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<document id="B04C9F91BA28E761F0718AFEBDBD899E" ID-CLB-Dataset="54097" ID-DOI="10.11646/megataxa.4.1.1" ID-GBIF-Dataset="7c9bc39d-9910-46f1-9558-c0647b9cbbdb" ID-ISSN="2703-3090" ID-Zenodo-Dep="4591138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614033145335" checkinUser="plazi" docAuthor="Kim, Il-Hoi &amp; Boxshall, Geoff A." docDate="2020" docId="03C487CBEE503B34FF4DFD46FC24F7D7" docLanguage="en" docName="megataxa.4.1.1.pdf.imf" docOrigin="Megataxa 4 (1)" docSource="http://zoobank.org/3fdd970e-62f1-4f67-8cce-10870bdb3c01" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.5:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="5" docTitle="Bonnierilla eurypodata Kim &amp; Boxshall 2020, sp. nov." docType="treatment" docVersion="10" lastPageNumber="258" masterDocId="FFFDFFB3EF513A35FFC5FFCEFFC5FFC1" masterDocTitle="Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)" masterLastPageNumber="6" masterPageNumber="1" pageNumber="258" updateTime="1698930853666" updateUser="ExternalLinkService">
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<mods:title id="05A2A7B35495D2C3DFF6ECFB4843471A">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
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<mods:namePart id="AE88CFE14C5AB573E9A0BDA33C04BF41">Kim, Il-Hoi</mods:namePart>
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<mods:affiliation id="88F3B3B36C7718DA738015B40C2C3237">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<mods:namePart id="EFB50F920C4F5AB87833B44AEB091A41">Boxshall, Geoff A.</mods:namePart>
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<mods:affiliation id="753ADB61C777E25F9032ABBCEB5E375B">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<treatment id="03C487CBEE503B34FF4DFD46FC24F7D7" ID-DOI="http://doi.org/10.5281/zenodo.5699811" ID-GBIF-Taxon="194475091" ID-Zenodo-Dep="5699811" LSID="urn:lsid:plazi:treatment:03C487CBEE503B34FF4DFD46FC24F7D7" httpUri="http://treatment.plazi.org/id/03C487CBEE503B34FF4DFD46FC24F7D7" lastPageNumber="258" pageId="257" pageNumber="258">
<subSubSection id="C3776556EE503B34FF4DFD46FDC3FD62" box="[136,518,648,675]" pageId="257" pageNumber="258" type="nomenclature">
<paragraph id="8BD236DDEE503B34FF4DFD46FDC3FD62" blockId="257.[136,518,648,709]" box="[136,518,648,675]" pageId="257" pageNumber="258">
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<emphasis id="B919EACFEE503B34FF4DFD46FDC3FD62" bold="true" box="[136,518,648,675]" pageId="257" pageNumber="258">
<taxonomicName id="4C6D4D5EEE503B34FF4DFD46FE64FD62" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[136,417,648,675]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="eurypodata" status="sp. nov.">
<emphasis id="B919EACFEE503B34FF4DFD46FE64FD62" bold="true" box="[136,417,648,675]" italics="true" pageId="257" pageNumber="258">Bonnierilla eurypodata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B4EE503B34FE6CFD47FDC3FD62" box="[425,518,649,675]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3776556EE503B34FF4DFD65FEFEFD04" box="[136,315,683,709]" pageId="257" pageNumber="258" type="description">
<paragraph id="8BD236DDEE503B34FF4DFD65FEFEFD04" blockId="257.[136,518,648,709]" box="[136,315,683,709]" pageId="257" pageNumber="258">
(
<figureCitation id="13562A58EE503B34FF55FD65FF39FD04" box="[144,252,683,709]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Figs. 172</figureCitation>
,
<figureCitation id="13562A58EE503B34FECDFD65FEF6FD04" box="[264,307,683,709]" captionStart="FIGURE 173" captionStartId="259.[136,245,1907,1933]" captionTargetBox="[241,1357,219,1840]" captionTargetId="figure-18@259.[196,1391,185,1870]" captionTargetPageId="259" captionText="FIGURE 173. Bonnierilla eurypodata sp. nov., female. A, paragnath; B, maxilliped; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591613" httpUri="https://zenodo.org/record/4591613/files/figure.png" pageId="257" pageNumber="258">173</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3776556EE503B34FF4DFD21FE53FC72" pageId="257" pageNumber="258" type="materials_examined">
<paragraph id="8BD236DDEE503B34FF4DFD21FE53FC72" blockId="257.[136,778,751,2070]" pageId="257" pageNumber="258">
<emphasis id="B919EACFEE503B34FF4DFD21FEF8FCC8" bold="true" box="[136,317,751,777]" pageId="257" pageNumber="258">Typematerial.</emphasis>
<materialsCitation id="3B053C80EE503B34FE89FD21FF1DFCEA" ID-GBIF-Occurrence="3748446342" pageId="257" pageNumber="258" specimenCode="MNHN-IU-2014- 21284" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54D6887FEE503B34FE89FD21FE71FCC8" box="[332,436,751,777]" pageId="257" pageNumber="258" type="holotype">Holotype</typeStatus>
(intact
<specimenCount id="9D6BFD54EE503B34FDDDFD3EFDE8FCCB" box="[536,557,752,778]" pageId="257" pageNumber="258" type="female"></specimenCount>
,
<specimenCode id="DBCB9EA6EE503B34FD86FD21FF15FCEA" pageId="257" pageNumber="258">MNHN-IU-2014- 21284</specimenCode>
)
</materialsCitation>
<typeStatus id="54D6887FEE503B34FF2AFCDFFDC8FCED" box="[239,525,785,812]" pageId="257" pageNumber="258">anddissectedparatype</typeStatus>
(
<specimenCount id="9D6BFD54EE503B34FDF5FCDFFD86FCEA" box="[560,579,785,811]" pageId="257" pageNumber="258"></specimenCount>
, figured) from
<taxonomicName id="4C6D4D5EEE503B34FF4DFCFDFDD1FC8F" authority="(Sluiter, 1886)" baseAuthorityName="Sluiter" baseAuthorityYear="1886" box="[136,532,819,846]" class="Ascidiacea" family="Styelidae" genus="Polycarpa" kingdom="Animalia" order="Pleurogona" pageId="257" pageNumber="258" phylum="Chordata" rank="species" species="papillata">
<emphasis id="B919EACFEE503B34FF4DFCFDFEACFC8C" box="[136,361,819,845]" italics="true" pageId="257" pageNumber="258">Polycarpa papillata</emphasis>
(Sluiter, 1886)
</taxonomicName>
, CORAIL 2, DW146,
<collectingCountry id="F37A764DEE503B34FF4DFC98FEF8FCAE" box="[136,317,853,880]" name="New Caledonia" pageId="257" pageNumber="258">New Caledonia</collectingCountry>
, Chesterfield Plateau (
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,
<geoCoordinate id="EE59501AEE503B34FF4DFCB9FEA7FC53" box="[136,354,887,914]" degrees="158" direction="east" minutes="16" orientation="longitude" pageId="257" pageNumber="258" precision="1" seconds="16.7952" value="158.27133">158°1616.7952E</geoCoordinate>
), depth
<quantity id="4C959B38EE503B34FE01FCB9FDC1FC53" box="[452,516,887,914]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="257" pageNumber="258" unit="m" value="44.0">44 m</quantity>
, B. Richer de Forges-IRD coll.,
<date id="FFD3101DEE503B34FF3BFC57FE57FC72" box="[254,402,921,947]" pageId="257" pageNumber="258" value="1988-07-30">30 July 1988</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EE503B34FF78FC75FBE5FA70" pageId="257" pageNumber="258" type="etymology">
<paragraph id="8BD236DDEE503B34FF78FC75FDAAFBDB" blockId="257.[136,778,751,2070]" pageId="257" pageNumber="258">
<emphasis id="B919EACFEE503B34FF78FC75FEFBFC14" bold="true" box="[189,318,955,981]" pageId="257" pageNumber="258">Etymology</emphasis>
. The specific name is derived from the Greek
<emphasis id="B919EACFEE503B34FF13FC10FECDFC36" box="[214,264,990,1015]" italics="true" pageId="257" pageNumber="258">eury</emphasis>
(=broad) and
<emphasis id="B919EACFEE503B34FE68FC13FE12FC36" box="[429,471,989,1015]" italics="true" pageId="257" pageNumber="258">pod</emphasis>
(=a foot) and refers to the relatively broad exopodal segment of leg 5.
</paragraph>
<paragraph id="8BD236DDEE503B34FC4FFC31FA5BFB60" pageId="257" pageNumber="258">
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<tr id="355D349FEE50C5CAFCEFFC31FA64FBD8" box="[810,1441,1023,1049]" gridrow="0" pageId="257" pageNumber="258" rowspan-0="1">
<th id="768C5DE3EE50C5CAFC4FFC31FC00FBD8" box="[906,965,1023,1049]" gridcol="1" gridrow="0" pageId="257" pageNumber="258">Coxa</th>
<th id="768C5DE3EE50C5CAFC1CFC31FBD2FBD8" box="[985,1047,1023,1049]" gridcol="2" gridrow="0" pageId="257" pageNumber="258">Basis</th>
<th id="768C5DE3EE50C5CAFBEEFC31FB1CFBD8" box="[1067,1241,1023,1049]" gridcol="3" gridrow="0" pageId="257" pageNumber="258">Exopod</th>
<th id="768C5DE3EE50C5CAFB2BFC31FA64FBD8" box="[1262,1441,1023,1049]" gridcol="4" gridrow="0" pageId="257" pageNumber="258">Endopod</th>
</tr>
<tr id="355D349FEE50C5CAFCEFFBEFFA64FBFD" box="[810,1441,1057,1084]" gridrow="1" pageId="257" pageNumber="258">
<th id="768C5DE3EE50C5CAFCEFFBEFFCAFFBFD" box="[810,874,1057,1084]" gridcol="0" gridrow="1" pageId="257" pageNumber="258">Leg 1</th>
<td id="768C5DE3EE50C5CAFC4FFBEFFC00FBFD" box="[906,965,1057,1084]" gridcol="1" gridrow="1" pageId="257" pageNumber="258">0-0</td>
<td id="768C5DE3EE50C5CAFC1CFBEFFBD2FBFD" box="[985,1047,1057,1084]" gridcol="2" gridrow="1" pageId="257" pageNumber="258">1-I</td>
<td id="768C5DE3EE50C5CAFBEEFBEFFB1CFBFD" box="[1067,1241,1057,1084]" gridcol="3" gridrow="1" pageId="257" pageNumber="258">I-1; I-1; III, I, 4</td>
<td id="768C5DE3EE50C5CAFB2BFBEFFA64FBFD" box="[1262,1441,1057,1084]" gridcol="4" gridrow="1" pageId="257" pageNumber="258">0-1; 0-1; 1, 2, 3</td>
</tr>
<tr id="355D349FEE50C5CAFCEFFB8DFA64FB9F" box="[810,1441,1091,1118]" gridrow="2" pageId="257" pageNumber="258">
<th id="768C5DE3EE50C5CAFCEFFB8DFCAFFB9F" box="[810,874,1091,1118]" gridcol="0" gridrow="2" pageId="257" pageNumber="258">Leg 2</th>
<td id="768C5DE3EE50C5CAFC4FFB8DFC00FB9F" box="[906,965,1091,1118]" gridcol="1" gridrow="2" pageId="257" pageNumber="258">0-0</td>
<td id="768C5DE3EE50C5CAFC1CFB8DFBD2FB9F" box="[985,1047,1091,1118]" gridcol="2" gridrow="2" pageId="257" pageNumber="258">1-0</td>
<td id="768C5DE3EE50C5CAFBEEFB8DFB1CFB9F" box="[1067,1241,1091,1118]" gridcol="3" gridrow="2" pageId="257" pageNumber="258">I-1; 0-1; I, I, 5</td>
<td id="768C5DE3EE50C5CAFB2BFB8DFA64FB9F" box="[1262,1441,1091,1118]" gridcol="4" gridrow="2" pageId="257" pageNumber="258">0-1; 0-2; 1, 2, 3</td>
</tr>
<tr id="355D349FEE50C5CAFCEFFBABFA64FB41" box="[810,1441,1125,1152]" gridrow="3" pageId="257" pageNumber="258">
<th id="768C5DE3EE50C5CAFCEFFBABFCAFFB41" box="[810,874,1125,1152]" gridcol="0" gridrow="3" pageId="257" pageNumber="258">Leg 3</th>
<td id="768C5DE3EE50C5CAFC4FFBABFC00FB41" box="[906,965,1125,1152]" gridcol="1" gridrow="3" pageId="257" pageNumber="258">0-1</td>
<td id="768C5DE3EE50C5CAFC1CFBABFBD2FB41" box="[985,1047,1125,1152]" gridcol="2" gridrow="3" pageId="257" pageNumber="258">1-0</td>
<td id="768C5DE3EE50C5CAFBEEFBABFB1CFB41" box="[1067,1241,1125,1152]" gridcol="3" gridrow="3" pageId="257" pageNumber="258">0-1; 0-1; I, I, 5</td>
<td id="768C5DE3EE50C5CAFB2BFBABFA64FB41" box="[1262,1441,1125,1152]" gridcol="4" gridrow="3" pageId="257" pageNumber="258">0-1; 0-2; 1, 2, 3</td>
</tr>
<tr id="355D349FEE50C5CAFCEFFB49FA64FB63" box="[810,1441,1159,1186]" gridrow="4" pageId="257" pageNumber="258">
<th id="768C5DE3EE50C5CAFCEFFB49FCAFFB63" box="[810,874,1159,1186]" gridcol="0" gridrow="4" pageId="257" pageNumber="258">Leg 4</th>
<td id="768C5DE3EE50C5CAFC4FFB49FC00FB63" box="[906,965,1159,1186]" gridcol="1" gridrow="4" pageId="257" pageNumber="258">0-1</td>
<td id="768C5DE3EE50C5CAFC1CFB49FBD2FB63" box="[985,1047,1159,1186]" gridcol="2" gridrow="4" pageId="257" pageNumber="258">1-0</td>
<td id="768C5DE3EE50C5CAFBEEFB49FB1CFB63" box="[1067,1241,1159,1186]" gridcol="3" gridrow="4" pageId="257" pageNumber="258">0-1; 0-1; I, I, 5</td>
<td id="768C5DE3EE50C5CAFB2BFB49FA64FB63" box="[1262,1441,1159,1186]" gridcol="4" gridrow="4" pageId="257" pageNumber="258">0-1; 0-2; 1, 2, 2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD236DDEE503B34FF78FBEFFF37FAAF" blockId="257.[136,778,751,2070]" pageId="257" pageNumber="258">
<emphasis id="B919EACFEE503B34FF78FBEFFE76FBFA" bold="true" box="[189,435,1057,1083]" pageId="257" pageNumber="258">Descriptionoffemale</emphasis>
.Body(
<figureCitation id="13562A58EE503B34FDC0FBEFFDB1FBFA" box="[517,628,1057,1083]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig.172A</figureCitation>
)compressed, inflated, 2.45 mm long. Cephalosome well-defined from metasome; metasomeunsegmented. Freeurosome (
<figureCitation id="13562A58EE503B34FD19FBABFF03FB60" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172B</figureCitation>
) stout, 5-segmented. Caudal rami widely separated from each other; each ramus (
<figureCitation id="13562A58EE503B34FE1FFB67FD88FB02" box="[474,589,1193,1219]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172C</figureCitation>
) fusiform, about 2.8 times longer than wide (110×40 μm) and about 1.3 times longer than anal somite, armed with 6 naked setae; 2 proximal setae as long as width of ramus, located at 40 and 64% of ramus length; distal 4 setae shorter than width of ramus.
</paragraph>
<paragraph id="8BD236DDEE503B34FF78FAB8FF28F8E9" blockId="257.[136,778,751,2070]" pageId="257" pageNumber="258">
Rostrum (
<figureCitation id="13562A58EE503B34FEFDFABBFE72FA51" box="[312,439,1397,1424]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172D</figureCitation>
) widerthan long, strongly tapering, withrounded apex. Antennule (
<figureCitation id="13562A58EE503B34FD42FA59FCC7FA73" box="[647,770,1431,1458]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172E</figureCitation>
) obscurely 8-segmented; armature formula 3, 15, 9+aesthetasc. 4, 3+aesthetasc, 1, 2+aesthetasc, and 7+aesthetasc; all setae naked; 1 seta on second segment rod-shaped. Antenna (
<figureCitation id="13562A58EE503B34FE53F9D1FDD4F9FB" box="[406,529,1567,1594]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172F</figureCitation>
) 4-segmented; coxa unarmed; basis with 2 minute vestigial setae distally; first endopodal segment with 1 small seta subdistally; compound distal endopodal segment 3.1 times longer than wide and about 1.5 times longer than first endopodal segment, armed with 8 setae (distal 3 blunt at tip) plus strongly curved terminal claw, less than half length of segment.
</paragraph>
<paragraph id="8BD236DDEE503B34FF78F8FEFBE3FE16" blockId="257.[136,778,751,2070]" lastBlockId="257.[810,1452,173,982]" pageId="257" pageNumber="258">
Labrum (
<figureCitation id="13562A58EE503B34FEF3F8FEFE7FF88B" box="[310,442,1840,1866]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172G</figureCitation>
) with patches of setules posterolaterally and spinules on posteromedian lobe. Mandible (
<figureCitation id="13562A58EE503B34FEC4F8BAFEB6F84F" box="[257,371,1908,1934]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172H</figureCitation>
) bearing 5 teeth on coxal gnathobase and 3 needle-like spinules between distal second and third teeth; palp armed as usual: 1 seta on basis, 5 on exopod, 4 on first endopodal segment and 9 on second. Paragnath (
<figureCitation id="13562A58EE503B34FF55F832FED7F7D7" box="[144,274,2044,2070]" captionStart="FIGURE 173" captionStartId="259.[136,245,1907,1933]" captionTargetBox="[241,1357,219,1840]" captionTargetId="figure-18@259.[196,1391,185,1870]" captionTargetPageId="259" captionText="FIGURE 173. Bonnierilla eurypodata sp. nov., female. A, paragnath; B, maxilliped; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591613" httpUri="https://zenodo.org/record/4591613/files/figure.png" pageId="257" pageNumber="258">Fig. 173A</figureCitation>
) with large dentiform process apically. Maxillule (
<figureCitation id="13562A58EE503B34FC62FF63FBD5FF06" box="[935,1040,173,199]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172I</figureCitation>
) not specialized, armedwith 9 setae on arthrite, 1 on coxal endite, 2 on coxal epipodite, 3 on basis, 4 each on exopod and endopod. Maxilla (
<figureCitation id="13562A58EE503B34FAFFFF3FFA66FECA" box="[1338,1443,241,267]" captionStart="FIGURE 172" captionStartId="258.[136,245,1907,1933]" captionTargetBox="[241,1345,214,1844]" captionTargetId="figure-19@258.[196,1392,185,1870]" captionTargetPageId="258" captionText="FIGURE 172. Bonnierilla eurypodata sp. nov., female. A, habitus, right; B, urosome, ventral; C, caudal ramus; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule; J, maxilla. Scale bars: A, 0.2 mm; B, 0.1 mm; C-J, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591611" httpUri="https://zenodo.org/record/4591611/files/figure.png" pageId="257" pageNumber="258">Fig. 172J</figureCitation>
) 5-segmented; syncoxawith 8 setae (3, 1, 2, and 2); basis with claw bearing setules on both margins, plus 1 seta and 1 rudimentaryseta; endopodwith 1, 1, and 3 setaeonfirst to third segments, respectively. Maxilliped (
<figureCitation id="13562A58EE503B34FAE8FEB7FA66FE52" box="[1325,1443,377,403]" captionStart="FIGURE 173" captionStartId="259.[136,245,1907,1933]" captionTargetBox="[241,1357,219,1840]" captionTargetId="figure-18@259.[196,1391,185,1870]" captionTargetPageId="259" captionText="FIGURE 173. Bonnierilla eurypodata sp. nov., female. A, paragnath; B, maxilliped; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591613" httpUri="https://zenodo.org/record/4591613/files/figure.png" pageId="257" pageNumber="258">Fig. 173B</figureCitation>
) incompletely 2-segmented with 10 setaeonfirst segment and 2 setaeon second.
</paragraph>
<paragraph id="8BD236DDEE503B34FC9AFE2EFC10FC14" blockId="257.[810,1452,173,982]" pageId="257" pageNumber="258">
Legs 14 (
<figureCitation id="13562A58EE503B34FC23FE11FB89FE38" box="[998,1100,479,505]" captionStart="FIGURE 173" captionStartId="259.[136,245,1907,1933]" captionTargetBox="[241,1357,219,1840]" captionTargetId="figure-18@259.[196,1391,185,1870]" captionTargetPageId="259" captionText="FIGURE 173. Bonnierilla eurypodata sp. nov., female. A, paragnath; B, maxilliped; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591613" httpUri="https://zenodo.org/record/4591613/files/figure.png" pageId="257" pageNumber="258">Fig. 173</figureCitation>
C-F) with 3-segmented rami. Inner coxal seta absent in legs 1 and 2, but present in legs 3 and 4. Outer seta on basis large and plumose in leg 1 but small and naked in legs 24. Inner distal spine on basis of leg 1 naked, 49 μm long, extending beyond distal border of first endopodal segment. Exopods of legs 24 each about 1.8 times longerthan endopods. First and second exopodal segments of legs 24 lacking outer spines or with rudimentary spines. Third exopodal segment of legs 24 bearing only 2 spines; with inner setae naked. Third exopodal segment of leg 2 with 2 spines and 5 setae (armature formula I, I, 5) in
<typeStatus id="54D6887FEE503B34FB43FC9BFB2EFCAE" box="[1158,1259,853,879]" pageId="257" pageNumber="258" type="holotype">holotype</typeStatus>
, but abnormally with 2 spines and 4 setae (armature formula I, I, 4;
<figureCitation id="13562A58EE503B34FABAFCB9FCACFC72" captionStart="FIGURE 173" captionStartId="259.[136,245,1907,1933]" captionTargetBox="[241,1357,219,1840]" captionTargetId="figure-18@259.[196,1391,185,1870]" captionTargetPageId="259" captionText="FIGURE 173. Bonnierilla eurypodata sp. nov., female. A, paragnath; B, maxilliped; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591613" httpUri="https://zenodo.org/record/4591613/files/figure.png" pageId="257" pageNumber="258">Fig. 173D</figureCitation>
) in dissected specimen. Armature formula for legs 14 as follows:
</paragraph>
<paragraph id="8BD236DDEE503B34FC9AFB02FB36FA4E" blockId="257.[810,1452,1227,2070]" pageId="257" pageNumber="258">
Leg 5 (
<figureCitation id="13562A58EE503B34FC79FB05FBF3FB24" box="[956,1078,1227,1253]" captionStart="FIGURE 173" captionStartId="259.[136,245,1907,1933]" captionTargetBox="[241,1357,219,1840]" captionTargetId="figure-18@259.[196,1391,185,1870]" captionTargetPageId="259" captionText="FIGURE 173. Bonnierilla eurypodata sp. nov., female. A, paragnath; B, maxilliped; C, leg 1; D, leg 2; E, leg 3; F, leg 4; G, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591613" httpUri="https://zenodo.org/record/4591613/files/figure.png" pageId="257" pageNumber="258">Fig. 173G</figureCitation>
) 2-segmented; coxa short with naked outer distal seta and row of minute spinules at inner distal region; exopodal segment stout, about 2.3 times longerthan wide (173×76 μm) and abruptly narrowing in distal quarter; armed with naked apical seta; 4 rows of minute spinules present on inner margin.
</paragraph>
<paragraph id="8BD236DDEE503B34FC9AFA56FBE5FA70" blockId="257.[810,1452,1227,2070]" box="[863,1056,1431,1458]" pageId="257" pageNumber="258">
<emphasis id="B919EACFEE503B34FC9AFA56FC59FA73" bold="true" box="[863,924,1432,1458]" pageId="257" pageNumber="258">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EE503B34FC9AFA74FC24F7D7" pageId="257" pageNumber="258" type="discussion">
<paragraph id="8BD236DDEE503B34FC9AFA74FC24F7D7" blockId="257.[810,1452,1227,2070]" pageId="257" pageNumber="258">
<emphasis id="B919EACFEE503B34FC9AFA74FC09FA15" bold="true" box="[863,972,1466,1492]" pageId="257" pageNumber="258">Remarks</emphasis>
.
<taxonomicName id="4C6D4D5EEE503B34FC21FA77FB30FA12" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[996,1269,1465,1491]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="eurypodata" status="sp. nov.">
<emphasis id="B919EACFEE503B34FC21FA77FB30FA12" box="[996,1269,1465,1491]" italics="true" pageId="257" pageNumber="258">Bonnierilla eurypodata</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FAC3FA74FAAFFA15" bold="true" box="[1286,1386,1466,1492]" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FAC3FA74FAAFFA15" box="[1286,1386,1466,1492]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a typical form of
<taxonomicName id="4C6D4D5EEE503B34FC25FA15FBA5FA34" authorityName="Canu" authorityYear="1891" box="[992,1120,1499,1525]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEE503B34FC25FA15FBA5FA34" box="[992,1120,1499,1525]" italics="true" pageId="257" pageNumber="258">Bonnierilla</emphasis>
</taxonomicName>
and is similar to
<taxonomicName id="4C6D4D5EEE503B34FAEBFA12FA69FA34" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1326,1452,1499,1526]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="iboensis" status="sp. nov.">
<emphasis id="B919EACFEE503B34FAEBFA12FAFAFA34" box="[1326,1343,1500,1525]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FA95FA15FA69FA34" box="[1360,1452,1499,1525]" italics="true" pageId="257" pageNumber="258">iboensis</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FCEFFA30FC46F9D9" bold="true" box="[810,899,1534,1560]" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FCEFFA30FC46F9D9" box="[810,899,1534,1560]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C6D4D5EEE503B34FC7DFA30FB8EF9D9" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[952,1099,1534,1560]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="reniformis" status="sp. nov.">
<emphasis id="B919EACFEE503B34FC7DFA30FC0CF9D6" box="[952,969,1534,1559]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FC12FA30FB8EF9D9" box="[983,1099,1534,1560]" italics="true" pageId="257" pageNumber="258">reniformis</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FB94FA30FB6FF9D9" bold="true" box="[1105,1194,1534,1560]" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FB94FA30FB6FF9D9" box="[1105,1194,1534,1560]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, both described above. The most obvious differences between
<taxonomicName id="4C6D4D5EEE503B34FB26F9EEFA46F9F8" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1251,1411,1567,1594]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="eurypodata" status="sp. nov.">
<emphasis id="B919EACFEE503B34FB26F9EEFB31F9F8" box="[1251,1268,1568,1593]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FAC6F9D1FA46F9F8" box="[1283,1411,1567,1593]" italics="true" pageId="257" pageNumber="258">eurypodata</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FA4FF9EEFC9EF99D" bold="true" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FA4FF9EEFC9EF99D" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and these other two species are in the dimensions of the caudal ramus and the exopod of leg
<quantity id="4C959B38EE503B34FB32F9ADFAE1F9BC" box="[1271,1316,1635,1662]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="257" pageNumber="258" unit="in" value="5.0">5 in</quantity>
the female. The caudal ramus of
<taxonomicName id="4C6D4D5EEE503B34FBDEF948FB79F95E" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1051,1212,1669,1696]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="eurypodata" status="sp. nov.">
<emphasis id="B919EACFEE503B34FBDEF948FBE9F95E" box="[1051,1068,1670,1695]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FBF9F94BFB79F95E" box="[1084,1212,1669,1695]" italics="true" pageId="257" pageNumber="258">eurypodata</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FB01F948FADAF961" bold="true" box="[1220,1311,1670,1696]" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FB01F948FADAF961" box="[1220,1311,1670,1696]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is about 2.8 times longer than wide (110×40 μm) in contrast to 4.6 timeslongerthan wide (206×45 μm) in
<taxonomicName id="4C6D4D5EEE503B34FAC4F904FA45F922" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1281,1408,1737,1764]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="iboensis" status="sp. nov.">
<emphasis id="B919EACFEE503B34FAC4F904FAD7F922" box="[1281,1298,1738,1763]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FAE1F907FA45F922" box="[1316,1408,1737,1763]" italics="true" pageId="257" pageNumber="258">iboensis</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FA4FF904FC9EF8C7" bold="true" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FA4FF904FC9EF8C7" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 3.4 timeslongerthan wide (211×63 μm) in
<taxonomicName id="4C6D4D5EEE503B34FA56F922FC5BF8E9" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="reniformis" status="sp. nov.">
<emphasis id="B919EACFEE503B34FA56F922FA61F8C4" box="[1427,1444,1772,1797]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FCEFF8C0FC5BF8E9" box="[810,926,1806,1832]" italics="true" pageId="257" pageNumber="258">reniformis</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FC61F8C0FC3BF8E9" bold="true" box="[932,1022,1806,1832]" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FC61F8C0FC3BF8E9" box="[932,1022,1806,1832]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The exopod of leg 5 of
<taxonomicName id="4C6D4D5EEE503B34FAC8F8C0FA69F8E6" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1293,1452,1805,1832]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="eurypodata" status="sp. nov.">
<emphasis id="B919EACFEE503B34FAC8F8C0FADBF8E6" box="[1293,1310,1806,1831]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FAE9F8C3FA69F8E6" box="[1324,1452,1805,1831]" italics="true" pageId="257" pageNumber="258">eurypodata</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FCEFF8FEFC41F88B" bold="true" box="[810,900,1840,1866]" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FCEFF8FEFC41F88B" box="[810,900,1840,1866]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is 2.3 times longer than wide (173×76 μm), with a single apical seta, in contrast to 2.5 times longer than wide (136×55 μm) in
<taxonomicName id="4C6D4D5EEE503B34FBE2F8BAFB61F84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1063,1188,1908,1934]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="iboensis" status="sp. nov.">
<emphasis id="B919EACFEE503B34FBE2F8BAFBFDF84C" box="[1063,1080,1908,1933]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FB8DF8BAFB61F84F" box="[1096,1188,1908,1934]" italics="true" pageId="257" pageNumber="258">iboensis</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FB68F8BAFACDF84F" bold="true" box="[1197,1288,1908,1934]" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FB68F8BAFACDF84F" box="[1197,1288,1908,1934]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 2.7 times longerthanwide (203×74 μm) in
<taxonomicName id="4C6D4D5EEE503B34FB68F858FA84F871" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1197,1345,1942,1968]" class="Maxillopoda" family="Notodelphyidae" genus="Bonnierilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="257" pageNumber="258" phylum="Arthropoda" rank="species" species="reniformis" status="sp. nov.">
<emphasis id="B919EACFEE503B34FB68F858FB7BF86E" box="[1197,1214,1942,1967]" italics="true" pageId="257" pageNumber="258">B</emphasis>
.
<emphasis id="B919EACFEE503B34FB08F858FA84F871" box="[1229,1345,1942,1968]" italics="true" pageId="257" pageNumber="258">reniformis</emphasis>
</taxonomicName>
<emphasis id="B919EACFEE503B34FA8CF858FA61F871" bold="true" box="[1353,1444,1942,1968]" pageId="257" pageNumber="258">
<taxonomicNameLabel id="A22A57B4EE503B34FA8CF858FA61F871" box="[1353,1444,1942,1968]" pageId="257" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, with 2 unequal distal setae in the latter two species. These differences are robust and support the establishment of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>