treatments-xml/data/03/C4/BE/03C4BE4AFFF70751FF19FB82FBBAFA4E.xml
2024-06-21 12:22:17 +02:00

325 lines
44 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="78CE9F5EB4D1B8CA00714B97458F4427" ID-DOI="10.5281/zenodo.194125" ID-GBIF-Dataset="6ae8ac53-d51b-4bdc-b3c9-856e0a3ef8a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="194125" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460191135421" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2010" docId="03C4BE4AFFF70751FF19FB82FBBAFA4E" docLanguage="en" docName="zt02403p041.pdf" docOrigin="Zootaxa 2403" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng 2006" docType="treatment" docVersion="12" lastPageNumber="27" masterDocId="FFFDC632FFF60755FF8EFF9DFFC2FFBC" masterDocTitle="New findings of rare or little-known alpheid shrimp genera (Crustacea, Decapoda) in Moorea, French Polynesia" masterLastPageNumber="41" masterPageNumber="23" pageNumber="24" updateTime="1698236446756" updateUser="plazi">
<mods:mods id="71E568FBC2D396BD4EFDE77505A13D45" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="82BDCF0175884DC60783350CC4928B4B">
<mods:title id="A40C2EE898C3297D5E42EFE0C37337E4">New findings of rare or little-known alpheid shrimp genera (Crustacea, Decapoda) in Moorea, French Polynesia</mods:title>
</mods:titleInfo>
<mods:name id="D457885EB4C7379C45A6996030A657C6" type="personal">
<mods:role id="5616EC228E994F00F62FA5D295D3A92E">
<mods:roleTerm id="E046B2CB1DCDB2F8A5ECB7E941735DD8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C8FB13F419F79AC216BB3AA286E8B469">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="CAC976AC9D373F558FC82D6D7F07C393">text</mods:typeOfResource>
<mods:relatedItem id="87295C38DC8073E02C05C2BE6A761F74" type="host">
<mods:titleInfo id="5127167DA1883797249A8590F656CFF3">
<mods:title id="1CFD93A0058921C0E38316756B146518">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AA7068ABE73B030937D1353B3C5ED4A4">
<mods:date id="210DED52B9BB99A4FA54B843888D3273">2010</mods:date>
<mods:detail id="5E97AB239538BFBACE9D470FD50A4359" type="volume">
<mods:number id="294D9AB9E5A40E19719829A6BC7E608F">2403</mods:number>
</mods:detail>
<mods:extent id="50D8A9B5AEC38747343742D552CA1FDE" unit="page">
<mods:start id="89FFCCA920468152D5E4576EBA24BB1C">23</mods:start>
<mods:end id="7A7A702C1FE5AF04EFBC18FBC3D53FB6">41</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FB7F74BB8291DB6086C6069D7E5F2545">journal article</mods:classification>
<mods:identifier id="12E10133E93CA48428A11B08422A0874" type="DOI">10.5281/zenodo.194125</mods:identifier>
<mods:identifier id="83DF5FF298D145532B646AC024E5EDB8" type="GBIF-Dataset">6ae8ac53-d51b-4bdc-b3c9-856e0a3ef8a0</mods:identifier>
<mods:identifier id="21F6CAFB304C478F46B04A22BB3F9DA5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="415883A4D31DCA3CC1224ED2B7AF62C7" type="Zenodo-Dep">194125</mods:identifier>
</mods:mods>
<treatment id="03C4BE4AFFF70751FF19FB82FBBAFA4E" ID-DOI="http://doi.org/10.5281/zenodo.4509732" ID-GBIF-Taxon="124956771" ID-Zenodo-Dep="4509732" LSID="urn:lsid:plazi:treatment:03C4BE4AFFF70751FF19FB82FBBAFA4E" httpUri="http://treatment.plazi.org/id/03C4BE4AFFF70751FF19FB82FBBAFA4E" lastPageId="4" lastPageNumber="27" pageId="1" pageNumber="24">
<subSubSection id="C3775CD7FFF70754FF19FB82FEE0FBE0" pageId="1" pageNumber="24" type="nomenclature">
<paragraph id="8BD20F5CFFF70754FF19FB82FCACFB85" blockId="1.[151,878,1055,1116]" box="[151,878,1055,1081]" pageId="1" pageNumber="24">
<heading id="D09AB830FFF70754FF19FB82FCACFB85" bold="true" box="[151,878,1055,1081]" fontSize="11" level="1" pageId="1" pageNumber="24" reason="1">
<taxonomicName id="4C6D74DFFFF70754FF19FB82FCACFB85" ID-CoL="85L2F" authority="Anker, Poddoubtchenko &amp; Jeng, 2006" authorityName="Anker, Poddoubtchenko &amp; Jeng" authorityYear="2006" box="[151,878,1055,1081]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF70754FF19FB82FCACFB85" bold="true" box="[151,878,1055,1081]" pageId="1" pageNumber="24">
<emphasis id="B919D34EFFF70754FF19FB82FE4FFB85" bold="true" box="[151,397,1055,1081]" italics="true" pageId="1" pageNumber="24">Acanthanas pusillus</emphasis>
<bibRefCitation id="EFFC72ADFFF70754FE1BFB82FCACFB85" author="Anker" box="[405,878,1055,1081]" pageId="1" pageNumber="40" refString="Anker, A., Poddoubtchenko, D. &amp; Jeng, M. S. (2006) Acanthanas pusillus, new genus, new species, a miniature alpheid shrimp with spiny eyes from the Philippines (Crustacea: Decapoda). Raffles Bulletin of Zoology, 54, 341 - 348." type="journal article" year="2006">Anker, Poddoubtchenko &amp; Jeng, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD20F5CFFF70754FF19FBDFFEE0FBE0" blockId="1.[151,878,1055,1116]" box="[151,290,1090,1116]" pageId="1" pageNumber="24">
<figureCitation id="135613D9FFF70754FF19FBDFFEC7FBE0" box="[151,261,1090,1116]" captionStart="FIGURE 1" captionStartId="2.[151,255,1858,1882]" captionTargetBox="[175,1410,477,1840]" captionTargetId="figure@2.[175,1412,477,1840]" captionTargetPageId="2" captionText="FIGURE 1. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A C, male (CL 2.30 mm), FLMNH UF Arthropoda 16055; D E, ovigerous female (CL 2.20 mm), FLMNH UF Arthropoda 15613; A, dorsal view; B, lateral view; C, frontal region, dorsolateral view; D, lateral view; E frontal region and chelipeds, lateral view." httpUri="https://zenodo.org/record/194126/files/figure.png" pageId="1" pageNumber="24">Figures 1</figureCitation>
,
<figureCitation id="135613D9FFF70754FE9DFBDFFEE0FBE0" box="[275,290,1090,1116]" captionStart="FIGURE 2" captionStartId="3.[151,255,1323,1347]" captionTargetBox="[151,1434,181,1270]" captionTargetId="figure@3.[151,1436,181,1302]" captionTargetPageId="3" captionText="FIGURE 2. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A D, larger male (CL 2.15 mm), FLMNH UF Arthropoda 23090; E, smaller male (CL 1.32 mm), OUMNH-ZC. 2010 - 01 - 001; A, lateral view; B, right cheliped, lateral view; C, same, chela; D, same, chela and carpus, mesial view; E, pleopod. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/194127/files/figure.png" pageId="1" pageNumber="24">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF70754FF19FB15FC6FFB23" box="[151,941,1160,1183]" pageId="1" pageNumber="24" type="reference_group">
<paragraph id="8BD20F5CFFF70754FF19FB15FC6FFB23" blockId="1.[151,941,1160,1183]" box="[151,941,1160,1183]" pageId="1" pageNumber="24">
<treatmentCitationGroup id="AB7D2872FFF70754FF19FB15FC6FFB23" box="[151,941,1160,1183]" pageId="1" pageNumber="24">
<taxonomicName id="4C6D74DFFFF70754FF19FB15FCFCFB23" ID-CoL="85L2F" authority="Anker, Poddoubtchenko &amp; Jeng 2006: 342" authorityName="Anker, Poddoubtchenko &amp; Jeng" authorityPageNumber="342" authorityYear="2006" box="[151,830,1160,1183]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF70754FF19FB15FEACFB23" box="[151,366,1160,1183]" italics="true" pageId="1" pageNumber="24">Acanthanas pusillus</emphasis>
<treatmentCitation id="0ACC294DFFF70754FEFAFB15FCFCFB23" author="Anker" box="[372,830,1160,1183]" httpUri="http://treatment.plazi.org/id/03F587BFFFE2FF8FD4B542200B9CDBE3" page="342" pageId="1" pageNumber="24" year="2006">
<bibRefCitation id="EFFC72ADFFF70754FEFAFB15FCCBFB23" author="Anker" box="[372,777,1160,1183]" pageId="1" pageNumber="40" refString="Anker, A., Poddoubtchenko, D. &amp; Jeng, M. S. (2006) Acanthanas pusillus, new genus, new species, a miniature alpheid shrimp with spiny eyes from the Philippines (Crustacea: Decapoda). Raffles Bulletin of Zoology, 54, 341 - 348." type="journal article" year="2006">Anker, Poddoubtchenko &amp; Jeng 2006</bibRefCitation>
: 342
</treatmentCitation>
</taxonomicName>
, figs. 15.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF70754FF19FB5AFEB8F97B" pageId="1" pageNumber="24" type="materials_examined">
<paragraph id="8BD20F5CFFF70754FF19FB5AFEB8F97B" blockId="1.[151,1437,1223,2034]" pageId="1" pageNumber="24">
<emphasis id="B919D34EFFF70754FF19FB5AFE4CFB5D" bold="true" box="[151,398,1223,1249]" pageId="1" pageNumber="24">Material examined</emphasis>
.
<collectingCountry id="F37A4FCCFFF70754FE11FB55FDB5FB5E" box="[415,631,1224,1250]" name="French Polynesia" pageId="1" pageNumber="24">
French
<taxonomicName id="4C6D74DFFFF70754FE73FB55FDB5FB5E" box="[509,631,1224,1250]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="29" phylum="Arthropoda" rank="species" species="polynesia">Polynesia</taxonomicName>
</collectingCountry>
, Society Islands.
<specimenCount id="9D6BC4D5FFF70754FCDEFB55FC61FB5E" box="[848,931,1224,1250]" pageId="1" pageNumber="24" type="male">1 male</specimenCount>
(CL
<quantity id="4C95A2B9FFF70754FC68FB55FB9AFB5E" box="[998,1112,1224,1250]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="24" unit="mm" value="2.3">2.30 mm</quantity>
),
<collectionCode id="ED7C9799FFF70754FBE0FB55FB1AFB5E" box="[1134,1240,1224,1250]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="24">FLMNH</collectionCode>
UF Arthropoda 16055, Moorea, between Temae and Afareaitu, 17.5145˚ S, 149.7616˚ W, outer reef slope, deep from within reef rubble, depth:
<quantity id="4C95A2B9FFF70754FEF0FA8FFE7DFA90" box="[382,447,1298,1324]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="24" unit="m" value="20.0">20 m</quantity>
, coll. C. Meyer, C. McKeon, J. Moore, S. Fay, G. Paulay, processed by A. Anker,
<date id="FFD3299CFFF70754FF19FAA5FED6FAEE" box="[151,276,1336,1362]" pageId="1" pageNumber="24" value="2008-10-23">23.X.2008</date>
[fcn BMOO-4077]; 1 ovigerous female (CL
<quantity id="4C95A2B9FFF70754FCA0FAA5FC5EFAEE" box="[814,924,1336,1362]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="24" unit="mm" value="2.2">2.20 mm</quantity>
),
<collectionCode id="ED7C9799FFF70754FC3FFAA5FBDAFAEE" box="[945,1048,1336,1362]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="24">FLMNH</collectionCode>
UF Arthropoda 15613, Moorea, east of Opunohu pass, at shark feeding buoy, 17.4817˚ S, 149.8558˚ W, outer reef slope, gentle sand and reef slope, from within coral rubble, outer reef slope, depth:
<quantity id="4C95A2B9FFF70754FCC4FA1FFC7BFA20" box="[842,953,1410,1436]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="1" pageNumber="24" unit="m" value="17.5" valueMax="18.0" valueMin="17.0">1718 m</quantity>
, coll. C. McKeon, J. Moore, G. Paulay, processed by A. Anker,
<date id="FFD3299CFFF70754FE4CFA35FD85FA7E" box="[450,583,1448,1474]" pageId="1" pageNumber="24" value="2008-10-16">16.X.2008</date>
, fcn BMOO-2811;
<specimenCount id="9D6BC4D5FFF70754FCB9FA35FC60FA7E" box="[823,930,1448,1474]" pageId="1" pageNumber="24" type="female">1 female</specimenCount>
(CL
<quantity id="4C95A2B9FFF70754FC6BFA35FB95FA7E" box="[997,1111,1448,1474]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="24" unit="mm" value="1.85">1.85 mm</quantity>
),
<collectionCode id="ED7C9799FFF70754FBE3FA35FB15FA7E" box="[1133,1239,1448,1474]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="24">FLMNH</collectionCode>
UF Arthropoda 23088, Moorea, off Temae, 17.4784˚ S, 149.7639˚ W, outer reef slope, deep from within rubble, rubble brushing, depth:
<quantity id="4C95A2B9FFF70754FEE0FA6FFE22F9B0" box="[366,480,1522,1548]" metricMagnitude="1" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="1" pageNumber="24" unit="m" value="20.5" valueMax="23.0" valueMin="18.0">1823 m</quantity>
, coll. G. Paulay, C. McKeon, S. Bush, S. Haddock, processed by A. Anker,
<date id="FFD3299CFFF70754FF19F985FED6F98E" box="[151,276,1560,1586]" pageId="1" pageNumber="24" value="2009-10-29">29.X.2009</date>
[fcn BMOO-6707];
<specimenCount id="9D6BC4D5FFF70754FD85F985FD99F98E" box="[523,603,1560,1586]" pageId="1" pageNumber="24" type="male">1 male</specimenCount>
(CL
<quantity id="4C95A2B9FFF70754FD15F985FCCBF98E" box="[667,777,1560,1586]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="1" pageNumber="24" unit="mm" value="2.15">2.15 mm</quantity>
),
<collectionCode id="ED7C9799FFF70754FC90F985FC47F98E" box="[798,901,1560,1586]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="24">FLMNH</collectionCode>
UF Arthropoda 23090, Moorea, Temae Bay, 149.7517˚ S, 17.4915˚ W, steep reef drop-off, under rubble, rubble brushing and cracking, depth:
<quantity id="4C95A2B9FFF70754FAB7F9A0FA55F9EB" box="[1337,1431,1597,1623]" metricMagnitude="1" metricUnit="m" metricValue="1.35" metricValueMax="2.0" metricValueMin="0.7" pageId="1" pageNumber="24" unit="m" value="13.5" valueMax="20.0" valueMin="7.0">720 m</quantity>
, coll. C. Meyer, S. McPherson, A. Collins, M. Parker, C. McKeon, processed by A. Anker,
<date id="FFD3299CFFF70754FB53F9FFFA9DF9C0" box="[1245,1375,1634,1660]" pageId="1" pageNumber="24" value="2009-10-22">22.X.2009</date>
; [fcn BMOO-6487];
<specimenCount id="9D6BC4D5FFF70754FEDAF915FE70F91E" box="[340,434,1672,1698]" pageId="1" pageNumber="24" type="male">2 males</specimenCount>
(CL
<quantity id="4C95A2B9FFF70754FE7BF915FDA7F91E" box="[501,613,1672,1698]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="24" unit="mm" value="1.3">1.30 mm</quantity>
,
<quantity id="4C95A2B9FFF70754FDFCF915FD26F91E" box="[626,740,1672,1698]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="1" pageNumber="24" unit="mm" value="1.32">1.32 mm</quantity>
), OUMNH-ZC.2010-01-001, same collection data as for previous specimen.
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF70751FF48F94FFF24FCEE" lastPageId="4" lastPageNumber="27" pageId="1" pageNumber="24" type="description">
<paragraph id="8BD20F5CFFF70754FF48F94FFD8CF83E" blockId="1.[151,1437,1223,2034]" pageId="1" pageNumber="24">
<emphasis id="B919D34EFFF70754FF48F94FFE95F950" bold="true" box="[198,343,1746,1772]" pageId="1" pageNumber="24">Description</emphasis>
. Anker
<emphasis id="B919D34EFFF70754FE32F94EFE38F950" box="[444,506,1747,1772]" italics="true" pageId="1" pageNumber="24">et al.</emphasis>
(2006) provided a detailed description of
<taxonomicName id="4C6D74DFFFF70754FC72F94EFBB9F950" box="[1020,1147,1747,1772]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF70754FC72F94EFBB9F950" box="[1020,1147,1747,1772]" italics="true" pageId="1" pageNumber="24">A. pusillus</emphasis>
</taxonomicName>
based on two complete females (one of which was dissected). The finding of several male specimens in Moorea, including an almost complete adult male, enables to complete the original description with the description of two important male features, viz. the cheliped and the pleopod. It also makes necessary a slight emendation of the generic diagnosis of
<taxonomicName id="4C6D74DFFFF70754FEA2F8F5FE75F83D" box="[300,439,1896,1921]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B919D34EFFF70754FEA2F8F5FE75F83D" box="[300,439,1896,1921]" italics="true" pageId="1" pageNumber="24">Acanthanas</emphasis>
</taxonomicName>
(see below).
</paragraph>
<paragraph id="8BD20F5CFFF70754FF48F810FB19F81B" blockId="1.[151,1437,1223,2034]" box="[198,1243,1933,1959]" pageId="1" pageNumber="24">
<emphasis id="B919D34EFFF70754FF48F810FCB4F81B" bold="true" box="[198,886,1933,1959]" pageId="1" pageNumber="24">
Description of the male cheliped of
<taxonomicName id="4C6D74DFFFF70754FDF1F810FCB4F81B" box="[639,886,1933,1959]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF70754FDF1F810FCB4F81B" bold="true" box="[639,886,1933,1959]" italics="true" pageId="1" pageNumber="24">Acanthanas pusillus</emphasis>
</taxonomicName>
</emphasis>
(cf. Anker
<emphasis id="B919D34EFFF70754FC73F810FBF5F81A" box="[1021,1079,1933,1958]" italics="true" pageId="1" pageNumber="24">et al.</emphasis>
2006, p. 343)
</paragraph>
<paragraph id="8BD20F5CFFF70757FF48F82FFD52FE0B" blockId="1.[151,1437,1223,2034]" lastBlockId="2.[151,1436,152,439]" lastPageId="2" lastPageNumber="25" pageId="1" pageNumber="24">
Male chelipeds symmetrical or subsymmetrical, polymorphic depending on age/size. Chelipeds of immature males slightly longer and more robust (
<figureCitation id="135613D9FFF70754FD52F845FCDCF84E" box="[732,798,2008,2034]" captionStart="FIGURE 1" captionStartId="2.[151,255,1858,1882]" captionTargetBox="[175,1410,477,1840]" captionTargetId="figure@2.[175,1412,477,1840]" captionTargetPageId="2" captionText="FIGURE 1. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A C, male (CL 2.30 mm), FLMNH UF Arthropoda 16055; D E, ovigerous female (CL 2.20 mm), FLMNH UF Arthropoda 15613; A, dorsal view; B, lateral view; C, frontal region, dorsolateral view; D, lateral view; E frontal region and chelipeds, lateral view." httpUri="https://zenodo.org/record/194126/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
B) than those of females (
<figureCitation id="135613D9FFF70754FBDFF845FB51F84E" box="[1105,1171,2008,2034]" captionStart="FIGURE 1" captionStartId="2.[151,255,1858,1882]" captionTargetBox="[175,1410,477,1840]" captionTargetId="figure@2.[175,1412,477,1840]" captionTargetPageId="2" captionText="FIGURE 1. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A C, male (CL 2.30 mm), FLMNH UF Arthropoda 16055; D E, ovigerous female (CL 2.20 mm), FLMNH UF Arthropoda 15613; A, dorsal view; B, lateral view; C, frontal region, dorsolateral view; D, lateral view; E frontal region and chelipeds, lateral view." httpUri="https://zenodo.org/record/194126/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
D). Chelipeds of adult males (CL&gt;
<quantity id="4C95A2B9FFF40757FEA5FF05FEB0FF0E" box="[299,370,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="25" unit="mm" value="2.0">2 mm</quantity>
) significantly enlarged relative to body size (
<figureCitation id="135613D9FFF40757FC07FF05FC12FF0E" box="[905,976,152,178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1323,1347]" captionTargetBox="[151,1434,181,1270]" captionTargetId="figure@3.[151,1436,181,1302]" captionTargetPageId="3" captionText="FIGURE 2. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A D, larger male (CL 2.15 mm), FLMNH UF Arthropoda 23090; E, smaller male (CL 1.32 mm), OUMNH-ZC. 2010 - 01 - 001; A, lateral view; B, right cheliped, lateral view; C, same, chela; D, same, chela and carpus, mesial view; E, pleopod. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/194127/files/figure.png" pageId="2" pageNumber="25">Fig. 2</figureCitation>
A), elongate, robust, with large chelae; basis with slender distal spine on dorsal margin; ischium with row of five slender spines on dorsal margin (
<figureCitation id="135613D9FFF40757FF2EFF7FFF2BFF40" box="[160,233,226,252]" captionStart="FIGURE 2" captionStartId="3.[151,255,1323,1347]" captionTargetBox="[151,1434,181,1270]" captionTargetId="figure@3.[151,1436,181,1302]" captionTargetPageId="3" captionText="FIGURE 2. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A D, larger male (CL 2.15 mm), FLMNH UF Arthropoda 23090; E, smaller male (CL 1.32 mm), OUMNH-ZC. 2010 - 01 - 001; A, lateral view; B, right cheliped, lateral view; C, same, chela; D, same, chela and carpus, mesial view; E, pleopod. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/194127/files/figure.png" pageId="2" pageNumber="25">Fig. 2</figureCitation>
B); merus robust, widening distally, both dorsal and ventral margins ending in sharp tooth (
<figureCitation id="135613D9FFF40757FAA1FF7FFABAFF40" box="[1327,1400,226,252]" captionStart="FIGURE 2" captionStartId="3.[151,255,1323,1347]" captionTargetBox="[151,1434,181,1270]" captionTargetId="figure@3.[151,1436,181,1302]" captionTargetPageId="3" captionText="FIGURE 2. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A D, larger male (CL 2.15 mm), FLMNH UF Arthropoda 23090; E, smaller male (CL 1.32 mm), OUMNH-ZC. 2010 - 01 - 001; A, lateral view; B, right cheliped, lateral view; C, same, chela; D, same, chela and carpus, mesial view; E, pleopod. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/194127/files/figure.png" pageId="2" pageNumber="25">Fig. 2</figureCitation>
B); carpus cup-shaped, embracing proximal portion of palm; palm smooth, somewhat compressed laterally, ovoid in cross-section, distinctly longer than wide, with numerous fine setae along ventral surface; fingers longer than half length of palm, somewhat twisted, dactylus strongly curved; cutting edges of fingers unarmed (on dactylus) or with inconspicuously irregular teeth (on pollex); lateral side of pollex shallowly depressed and bearing strong subdistal tooth (
<figureCitation id="135613D9FFF40757FD8CFE00FD8BFE0B" box="[514,585,413,439]" captionStart="FIGURE 2" captionStartId="3.[151,255,1323,1347]" captionTargetBox="[151,1434,181,1270]" captionTargetId="figure@3.[151,1436,181,1302]" captionTargetPageId="3" captionText="FIGURE 2. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A D, larger male (CL 2.15 mm), FLMNH UF Arthropoda 23090; E, smaller male (CL 1.32 mm), OUMNH-ZC. 2010 - 01 - 001; A, lateral view; B, right cheliped, lateral view; C, same, chela; D, same, chela and carpus, mesial view; E, pleopod. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/194127/files/figure.png" pageId="2" pageNumber="25">Fig. 2</figureCitation>
C, D).
</paragraph>
<caption id="DF125FD4FFF40757FF19F8DFFF13F805" httpUri="https://zenodo.org/record/194126/files/figure.png" pageId="2" pageNumber="25" targetBox="[175,1410,477,1840]" targetPageId="2">
<paragraph id="8BD20F5CFFF40757FF19F8DFFF13F805" blockId="2.[151,1436,1858,1977]" pageId="2" pageNumber="25">
<emphasis id="B919D34EFFF40757FF19F8DFFED1F8E6" bold="true" box="[151,275,1858,1882]" pageId="2" pageNumber="25">FIGURE 1</emphasis>
.
<taxonomicName id="4C6D74DFFFF40757FEACF8DFFC66F8E5" authority="Anker, Poddoubtchenko &amp; Jeng, 2006" authorityName="Anker, Poddoubtchenko &amp; Jeng" authorityYear="2006" box="[290,932,1858,1881]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF40757FEACF8DFFE3EF8E5" box="[290,508,1858,1881]" italics="true" pageId="2" pageNumber="25">Acanthanas pusillus</emphasis>
Anker, Poddoubtchenko &amp; Jeng, 2006
</taxonomicName>
, two specimens from Moorea, AC, male (CL 2.30 mm), FLMNH UF Arthropoda 16055; DE, ovigerous female (CL 2.20 mm), FLMNH UF Arthropoda 15613; A, dorsal view; B, lateral view; C, frontal region, dorsolateral view; D, lateral view; E frontal region and chelipeds, lateral view.
</paragraph>
</caption>
<caption id="DF125FD4FFF50756FF19FAB6FF32FA1F" httpUri="https://zenodo.org/record/194127/files/figure.png" pageId="3" pageNumber="26" targetBox="[151,1434,181,1270]" targetPageId="3">
<paragraph id="8BD20F5CFFF50756FF19FAB6FF32FA1F" blockId="3.[151,1436,1323,1443]" pageId="3" pageNumber="26">
<emphasis id="B919D34EFFF50756FF19FAB6FED0FAFF" bold="true" box="[151,274,1323,1347]" pageId="3" pageNumber="26">FIGURE 2</emphasis>
.
<taxonomicName id="4C6D74DFFFF50756FEAEFAB1FC5AFAFF" authority="Anker, Poddoubtchenko &amp; Jeng, 2006" authorityName="Anker, Poddoubtchenko &amp; Jeng" authorityYear="2006" box="[288,920,1324,1347]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF50756FEAEFAB1FE35FAFF" box="[288,503,1324,1347]" italics="true" pageId="3" pageNumber="26">Acanthanas pusillus</emphasis>
Anker, Poddoubtchenko &amp; Jeng, 2006
</taxonomicName>
, two specimens from Moorea, AD, larger male (CL 2.15 mm), FLMNH UF Arthropoda 23090; E, smaller male (CL 1.32 mm), OUMNH-ZC.2010-01-001; A, lateral view; B, right cheliped, lateral view; C, same, chela; D, same, chela and carpus, mesial view; E, pleopod. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD20F5CFFF50756FF48FA4DFB14FA57" blockId="3.[151,1437,1488,2038]" box="[198,1238,1488,1515]" pageId="3" pageNumber="26">
<emphasis id="B919D34EFFF50756FF48FA4DFCB3FA57" bold="true" box="[198,881,1488,1515]" pageId="3" pageNumber="26">
Description of the male pleopod of
<taxonomicName id="4C6D74DFFFF50756FDF4FA4CFCB3FA57" box="[634,881,1489,1515]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF50756FDF4FA4CFCB3FA57" bold="true" box="[634,881,1489,1515]" italics="true" pageId="3" pageNumber="26">Acanthanas pusillus</emphasis>
</taxonomicName>
</emphasis>
(cf. Anker
<emphasis id="B919D34EFFF50756FC76FA4CFBF1FA56" box="[1016,1075,1489,1514]" italics="true" pageId="3" pageNumber="26">et al.</emphasis>
2006, p. 344)
</paragraph>
<paragraph id="8BD20F5CFFF50756FF48FA6BFD63F98A" blockId="3.[151,1437,1488,2038]" pageId="3" pageNumber="26">
Male pleopod with endopod smaller than exopod, bearing appendix interna and appendix masculine, latter reaching distal margin of endopod (
<figureCitation id="135613D9FFF50756FDB6F981FDBDF98A" box="[568,639,1564,1590]" captionStart="FIGURE 2" captionStartId="3.[151,255,1323,1347]" captionTargetBox="[151,1434,181,1270]" captionTargetId="figure@3.[151,1436,181,1302]" captionTargetPageId="3" captionText="FIGURE 2. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A D, larger male (CL 2.15 mm), FLMNH UF Arthropoda 23090; E, smaller male (CL 1.32 mm), OUMNH-ZC. 2010 - 01 - 001; A, lateral view; B, right cheliped, lateral view; C, same, chela; D, same, chela and carpus, mesial view; E, pleopod. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/194127/files/figure.png" pageId="3" pageNumber="26">Fig. 2</figureCitation>
E).
</paragraph>
<paragraph id="8BD20F5CFFF50756FF48F9DDFC02F9E7" blockId="3.[151,1437,1488,2038]" box="[198,960,1600,1627]" pageId="3" pageNumber="26">
<emphasis id="B919D34EFFF50756FF48F9DDFD99F9E7" bold="true" box="[198,603,1600,1627]" pageId="3" pageNumber="26">
Revised diagnosis of
<taxonomicName id="4C6D74DFFFF50756FE44F9DCFD99F9E7" box="[458,603,1601,1627]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B919D34EFFF50756FE44F9DCFD99F9E7" bold="true" box="[458,603,1601,1627]" italics="true" pageId="3" pageNumber="26">Acanthanas</emphasis>
</taxonomicName>
</emphasis>
(cf. Anker
<emphasis id="B919D34EFFF50756FD6CF9DCFCDEF9E6" box="[738,796,1601,1626]" italics="true" pageId="3" pageNumber="26">et al.</emphasis>
2006, p. 342)
</paragraph>
<paragraph id="8BD20F5CFFF50751FF4AF9FBFDA2FDCE" blockId="3.[151,1437,1488,2038]" lastBlockId="4.[151,1437,152,1522]" lastPageId="4" lastPageNumber="27" pageId="3" pageNumber="26">Body bulky stout. Frontal margin with long, distally acute rostrum and acute orbital teeth. Pterygostomial angle bluntly angular to rounded, protruding. Eyes mostly concealed in dorsal and lateral view, visible in frontal view; anteromesial margin of cornea and base of eyestalk with spiny projections. Antennular peduncle short, stout; first segment with large tooth on ventromesial carina; lateral antennular flagellum feebly biramous. Antenna with moderately robust basicerite; carpocerite short, not overreaching scaphocerite, latter with well-developed blade. Mouthparts typical for family; mandible with two-segmented palp and expanded incisor process. Third maxilliped pediform, lateral plate ear-shaped, acute distally; tip of ultimate segment with spines. First pereiopods (= chelipeds) equal, symmetrical or subsymmetrical, carried extended, sexually dimorphic, feebly enlarged in females and immature males, greatly enlarged, robust in adult males; ischium unarmed in females, armed with slender spines on dorsal margin in adult males; merus ending in acute or subacute distal tooth dorsally and in adult males also ventrally; carpus somewhat elongate, subcylindrical in females, short, cup-shaped, distally widening in adult males, mesial side without setal rows; chelae simple, smooth, feebly enlarged in females, conspicuously enlarged in adult males; linea impressa and adhesive disks absent; fingers without snapping mechanism, unarmed in females, pollex armed with small teeth in adult males. Second pereiopod with five-segmented carpus. Third pereiopod with ischium bearing spine on ventrolateral surface; merus and carpus unarmed; propodus with spinules on ventral margin; dactylus biunguiculate. Fifth pereiopod with cleaning brush consisting of several setal rows on propodus. Sixth abdominal somite with articulated plate at posteroventral angle. Second pleopod with appendices masculina and interna in both males and females (including ovigerous females). Uropod with lateral lobe of protopod [formerly called “sympodite”] bearing two teeth; diaeresis of exopod sinuous. Telson with two pairs of dorsal spines and two pairs of posterolateral spines; posterior margin feebly rounded; anal tubercles absent. Gill/ exopod formula: 5 pleurobranchs (above P15); 0 arthrobranch; 0 podobranch; 2 lobe-shaped epipods (Mxp1- 2); 3 strap-like epipods = mastigobranchs (Mxp3 and P12), 3 setobranchs (P13), 3 fully developed exopods (Mxp13), 1 rudimentary exopod (P1).</paragraph>
<paragraph id="8BD20F5CFFF20751FF48FDE0FF24FCEE" blockId="4.[151,1437,152,1522]" pageId="4" pageNumber="27">
<emphasis id="B919D34EFFF20751FF48FDE0FE43FD2B" bold="true" box="[198,385,637,663]" pageId="4" pageNumber="27">Colour pattern</emphasis>
. As the
<typeStatus id="54D6B1FEFFF20751FE6CFDE0FDD7FD2B" box="[482,533,637,663]" pageId="4" pageNumber="27">type</typeStatus>
specimens, some individuals from Moorea had an intricate pattern of red and white marbling, white and red patches and small red dots (
<figureCitation id="135613D9FFF20751FCDBFD3FFC5AFD00" box="[853,920,674,700]" captionStart="FIGURE 1" captionStartId="2.[151,255,1858,1882]" captionTargetBox="[175,1410,477,1840]" captionTargetId="figure@2.[175,1412,477,1840]" captionTargetPageId="2" captionText="FIGURE 1. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A C, male (CL 2.30 mm), FLMNH UF Arthropoda 16055; D E, ovigerous female (CL 2.20 mm), FLMNH UF Arthropoda 15613; A, dorsal view; B, lateral view; C, frontal region, dorsolateral view; D, lateral view; E frontal region and chelipeds, lateral view." httpUri="https://zenodo.org/record/194126/files/figure.png" pageId="4" pageNumber="27">Fig. 1</figureCitation>
D, E); other specimens had a more distinct pattern consisting of a white carapace speckled with tiny red dots, a mostly red abdomen with some whitish or transparent dots (
<figureCitation id="135613D9FFF20751FEE7FD70FE6CFCBB" box="[361,430,749,775]" captionStart="FIGURE 1" captionStartId="2.[151,255,1858,1882]" captionTargetBox="[175,1410,477,1840]" captionTargetId="figure@2.[175,1412,477,1840]" captionTargetPageId="2" captionText="FIGURE 1. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A C, male (CL 2.30 mm), FLMNH UF Arthropoda 16055; D E, ovigerous female (CL 2.20 mm), FLMNH UF Arthropoda 15613; A, dorsal view; B, lateral view; C, frontal region, dorsolateral view; D, lateral view; E frontal region and chelipeds, lateral view." httpUri="https://zenodo.org/record/194126/files/figure.png" pageId="4" pageNumber="27">Fig. 1</figureCitation>
A, B; 2A); chelipeds, walking legs and frontal appendages mostly semitransparent whitish; chelipeds speckled with a few red dots; see also description and photographs in Anker
<emphasis id="B919D34EFFF20751FA82FC8FFA8AFC97" box="[1292,1352,786,811]" italics="true" pageId="4" pageNumber="27">et al.</emphasis>
(2006, fig. 5).
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF20751FF48FCC0FA4DFC20" pageId="4" pageNumber="27" type="materials_examined">
<paragraph id="8BD20F5CFFF20751FF48FCC0FD55FCCB" blockId="4.[151,1437,152,1522]" box="[198,663,861,887]" pageId="4" pageNumber="27">
<emphasis id="B919D34EFFF20751FF48FCC0FEA6FCCB" bold="true" box="[198,356,861,887]" pageId="4" pageNumber="27">
<typeStatus id="54D6B1FEFFF20751FF48FCC0FEC6FCCB" box="[198,260,861,887]" pageId="4" pageNumber="27">Type</typeStatus>
locality
</emphasis>
. Panglao, the
<collectingCountry id="F37A4FCCFFF20751FD85FCC0FD51FCCB" box="[523,659,861,887]" name="Philippines" pageId="4" pageNumber="27">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BD20F5CFFF20751FF48FC1FFA4DFC20" blockId="4.[151,1437,152,1522]" box="[198,1423,898,924]" pageId="4" pageNumber="27">
<emphasis id="B919D34EFFF20751FF48FC1FFEEAFC20" bold="true" box="[198,296,898,924]" pageId="4" pageNumber="27">Ecology</emphasis>
. Dwelling deep in crevices of dead corals, at depths of
<quantity id="4C95A2B9FFF20751FC3FFC1FFBC8FC20" box="[945,1034,898,924]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.3" metricValueMin="0.7" pageId="4" pageNumber="27" unit="m" value="15.0" valueMax="23.0" valueMin="7.0">723 m</quantity>
(Moorea) or
<quantity id="4C95A2B9FFF20751FB29FC1FFACCFC20" box="[1191,1294,898,924]" metricMagnitude="1" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="4" pageNumber="27" unit="m" value="22.0" valueMax="24.0" valueMin="20.0">2024 m</quantity>
(Panglao).
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF20751FF48FC3AFA5AFBB0" pageId="4" pageNumber="27" type="distribution">
<paragraph id="8BD20F5CFFF20751FF48FC3AFA5AFBB0" blockId="4.[151,1437,152,1522]" pageId="4" pageNumber="27">
<emphasis id="B919D34EFFF20751FF48FC3AFE9DFC7D" bold="true" box="[198,351,935,961]" pageId="4" pageNumber="27">Distribution</emphasis>
. Previously known only from the two
<typeStatus id="54D6B1FEFFF20751FCA9FC35FC9BFC7E" box="[807,857,936,962]" pageId="4" pageNumber="27">type</typeStatus>
specimens from Panglao, the
<collectingCountry id="F37A4FCCFFF20751FB33FC35FA83FC7E" box="[1213,1345,936,962]" name="Philippines" pageId="4" pageNumber="27">Philippines</collectingCountry>
(Anker
<emphasis id="B919D34EFFF20751FF19FC50FF13FC5A" box="[151,209,973,998]" italics="true" pageId="4" pageNumber="27">et al.</emphasis>
2006); now recorded for the first time from Moorea,
<collectingCountry id="F37A4FCCFFF20751FCB3FC50FBC8FC5B" box="[829,1034,973,999]" name="French Polynesia" pageId="4" pageNumber="27">
French
<taxonomicName id="4C6D74DFFFF20751FC1BFC50FBC8FC5B" box="[917,1034,973,999]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="polynesia">Polynesia</taxonomicName>
</collectingCountry>
, representing a considerable range extension of both the genus and the species from the “Coral Triangle” into the central-western Pacific Ocean.
</paragraph>
</subSubSection>
<subSubSection id="C3775CD7FFF20751FF48FB8AFBBAFA4E" pageId="4" pageNumber="27" type="discussion">
<paragraph id="8BD20F5CFFF20751FF48FB8AFD8FFAE0" blockId="4.[151,1437,152,1522]" pageId="4" pageNumber="27">
<emphasis id="B919D34EFFF20751FF48FB8AFEFAFB8D" bold="true" box="[198,312,1047,1073]" pageId="4" pageNumber="27">Remarks</emphasis>
. This remarkable dwarf alpheid differs from all other alpheids by having two sharp spine-like projections on each eye (dorsal projection of left eye is visible in
<figureCitation id="135613D9FFF20751FC45FBA0FBD3FBEB" box="[971,1041,1085,1111]" captionStart="FIGURE 1" captionStartId="2.[151,255,1858,1882]" captionTargetBox="[175,1410,477,1840]" captionTargetId="figure@2.[175,1412,477,1840]" captionTargetPageId="2" captionText="FIGURE 1. Acanthanas pusillus Anker, Poddoubtchenko &amp; Jeng, 2006, two specimens from Moorea, A C, male (CL 2.30 mm), FLMNH UF Arthropoda 16055; D E, ovigerous female (CL 2.20 mm), FLMNH UF Arthropoda 15613; A, dorsal view; B, lateral view; C, frontal region, dorsolateral view; D, lateral view; E frontal region and chelipeds, lateral view." httpUri="https://zenodo.org/record/194126/files/figure.png" pageId="4" pageNumber="27">Fig. 1</figureCitation>
C). This feature, along with the presence of small, unspecialised chelipeds in the two
<typeStatus id="54D6B1FEFFF20751FC88FBFFFCFBFBC0" box="[774,825,1122,1148]" pageId="4" pageNumber="27">type</typeStatus>
specimens (both females) were used by Anker
<emphasis id="B919D34EFFF20751FAECFBFEFA5EFBC0" box="[1378,1436,1123,1148]" italics="true" pageId="4" pageNumber="27">et al.</emphasis>
(2006) to separate
<taxonomicName id="4C6D74DFFFF20751FEFAFB15FE3DFB1D" box="[372,511,1160,1185]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B919D34EFFF20751FEFAFB15FE3DFB1D" box="[372,511,1160,1185]" italics="true" pageId="4" pageNumber="27">Acanthanas</emphasis>
</taxonomicName>
from the undoubtedly closest genus,
<taxonomicName id="4C6D74DFFFF20751FC35FB15FB7BFB1E" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[955,1209,1160,1186]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B919D34EFFF20751FC35FB15FBD9FB1D" box="[955,1051,1160,1185]" italics="true" pageId="4" pageNumber="27">Athanas</emphasis>
<bibRefCitation id="EFFC72ADFFF20751FBADFB15FB7BFB1E" author="Leach" box="[1059,1209,1160,1186]" pageId="4" pageNumber="41" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (ed.), The Edinburgh Encyclopaedia, 7, pp. 383 - 437. Sir Brewster, Edinburgh." type="journal article" year="1814">Leach, 1814</bibRefCitation>
</taxonomicName>
. The finding of the first adult male specimen of
<taxonomicName id="4C6D74DFFFF20751FE7FFB30FDB2FB7A" box="[497,624,1197,1222]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF20751FE7FFB30FDB2FB7A" box="[497,624,1197,1222]" italics="true" pageId="4" pageNumber="27">A. pusillus</emphasis>
</taxonomicName>
in Moorea (FLMNH 23090) is particularly interesting for the right cheliped (the left is missing) is similar to the chelipeds defining the
<taxonomicName id="4C6D74DFFFF20751FC58FB4EFA84FB50" authority="Leach 1814" authorityName="Leach" authorityYear="1814" box="[982,1350,1234,1260]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="nitescens">
<emphasis id="B919D34EFFF20751FC58FB4EFB72FB50" box="[982,1200,1235,1260]" italics="true" pageId="4" pageNumber="27">Athanas nitescens</emphasis>
<bibRefCitation id="EFFC72ADFFF20751FB36FB4FFA84FB50" author="Leach" box="[1208,1350,1234,1260]" pageId="4" pageNumber="41" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (ed.), The Edinburgh Encyclopaedia, 7, pp. 383 - 437. Sir Brewster, Edinburgh." type="journal article" year="1814">Leach 1814</bibRefCitation>
</taxonomicName>
group, which may further link
<taxonomicName id="4C6D74DFFFF20751FE25FB65FDF4FAAD" box="[427,566,1272,1297]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B919D34EFFF20751FE25FB65FDF4FAAD" box="[427,566,1272,1297]" italics="true" pageId="4" pageNumber="27">Acanthanas</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6D74DFFFF20751FDE0FB65FD0CFAAD" box="[622,718,1272,1297]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B919D34EFFF20751FDE0FB65FD0CFAAD" box="[622,718,1272,1297]" italics="true" pageId="4" pageNumber="27">Athanas</emphasis>
</taxonomicName>
. The latter genus is a fairly heterogeneous group in need of a revision (
<bibRefCitation id="EFFC72ADFFF20751FE84FA80FDC3FA8B" author="Anker" box="[266,513,1309,1335]" pageId="4" pageNumber="40" refString="Anker, A. &amp; Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinition of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker &amp; Jeng 2007</bibRefCitation>
), especially because based on this new evidence,
<taxonomicName id="4C6D74DFFFF20751FBECFA80FB33FA8A" box="[1122,1265,1309,1334]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B919D34EFFF20751FBECFA80FB33FA8A" box="[1122,1265,1309,1334]" italics="true" pageId="4" pageNumber="27">Acanthanas</emphasis>
</taxonomicName>
appears to be embedded within
<emphasis id="B919D34EFFF20751FEE7FADEFD84FAE0" box="[361,582,1347,1372]" italics="true" pageId="4" pageNumber="27">
<taxonomicName id="4C6D74DFFFF20751FEE7FADEFDD3FAE0" box="[361,529,1347,1372]" class="Malacostraca" family="Alpheidae" genus="Athanas" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="sensu">
Athanas
<taxonomicNameLabel id="A22A6E35FFF20751FE5EFADEFDD3FAE0" box="[464,529,1347,1372]" pageId="4" pageNumber="27" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
.
</paragraph>
<paragraph id="8BD20F5CFFF20751FF4AFAF5FBBAFA4E" blockId="4.[151,1437,152,1522]" pageId="4" pageNumber="27">
The very small size of
<taxonomicName id="4C6D74DFFFF20751FE40FAF5FD8BFA3D" box="[462,585,1384,1409]" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B919D34EFFF20751FE40FAF5FD8BFA3D" box="[462,585,1384,1409]" italics="true" pageId="4" pageNumber="27">A. pusillus</emphasis>
</taxonomicName>
(CL
<quantity id="4C95A2B9FFF20751FD08FAF5FCD4FA3E" box="[646,790,1384,1410]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="2.3" metricValueMin="1.3" pageId="4" pageNumber="27" unit="mm" value="1.7999999999999998" valueMax="2.3" valueMin="1.3">1.32.3 mm</quantity>
, TL approximately 5.08.0 mm) and highly cryptic life style makes it impossible to spot underwater; the only way to collect it is to break and brush off large pieces of dead corals collected on outer reef slopes or in deeper rubble beds. It appears to be fairly common in rubble beds off Moorea, with multiple specimens collected over a period of several months.
</paragraph>
</subSubSection>
</treatment>
</document>