007E5EF595190637FBAA" box="[426,526,1035,1137]" gridcol="2" gridrow="9" pageId="43" pageNumber="1318">
|
||
0.017 0.967 14.017
|
||
<emphasis id="B9173E7EF926FFAA5E9D954905EBFBAB" box="[450,466,1115,1136]" italics="true" pageId="43" pageNumber="1318">P</emphasis>
|
||
51
|
||
</td>
|
||
</tr>
|
||
<tr id="3553E02EF926007E5F02956707C2FB50" box="[93,1019,1141,1163]" gridrow="10" pageId="43" pageNumber="1318" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76828952F926007E5F029567054EFB50" box="[93,375,1141,1163]" colspan="2" colspanRight="1" gridcol="0" gridrow="10" pageId="43" pageNumber="1318">IDH, ODH: monomorphic</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDCE26CF926FFAA5FD995AA04F0FAF4" blockId="43.[93,1049,1208,1392]" pageId="43" pageNumber="1318">Aperture strongly thickened and expanded into a collar, with a very distinct rim forming a sharp angle inside the outer lip. Peristome distinctly auriculated, gradually narrowing at the columellar insertion; collar usually contained in a plane, not distorted.</paragraph>
|
||
<paragraph id="8BDCE26CF926FFAA5FD994250776FA94" blockId="43.[93,1049,1208,1392]" box="[134,847,1335,1359]" pageId="43" pageNumber="1318">Shell colour of a uniform pale brown to tan. Peristome white.</paragraph>
|
||
<paragraph id="8BDCE26CF926FFAA5FD99445071CFAB4" blockId="43.[93,1049,1208,1392]" box="[134,805,1367,1392]" pageId="43" pageNumber="1318">Female and male genital systems as in general description.</paragraph>
|
||
<paragraph id="8BDCE26CF926FFAA5F02949504F9FA44" blockId="43.[93,1049,1415,1661]" box="[93,192,1415,1439]" pageId="43" pageNumber="1318">
|
||
<emphasis id="B9173E7EF926FFAA5F02949504F9FA44" box="[93,192,1415,1439]" italics="true" pageId="43" pageNumber="1318">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDCE26CF926FFAA5FD994B507C4FA25" blockId="43.[93,1049,1415,1661]" pageId="43" pageNumber="1318">
|
||
This species is easily recognized by its pattern of alternating strong and thin ribs.
|
||
<taxonomicName id="4C6399EFF926FFAA5F0294D4055AFA05" box="[93,355,1478,1502]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="1318" phylum="Mollusca" rank="species" species="asturicum">
|
||
<emphasis id="B9173E7EF926FFAA5F0294D4055AFA05" box="[93,355,1478,1502]" italics="true" pageId="43" pageNumber="1318">Cochlostoma asturicum</emphasis>
|
||
</taxonomicName>
|
||
, which is found sympatrically in the Picos de Europa area, is similar in general shape and size, but has equal ribs and a darker shell colour.
|
||
</paragraph>
|
||
<paragraph id="8BDCE26CF926FFAA5FD9971404C7F9A6" blockId="43.[93,1049,1415,1661]" pageId="43" pageNumber="1318">
|
||
Several characters are shared with the eastern Pyrenean species
|
||
<taxonomicName id="4C6399EFF926FFAA5C2B971407DAF9C5" baseAuthorityName="Dupuy" baseAuthorityYear="1851" box="[884,995,1542,1566]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="1318" phylum="Mollusca" rank="species" species="nouleti">
|
||
<emphasis id="B9173E7EF926FFAA5C2B971407DAF9C5" box="[884,995,1542,1566]" italics="true" pageId="43" pageNumber="1318">C. nouleti</emphasis>
|
||
</taxonomicName>
|
||
: the alternating sculpture, the wrinkled pattern on the very early part of the spire, the de®nite inner rim of the aperture, and among the allozymic characters the SOD and MPI patterns.
|
||
</paragraph>
|
||
<caption id="DF1CB2E4F921FFAD5F0292BB05EAFC2F" ID-DOI="http://doi.org/10.5281/zenodo.5277353" ID-Zenodo-Dep="5277353" httpUri="https://zenodo.org/record/5277353/files/figure.png" pageId="44" pageNumber="1319" startId="44.[93,109,937,958]" targetBox="[93,1050,105,911]" targetPageId="44">
|
||
<paragraph id="8BDCE26CF921FFAD5F0292BB05EAFC2F" blockId="44.[93,1049,937,1012]" pageId="44" pageNumber="1319">
|
||
FIG. 29. Shells of
|
||
<taxonomicName id="4C6399EFF921FFAD5E7E92BB05CEFC64" authorityName="Gofas" authorityYear="1989" box="[289,503,937,959]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="44" pageNumber="1319" phylum="Mollusca" rank="species" species="oscitans">
|
||
<emphasis id="B9173E7EF921FFAD5E7E92BB05CEFC64" box="[289,503,937,959]" italics="true" pageId="44" pageNumber="1319">Cochlostoma oscitans</emphasis>
|
||
</taxonomicName>
|
||
from Alisas. a: female (11.0 mm). b,c: male (9.9 mm). d: female (10.0 mm) with a very distorted outer lip. e,f,g: apical part showing embryonic whorls (scale bars are 1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDCE26CF921FFAD5F029530048AFBE1" blockId="44.[93,1049,1058,1242]" box="[93,179,1058,1082]" pageId="44" pageNumber="1319">
|
||
<emphasis id="B9173E7EF921FFAD5F029530048AFBE1" box="[93,179,1058,1082]" italics="true" pageId="44" pageNumber="1319">Habitat</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDCE26CF921FFAD5FD995500588FB01" blockId="44.[93,1049,1058,1242]" pageId="44" pageNumber="1319">
|
||
Among limestone, mostly among mounds of loose large stones in shaded places. The partitioning of the habitat is clearcut where the species is sympatric with
|
||
<taxonomicName id="4C6399EFF921FFAD5F02959004E3FB41" box="[93,218,1154,1178]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="44" pageNumber="1319" phylum="Mollusca" rank="species" species="hidalgoi">
|
||
<emphasis id="B9173E7EF921FFAD5F02959004E3FB41" box="[93,218,1154,1178]" italics="true" pageId="44" pageNumber="1319">C. hidalgoi</emphasis>
|
||
</taxonomicName>
|
||
; the latter living higher on exposed surfaces (®gure 34).
|
||
<taxonomicName id="4C6399EFF921FFAD5CD6959004D1FB61" authorityName="Gofas" authorityYear="1989" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="44" pageNumber="1319" phylum="Mollusca" rank="species" species="bicostulatum">
|
||
<emphasis id="B9173E7EF921FFAD5CD6959004D1FB61" italics="true" pageId="44" pageNumber="1319">Cochlostoma bicostulatum</emphasis>
|
||
</taxonomicName>
|
||
is more ubiquitous and sometimes very abundant in the coastal area, where it is often found alone.
|
||
</paragraph>
|
||
<paragraph id="8BDCE26CF921FFAD5F02941004D8FAC1" blockId="44.[93,1049,1282,1402]" box="[93,225,1282,1306]" pageId="44" pageNumber="1319">
|
||
<emphasis id="B9173E7EF921FFAD5F02941004D8FAC1" box="[93,225,1282,1306]" italics="true" pageId="44" pageNumber="1319">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDCE26CF921FFAD5FD9943006FFFAA1" blockId="44.[93,1049,1282,1402]" pageId="44" pageNumber="1319">
|
||
The coastal areas of the
|
||
<collectingRegion id="49A72C8EF921FFAD5EF794310662FAE0" box="[424,603,1315,1339]" country="Spain" name="Pais Vasco" pageId="44" pageNumber="1319">Basque country</collectingRegion>
|
||
, Cantabrian range and
|
||
<collectingRegion id="49A72C8EF921FFAD5C29943007EDFAE1" box="[886,980,1314,1338]" country="Spain" name="Asturias" pageId="44" pageNumber="1319">Asturias</collectingRegion>
|
||
. This species is predominant at lower altitudes, and can be found only a few metres away from the maritime splash zone if limestone is present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |