423 lines
49 KiB
XML
423 lines
49 KiB
XML
<document ID-DOI="10.5733/afin.051.0102" ID-ISSN="2305-2562" ID-Zenodo-Dep="7911071" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683578878403" checkinUser="felipe" docAuthor="Wesener, Thomas, Bespalova, Ioulia & Sierwald, Petra" docDate="2010" docId="03CA1219FFFF9E6DFEBBFD3DF37F1AC5" docLanguage="en" docName="AfrInvertebr.51.1.133-161.pdf" docOrigin="African Invertebrates 51 (1)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.051.0102" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.3:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="3" docTitle="Zoosphaerium tigrioculatum Wesener & Bespalova & Sierwald 2010, sp. n." docType="treatment" docVersion="4" lastPageNumber="148" masterDocId="FFF36A61FFF39E7DFFFEFFDBF0581B4A" masterDocTitle="Madagascar’s living giants: discovery of five new species of endemic giant pill-millipedes from Madagascar (Diplopoda: Sphaerotheriida: Arthrosphaeridae: Zoosphaerium)" masterLastPageNumber="133" masterPageNumber="133" pageNumber="144" updateTime="1683605769834" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Madagascar’s living giants: discovery of five new species of endemic giant pill-millipedes from Madagascar (Diplopoda: Sphaerotheriida: Arthrosphaeridae: Zoosphaerium)</mods:title>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bespalova, Ioulia</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-05-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>51</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>133</mods:start>
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<mods:end>133</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.051.0102</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5733/afin.051.0102</mods:identifier>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7911071</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03CA1219FFFF9E6DFEBBFD3DF37F1AC5" httpUri="http://treatment.plazi.org/id/03CA1219FFFF9E6DFEBBFD3DF37F1AC5" lastPageId="16" lastPageNumber="148" pageId="12" pageNumber="144">
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<subSubSection box="[325,998,742,767]" pageId="12" pageNumber="144" type="nomenclature">
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<paragraph blockId="12.[181,1143,742,1671]" box="[325,998,742,767]" pageId="12" pageNumber="144">
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<heading box="[325,998,742,767]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="144" reason="2">
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<taxonomicName ID-CoL="7GH33" authority="Wesener & Bespalova" authorityName="Wesener & Bespalova & Sierwald" authorityYear="2010" box="[325,926,742,767]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="144" phylum="Arthropoda" rank="species" species="tigrioculatum" status="sp. n.">
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<emphasis bold="true" box="[325,669,742,767]" pageId="12" pageNumber="144">Zoosphaerium tigrioculatum</emphasis>
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Wesener & Bespalova
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</taxonomicName>
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,
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<taxonomicNameLabel box="[938,998,742,767]" pageId="12" pageNumber="144" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="148" pageId="12" pageNumber="144" type="description">
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<paragraph blockId="12.[181,1143,742,1671]" box="[576,748,782,807]" pageId="12" pageNumber="144">
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||
<figureCitation box="[576,647,782,807]" captionStart="Fig" captionStartId="14.[182,215,1551,1571]" captionTargetBox="[182,1142,203,1528]" captionTargetId="figure-87@14.[182,1142,203,1528]" captionTargetPageId="14" captionText="Fig. 9. Zoosphaerium tigrioculatum sp. n., holotype: (A) 9th left leg; (B–D) left anterior telopod: (B) anterior view, (C) posterior view, (D) lateral view; (E–G) posterior telopods: (E) anterior view, (F) posterior view, (G) right posterior telopod, posterior view, detail. Roman numerals refer to number of podomere. Abbreviations: cr-t – crenulated, sclerotised teeth, ml – membranous lobe, IH – inner horns, IL – inner lobes, s-p – sclerotised spot, SR – stridulation rib. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7911459" httpUri="https://zenodo.org/record/7911459/files/figure.png" pageId="12" pageNumber="144">Figs 9</figureCitation>
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,
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<figureCitation box="[659,689,782,807]" captionStart="Fig" captionStartId="15.[144,177,905,925]" captionTargetBox="[154,1099,204,888]" captionTargetId="figure-302@15.[154,1100,203,888]" captionTargetPageId="15" captionText="Fig. 10. Zoosphaerium tigrioculatum sp. n., holotype, left antenna, SEM: (A) lateral view; (B) sensilla basiconica on first antennomere; (C) sensilla basiconica on second antennomere (apomorphy of the species); (D) apical view on disc. Arrows point to sensilla. Abbreviation: sb – sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.7911095" httpUri="https://zenodo.org/record/7911095/files/figure.png" pageId="12" pageNumber="144">10</figureCitation>
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,
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<figureCitation box="[701,748,782,807]" captionStart="Fig" captionStartId="24.[182,215,1061,1081]" captionTargetBox="[188,1134,203,1038]" captionTargetId="figure-244@24.[188,1134,203,1038]" captionTargetPageId="24" captionText="Fig. 17. Endoterga of tergites 10, arrows point to cuticular impressions: (A) holotype of Zoosphaerium muscorum sp. n., (B) holotype of Z. bambusoides sp. n., (C) holotype of Z. tigrioculatum sp. n., (D) holotype of Z. darthvaderi sp. n., (E) holotype of Z. heleios sp. n.Abbreviations: ci – cuticular impressions, IA – inner area with spines and long setae, mr – marginal ridge, mb – marginal bristles." figureDoi="http://doi.org/10.5281/zenodo.7911109" httpUri="https://zenodo.org/record/7911109/files/figure.png" pageId="12" pageNumber="144">17C</figureCitation>
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</paragraph>
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<paragraph blockId="12.[181,1143,742,1671]" box="[181,857,822,847]" pageId="12" pageNumber="144">
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Etymology: From Latin
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<emphasis box="[455,514,822,847]" italics="true" pageId="12" pageNumber="144">tigris</emphasis>
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(tiger) and
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<emphasis box="[645,718,822,847]" italics="true" pageId="12" pageNumber="144">oculus</emphasis>
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(ornament).
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</paragraph>
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<paragraph blockId="12.[181,1143,742,1671]" pageId="12" pageNumber="144">
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Differential diagnosis: The shape of the posterior telopods (
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<figureCitation box="[846,970,862,887]" captionStart="Fig" captionStartId="14.[182,215,1551,1571]" captionTargetBox="[182,1142,203,1528]" captionTargetId="figure-87@14.[182,1142,203,1528]" captionTargetPageId="14" captionText="Fig. 9. Zoosphaerium tigrioculatum sp. n., holotype: (A) 9th left leg; (B–D) left anterior telopod: (B) anterior view, (C) posterior view, (D) lateral view; (E–G) posterior telopods: (E) anterior view, (F) posterior view, (G) right posterior telopod, posterior view, detail. Roman numerals refer to number of podomere. Abbreviations: cr-t – crenulated, sclerotised teeth, ml – membranous lobe, IH – inner horns, IL – inner lobes, s-p – sclerotised spot, SR – stridulation rib. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7911459" httpUri="https://zenodo.org/record/7911459/files/figure.png" pageId="12" pageNumber="144">Figs 9E–G</figureCitation>
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) of
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<taxonomicName authorityName="Wesener & Bespalova & Sierwald" authorityYear="2010" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="144" phylum="Arthropoda" rank="species" species="tigrioculatum">
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<emphasis italics="true" pageId="12" pageNumber="144">Z. tigrioculatum</emphasis>
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</taxonomicName>
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clearly identifies this species as a member of the
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<taxonomicName baseAuthorityName="De Saussure & Zehntner" baseAuthorityYear="1902" box="[780,974,894,919]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="144" phylum="Arthropoda" rank="species" species="coquerelianum">
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<emphasis box="[780,974,894,919]" italics="true" pageId="12" pageNumber="144">Z. coquerelianum</emphasis>
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</taxonomicName>
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species-group. The presence of one large triangular membranous lobe on the fixed finger and two large membranous lobes on the movable finger (
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<figureCitation box="[663,752,958,983]" captionStart="Fig" captionStartId="14.[182,215,1551,1571]" captionTargetBox="[182,1142,203,1528]" captionTargetId="figure-87@14.[182,1142,203,1528]" captionTargetPageId="14" captionText="Fig. 9. Zoosphaerium tigrioculatum sp. n., holotype: (A) 9th left leg; (B–D) left anterior telopod: (B) anterior view, (C) posterior view, (D) lateral view; (E–G) posterior telopods: (E) anterior view, (F) posterior view, (G) right posterior telopod, posterior view, detail. Roman numerals refer to number of podomere. Abbreviations: cr-t – crenulated, sclerotised teeth, ml – membranous lobe, IH – inner horns, IL – inner lobes, s-p – sclerotised spot, SR – stridulation rib. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7911459" httpUri="https://zenodo.org/record/7911459/files/figure.png" pageId="12" pageNumber="144">Fig. 9G</figureCitation>
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) is a character shared only with
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<taxonomicName authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="144" phylum="Arthropoda" rank="species" species="isalo">
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<emphasis italics="true" pageId="12" pageNumber="144">Z. isalo</emphasis>
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<bibRefCitation author="WESENER, T." box="[246,415,990,1015]" pageId="12" pageNumber="144" pagination="1 - 134" refId="ref12130" refString="WESENER, T. 2009. Unexplored richness: discovery of 31 new species of giant pill-millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida). Zootaxa 2097: 1 - 134." type="journal article" year="2009">Wesener, 2009</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[476,784,990,1015]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="144" phylum="Arthropoda" rank="species" species="bilobum">
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<emphasis box="[476,601,990,1015]" italics="true" pageId="12" pageNumber="144">Z. bilobum</emphasis>
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<bibRefCitation author="WESENER, T." box="[611,784,990,1015]" pageId="12" pageNumber="144" pagination="1 - 134" refId="ref12130" refString="WESENER, T. 2009. Unexplored richness: discovery of 31 new species of giant pill-millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida). Zootaxa 2097: 1 - 134." type="journal article" year="2009">Wesener, 2009</bibRefCitation>
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</taxonomicName>
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.
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<taxonomicName authorityName="Wesener & Bespalova & Sierwald" authorityYear="2010" box="[798,984,990,1015]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="144" phylum="Arthropoda" rank="species" species="tigrioculatum">
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<emphasis box="[798,984,990,1015]" italics="true" pageId="12" pageNumber="144">Z. tigrioculatum</emphasis>
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</taxonomicName>
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has a unique combination of characters, like crenulated teeth on the third podomere of the anterior telopods (
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<figureCitation box="[292,380,1054,1079]" captionStart="Fig" captionStartId="14.[182,215,1551,1571]" captionTargetBox="[182,1142,203,1528]" captionTargetId="figure-87@14.[182,1142,203,1528]" captionTargetPageId="14" captionText="Fig. 9. Zoosphaerium tigrioculatum sp. n., holotype: (A) 9th left leg; (B–D) left anterior telopod: (B) anterior view, (C) posterior view, (D) lateral view; (E–G) posterior telopods: (E) anterior view, (F) posterior view, (G) right posterior telopod, posterior view, detail. Roman numerals refer to number of podomere. Abbreviations: cr-t – crenulated, sclerotised teeth, ml – membranous lobe, IH – inner horns, IL – inner lobes, s-p – sclerotised spot, SR – stridulation rib. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7911459" httpUri="https://zenodo.org/record/7911459/files/figure.png" pageId="12" pageNumber="144">Fig. 9C</figureCitation>
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), separating it clearly from the latter two (Table 1).
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</paragraph>
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<paragraph blockId="12.[181,1143,742,1671]" box="[181,321,1094,1119]" pageId="12" pageNumber="144">Description:</paragraph>
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<paragraph blockId="12.[181,1143,742,1671]" box="[181,246,1134,1159]" pageId="12" pageNumber="144">
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<emphasis box="[181,239,1134,1159]" italics="true" pageId="12" pageNumber="144">Male</emphasis>
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.
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</paragraph>
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<paragraph blockId="12.[181,1143,742,1671]" pageId="12" pageNumber="144">
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<emphasis box="[181,317,1174,1199]" italics="true" pageId="12" pageNumber="144">Body length</emphasis>
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:
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<typeStatus box="[333,442,1174,1199]" pageId="12" pageNumber="144">Holotype</typeStatus>
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: length
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<quantity box="[532,635,1174,1199]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="12" pageNumber="144" unit="mm" value="20.7">20.7 mm</quantity>
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, width of thoracic shield
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<quantity box="[925,1027,1174,1199]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="12" pageNumber="144" unit="mm" value="11.8">11.8 mm</quantity>
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, height of thoracic shield
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<quantity box="[353,442,1206,1231]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="12" pageNumber="144" unit="mm" value="6.3">6.3 mm</quantity>
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.
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</paragraph>
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<paragraph blockId="12.[181,1143,742,1671]" box="[181,628,1246,1271]" pageId="12" pageNumber="144">
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<emphasis box="[181,270,1246,1271]" italics="true" pageId="12" pageNumber="144">Habitus</emphasis>
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: Small, glabrous pill-millipede.
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</paragraph>
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<paragraph blockId="12.[181,1143,742,1671]" pageId="12" pageNumber="144">
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<emphasis box="[181,315,1286,1311]" italics="true" pageId="12" pageNumber="144">Colouration</emphasis>
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: Some discolouration may have occurred because of preservation in alcohol. Tergites anteriorly khaki coloured, followed by seaweed-green stripe, followed by dark brown margin at posterior. Dorsal side of anal shield same colour as tergites, ventral side of anal shield has light green tint. Legs bright teal under light. Head, collum and antennae dark olive-green.
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</paragraph>
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<paragraph blockId="12.[181,1143,742,1671]" pageId="12" pageNumber="144">
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<emphasis box="[181,241,1454,1479]" italics="true" pageId="12" pageNumber="144">Head</emphasis>
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: Eyes with
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<emphasis box="[368,394,1454,1479]" italics="true" pageId="12" pageNumber="144">ca</emphasis>
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65 ocelli.Antennae long and thin, with cylindrical joints (
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<figureCitation box="[1026,1129,1454,1479]" captionStart="Fig" captionStartId="15.[144,177,905,925]" captionTargetBox="[154,1099,204,888]" captionTargetId="figure-302@15.[154,1100,203,888]" captionTargetPageId="15" captionText="Fig. 10. Zoosphaerium tigrioculatum sp. n., holotype, left antenna, SEM: (A) lateral view; (B) sensilla basiconica on first antennomere; (C) sensilla basiconica on second antennomere (apomorphy of the species); (D) apical view on disc. Arrows point to sensilla. Abbreviation: sb – sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.7911095" httpUri="https://zenodo.org/record/7911095/files/figure.png" pageId="12" pageNumber="144">Fig. 10A</figureCitation>
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). Length of antennomeres: 1=2>3>4=5<6. Small shallow concavity at base of first antennomere (
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<figureCitation box="[252,354,1518,1543]" captionStart="Fig" captionStartId="15.[144,177,905,925]" captionTargetBox="[154,1099,204,888]" captionTargetId="figure-302@15.[154,1100,203,888]" captionTargetPageId="15" captionText="Fig. 10. Zoosphaerium tigrioculatum sp. n., holotype, left antenna, SEM: (A) lateral view; (B) sensilla basiconica on first antennomere; (C) sensilla basiconica on second antennomere (apomorphy of the species); (D) apical view on disc. Arrows point to sensilla. Abbreviation: sb – sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.7911095" httpUri="https://zenodo.org/record/7911095/files/figure.png" pageId="12" pageNumber="144">Fig. 10B</figureCitation>
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). Many sclerotised teeth at base of antennomere 1–3. Two rows of sensilla basiconica present on first antennomere (
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<figureCitation box="[698,801,1550,1575]" captionStart="Fig" captionStartId="15.[144,177,905,925]" captionTargetBox="[154,1099,204,888]" captionTargetId="figure-302@15.[154,1100,203,888]" captionTargetPageId="15" captionText="Fig. 10. Zoosphaerium tigrioculatum sp. n., holotype, left antenna, SEM: (A) lateral view; (B) sensilla basiconica on first antennomere; (C) sensilla basiconica on second antennomere (apomorphy of the species); (D) apical view on disc. Arrows point to sensilla. Abbreviation: sb – sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.7911095" httpUri="https://zenodo.org/record/7911095/files/figure.png" pageId="12" pageNumber="144">Fig. 10B</figureCitation>
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), isolated sensilla also present on second antennomere (
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<figureCitation box="[467,573,1582,1607]" captionStart="Fig" captionStartId="15.[144,177,905,925]" captionTargetBox="[154,1099,204,888]" captionTargetId="figure-302@15.[154,1100,203,888]" captionTargetPageId="15" captionText="Fig. 10. Zoosphaerium tigrioculatum sp. n., holotype, left antenna, SEM: (A) lateral view; (B) sensilla basiconica on first antennomere; (C) sensilla basiconica on second antennomere (apomorphy of the species); (D) apical view on disc. Arrows point to sensilla. Abbreviation: sb – sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.7911095" httpUri="https://zenodo.org/record/7911095/files/figure.png" pageId="12" pageNumber="144">Fig. 10C</figureCitation>
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). Apical disc bearing 4 sensory cones (
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<figureCitation box="[1023,1129,1582,1607]" captionStart="Fig" captionStartId="15.[144,177,905,925]" captionTargetBox="[154,1099,204,888]" captionTargetId="figure-302@15.[154,1100,203,888]" captionTargetPageId="15" captionText="Fig. 10. Zoosphaerium tigrioculatum sp. n., holotype, left antenna, SEM: (A) lateral view; (B) sensilla basiconica on first antennomere; (C) sensilla basiconica on second antennomere (apomorphy of the species); (D) apical view on disc. Arrows point to sensilla. Abbreviation: sb – sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.7911095" httpUri="https://zenodo.org/record/7911095/files/figure.png" pageId="12" pageNumber="144">Fig. 10D</figureCitation>
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). Mouthparts not dissected. Collum glabrous with few setae at corners on either side of head.
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</paragraph>
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<tableNote pageId="13" pageNumber="145" targetBox="[153,1101,289,678]" targetPageId="13">
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<paragraph blockId="13.[203,1046,204,272]" box="[579,667,204,224]" pageId="13" pageNumber="145">TABLE 1</paragraph>
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<paragraph blockId="13.[203,1046,204,272]" pageId="13" pageNumber="145">
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Comparison of
|
||
<taxonomicName authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[344,553,228,249]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="isalo">
|
||
<emphasis box="[344,412,229,249]" italics="true" pageId="13" pageNumber="145">Z. isalo</emphasis>
|
||
<bibRefCitation author="WESENER, T." box="[417,553,228,248]" pageId="13" pageNumber="145" pagination="1 - 134" refId="ref12130" refString="WESENER, T. 2009. Unexplored richness: discovery of 31 new species of giant pill-millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida). Zootaxa 2097: 1 - 134." type="journal article" year="2009">Wesener, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[562,798,228,249]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="bilobum">
|
||
<emphasis box="[562,660,229,249]" italics="true" pageId="13" pageNumber="145">Z. bilobum</emphasis>
|
||
<bibRefCitation author="WESENER, T." box="[665,798,228,248]" pageId="13" pageNumber="145" pagination="1 - 134" refId="ref12130" refString="WESENER, T. 2009. Unexplored richness: discovery of 31 new species of giant pill-millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida). Zootaxa 2097: 1 - 134." type="journal article" year="2009">Wesener, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Wesener & Bespalova & Sierwald, 2010" authorityName="Wesener & Bespalova & Sierwald" authorityYear="2010" box="[843,990,229,249]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="tigrioculatum" status="sp. n.">
|
||
<emphasis box="[843,990,229,249]" italics="true" pageId="13" pageNumber="145">Z. tigrioculatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[996,1045,228,248]" pageId="13" pageNumber="145" rank="species">sp. n.</taxonomicNameLabel>
|
||
Abbreviations: ANT – antenna, aT – anterior telopod, bas. – basiconica, Endo – endotergum.
|
||
</paragraph>
|
||
</tableNote>
|
||
<paragraph pageId="13" pageNumber="145">
|
||
<table box="[153,1101,289,678]" gridcols="4" gridrows="11" pageId="13" pageNumber="145">
|
||
<tr box="[153,1101,289,310]" gridrow="0" pageId="13" pageNumber="145">
|
||
<th box="[153,370,289,310]" gridcol="0" gridrow="0" pageId="13" pageNumber="145">
|
||
<emphasis bold="true" box="[153,252,289,309]" pageId="13" pageNumber="145">Character</emphasis>
|
||
</th>
|
||
<th box="[404,593,289,310]" gridcol="1" gridrow="0" pageId="13" pageNumber="145">
|
||
<taxonomicName authorityName="Wesener" authorityYear="2009" box="[404,473,290,310]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="isalo">
|
||
<emphasis bold="true" box="[404,473,290,310]" italics="true" pageId="13" pageNumber="145">Z. isalo</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th box="[631,862,289,310]" gridcol="2" gridrow="0" pageId="13" pageNumber="145">
|
||
<taxonomicName authorityName="Wesener" authorityYear="2009" box="[631,733,290,310]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="bilobum">
|
||
<emphasis bold="true" box="[631,733,290,310]" italics="true" pageId="13" pageNumber="145">Z. bilobum</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th box="[892,1101,289,310]" gridcol="3" gridrow="0" pageId="13" pageNumber="145">
|
||
<emphasis bold="true" box="[892,1101,289,310]" pageId="13" pageNumber="145">
|
||
<taxonomicName authority="Wesener & Bespalova & Sierwald, 2010" authorityName="Wesener & Bespalova & Sierwald" authorityYear="2010" box="[892,1044,290,310]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="tigrioculatum" status="sp. n.">
|
||
<emphasis bold="true" box="[892,1044,290,310]" italics="true" pageId="13" pageNumber="145">Z. tigrioculatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1050,1101,289,309]" pageId="13" pageNumber="145" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[153,1101,322,342]" gridrow="1" pageId="13" pageNumber="145">
|
||
<th box="[153,370,322,342]" gridcol="0" gridrow="1" pageId="13" pageNumber="145">shape of anal shield</th>
|
||
<td box="[404,593,322,342]" gridcol="1" gridrow="1" pageId="13" pageNumber="145">tapering</td>
|
||
<td box="[631,862,322,342]" gridcol="2" gridrow="1" pageId="13" pageNumber="145">tapering</td>
|
||
<td box="[892,1101,322,342]" gridcol="3" gridrow="1" pageId="13" pageNumber="145">weakly bell-shaped</td>
|
||
</tr>
|
||
<tr box="[153,1101,351,374]" gridrow="2" pageId="13" pageNumber="145">
|
||
<th box="[153,370,351,374]" gridcol="0" gridrow="2" pageId="13" pageNumber="145">locking carinae</th>
|
||
<td box="[404,593,351,374]" gridcol="1" gridrow="2" pageId="13" pageNumber="145">2nd 3× 1st</td>
|
||
<td box="[631,862,351,374]" gridcol="2" gridrow="2" pageId="13" pageNumber="145">2nd 2.5× 1st</td>
|
||
<td box="[892,1101,351,374]" gridcol="3" gridrow="2" pageId="13" pageNumber="145">2nd 4× 1st</td>
|
||
</tr>
|
||
<tr box="[153,1101,384,430]" gridrow="3" pageId="13" pageNumber="145">
|
||
<th box="[153,370,384,430]" gridcol="0" gridrow="3" pageId="13" pageNumber="145">1st leg, no. of ventral spines</th>
|
||
<td box="[404,593,384,430]" gridcol="1" gridrow="3" pageId="13" pageNumber="145">3 or 4</td>
|
||
<td box="[631,862,384,430]" gridcol="2" gridrow="3" pageId="13" pageNumber="145">6 or 7</td>
|
||
<td box="[892,1101,384,430]" gridcol="3" gridrow="3" pageId="13" pageNumber="145">4 or 5</td>
|
||
</tr>
|
||
<tr box="[153,1101,440,486]" gridrow="4" pageId="13" pageNumber="145">
|
||
<th box="[153,370,440,486]" gridcol="0" gridrow="4" pageId="13" pageNumber="145">2nd leg, no. of ventral spines</th>
|
||
<td box="[404,593,440,486]" gridcol="1" gridrow="4" pageId="13" pageNumber="145">4 or 5</td>
|
||
<td box="[631,862,440,486]" gridcol="2" gridrow="4" pageId="13" pageNumber="145">8 or 9</td>
|
||
<td box="[892,1101,440,486]" gridcol="3" gridrow="4" pageId="13" pageNumber="145">6 or 7</td>
|
||
</tr>
|
||
<tr box="[153,1101,496,518]" gridrow="5" pageId="13" pageNumber="145">
|
||
<th box="[153,370,496,518]" gridcol="0" gridrow="5" pageId="13" pageNumber="145">aT, 2nd podomere in av</th>
|
||
<td box="[404,593,496,518]" gridcol="1" gridrow="5" pageId="13" pageNumber="145">visible</td>
|
||
<td box="[631,862,496,518]" gridcol="2" gridrow="5" pageId="13" pageNumber="145">not visible</td>
|
||
<td box="[892,1101,496,518]" gridcol="3" gridrow="5" pageId="13" pageNumber="145">visible</td>
|
||
</tr>
|
||
<tr box="[153,1101,528,550]" gridrow="6" pageId="13" pageNumber="145">
|
||
<th box="[153,370,528,550]" gridcol="0" gridrow="6" pageId="13" pageNumber="145">aT, 3rd podomere</th>
|
||
<td box="[404,593,528,550]" gridcol="1" gridrow="6" pageId="13" pageNumber="145">w.o. crenulated teeth</td>
|
||
<td box="[631,862,528,550]" gridcol="2" gridrow="6" pageId="13" pageNumber="145">w.o. crenulated teeth</td>
|
||
<td box="[892,1101,528,550]" gridcol="3" gridrow="6" pageId="13" pageNumber="145">w. crenulated teeth</td>
|
||
</tr>
|
||
<tr box="[153,1101,562,582]" gridrow="7" pageId="13" pageNumber="145">
|
||
<th box="[153,370,562,582]" gridcol="0" gridrow="7" pageId="13" pageNumber="145">ANT, sclerotised teeth</th>
|
||
<td box="[404,593,562,582]" gridcol="1" gridrow="7" pageId="13" pageNumber="145">on antennomere 1–4</td>
|
||
<td box="[631,862,562,582]" gridcol="2" gridrow="7" pageId="13" pageNumber="145">on antennomere 1–5</td>
|
||
<td box="[892,1101,562,582]" gridcol="3" gridrow="7" pageId="13" pageNumber="145">on antennomere 1–3</td>
|
||
</tr>
|
||
<tr box="[153,1101,592,614]" gridrow="8" pageId="13" pageNumber="145">
|
||
<th box="[153,370,592,614]" gridcol="0" gridrow="8" pageId="13" pageNumber="145">ANT, sensilla bas.</th>
|
||
<td box="[404,593,592,614]" gridcol="1" gridrow="8" pageId="13" pageNumber="145">only on 1st</td>
|
||
<td box="[631,862,592,614]" gridcol="2" gridrow="8" pageId="13" pageNumber="145">on 1st and 5th</td>
|
||
<td box="[892,1101,592,614]" gridcol="3" gridrow="8" pageId="13" pageNumber="145">on 1st and 2nd</td>
|
||
</tr>
|
||
<tr box="[153,1101,626,646]" gridrow="9" pageId="13" pageNumber="145">
|
||
<th box="[153,370,626,646]" gridcol="0" gridrow="9" pageId="13" pageNumber="145">Endo, marginal bristle</th>
|
||
<td box="[404,593,626,646]" gridcol="1" gridrow="9" pageId="13" pageNumber="145">protruding to margin</td>
|
||
<td box="[631,862,626,646]" gridcol="2" gridrow="9" pageId="13" pageNumber="145">extending beyond margin</td>
|
||
<td box="[892,1101,626,646]" gridcol="3" gridrow="9" pageId="13" pageNumber="145">extending beyond</td>
|
||
</tr>
|
||
<tr box="[153,1101,658,678]" gridrow="10" pageId="13" pageNumber="145">
|
||
<th box="[153,370,658,678]" gridcol="0" gridrow="10" pageId="13" pageNumber="145">Endo, cuticular patterns</th>
|
||
<td box="[404,593,658,678]" gridcol="1" gridrow="10" pageId="13" pageNumber="145">single row</td>
|
||
<td box="[631,862,658,678]" gridcol="2" gridrow="10" pageId="13" pageNumber="145">two rows</td>
|
||
<td box="[892,1101,658,678]" gridcol="3" gridrow="10" pageId="13" pageNumber="145">two rows</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" box="[144,565,749,774]" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,319,749,774]" italics="true" pageId="13" pageNumber="145">Thoracic shield</emphasis>
|
||
: Surface like tergites.
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,231,789,814]" italics="true" pageId="13" pageNumber="145">Tergites</emphasis>
|
||
: Surface glabrous, shiny and smooth. Under magnification (50×) tergites display leathery pattern. Under SEM, surface covered by miniscule pits holding stubby sensory hair. Tips of paratergites projecting posteriorly and fairly rounded.
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,281,893,918]" italics="true" pageId="13" pageNumber="145">Endotergum</emphasis>
|
||
: Double row of marginal bristles of varying lengths, some extending far past edge of margin. Two rows of regularly distributed cuticular impressions between marginal ridge and inner area. Internal section with short spines and very few isolated long bristles (
|
||
<figureCitation box="[297,399,989,1014]" captionStart="Fig" captionStartId="24.[182,215,1061,1081]" captionTargetBox="[188,1134,203,1038]" captionTargetId="figure-244@24.[188,1134,203,1038]" captionTargetPageId="24" captionText="Fig. 17. Endoterga of tergites 10, arrows point to cuticular impressions: (A) holotype of Zoosphaerium muscorum sp. n., (B) holotype of Z. bambusoides sp. n., (C) holotype of Z. tigrioculatum sp. n., (D) holotype of Z. darthvaderi sp. n., (E) holotype of Z. heleios sp. n.Abbreviations: ci – cuticular impressions, IA – inner area with spines and long setae, mr – marginal ridge, mb – marginal bristles." figureDoi="http://doi.org/10.5281/zenodo.7911109" httpUri="https://zenodo.org/record/7911109/files/figure.png" pageId="13" pageNumber="145">Fig. 17C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,272,1029,1054]" italics="true" pageId="13" pageNumber="145">Anal shield</emphasis>
|
||
: Fairly well-rounded with very slight indication of bell shape because of some bulge toward dorsal side. Surface like tergites. Underside carrying 2 black locking carinae, anterior one short, about 1/4 length of posterior one. Carinae separated by a space about as long as 1/2 anterior carina.Anterior carina slightly curving towards margin of anal shield.
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,197,1197,1222]" italics="true" pageId="13" pageNumber="145">Legs</emphasis>
|
||
: Tarsi 1 with 4 or 5, 2 with 6 or 7, 3 with 8 or 9, and 4–21 with 8–10 ventral spines. First 2 leg pairs with an apical spine. Tarsi of legs 4–21 with an apical spine. In leg 10 femur 1.89×, tarsi 4.70× longer than wide (
|
||
<figureCitation box="[626,716,1261,1286]" captionStart="Fig" captionStartId="14.[182,215,1551,1571]" captionTargetBox="[182,1142,203,1528]" captionTargetId="figure-87@14.[182,1142,203,1528]" captionTargetPageId="14" captionText="Fig. 9. Zoosphaerium tigrioculatum sp. n., holotype: (A) 9th left leg; (B–D) left anterior telopod: (B) anterior view, (C) posterior view, (D) lateral view; (E–G) posterior telopods: (E) anterior view, (F) posterior view, (G) right posterior telopod, posterior view, detail. Roman numerals refer to number of podomere. Abbreviations: cr-t – crenulated, sclerotised teeth, ml – membranous lobe, IH – inner horns, IL – inner lobes, s-p – sclerotised spot, SR – stridulation rib. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7911459" httpUri="https://zenodo.org/record/7911459/files/figure.png" pageId="13" pageNumber="145">Fig. 9A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,319,1301,1326]" italics="true" pageId="13" pageNumber="145">Stigmatic plates</emphasis>
|
||
: First plate with triangular lobe. Lobe short and thin, pinched, triangular area at apex projecting towards coxa.
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" box="[144,360,1373,1398]" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,235,1373,1398]" italics="true" pageId="13" pageNumber="145">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,409,1413,1438]" italics="true" pageId="13" pageNumber="145">Male sexual characters</emphasis>
|
||
: Gonopore covered with a single, medially divided, elliptical sclerotised plate. Small apical portion of plate membranous. Covers 1/2 height and 2/3 width of coxa. Anal shield with slight bell shape, covered with very short setae.
|
||
</paragraph>
|
||
<paragraph blockId="13.[144,1105,749,1670]" lastBlockId="15.[143,1103,1037,1670]" lastPageId="15" lastPageNumber="147" pageId="13" pageNumber="145">
|
||
<emphasis box="[144,343,1517,1542]" italics="true" pageId="13" pageNumber="145">Anterior telopods</emphasis>
|
||
(
|
||
<figureCitation box="[360,491,1517,1542]" captionStart="Fig" captionStartId="14.[182,215,1551,1571]" captionTargetBox="[182,1142,203,1528]" captionTargetId="figure-87@14.[182,1142,203,1528]" captionTargetPageId="14" captionText="Fig. 9. Zoosphaerium tigrioculatum sp. n., holotype: (A) 9th left leg; (B–D) left anterior telopod: (B) anterior view, (C) posterior view, (D) lateral view; (E–G) posterior telopods: (E) anterior view, (F) posterior view, (G) right posterior telopod, posterior view, detail. Roman numerals refer to number of podomere. Abbreviations: cr-t – crenulated, sclerotised teeth, ml – membranous lobe, IH – inner horns, IL – inner lobes, s-p – sclerotised spot, SR – stridulation rib. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7911459" httpUri="https://zenodo.org/record/7911459/files/figure.png" pageId="13" pageNumber="145">Figs 9B–D</figureCitation>
|
||
): Harp with one strongly pronounced stridulation rib. Process of second podomere only slightly bent towards third podomere; apical portion facing third podomere covered with sclerotised spots. Below spots at base of projection with a sclerotised spine. Third podomere basally also including sclerotised spine. Large lobe covered by a mound of sclerotised spots located on mesal half of process, concavity on lateral half. A brown sclerotised spot located at very apex of podomere. Tiny sclerotised spine lateral to brown sclerotised spot, 3 or 4 sclerotised spines meso-apically, small sclerotised spine close to lateral margin of mound of sclerotised spots. Two sclerotised, crenulated teeth on apicolateral margin of concavity.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7911459" ID-Zenodo-Dep="7911459" httpUri="https://zenodo.org/record/7911459/files/figure.png" pageId="14" pageNumber="146" startId="14.[182,215,1551,1571]" targetBox="[182,1142,203,1528]" targetPageId="14" targetType="figure">
|
||
<paragraph blockId="14.[182,1143,1549,1667]" pageId="14" pageNumber="146">
|
||
Fig. 9.
|
||
<taxonomicName authority="Wesener & Bespalova & Sierwald, 2010" authorityName="Wesener & Bespalova & Sierwald" authorityYear="2010" box="[243,501,1551,1571]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="146" phylum="Arthropoda" rank="species" species="tigrioculatum" status="sp. n.">
|
||
<emphasis box="[243,501,1551,1571]" italics="true" pageId="14" pageNumber="146">Zoosphaerium tigrioculatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[506,554,1551,1571]" pageId="14" pageNumber="146" rank="species">sp. n.</taxonomicNameLabel>
|
||
, holotype: (A) 9
|
||
<superScript attach="left" box="[699,709,1549,1561]" fontSize="5" pageId="14" pageNumber="146">th</superScript>
|
||
left leg; (B–D) left anterior telopod: (B) anterior view, (C) posterior view, (D) lateral view; (E–G) posterior telopods: (E) anterior view, (F) posterior view, (G) right posterior telopod, posterior view, detail. Roman numerals refer to number of podomere. Abbreviations: cr-t – crenulated, sclerotised teeth, ml – membranous lobe, IH – inner horns, IL – inner lobes, s-p – sclerotised spot, SR – stridulation rib. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7911095" ID-Zenodo-Dep="7911095" httpUri="https://zenodo.org/record/7911095/files/figure.png" pageId="15" pageNumber="147" startId="15.[144,177,905,925]" targetBox="[154,1099,204,888]" targetPageId="15" targetType="figure">
|
||
<paragraph blockId="15.[144,1104,905,998]" pageId="15" pageNumber="147">
|
||
Fig. 10.
|
||
<taxonomicName authority="Wesener & Bespalova & Sierwald, 2010" authorityName="Wesener & Bespalova & Sierwald" authorityYear="2010" box="[224,486,906,926]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="tigrioculatum" status="sp. n.">
|
||
<emphasis box="[224,486,906,926]" italics="true" pageId="15" pageNumber="147">Zoosphaerium tigrioculatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[494,547,905,925]" pageId="15" pageNumber="147" rank="species">sp. n.</taxonomicNameLabel>
|
||
, holotype, left antenna, SEM: (A) lateral view; (B)
|
||
<emphasis italics="true" pageId="15" pageNumber="147">sensilla basiconica</emphasis>
|
||
on first antennomere; (C)
|
||
<emphasis box="[574,747,930,950]" italics="true" pageId="15" pageNumber="147">sensilla basiconica</emphasis>
|
||
on second antennomere (apomorphy of the species); (D) apical view on disc. Arrows point to
|
||
<emphasis box="[729,798,954,974]" italics="true" pageId="15" pageNumber="147">sensilla</emphasis>
|
||
. Abbreviation: sb –
|
||
<emphasis italics="true" pageId="15" pageNumber="147">sensilla basiconica</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="15.[143,1103,1037,1670]" pageId="15" pageNumber="147">
|
||
<emphasis box="[143,351,1173,1198]" italics="true" pageId="15" pageNumber="147">Posterior telopods</emphasis>
|
||
(
|
||
<figureCitation box="[367,495,1173,1198]" captionStart="Fig" captionStartId="14.[182,215,1551,1571]" captionTargetBox="[182,1142,203,1528]" captionTargetId="figure-87@14.[182,1142,203,1528]" captionTargetPageId="14" captionText="Fig. 9. Zoosphaerium tigrioculatum sp. n., holotype: (A) 9th left leg; (B–D) left anterior telopod: (B) anterior view, (C) posterior view, (D) lateral view; (E–G) posterior telopods: (E) anterior view, (F) posterior view, (G) right posterior telopod, posterior view, detail. Roman numerals refer to number of podomere. Abbreviations: cr-t – crenulated, sclerotised teeth, ml – membranous lobe, IH – inner horns, IL – inner lobes, s-p – sclerotised spot, SR – stridulation rib. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7911459" httpUri="https://zenodo.org/record/7911459/files/figure.png" pageId="15" pageNumber="147">Figs 9E–G</figureCitation>
|
||
): Movable finger moderately thick (2.32× longer than wide), weakly hooking towards fixed finger. One spine located apically, second slightly below first, third at midpoint of finger. Two membranous lobes present, one at midpoint, posteriorly to spine, second between midpoint and apex. On apical posterior margin are 11 sclerotised, crenulated teeth. Fixed finger slightly shorter than movable finger, curving towards latter. Membranous lobe located basally. Tip of fixed finger covered by some sclerotised spots. Telopods shiny and glabrous except for a small patch anteriorly and laterally on first podomere. Inner lobes fairly thin. Tips of inner horns slightly curving inward.
|
||
</paragraph>
|
||
<paragraph blockId="15.[143,1103,1037,1670]" pageId="15" pageNumber="147">
|
||
<materialsCitation collectingDate="2006-04-15" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" county="Parc National d'Andringitra" elevation="2000" latitude="-22.152222" location="Ambalavao" longLatPrecision="21" longitude="46.89917" municipality="Plateau d'Andohariana" pageId="15" pageNumber="147" specimenCode="BLF 13757, CASENT 9032798" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
|
||
<typeStatus box="[143,230,1467,1487]" pageId="15" pageNumber="147">Holotype</typeStatus>
|
||
: ơ (
|
||
<collectionCode box="[260,320,1467,1487]" pageId="15" pageNumber="147">CASC</collectionCode>
|
||
<specimenCode box="[324,425,1467,1487]" collectionCode="BLF" pageId="15" pageNumber="147">BLF 13757</specimenCode>
|
||
(
|
||
<specimenCode box="[435,609,1467,1487]" collectionCode="CASENT" pageId="15" pageNumber="147">CASENT 9032798</specimenCode>
|
||
)).
|
||
<collectingCountry box="[630,793,1467,1487]" name="Madagascar" pageId="15" pageNumber="147">MADAGASCAR</collectingCountry>
|
||
:
|
||
<emphasis box="[800,973,1467,1487]" italics="true" pageId="15" pageNumber="147">
|
||
<collectingRegion box="[800,973,1467,1487]" country="Madagascar" name="Fianarantsoa" pageId="15" pageNumber="147">Fianarantsoa Prov.</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingCounty pageId="15" pageNumber="147">Parc National d’Andringitra</collectingCounty>
|
||
,
|
||
<collectingMunicipality box="[281,495,1491,1511]" pageId="15" pageNumber="147">Plateau d’Andohariana</collectingMunicipality>
|
||
,
|
||
<quantity box="[507,584,1491,1511]" metricMagnitude="4" metricUnit="m" metricValue="3.59" pageId="15" pageNumber="147" unit="km" value="35.9">35.9 km</quantity>
|
||
205°
|
||
<location LSID="urn:lsid:plazi:treatment:03CA1219FFFF9E6DFEBBFD3DF37F1AC5:8EBCF5D4FFFC9E72FD7CFA08F2B51EAD" box="[642,749,1491,1511]" country="Madagascar" county="Parc National d'Andringitra" latitude="-22.152222" longLatPrecision="21" longitude="46.89917" municipality="Plateau d'Andohariana" name="Ambalavao" pageId="15" pageNumber="147" stateProvince="Fianarantsoa">Ambalavao</location>
|
||
, ericoid thicket,
|
||
<quantity box="[908,982,1491,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="147" unit="m" value="2000.0">
|
||
<elevation box="[908,982,1491,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="147" unit="m" value="2000.0">2000 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate box="[994,1100,1488,1511]" degrees="22" direction="south" minutes="09" orientation="latitude" pageId="15" pageNumber="147" precision="15" seconds="08" value="-22.152222">22°09'08"S</geoCoordinate>
|
||
:
|
||
<geoCoordinate box="[143,249,1512,1535]" degrees="46" direction="east" minutes="53" orientation="longitude" pageId="15" pageNumber="147" precision="15" seconds="57" value="46.89917">46°53'57"E</geoCoordinate>
|
||
,
|
||
<collectorName box="[258,361,1515,1535]" pageId="15" pageNumber="147">B.L. Fisher</collectorName>
|
||
<emphasis box="[367,413,1515,1535]" italics="true" pageId="15" pageNumber="147">et al.</emphasis>
|
||
,
|
||
<date box="[423,521,1515,1535]" pageId="15" pageNumber="147" value="2006-04-15">
|
||
<collectingDate box="[423,521,1515,1535]" pageId="15" pageNumber="147" value="2006-04-15">15.iv.2006</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="15.[143,1103,1037,1670]" pageId="15" pageNumber="147">
|
||
Distribution & Ecology:
|
||
<taxonomicName authorityName="Wesener & Bespalova & Sierwald" authorityYear="2010" box="[423,606,1549,1574]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="tigrioculatum">
|
||
<emphasis box="[423,606,1549,1574]" italics="true" pageId="15" pageNumber="147">Z. tigrioculatum</emphasis>
|
||
</taxonomicName>
|
||
is a giant pill-millipede species found at the highest elevation (
|
||
<quantity box="[349,437,1581,1606]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="147" unit="m" value="2000.0">2000 m</quantity>
|
||
) on
|
||
<collectingCountry box="[486,625,1581,1606]" name="Madagascar" pageId="15" pageNumber="147">Madagascar</collectingCountry>
|
||
. It seems to live in the ericoid shrub above the tree line. This species has only been recorded from the summit of Andringitra (
|
||
<figureCitation box="[151,221,1645,1670]" captionStart="Fig" captionStartId="3.[374,407,1647,1667]" captionTargetBox="[298,946,604,1625]" captionTargetId="figure-126@3.[298,947,604,1627]" captionTargetPageId="3" captionText="Fig. 2. Distribution map of new Zoosphaerium species." figureDoi="http://doi.org/10.5281/zenodo.7911075" httpUri="https://zenodo.org/record/7911075/files/figure.png" pageId="15" pageNumber="147">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="16.[181,1142,206,399]" box="[181,980,206,231]" pageId="16" pageNumber="148">
|
||
<emphasis box="[181,980,206,231]" italics="true" pageId="16" pageNumber="148">
|
||
New species not assigned to any existing
|
||
<taxonomicName box="[644,980,206,231]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="species-groups">Zoosphaerium species-groups</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="16.[181,1142,206,399]" pageId="16" pageNumber="148">
|
||
Currently, there are only three species,
|
||
<taxonomicName authority="Jeekel, 1999" authorityName="Jeekel" authorityYear="1999" box="[628,931,246,271]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="subreflexum">
|
||
<emphasis box="[628,788,246,271]" italics="true" pageId="16" pageNumber="148">Z. subreflexum</emphasis>
|
||
Jeekel, 1999
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="fisheri">
|
||
<emphasis box="[941,1037,246,271]" italics="true" pageId="16" pageNumber="148">Z. fisheri</emphasis>
|
||
<bibRefCitation author="WESENER, T." pageId="16" pageNumber="148" pagination="1 - 134" refId="ref12130" refString="WESENER, T. 2009. Unexplored richness: discovery of 31 new species of giant pill-millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida). Zootaxa 2097: 1 - 134." type="journal article" year="2009">Wesener, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Wesener, 2009" authorityName=": Wesener" authorityYear="2009" box="[297,647,278,303]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="analavelona">
|
||
<emphasis box="[297,467,278,303]" italics="true" pageId="16" pageNumber="148">Z. analavelona</emphasis>
|
||
<bibRefCitation author="WESENER, T." box="[477,647,278,303]" pageId="16" pageNumber="148" pagination="1 - 134" refId="ref12130" refString="WESENER, T. 2009. Unexplored richness: discovery of 31 new species of giant pill-millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida). Zootaxa 2097: 1 - 134." type="journal article" year="2009">Wesener, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, which could not be assigned to a species-group. The two newly described species do not share any special characters with each other or with the three species mentioned above. They feature unusual characters which place them in an isolated position inside
|
||
<taxonomicName authorityName="Pocock" authorityYear="1895" box="[638,800,374,399]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[638,800,374,399]" italics="true" pageId="16" pageNumber="148">Zoosphaerium</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |