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<document id="B31EAC8D50DA964234B6F33CD85CE111" ID-DOI="10.5281/zenodo.194125" ID-GBIF-Dataset="6ae8ac53-d51b-4bdc-b3c9-856e0a3ef8a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="194125" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460191135421" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2010" docId="03C4BE4AFFF90745FF19FC1FFCDBFE9E" docLanguage="en" docName="zt02403p041.pdf" docOrigin="Zootaxa 2403" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptalpheus denticulatus Anker &amp; Marin 2009" docType="treatment" docVersion="11" lastPageNumber="38" masterDocId="FFFDC632FFF60755FF8EFF9DFFC2FFBC" masterDocTitle="New findings of rare or little-known alpheid shrimp genera (Crustacea, Decapoda) in Moorea, French Polynesia" masterLastPageNumber="41" masterPageNumber="23" pageNumber="38" updateTime="1698236446756" updateUser="plazi">
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<mods:title id="538E1160B5B600276EDF8C5ACBA4C330">New findings of rare or little-known alpheid shrimp genera (Crustacea, Decapoda) in Moorea, French Polynesia</mods:title>
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<treatment id="03C4BE4AFFF90745FF19FC1FFCDBFE9E" ID-DOI="http://doi.org/10.5281/zenodo.3504079" ID-GBIF-Taxon="124956769" ID-Zenodo-Dep="3504079" LSID="urn:lsid:plazi:treatment:03C4BE4AFFF90745FF19FC1FFCDBFE9E" httpUri="http://treatment.plazi.org/id/03C4BE4AFFF90745FF19FC1FFCDBFE9E" lastPageId="16" lastPageNumber="38" pageId="15" pageNumber="38">
<subSubSection id="C3775CD7FFF9075AFF19FC1FFEC5FC03" pageId="15" pageNumber="38" type="nomenclature">
<paragraph id="8BD20F5CFFF9075AFF19FC1FFD1FFC20" blockId="15.[151,733,898,959]" box="[151,733,898,924]" pageId="15" pageNumber="38">
<heading id="D09AB830FFF9075AFF19FC1FFD1FFC20" bold="true" box="[151,733,898,924]" fontSize="11" level="1" pageId="15" pageNumber="38" reason="1">
<taxonomicName id="4C6D74DFFFF9075AFF19FC1FFD1FFC20" ID-CoL="85TJH" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[151,733,898,924]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B919D34EFFF9075AFF19FC1FFD1FFC20" bold="true" box="[151,733,898,924]" pageId="15" pageNumber="38">
<emphasis id="B919D34EFFF9075AFF19FC1FFE04FC20" bold="true" box="[151,454,898,924]" italics="true" pageId="15" pageNumber="38">Leptalpheus denticulatus</emphasis>
<bibRefCitation id="EFFC72ADFFF9075AFE43FC1FFD1FFC20" author="Anker" box="[461,733,898,924]" pageId="15" pageNumber="40" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin, 2009</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD20F5CFFF9075AFF19FC38FEC5FC03" blockId="15.[151,733,898,959]" box="[151,263,933,959]" pageId="15" pageNumber="38">
<figureCitation id="135613D9FFF9075AFF19FC38FEC5FC03" box="[151,263,933,959]" captionStart="FIGURE 10" captionStartId="16.[151,256,1827,1851]" captionTargetBox="[151,1436,346,1805]" captionTargetId="figure@16.[151,1436,346,1805]" captionTargetPageId="16" captionText="FIGURE 10. Leptalpheus denticulatus Anker &amp; Marin, 2009, three specimens from Moorea: A-D, ovigerous female (CL 8.90 mm), FLMNH UF Arthropoda 16463; E, ovigerous female (CL 5.40 mm), FLMNH UF Arthropoda 16532; F, female (CL 4.60 mm), FLMNH UF Arthropoda 16528; A, dorsal view; B, lateral view; C, frontal region, dorsal view; D, major chela, detail of fingers; E, tail fan, dorsal view; F, major cheliped in situ, lateral view." httpUri="https://zenodo.org/record/194135/files/figure.png" pageId="15" pageNumber="38">Figure 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF9075AFF19FC6DFC42FBBB" box="[151,896,1008,1031]" pageId="15" pageNumber="38" type="reference_group">
<paragraph id="8BD20F5CFFF9075AFF19FC6DFC42FBBB" blockId="15.[151,896,1008,1031]" box="[151,896,1008,1031]" pageId="15" pageNumber="38">
<treatmentCitationGroup id="AB7D2872FFF9075AFF19FC6DFC42FBBB" box="[151,896,1008,1031]" pageId="15" pageNumber="38">
<taxonomicName id="4C6D74DFFFF9075AFF19FC6DFD74FBBB" ID-CoL="85TJH" authority="Anker &amp; Marin 2009: 92" authorityName="Anker &amp; Marin" authorityPageNumber="92" authorityYear="2009" box="[151,694,1008,1031]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B919D34EFFF9075AFF19FC6DFE61FBBB" box="[151,419,1008,1031]" italics="true" pageId="15" pageNumber="38">Leptalpheus denticulatus</emphasis>
<treatmentCitation id="0ACC294DFFF9075AFE27FC6DFD74FBBB" author="Anker" box="[425,694,1008,1031]" httpUri="http://treatment.plazi.org/id/03AAE9259F1EFFE0FF7EF85168F54F55" page="92" pageId="15" pageNumber="38" year="2009">
<bibRefCitation id="EFFC72ADFFF9075AFE27FC6DFD4CFBBB" author="Anker" box="[425,654,1008,1031]" pageId="15" pageNumber="40" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin 2009</bibRefCitation>
: 92
</treatmentCitation>
</taxonomicName>
, figs. 16, 7a, b, 8.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF9075AFF19FBA8FF21FA23" pageId="15" pageNumber="38" type="materials_examined">
<paragraph id="8BD20F5CFFF9075AFF19FBA8FF21FA23" blockId="15.[151,1437,1077,2036]" pageId="15" pageNumber="38">
<emphasis id="B919D34EFFF9075AFF19FBA8FE4FFBF3" bold="true" box="[151,397,1077,1103]" pageId="15" pageNumber="38">Material examined</emphasis>
.
<collectingCountry id="F37A4FCCFFF9075AFE10FBA8FDB7FBF3" box="[414,629,1077,1103]" name="French Polynesia" pageId="15" pageNumber="38">
French
<taxonomicName id="4C6D74DFFFF9075AFE75FBA8FDB7FBF3" box="[507,629,1077,1103]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="29" phylum="Arthropoda" rank="species" species="polynesia">Polynesia</taxonomicName>
</collectingCountry>
, Society Islands. 1 ovigerous female (CL
<quantity id="4C95A2B9FFF9075AFBF2FBA8FB2CFBF3" box="[1148,1262,1077,1103]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="15" pageNumber="38" unit="mm" value="8.9">8.90 mm</quantity>
),
<collectionCode id="ED7C9799FFF9075AFA8DFBA8FAAEFBF3" box="[1283,1388,1077,1103]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="15" pageNumber="38">FLMNH</collectionCode>
UF Arthropoda 16463, Moorea, lagoon near Nihimaru River estuary, near-shore sand flat with some rocks, algae, holothurians (
<taxonomicName id="4C6D74DFFFF9075AFEB3FB1DFE3BFB25" box="[317,505,1152,1177]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="15" pageNumber="38" phylum="Echinodermata" rank="species" species="atra">
<emphasis id="B919D34EFFF9075AFEB3FB1DFE3BFB25" box="[317,505,1152,1177]" italics="true" pageId="15" pageNumber="38">Holothuria atra</emphasis>
</taxonomicName>
) from burrow, suction pump, depth:
<quantity id="4C95A2B9FFF9075AFC3DFB1DFBDAFB26" box="[947,1048,1152,1178]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="15" pageNumber="38" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.51 m</quantity>
, coll. A. Anker,
<date id="FFD3299CFFF9075AFB53FB1DFAA7FB26" box="[1245,1381,1152,1178]" pageId="15" pageNumber="38" value="2008-11-17">17.XI.2008</date>
[fcn BMOO-5519];
<specimenCount id="9D6BC4D5FFF9075AFEC0FB38FE77FB03" box="[334,437,1189,1215]" pageId="15" pageNumber="38" type="female">1 female</specimenCount>
(CL
<quantity id="4C95A2B9FFF9075AFE7BFB38FDA1FB03" box="[501,611,1189,1215]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="15" pageNumber="38" unit="mm" value="4.6">4.60 mm</quantity>
),
<collectionCode id="ED7C9799FFF9075AFDF7FB38FD22FB03" box="[633,736,1189,1215]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="15" pageNumber="38">FLMNH</collectionCode>
UF Arthropoda 16528, Moorea, lagoon, between Papetoai and Hotel Intercontinental, near-shore sand flat with intense burrowing activity, silt and sand, mounds, holothurians (
<taxonomicName id="4C6D74DFFFF9075AFEB4FB6DFE36FAB5" box="[314,500,1264,1289]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Aspidochirotida" pageId="15" pageNumber="38" phylum="Echinodermata" rank="species" species="atra">
<emphasis id="B919D34EFFF9075AFEB4FB6DFE36FAB5" box="[314,500,1264,1289]" italics="true" pageId="15" pageNumber="38">Holothuria atra</emphasis>
</taxonomicName>
), from burrow, suction pump, depth:
<quantity id="4C95A2B9FFF9075AFC27FB6DFBE3FAB6" box="[937,1057,1264,1290]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="15" pageNumber="38" unit="m" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.30.6 m</quantity>
, coll. A. Anker,
<date id="FFD3299CFFF9075AFB6EFB6DFAA4FAB6" box="[1248,1382,1264,1290]" pageId="15" pageNumber="38" value="2008-11-23">23.XI.2008</date>
[fcn BMOO-5636]; ovigerous female (CL
<quantity id="4C95A2B9FFF9075AFDD6FA88FD04FA93" box="[600,710,1301,1327]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="38" unit="mm" value="5.4">5.40 mm</quantity>
),
<collectionCode id="ED7C9799FFF9075AFD55FA88FC80FA93" box="[731,834,1301,1327]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="15" pageNumber="38">FLMNH</collectionCode>
UF Arthropoda 16532, same collection data as for previous specimen, fcn 5641;
<specimenCount id="9D6BC4D5FFF9075AFE79FAA7FD84FAE8" box="[503,582,1338,1364]" pageId="15" pageNumber="38" type="male">1 male</specimenCount>
(CL
<quantity id="4C95A2B9FFF9075AFD0DFAA7FD32FAE8" box="[643,752,1338,1364]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="15" pageNumber="38" unit="mm" value="4.9">4.90 mm</quantity>
),
<specimenCount id="9D6BC4D5FFF9075AFC8AFAA7FCA8FAE8" box="[772,874,1338,1364]" pageId="15" pageNumber="38" type="female">1 female</specimenCount>
(CL 7.00 mm),
<specimenCount id="9D6BC4D5FFF9075AFBA6FAA7FB5EFAE8" box="[1064,1180,1338,1364]" pageId="15" pageNumber="38" type="juvenile">1 juvenile</specimenCount>
(CL not determined),
<collectionCode id="ED7C9799FFF9075AFF19FAFDFEC2FAC6" box="[151,256,1376,1402]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="15" pageNumber="38">FLMNH</collectionCode>
UF Arthropoda 16526, same collection data as for previous specimens [
<collectionCode id="ED7C9799FFF9075AFBF8FAFDFB11FAC6" box="[1142,1235,1376,1402]" pageId="15" pageNumber="38">BMOO</collectionCode>
fcn 5634, 5635, 5633].
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF9075AFF48FA37FC96F9E6" pageId="15" pageNumber="38" type="description">
<paragraph id="8BD20F5CFFF9075AFF48FA37FD69FA78" blockId="15.[151,1437,1077,2036]" box="[198,683,1450,1476]" pageId="15" pageNumber="38">
<emphasis id="B919D34EFFF9075AFF48FA37FE95FA78" bold="true" box="[198,343,1450,1476]" pageId="15" pageNumber="38">Description</emphasis>
. See
<bibRefCitation id="EFFC72ADFFF9075AFE19FA37FD65FA78" author="Anker" box="[407,679,1450,1476]" pageId="15" pageNumber="40" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin (2009)</bibRefCitation>
.
</paragraph>
<paragraph id="8BD20F5CFFF9075AFF48FA52FC96F9E6" blockId="15.[151,1437,1077,2036]" pageId="15" pageNumber="38">
<emphasis id="B919D34EFFF9075AFF48FA52FE4AFA55" bold="true" box="[198,392,1487,1513]" pageId="15" pageNumber="38">Colour pattern</emphasis>
. Uniform pinkish, speckled with reddish chromatophores (
<figureCitation id="135613D9FFF9075AFBD1FA4DFB7EFA56" box="[1119,1212,1488,1514]" captionStart="FIGURE 10" captionStartId="16.[151,256,1827,1851]" captionTargetBox="[151,1436,346,1805]" captionTargetId="figure@16.[151,1436,346,1805]" captionTargetPageId="16" captionText="FIGURE 10. Leptalpheus denticulatus Anker &amp; Marin, 2009, three specimens from Moorea: A-D, ovigerous female (CL 8.90 mm), FLMNH UF Arthropoda 16463; E, ovigerous female (CL 5.40 mm), FLMNH UF Arthropoda 16532; F, female (CL 4.60 mm), FLMNH UF Arthropoda 16528; A, dorsal view; B, lateral view; C, frontal region, dorsal view; D, major chela, detail of fingers; E, tail fan, dorsal view; F, major cheliped in situ, lateral view." httpUri="https://zenodo.org/record/194135/files/figure.png" pageId="15" pageNumber="38">Fig. 10</figureCitation>
), more intense on antennular peduncles and tail fan (
<figureCitation id="135613D9FFF9075AFDBCFA68FD4BF9B3" box="[562,649,1525,1551]" captionStart="FIGURE 10" captionStartId="16.[151,256,1827,1851]" captionTargetBox="[151,1436,346,1805]" captionTargetId="figure@16.[151,1436,346,1805]" captionTargetPageId="16" captionText="FIGURE 10. Leptalpheus denticulatus Anker &amp; Marin, 2009, three specimens from Moorea: A-D, ovigerous female (CL 8.90 mm), FLMNH UF Arthropoda 16463; E, ovigerous female (CL 5.40 mm), FLMNH UF Arthropoda 16532; F, female (CL 4.60 mm), FLMNH UF Arthropoda 16528; A, dorsal view; B, lateral view; C, frontal region, dorsal view; D, major chela, detail of fingers; E, tail fan, dorsal view; F, major cheliped in situ, lateral view." httpUri="https://zenodo.org/record/194135/files/figure.png" pageId="15" pageNumber="38">Fig. 10</figureCitation>
C, D), and in large individuals forming diffuse bands on abdomen (
<figureCitation id="135613D9FFF9075AFF2EF987FF3AF988" box="[160,248,1562,1588]" captionStart="FIGURE 10" captionStartId="16.[151,256,1827,1851]" captionTargetBox="[151,1436,346,1805]" captionTargetId="figure@16.[151,1436,346,1805]" captionTargetPageId="16" captionText="FIGURE 10. Leptalpheus denticulatus Anker &amp; Marin, 2009, three specimens from Moorea: A-D, ovigerous female (CL 8.90 mm), FLMNH UF Arthropoda 16463; E, ovigerous female (CL 5.40 mm), FLMNH UF Arthropoda 16532; F, female (CL 4.60 mm), FLMNH UF Arthropoda 16528; A, dorsal view; B, lateral view; C, frontal region, dorsal view; D, major chela, detail of fingers; E, tail fan, dorsal view; F, major cheliped in situ, lateral view." httpUri="https://zenodo.org/record/194135/files/figure.png" pageId="15" pageNumber="38">Fig. 10</figureCitation>
A, B; see also
<bibRefCitation id="EFFC72ADFFF9075AFE25F987FD6DF988" author="Anker" box="[427,687,1562,1588]" pageId="15" pageNumber="40" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin 2009</bibRefCitation>
, fig. 8); walking legs mostly colourless, major chela hyalinewhitish, with a few red chromatophores on palm (
<figureCitation id="135613D9FFF9075AFD51F9DDFCF6F9E6" box="[735,820,1600,1626]" captionStart="FIGURE 10" captionStartId="16.[151,256,1827,1851]" captionTargetBox="[151,1436,346,1805]" captionTargetId="figure@16.[151,1436,346,1805]" captionTargetPageId="16" captionText="FIGURE 10. Leptalpheus denticulatus Anker &amp; Marin, 2009, three specimens from Moorea: A-D, ovigerous female (CL 8.90 mm), FLMNH UF Arthropoda 16463; E, ovigerous female (CL 5.40 mm), FLMNH UF Arthropoda 16532; F, female (CL 4.60 mm), FLMNH UF Arthropoda 16528; A, dorsal view; B, lateral view; C, frontal region, dorsal view; D, major chela, detail of fingers; E, tail fan, dorsal view; F, major cheliped in situ, lateral view." httpUri="https://zenodo.org/record/194135/files/figure.png" pageId="15" pageNumber="38">Fig. 10</figureCitation>
F).
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF9075AFF48F9F8FC08F976" pageId="15" pageNumber="38" type="materials_examined">
<paragraph id="8BD20F5CFFF9075AFF48F9F8FD4CF9C3" blockId="15.[151,1437,1077,2036]" box="[198,654,1637,1663]" pageId="15" pageNumber="38">
<emphasis id="B919D34EFFF9075AFF48F9F8FEA6F9C3" bold="true" box="[198,356,1637,1663]" pageId="15" pageNumber="38">
<typeStatus id="54D6B1FEFFF9075AFF48F9F8FEC6F9C3" box="[198,260,1637,1663]" pageId="15" pageNumber="38">Type</typeStatus>
locality
</emphasis>
. Nhatrang Bay,
<collectingCountry id="F37A4FCCFFF9075AFDADF9F8FD48F9C3" box="[547,650,1637,1663]" name="Vietnam" pageId="15" pageNumber="38">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BD20F5CFFF9075AFF48F917FC08F976" blockId="15.[151,1437,1077,2036]" pageId="15" pageNumber="38">
<emphasis id="B919D34EFFF9075AFF48F917FEEAF918" bold="true" box="[198,296,1674,1700]" pageId="15" pageNumber="38">Ecology</emphasis>
. All Moorea specimens were collected on shallow near-shore sand flats, at depths ranging from
<quantity id="4C95A2B9FFF9075AFF19F92DFED2F976" box="[151,272,1712,1738]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="15" pageNumber="38" unit="m" value="0.65" valueMax="1.0" valueMin="0.3">0.3 to 1 m</quantity>
, from burrows of unknown, presumably callianassid, hosts.
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF9075AFF48F948FC23F8A8" pageId="15" pageNumber="38" type="distribution">
<paragraph id="8BD20F5CFFF9075AFF48F948FC23F8A8" blockId="15.[151,1437,1077,2036]" pageId="15" pageNumber="38">
<emphasis id="B919D34EFFF9075AFF48F948FE9DF953" bold="true" box="[198,351,1749,1775]" pageId="15" pageNumber="38">Distribution</emphasis>
. Nhatrang,
<collectingCountry id="F37A4FCCFFF9075AFE7EF948FD95F953" box="[496,599,1749,1775]" name="Vietnam" pageId="15" pageNumber="38">Vietnam</collectingCountry>
; Panglao, the
<collectingCountry id="F37A4FCCFFF9075AFC8DF948FC48F953" box="[771,906,1749,1775]" name="Philippines" pageId="15" pageNumber="38">Philippines</collectingCountry>
;
<collectingCountry id="F37A4FCCFFF9075AFC14F948FC04F953" box="[922,966,1749,1775]" name="Fiji" pageId="15" pageNumber="38">Viti</collectingCountry>
Levu,
<collectingCountry id="F37A4FCCFFF9075AFB90F948FB85F953" box="[1054,1095,1749,1775]" name="Fiji" pageId="15" pageNumber="38">Fiji</collectingCountry>
(
<bibRefCitation id="EFFC72ADFFF9075AFBD6F948FAA2F953" author="Anker" box="[1112,1376,1749,1775]" pageId="15" pageNumber="40" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin 2009</bibRefCitation>
); the present report extends its range eastwards to Moorea,
<collectingCountry id="F37A4FCCFFF9075AFC81F967FC1FF8A8" box="[783,989,1786,1812]" name="French Polynesia" pageId="15" pageNumber="38">
French
<taxonomicName id="4C6D74DFFFF9075AFCE9F967FC1FF8A8" box="[871,989,1786,1812]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="29" phylum="Arthropoda" rank="species" species="polynesia">Polynesia</taxonomicName>
</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF90745FF48F882FCDBFE9E" lastPageId="16" lastPageNumber="39" pageId="15" pageNumber="38" type="discussion">
<paragraph id="8BD20F5CFFF90745FF48F882FCDBFE9E" blockId="15.[151,1437,1077,2036]" lastBlockId="16.[151,1437,152,290]" lastPageId="16" lastPageNumber="39" pageId="15" pageNumber="38">
<emphasis id="B919D34EFFF9075AFF48F882FEFAF885" bold="true" box="[198,312,1823,1849]" pageId="15" pageNumber="38">Remarks</emphasis>
. Moorea is presently the only locality where
<taxonomicName id="4C6D74DFFFF9075AFCD1F8BDFBD2F885" box="[863,1040,1824,1849]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B919D34EFFF9075AFCD1F8BDFBD2F885" box="[863,1040,1824,1849]" italics="true" pageId="15" pageNumber="38">L. denticulatus</emphasis>
</taxonomicName>
is known to occur sympatrically with
<taxonomicName id="4C6D74DFFFF9075AFF5DF8D8FE9EF8E2" box="[211,348,1861,1886]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B919D34EFFF9075AFF5DF8D8FE9EF8E2" box="[211,348,1861,1886]" italics="true" pageId="15" pageNumber="38">L. pacificus</emphasis>
</taxonomicName>
. These two species are very closely related (
<bibRefCitation id="EFFC72ADFFF9075AFCE7F8D8FBABF8E3" author="Anker" box="[873,1129,1861,1887]" pageId="15" pageNumber="40" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin 2009</bibRefCitation>
), but as mentioned above,
<taxonomicName id="4C6D74DFFFF9075AFF19F8F6FE91F838" box="[151,339,1899,1924]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B919D34EFFF9075AFF19F8F6FE91F838" box="[151,339,1899,1924]" italics="true" pageId="15" pageNumber="38">L. denticulatus</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="4C6D74DFFFF9075AFD39F8F6FC88F838" box="[695,842,1899,1924]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B919D34EFFF9075AFD39F8F6FC88F838" box="[695,842,1899,1924]" italics="true" pageId="15" pageNumber="38">L. pacificus</emphasis>
</taxonomicName>
by the presence of a small tooth on the mesial emargination of the uropodal diaeresis (cf.
<figureCitation id="135613D9FFF9075AFD16F80DFD1CF816" box="[664,734,1936,1962]" captionStart="FIGURE 9" captionStartId="14.[151,255,1934,1958]" captionTargetBox="[151,1436,343,1911]" captionTargetId="figure@14.[151,1436,343,1912]" captionTargetPageId="14" captionText="FIGURE 9. Leptalpheus pacificus Banner &amp; Banner, 1974, male (CL 7.50 mm) from Moorea, FLMNH UF Arthropoda 16489; A, dorsal view; B, lateral view; C, frontal region, dorsal view; D, uropodal exopod, dorsal view; E, major cheliped in situ, lateral view; E, same, detail of fingers." httpUri="https://zenodo.org/record/194134/files/figure.png" pageId="15" pageNumber="38">Fig. 9</figureCitation>
D, 10E) and the presence of numerous red chromatophores that are evenly scattered over the body and together provide a uniform pinkish-reddish colour, more intense on the antennular peduncles (
<figureCitation id="135613D9FFF9075AFE42F847FDDFF848" box="[460,541,2010,2036]" captionStart="FIGURE 9" captionStartId="14.[151,255,1934,1958]" captionTargetBox="[151,1436,343,1911]" captionTargetId="figure@14.[151,1436,343,1912]" captionTargetPageId="14" captionText="FIGURE 9. Leptalpheus pacificus Banner &amp; Banner, 1974, male (CL 7.50 mm) from Moorea, FLMNH UF Arthropoda 16489; A, dorsal view; B, lateral view; C, frontal region, dorsal view; D, uropodal exopod, dorsal view; E, major cheliped in situ, lateral view; E, same, detail of fingers." httpUri="https://zenodo.org/record/194134/files/figure.png" pageId="15" pageNumber="38">Figs. 9</figureCitation>
AC, 10AC). Interestingly, all specimens of
<taxonomicName id="4C6D74DFFFF9075AFBB5F846FB2BF848" box="[1083,1257,2011,2036]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B919D34EFFF9075AFBB5F846FB2BF848" box="[1083,1257,2011,2036]" italics="true" pageId="15" pageNumber="38">L. denticulatus</emphasis>
</taxonomicName>
were collected from burrows near a small estuary or in very shallow silt-covered parts of the inshore lagoon, whereas
<taxonomicName id="4C6D74DFFFE60745FA0AFF05FEC1FF6A" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="39" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B919D34EFFE60745FA0AFF05FEC1FF6A" italics="true" pageId="16" pageNumber="39">L. pacificus</emphasis>
</taxonomicName>
was collected from burrows in fine white sand at some distance from the main coast, for instance near the Motu Tiahura Moorea channel, where water is cleaner and more dynamic, indicating a possible ecological niche separation between these two species.
</paragraph>
</subSubSection>
</treatment>
</document>