219 lines
26 KiB
XML
219 lines
26 KiB
XML
<document id="A415B299AB2E01A195B7DCC51DDE8997" ID-DOI="10.1111/j.1096-3642.2008.00400.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5446145" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630460512469" checkinUser="felipe" docAuthor="Savage, Jay M. & Watling, James I." docDate="2008" docId="03C487E43572C156FC66FAD4FC85FC56" docLanguage="en" docName="j.1096-3642.2008.00400.x.pdf" docOrigin="Zoological Journal of the Linnean Society 153 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00400.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Geophis bellus MYERS 2003" docType="treatment" docVersion="3" lastPageNumber="587" masterDocId="FFFDFF9C356BC14CFFDCFF96FF09FF96" masterDocTitle="Not so rare snakes: a revision of the Geophis sieboldi group (Colubridae: Dipsadinae) in lower Central America and Colombia" masterLastPageNumber="599" masterPageNumber="561" pageNumber="586" updateTime="1699279013921" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="A1A09A3452EACC9EB2BBEEA2FC2209C8">Not so rare snakes: a revision of the Geophis sieboldi group (Colubridae: Dipsadinae) in lower Central America and Colombia</mods:title>
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<mods:namePart id="54C1092846B310A289B6D3745552C4B3">Savage, Jay M.</mods:namePart>
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<mods:namePart id="656D542166DB553AFCF5282C60C12BE5">Watling, James I.</mods:namePart>
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<mods:typeOfResource id="508A804838600388ACE8BA198E97EDB7">text</mods:typeOfResource>
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<mods:title id="776108CEFC6E4D7B036D630B08B17C8F">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="548FD45F318B278EC6B711818916FA50">
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<mods:date id="2794DBBCF7510CF32D81D724D6EDA4B9">2008</mods:date>
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<mods:number id="DE6D6F73EEC39B39F436DEC37067B2DC">2008-07-31</mods:number>
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<mods:detail id="B22CABD4C80C114BCF7EAAD47741E107" type="volume">
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<mods:number id="67CCFB07C38D46F12853F378A0507EA9">153</mods:number>
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<mods:url id="77869EC37A61D5B3115EB21708F83462">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00400.x</mods:url>
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<mods:identifier id="5D3E3206180C9E09D139B81B4A402234" type="DOI">10.1111/j.1096-3642.2008.00400.x</mods:identifier>
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<mods:identifier id="CB1906CF1A9E9CD8DB593B1783EBD756" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="7FE15D0992156A1AB08A43BE961C3CA9" type="Zenodo-Dep">5446145</mods:identifier>
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<treatment id="03C487E43572C156FC66FAD4FC85FC56" LSID="urn:lsid:plazi:treatment:03C487E43572C156FC66FAD4FC85FC56" httpUri="http://treatment.plazi.org/id/03C487E43572C156FC66FAD4FC85FC56" lastPageId="26" lastPageNumber="587" pageId="25" pageNumber="586">
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<subSubSection id="C37765793572C155FC66FAD4FA28FAC1" box="[954,1313,1346,1367]" pageId="25" pageNumber="586" type="nomenclature">
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<paragraph id="8BD236F23572C155FC66FAD4FA28FAC1" blockId="25.[954,1313,1346,1367]" box="[954,1313,1346,1367]" pageId="25" pageNumber="586">
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<heading id="D09A819E3572C155FC66FAD4FA28FAC1" box="[954,1313,1346,1367]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="586" reason="2">
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<taxonomicName id="4C6D4D713572C155FC66FAD4FA28FAC1" authority="MYERS, 2003" authorityName="MYERS" authorityYear="2003" box="[954,1313,1346,1367]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="586" phylum="Chordata" rank="species" species="bellus">
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<emphasis id="B919EAE03572C155FC66FAD4FB77FAC0" box="[954,1150,1346,1367]" italics="true" pageId="25" pageNumber="586">GEOPHIS BELLUS</emphasis>
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MYERS, 2003
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37765793572C156FCE6FAFEFE30FEC0" lastPageId="26" lastPageNumber="587" pageId="25" pageNumber="586" type="reference_group">
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<paragraph id="8BD236F23572C155FCE6FAFEFCCEFA4C" blockId="25.[826,1441,1384,1498]" pageId="25" pageNumber="586">
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<taxonomicName id="4C6D4D713572C155FCE6FAFEFBCBFAE8" authority="Myers, 2003: 30" authorityName="MYERS" authorityPageNumber="30" authorityYear="2003" box="[826,1218,1384,1406]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="586" phylum="Chordata" rank="species" species="bellus">
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<emphasis id="B919EAE03572C155FCE6FAFEFCE1FAEB" box="[826,1000,1384,1405]" italics="true" pageId="25" pageNumber="586">Geophis bellus</emphasis>
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<materialsCitation id="3B053CAF3572C155FC24FAFFFBCBFAE8" accessionNumber="KU 110703" box="[1016,1218,1385,1406]" collectionCode="KU" collectorName="de Pacora" country="Panama" elevation="700" location="Cerro Jefe" municipality="Altos de Pacora" pageId="25" pageNumber="586" specimenCount="1" typeStatus="holotype">Myers, 2003: 30</materialsCitation>
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</taxonomicName>
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, figs 11–14 (Type locality:
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<collectingCountry id="F37A76623572C155FC40FA11FCF7FA0A" box="[924,1022,1415,1436]" name="Panama" pageId="25" pageNumber="586">Panama</collectingCountry>
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:
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<collectingCountry id="F37A76623572C155FBD4FA11FB63FA0A" box="[1032,1130,1415,1436]" name="Panama" pageId="25" pageNumber="586">Panama</collectingCountry>
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: Pacora: east of
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<location id="8EB260293572C155FAF8FA11FA97FA0B" LSID="urn:lsid:plazi:treatment:03C487E43572C156FC66FAD4FC85FC56:8EB260293572C155FAF8FA11FA97FA0B" box="[1316,1438,1415,1437]" country="Panama" municipality="Altos de Pacora" name="Cerro Jefe" pageId="25" pageNumber="586">Cerro Jefe</location>
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: near
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<collectingMunicipality id="6BB6AC883572C155FCAFFA30FB2EFA2D" box="[883,1063,1446,1468]" pageId="25" pageNumber="586">
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Altos
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<collectorName id="269853243572C155FC69FA30FB2EFA2D" box="[949,1063,1446,1468]" pageId="25" pageNumber="586">de Pacora</collectorName>
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</collectingMunicipality>
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,
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<quantity id="4C959B173572C155FBEDFA30FB70FA2D" box="[1073,1145,1446,1467]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="586" unit="m" value="700.0">
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<elevation id="0040D1C13572C155FBEDFA30FB70FA2D" box="[1073,1145,1446,1467]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="586" unit="m" value="700.0">700 m</elevation>
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</quantity>
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;
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<typeStatus id="54D688503572C155FB59FA30FBE6FA2A" box="[1157,1263,1446,1468]" pageId="25" pageNumber="586" type="holotype">Holotype</typeStatus>
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: KU 110703, an adult male).
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</paragraph>
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<paragraph id="8BD236F23572C156FCE6F99FFE30FEC0" blockId="25.[826,1441,1544,1903]" lastBlockId="26.[144,759,198,342]" lastPageId="26" lastPageNumber="587" pageId="25" pageNumber="586">
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<emphasis id="B919EAE03572C155FCE6F99FFCCCF988" box="[826,965,1545,1566]" italics="true" pageId="25" pageNumber="586">Diagnostics:</emphasis>
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The single known male specimen is distinguished from other members of the
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<taxonomicName id="4C6D4D713572C155FACFF9B1FC7AF9CC" baseAuthorityName="Jan" baseAuthorityYear="1862" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="586" phylum="Chordata" rank="species" species="sieboldi">
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<emphasis id="B919EAE03572C155FACFF9B1FC7AF9CC" italics="true" pageId="25" pageNumber="586">Geophis sieboldi</emphasis>
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</taxonomicName>
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species group by the following combination of characters (see
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<figureCitation id="13562A773572C155FC2EF9F2FB4FF9EC" box="[1010,1094,1636,1658]" captionStart="Figure 1" captionStartId="2.[144,223,956,975]" captionTargetBox="[370,1197,200,893]" captionTargetId="figure-351@2.[367,1199,198,897]" captionTargetPageId="2" captionText="Figure 1. Lateral head scalation in Geophis: A, pattern typical of G. zeledoni; B, pattern typical of G. brachycephalus complex; C, variant pattern; D, pattern typical of G. hoffmanni. PT, posterior temporal; U, upper postsupralabial; L, lower postsupralabial; PI, postinfralabial; number of supralabials and infralabials indicated by Arabic numerals; number of jubals indicated by Roman numerals." figureDoi="http://doi.org/10.5281/zenodo.5446147" httpUri="https://zenodo.org/record/5446147/files/figure.png" pageId="25" pageNumber="586">Fig. 1B</figureCitation>
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): (1) six supralabials; (2) two supralabials posterior to the orbit; (3) no anterior temporal or postlabials; (4) last supralabial separated from the parietal by an elongate posterior temporal; (5) two jubals posterior to posterior temporal and last supralabial, upper separated from contact with the parietal by the elongate posterior temporal; (6) dorsal scales in 15–15–15 rows; (7) dorsal scales on posterior 50% of body keeled; (8) anterior tip of maxilla pointed and toothless; posterior end depressed, curving slightly ventrad posteriorly; (9) ventrals 131; subcaudals 33; ventrals plus subcaudals 164; (10) upper surfaces glossy black with a white head band, venter uniform black.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37765793571C156FF4CFEE9FD41FBCE" pageId="26" pageNumber="587" type="description">
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||
<paragraph id="8BD236F23571C156FF4CFEE9FDCDFD51" blockId="26.[144,759,383,711]" pageId="26" pageNumber="587">
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<emphasis id="B919EAE03571C156FF4CFEE9FE35FE02" box="[144,316,383,404]" italics="true" pageId="26" pageNumber="587">Other features:</emphasis>
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The
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<typeStatus id="54D688503571C156FEA2FEE9FED6FE03" box="[382,479,383,405]" pageId="26" pageNumber="587" type="holotype">holotype</typeStatus>
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||
is
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<quantity id="4C959B173571C156FDDBFEE9FD6AFE02" box="[519,611,383,405]" metricMagnitude="-1" metricUnit="m" metricValue="1.69" pageId="26" pageNumber="587" unit="mm" value="169.0">169 mm</quantity>
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in standard length with a tail length of
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<quantity id="4C959B173571C156FE30FE08FD33FE25" box="[492,570,414,435]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="26" pageNumber="587" unit="mm" value="32.0">32 mm</quantity>
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||
(15.9% of total length); total length
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||
<quantity id="4C959B173571C156FE4CFE2AFEE6FE44" box="[400,495,444,466]" metricMagnitude="-1" metricUnit="m" metricValue="2.01" pageId="26" pageNumber="587" unit="mm" value="201.0">201 mm</quantity>
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. Preoculars 0–0. Postoculars 2–2. In the
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<typeStatus id="54D688503571C156FEA3FE4DFEEBFE67" box="[383,482,475,497]" pageId="26" pageNumber="587" type="holotype">holotype</typeStatus>
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, supralabial five on the right side of the head is split with the upper portion (anterior temporal of Myers, 2003) bordering the parietal and posterior temporal and the latter is shorter than its equivalent on the left side. Supralabials 6–6; supralabial 4 bordering the orbit on the right side of the head with 3–4 bordering the orbit on the left side. Infralabials 7–7, 5–5 bordering the chin shields.
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</paragraph>
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<paragraph id="8BD236F23571C156FF4CFD66FFDFFC28" blockId="26.[144,759,752,958]" pageId="26" pageNumber="587">
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<emphasis id="B919EAE03571C156FF4CFD66FE1BFC93" box="[144,274,752,773]" italics="true" pageId="26" pageNumber="587">Hemipenes:</emphasis>
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Based on an everted hemipenis (KU 110703). Organ bilobed. Sulcus spermaticus apparently centrifugal. Pedicel covered with spinules and small spines and two larger spines distally. Capitulum about twice as long as the truncus on asulcate side and covered mostly with small spines (Myers, 2003).
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</paragraph>
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<paragraph id="8BD236F23571C156FF4CFC70FD41FBCE" blockId="26.[144,759,998,1112]" pageId="26" pageNumber="587">
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<emphasis id="B919EAE03571C156FF4CFC70FE28FC6D" box="[144,289,998,1019]" italics="true" pageId="26" pageNumber="587">Distribution:</emphasis>
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Known from a tropical premontane wet forest site east of the
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<collectingCountry id="F37A76623571C156FE45FB93FEFEFB8C" box="[409,503,1029,1050]" name="Panama" pageId="26" pageNumber="587">Panama</collectingCountry>
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Canal and a tropical premontane rainforest site west of the Canal in central
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<collectingCountry id="F37A76623571C156FF36FBD4FE42FBC1" box="[234,331,1090,1111]" name="Panama" pageId="26" pageNumber="587">Panama</collectingCountry>
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,
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<quantity id="4C959B173571C156FE85FBD4FED1FBC1" box="[345,472,1090,1112]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="26" pageNumber="587" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600–800 m</quantity>
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(
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<figureCitation id="13562A773571C156FE36FBD4FD32FBCE" box="[490,571,1090,1112]" captionStart="Figure 16" captionStartId="24.[144,223,1108,1127]" captionTargetBox="[402,1168,195,1076]" captionTargetId="figure-333@24.[399,1168,195,1078]" captionTargetPageId="24" captionText="Figure 16. Geographical distribution of four species of Geophis in Panama and western Colombia. G. betanensis and G. nigroalbus are sympatric in Colombia." figureDoi="http://doi.org/10.5281/zenodo.5446179" httpUri="https://zenodo.org/record/5446179/files/figure.png" pageId="26" pageNumber="587">Fig. 16</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37765793571C156FF40FB18FB67FEC0" pageId="26" pageNumber="587" type="reference_group">
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<paragraph id="8BD236F23571C156FF40FB18FDE3FB35" blockId="26.[156,746,1166,1220]" box="[156,746,1166,1188]" pageId="26" pageNumber="587">
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<taxonomicName id="4C6D4D713571C156FF40FB18FDE3FB35" authority="RESTREPO & WRIGHT, 1987" authorityName="Restrepo & Wright" authorityYear="1987" box="[156,746,1166,1188]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="587" phylum="Chordata" rank="species" species="betaniensis">
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<emphasis id="B919EAE03571C156FF40FB18FE94FB34" box="[156,413,1166,1187]" italics="true" pageId="26" pageNumber="587">GEOPHIS BETANIENSIS</emphasis>
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||
<bibRefCitation id="EFFC4B033571C156FE7BFB19FDE3FB35" author="Restrepo JH & Wright JW" box="[423,746,1166,1188]" pageId="26" pageNumber="587" pagination="191 - 196" refId="ref27430" refString="Restrepo JH, Wright JW. 1987. A new species of colubrid snake genus Geophis from Colombia. Journal of Herpetology 21: 191 - 196." type="journal article" year="1987">RESTREPO & WRIGHT, 1987</bibRefCitation>
|
||
</taxonomicName>
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||
</paragraph>
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||
<paragraph id="8BD236F23571C156FE53FB38FEFFFB52" blockId="26.[156,746,1166,1220]" box="[399,502,1198,1220]" pageId="26" pageNumber="587">
|
||
(
|
||
<figureCitation id="13562A773571C156FE4BFB38FEE6FB52" box="[407,495,1198,1220]" captionStart="Figure 4" captionStartId="5.[164,243,1472,1491]" captionTargetBox="[169,763,910,1410]" captionTargetId="figure-377@5.[167,776,910,1412]" captionTargetPageId="5" captionText="Figure 4. Dorsal patterns in Geophis: A, dorsal pattern of G. betaniensis; B, dark transverse bands typical of Geophis dunni." figureDoi="http://doi.org/10.5281/zenodo.5446153" httpUri="https://zenodo.org/record/5446153/files/figure.png" pageId="26" pageNumber="587">FIG. 4A</figureCitation>
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||
)
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||
</paragraph>
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||
<paragraph id="8BD236F23571C156FF4CFB43FEDBFA12" blockId="26.[144,759,1237,1412]" pageId="26" pageNumber="587">
|
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<treatmentCitationGroup id="AB7D11DC3571C156FF4CFB43FEDBFA12" pageId="26" pageNumber="587">
|
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<treatmentCitation id="0ACC10E33571C156FF4CFB43FDFBFB7C" author="Restrepo JH & Wright JW" box="[144,754,1237,1259]" page="191" pageId="26" pageNumber="587" year="1987">
|
||
<taxonomicName id="4C6D4D713571C156FF4CFB43FDFBFB7C" authority="Restrepo & Wright, 1987: 191" authorityName="Restrepo & Wright" authorityPageNumber="191" authorityYear="1987" box="[144,754,1237,1259]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="587" phylum="Chordata" rank="species" species="betaniensis">
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||
<emphasis id="B919EAE03571C156FF4CFB43FE7EFB7C" box="[144,375,1237,1258]" italics="true" pageId="26" pageNumber="587">Geophis betaniensis</emphasis>
|
||
<bibRefCitation id="EFFC4B033571C156FE58FB43FDFBFB7C" author="Restrepo JH & Wright JW" box="[388,754,1237,1259]" pageId="26" pageNumber="587" pagination="191 - 196" refId="ref27430" refString="Restrepo JH, Wright JW. 1987. A new species of colubrid snake genus Geophis from Colombia. Journal of Herpetology 21: 191 - 196." type="journal article" year="1987">Restrepo & Wright, 1987: 191</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
, figs 1–3 (Type locality:
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||
<collectingCountry id="F37A76623571C156FE46FB65FD00FA9F" box="[410,521,1267,1289]" name="Colombia" pageId="26" pageNumber="587">Colombia</collectingCountry>
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||
: eastern slope of the Cordillera Occidental:
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||
<collectingRegion id="49A9F8103571C156FE44FA84FD5AFAB1" box="[408,595,1298,1320]" country="Colombia" name="Valle del Cauca" pageId="26" pageNumber="587">Valle del Cauca</collectingRegion>
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||
: Bolivar: Corregimiento de Betania,
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<geoCoordinate id="EE5950353571C156FE63FAA7FD07FAD0" box="[447,526,1327,1350]" degrees="4" direction="north" minutes="23" orientation="latitude" pageId="26" pageNumber="587" precision="925" value="4.383333">4°23′N</geoCoordinate>
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,
|
||
<geoCoordinate id="EE5950353571C156FDFAFAA6FD8FFAD0" box="[550,646,1327,1350]" degrees="76" direction="west" minutes="21" orientation="longitude" pageId="26" pageNumber="587" precision="925" value="-76.35">76°21′W</geoCoordinate>
|
||
,
|
||
<quantity id="4C959B173571C156FD40FAA6FDFBFAD0" box="[668,754,1328,1350]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="26" pageNumber="587" unit="m" value="1680.0">1680 m</quantity>
|
||
;
|
||
<typeStatus id="54D688503571C156FF4CFAD9FFF3FAF3" box="[144,250,1359,1381]" pageId="26" pageNumber="587" type="holotype">Holotype</typeStatus>
|
||
: UVC 7360, an adult female;
|
||
<treatmentCitation id="0ACC10E33571C156FD96FAD9FE0AFA15" author="Lips KR & Savage JM" page="414" pageId="26" pageNumber="587" year="1994">
|
||
<bibRefCitation id="EFFC4B033571C156FD96FAD9FE0AFA15" author="Lips KR & Savage JM" pageId="26" pageNumber="587" pagination="410 - 416" refId="ref27169" refString="Lips KR, Savage JM. 1994. A new fossorial snake of the genus Geophis (Reptilia: Serpentes: Colubridae) from the Cordillera de Talamanca of Costa Rica. Proceedings of the Biological Society of Washington 107: 410 - 416." type="journal article" year="1994">Lips & Savage, 1994: 414</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Myers, 2003: 36.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD236F23571C156FF4CFA3BFB67FEC0" blockId="26.[144,759,1452,1903]" lastBlockId="26.[806,1421,198,342]" pageId="26" pageNumber="587">
|
||
<emphasis id="B919EAE03571C156FF4CFA3BFE13FA54" box="[144,282,1453,1474]" italics="true" pageId="26" pageNumber="587">Diagnostics:</emphasis>
|
||
A member of the
|
||
<taxonomicName id="4C6D4D713571C156FE39FA3AFD97FA57" baseAuthorityName="Jan" baseAuthorityYear="1862" box="[485,670,1452,1473]" class="Reptilia" family="Dipsadidae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="587" phylum="Chordata" rank="species" species="sieboldi">
|
||
<emphasis id="B919EAE03571C156FE39FA3AFD97FA57" box="[485,670,1452,1473]" italics="true" pageId="26" pageNumber="587">Geophis sieboldi</emphasis>
|
||
</taxonomicName>
|
||
species group characterized by the following combination of features (see
|
||
<figureCitation id="13562A773571C156FEE9FA7CFE80F996" box="[309,393,1514,1536]" captionStart="Figure 1" captionStartId="2.[144,223,956,975]" captionTargetBox="[370,1197,200,893]" captionTargetId="figure-351@2.[367,1199,198,897]" captionTargetPageId="2" captionText="Figure 1. Lateral head scalation in Geophis: A, pattern typical of G. zeledoni; B, pattern typical of G. brachycephalus complex; C, variant pattern; D, pattern typical of G. hoffmanni. PT, posterior temporal; U, upper postsupralabial; L, lower postsupralabial; PI, postinfralabial; number of supralabials and infralabials indicated by Arabic numerals; number of jubals indicated by Roman numerals." figureDoi="http://doi.org/10.5281/zenodo.5446147" httpUri="https://zenodo.org/record/5446147/files/figure.png" pageId="26" pageNumber="587">Fig. 1B</figureCitation>
|
||
): (1) six supralabials; (2) two supralabials posterior to orbit; (3) no anterior temporal or postlabials; (4) last supralabial separated from contact with the parietal by a short posterior temporal; (5) three jubal scales posterior to the posterior temporal and last supralabial with the upper jubal in contact with the parietal; (6) dorsal scales in 15–15–15 scale rows; (7) dorsal and caudal scales smooth; (8) anterior tip of maxilla with first tooth at tip, posterior end of maxilla not depressed, not curving ventrad; (9) 140–143 ventrals in female; 20–28 subcaudals in females; ventrals plus subcaudals 168–173; (10) upper surfaces of body and tail reddish-brown with posterior margins of each scale black; black longitudinal ventrolateral stripes on tips of ventral from postoccipital region to tip of tail; venter yellow laterally, midventral area greenishyellow; subcloacal plate red.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37765793571C156FCFAFEE9FC85FC56" pageId="26" pageNumber="587" type="description">
|
||
<paragraph id="8BD236F23571C156FCFAFEE9FC76FDDB" blockId="26.[806,1422,383,742]" pageId="26" pageNumber="587">
|
||
<emphasis id="B919EAE03571C156FCFAFEE9FCD9FE02" box="[806,976,383,404]" italics="true" pageId="26" pageNumber="587">Other features:</emphasis>
|
||
The largest female is
|
||
<quantity id="4C959B173571C156FB0DFEE9FA24FE03" box="[1233,1325,383,405]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" pageId="26" pageNumber="587" unit="mm" value="260.0">260 mm</quantity>
|
||
in standard length with a tail length of
|
||
<quantity id="4C959B173571C156FB18FE08FA1BFE25" box="[1220,1298,414,436]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="26" pageNumber="587" unit="mm" value="36.0">36 mm</quantity>
|
||
(12.2% of total length); total length
|
||
<quantity id="4C959B173571C156FBBCFE2AFBB6FE44" box="[1120,1215,444,466]" metricMagnitude="-1" metricUnit="m" metricValue="2.96" pageId="26" pageNumber="587" unit="mm" value="296.0">296 mm</quantity>
|
||
. Tail length as a percentage of total length 12.2–12.3%. Preoculars 0–0. Postoculars 2–2. Supralabials 6–6, 3 and 4 bordering orbit. Infralabials 6–6, 4 in contact with genials.
|
||
</paragraph>
|
||
<paragraph id="8BD236F23571C156FCE3FDC0FB41FD73" blockId="26.[806,1422,383,742]" pageId="26" pageNumber="587">
|
||
<materialsCitation id="3B053CAF3571C156FCE3FDC0FB4DFD73" collectionCode="LACM" pageId="26" pageNumber="587" specimenCode="LACM 136189" specimenCount="3" typeStatus="holotype">
|
||
<bibRefCitation id="EFFC4B033571C156FCE3FDC0FB67FDFA" author="Restrepo JH & Wright JW" box="[831,1134,597,620]" pageId="26" pageNumber="587" pagination="191 - 196" refId="ref27430" refString="Restrepo JH, Wright JW. 1987. A new species of colubrid snake genus Geophis from Colombia. Journal of Herpetology 21: 191 - 196." type="journal article" year="1987">Restrepo & Wright (1987)</bibRefCitation>
|
||
state that the
|
||
<typeStatus id="54D688503571C156FAF9FDC3FA81FDFD" box="[1317,1416,597,619]" pageId="26" pageNumber="587" type="holotype">holotype</typeStatus>
|
||
, and presumably the
|
||
<typeStatus id="54D688503571C156FBFCFDE3FB9DFD1C" box="[1056,1172,629,650]" pageId="26" pageNumber="587" type="paratype">paratypes</typeStatus>
|
||
, had a pair of black stripes on scale rows 1 and 2 on each side in life. These stripes are not evident in the
|
||
<typeStatus id="54D688503571C156FB10FD24FA3BFD51" box="[1228,1330,690,711]" pageId="26" pageNumber="587" type="paratype">paratype</typeStatus>
|
||
(
|
||
<specimenCode id="DBCB9E893571C156FA9CFD27FC70FD70" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="26" pageNumber="587" type="Museum">LACM 136189</specimenCode>
|
||
) examined by us
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD236F23571C156FCFAFC86FB6BFCD5" blockId="26.[806,1421,783,835]" pageId="26" pageNumber="587">
|
||
<emphasis id="B919EAE03571C156FCFAFC86FCA1FCB3" box="[806,936,784,805]" italics="true" pageId="26" pageNumber="587">Hemipenes:</emphasis>
|
||
No information as both known specimens of this species are females.
|
||
</paragraph>
|
||
<paragraph id="8BD236F23571C156FCFAFCFBFC85FC56" blockId="26.[806,1421,877,960]" pageId="26" pageNumber="587">
|
||
<emphasis id="B919EAE03571C156FCFAFCFBFCB1FC14" box="[806,952,877,898]" italics="true" pageId="26" pageNumber="587">Distribution:</emphasis>
|
||
In the tropical premontane wet forest zone of the Cordillera Occidental of
|
||
<collectingCountry id="F37A76623571C156FB1CFC1DFA27FC37" box="[1216,1326,907,929]" name="Colombia" pageId="26" pageNumber="587">Colombia</collectingCountry>
|
||
,
|
||
<quantity id="4C959B173571C156FAE6FC1DFA84FC37" box="[1338,1421,907,929]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="26" pageNumber="587" unit="m" value="1680.0">1680 m</quantity>
|
||
(
|
||
<figureCitation id="13562A773571C156FCF2FC3CFC76FC56" box="[814,895,938,960]" captionStart="Figure 16" captionStartId="24.[144,223,1108,1127]" captionTargetBox="[402,1168,195,1076]" captionTargetId="figure-333@24.[399,1168,195,1078]" captionTargetPageId="24" captionText="Figure 16. Geographical distribution of four species of Geophis in Panama and western Colombia. G. betanensis and G. nigroalbus are sympatric in Colombia." figureDoi="http://doi.org/10.5281/zenodo.5446179" httpUri="https://zenodo.org/record/5446179/files/figure.png" pageId="26" pageNumber="587">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |