treatments-xml/data/03/C4/7D/03C47D7ADB1B5B7DCEC0FBA08E5FF830.xml
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<document id="B509A86FE7A21AC87EA50867D10CEE7A" ID-CLB-Dataset="32376" ID-DOI="10.11646/zootaxa.4294.3.2" ID-GBIF-Dataset="1ffe5af4-24f3-42bb-be14-c9df7e7f3191" ID-ISSN="1175-5326" ID-Zenodo-Dep="844788" ID-ZooBank="A8B7FFA1-4BA3-4619-AB68-BF1E25680516" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502974146766" checkinUser="plazi" docAuthor="Sumontha, Montri, Kunya, Kirati, Dangsri, Siriwat &amp; Pauwels, Olivier S. G." docDate="2017" docId="03C47D7ADB1B5B7DCEC0FBA08E5FF830" docLanguage="en" docName="zootaxa.4294.3.2.pdf.imf" docOrigin="Zootaxa 4294 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oligodon saiyok Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="319" masterDocId="FFFD0502DB1A5B7ECE57FFC58D28FFF6" masterDocTitle="Oligodon saiyok, a new limestone-dwelling kukri snake (Serpentes: Colubridae) from Kanchanaburi Province, western Thailand" masterLastPageNumber="328" masterPageNumber="316" pageNumber="317" updateTime="1698466799339" updateUser="ExternalLinkService">
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<mods:title id="52E33AFCA3639C9A0A41F5CA4C0F8045">Oligodon saiyok, a new limestone-dwelling kukri snake (Serpentes: Colubridae) from Kanchanaburi Province, western Thailand</mods:title>
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<mods:namePart id="5714C1F9632FEAD056F50897CC278757">Sumontha, Montri</mods:namePart>
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<mods:namePart id="23DFE548437D873C9B3FD69A8DB6CB8A">Kunya, Kirati</mods:namePart>
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<mods:namePart id="C5C7996495A50A04C38364D0474BB44C">Dangsri, Siriwat</mods:namePart>
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<mods:namePart id="EBFC5814B3232496E4F752BDFD54C81E">Pauwels, Olivier S. G.</mods:namePart>
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<treatment id="03C47D7ADB1B5B7DCEC0FBA08E5FF830" ID-DOI="http://doi.org/10.5281/zenodo.5698098" ID-GBIF-Taxon="132785379" ID-Zenodo-Dep="5698098" LSID="urn:lsid:plazi:treatment:03C47D7ADB1B5B7DCEC0FBA08E5FF830" httpUri="http://treatment.plazi.org/id/03C47D7ADB1B5B7DCEC0FBA08E5FF830" lastPageId="3" lastPageNumber="319" pageId="1" pageNumber="317">
<subSubSection id="C3779FE7DB1B5B7FCEC0FBA08F50FB89" box="[151,632,1125,1151]" pageId="1" pageNumber="317" type="nomenclature">
<paragraph id="8BD2CC6CDB1B5B7FCEC0FBA08F50FB89" blockId="1.[151,632,1125,1185]" box="[151,632,1125,1151]" pageId="1" pageNumber="317">
<heading id="D09A7B00DB1B5B7FCEC0FBA08F50FB89" bold="true" box="[151,632,1125,1151]" fontSize="11" level="1" pageId="1" pageNumber="317" reason="1">
<emphasis id="B919107EDB1B5B7FCEC0FBA08F50FB89" bold="true" box="[151,632,1125,1151]" pageId="1" pageNumber="317">
Description of
<taxonomicName id="4C6DB7EFDB1B5B7FCF18FBA08F3BFB89" authority="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels, 2017" authorityName="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels" authorityYear="2017" box="[335,531,1125,1151]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="317" phylum="Chordata" rank="species" species="saiyok" status="sp. nov.">
<emphasis id="B919107EDB1B5B7FCF18FBA08F3BFB89" bold="true" box="[335,531,1125,1151]" italics="true" pageId="1" pageNumber="317">Oligodon saiyok</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22AAD05DB1B5B7FCC4DFBA08F50FB89" box="[538,632,1125,1151]" pageId="1" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3779FE7DB1B5B7FCEC0FB4D8C23FB56" box="[151,267,1160,1185]" pageId="1" pageNumber="317" type="description">
<paragraph id="8BD2CC6CDB1B5B7FCEC0FB4D8C23FB56" blockId="1.[151,632,1125,1185]" box="[151,267,1160,1185]" pageId="1" pageNumber="317">
(
<figureCitation id="1356D0E9DB1B5B7FCEC8FB4D8C2AFB56" box="[159,258,1160,1185]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="2.[151,250,1218,1240]" captionStartId-1="3.[151,250,1358,1380]" captionStartId-2="4.[151,250,1041,1063]" captionStartId-3="4.[151,250,1986,2008]" captionStartId-4="5.[151,250,1029,1051]" captionStartId-5="5.[151,250,1991,2013]" captionStartId-6="6.[151,250,1089,1111]" captionTargetBox-0="[151,1436,342,1196]" captionTargetBox-1="[151,1436,373,1337]" captionTargetBox-2="[151,1436,297,1019]" captionTargetBox-3="[151,1436,1112,1964]" captionTargetBox-4="[172,1416,193,1006]" captionTargetBox-5="[203,1371,1101,1945]" captionTargetBox-6="[151,1436,193,1067]" captionTargetId-0="figure@2.[151,1436,342,1196]" captionTargetId-1="figure@3.[151,1436,373,1337]" captionTargetId-2="figure@4.[151,1436,193,1019]" captionTargetId-3="figure@4.[151,1436,1112,1964]" captionTargetId-4="figure@5.[172,1416,193,1006]" captionTargetId-5="figure@5.[203,1371,1101,1977]" captionTargetId-6="figure@6.[151,1436,193,1067]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="5" captionTargetPageId-6="6" captionText-0="FIGURE 1. Live holotype of Oligodon saiyok sp. nov. Photograph by K. Kunya." captionText-1="FIGURE 2. Ventral view of the preserved holotype of Oligodon saiyok sp. nov. Photograph by M. Sumontha." captionText-2="FIGURE 3. Dorsal view of the head of the preserved holotype of Oligodon saiyok sp. nov. Photograph by M. Sumontha." captionText-3="FIGURE 4. Left view of the head of the preserved holotype of Oligodon saiyok sp. nov. Photograph by M. Sumontha." captionText-4="FIGURE 5. Live paratype of Oligodon saiyok sp. nov. Photograph by K. Kunya." captionText-5="FIGURE 6. Ventral view of the preserved paratype of Oligodon saiyok sp. nov. Photograph by M. Sumontha." captionText-6="FIGURE 7. Left view of the head of the preserved paratype of Oligodon saiyok sp. nov. Photograph by M. Sumontha." httpUri-0="https://zenodo.org/record/844790/files/figure.png" httpUri-1="https://zenodo.org/record/844792/files/figure.png" httpUri-2="https://zenodo.org/record/844794/files/figure.png" httpUri-3="https://zenodo.org/record/844796/files/figure.png" httpUri-4="https://zenodo.org/record/844798/files/figure.png" httpUri-5="https://zenodo.org/record/844800/files/figure.png" httpUri-6="https://zenodo.org/record/844802/files/figure.png" pageId="1" pageNumber="317">Figs 17</figureCitation>
)
</paragraph>
</subSubSection>
<paragraph id="8BD2CC6CDB1B5B7FCEC0FB158F55FAC6" blockId="1.[151,1437,1232,2021]" pageId="1" pageNumber="317">
<materialsCitation id="3B05C631DB1B5B7FCEC0FB158F55FAC6" ID-GBIF-Occurrence="1586201938" collectingDate="2009-10-07" collectionCode="QSMI" collectorName="K. Kunya" country="Thailand" location="Benjarat Nakhon Cave Temple" municipality="Sai Yok District" pageId="1" pageNumber="317" specimenCode="QSMI 1506" specimenCount="1" stateProvince="Kanchanaburi Province" typeStatus="holotype">
<emphasis id="B919107EDB1B5B7FCEC0FB158C23FB1F" bold="true" box="[151,267,1232,1257]" pageId="1" pageNumber="317">
<typeStatus id="54D672CEDB1B5B7FCEC0FB158C2EFB1F" box="[151,262,1232,1257]" pageId="1" pageNumber="317" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBCB6417DB1B5B7FCF44FB158CB1FB1F" box="[275,409,1232,1257]" collectionCode="QSMI" pageId="1" pageNumber="317">QSMI 1506</specimenCode>
(field number MS 469); adult male from Wat Tham Benjarat Nakhon (=
<location id="8EB29AB7DB1B5B7FCA8BFB158C18FAFB" LSID="urn:lsid:plazi:treatment:03C47D7ADB1B5B7DCEC0FBA08E5FF830:8EB29AB7DB1B5B7FCA8BFB158C18FAFB" country="Thailand" municipality="Sai Yok District" name="Benjarat Nakhon Cave Temple" pageId="1" pageNumber="317" stateProvince="Kanchanaburi Province">Benjarat Nakhon Cave Temple</location>
),
<collectingMunicipality id="6BB65616DB1B5B7FCF12FB318CD5FAFB" box="[325,509,1268,1293]" pageId="1" pageNumber="317">Sai Yok District</collectingMunicipality>
,
<collectingRegion id="49A9028EDB1B5B7FCC5EFB318E3EFAFB" box="[521,790,1268,1293]" country="Thailand" name="Kanchanaburi" pageId="1" pageNumber="317">Kanchanaburi Province</collectingRegion>
, western
<collectingCountry id="F37A8CFCDB1B5B7FCDD7FB318ECFFAFB" box="[896,999,1268,1293]" name="Thailand" pageId="1" pageNumber="317">Thailand</collectingCountry>
; collected by
<collectorName id="2698A9BADB1B5B7FCAD2FB3089DEFAFB" box="[1157,1270,1269,1293]" pageId="1" pageNumber="317">K. Kunya</collectorName>
, M. Sumontha and S. Dangsri on
<date id="FFD3EAACDB1B5B7FCF31FADC8F3DFAC6" box="[358,533,1304,1329]" pageId="1" pageNumber="317" value="2009-10-07">
<collectingDate id="EF971344DB1B5B7FCF31FADC8F3DFAC6" box="[358,533,1304,1329]" pageId="1" pageNumber="317" value="2009-10-07">7 October 2009</collectingDate>
</date>
at 18.00.
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</paragraph>
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<materialsCitation id="3B05C631DB1B5B7FCE90FAF98863FAA3" ID-GBIF-Occurrence="1586201937" box="[199,1355,1340,1365]" collectionCode="QSMI" pageId="1" pageNumber="317" specimenCode="QSMI 1" specimenCount="2" typeStatus="paratype">
<emphasis id="B919107EDB1B5B7FCE90FAF98C13FAA3" bold="true" box="[199,315,1340,1365]" pageId="1" pageNumber="317">
<typeStatus id="54D672CEDB1B5B7FCE90FAF98C1EFAA3" box="[199,310,1340,1365]" pageId="1" pageNumber="317" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCode id="DBCB6417DB1B5B7FCF13FAF98C89FAA2" box="[324,417,1340,1364]" collectionCode="QSMI" pageId="1" pageNumber="317">QSMI 1</specimenCode>
507 (field number MS 470), adult female, same locality and collector as
<typeStatus id="54D672CEDB1B5B7FCAB2FAF98863FAA3" box="[1253,1355,1340,1365]" pageId="1" pageNumber="317" type="holotype">holotype</typeStatus>
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. It was found two hours later than the holotype.
</paragraph>
<paragraph id="8BD2CC6CDB1B5B7FCE90FA41883DFA13" blockId="1.[151,1437,1232,2021]" pageId="1" pageNumber="317">
<emphasis id="B919107EDB1B5B7FCE90FA418C69FA6B" bold="true" box="[199,321,1412,1437]" pageId="1" pageNumber="317">Diagnosis.</emphasis>
<taxonomicName id="4C6DB7EFDB1B5B7FCF1CFA418F2DFA6B" authority="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels, 2017" authorityName="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels" authorityYear="2017" box="[331,517,1412,1437]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="317" phylum="Chordata" rank="species" species="saiyok" status="sp. nov.">
<emphasis id="B919107EDB1B5B7FCF1CFA418F2DFA6B" box="[331,517,1412,1437]" italics="true" pageId="1" pageNumber="317">Oligodon saiyok</emphasis>
</taxonomicName>
<emphasis id="B919107EDB1B5B7FCC58FA418F43FA6B" bold="true" box="[527,619,1412,1437]" pageId="1" pageNumber="317">
<taxonomicNameLabel id="A22AAD05DB1B5B7FCC58FA418F43FA6B" box="[527,619,1412,1437]" pageId="1" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all other congeneric species by its maximal known SVL of
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, 8 SL, 17-17-15 DSR, 181187 VEN, single anal, 3843 SC, 13 MT, a dorsal pattern with 21 or 22 large blotches or rings, and venter with a dense network of subrectangular dark blotches.
</paragraph>
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Description of
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.
</emphasis>
Adult male. Body robust but elongate. Head short, neck moderately marked. Pupil round. SVL
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. TaL
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. Tail robust, tapering, accounting for 12.8 % of the TL (
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). HL
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; HW
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; HD
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. SnL
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. Snout short (27 % of HL, 1.9 times as long as ED). ED
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.
</paragraph>
<paragraph id="8BD2CC6CDB1B5B7FCE90F99B8C3DF949" blockId="1.[151,1437,1232,2021]" pageId="1" pageNumber="317">Body scalation. DSR 17-17-15, all smooth. DSR reduction from 17 to 15 occurs above the 133rd VEN by fusion of rows 4 and 5 (left) and 3 and 4 (right). Two PV + 187 VEN, laterally angulated. Anal plate single. SC 43, all divided.</paragraph>
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Head scalation. Rostral thick, curved onto upper snout surface, well visible from above, separating internasals by about one third of their length. Rostral width
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, rostral height
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. Nasals vertically divided. Nostril large, piercing top of middle of nasal. Internasals in broad contact, shorter than prefrontals. Prefrontals subrectangular, distinctly wider than long. Length of suture between internasals shorter than length of suture between prefrontals. Frontal pentagonal, 1.5 times as long as wide (frontal length
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, frontal width
<quantity id="4C956189DB1B5B7FCB62F89E88B8F885" box="[1333,1424,1883,1907]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="317" unit="mm" value="3.3">3.3 mm</quantity>
); 1/1 supraoculars, distinctly longer than wide; SL 8/8, 2nd and 3rd in contact with LOR, 4th and 5th in contact with orbit. LOR 1/1, distinctly longer (
<quantity id="4C956189DB1B5B7FCC46F8638F40F848" box="[529,616,1958,1982]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="317" unit="mm" value="1.8">1.8 mm</quantity>
) than high (
<quantity id="4C956189DB1B5B7FCCB8F8638E62F848" box="[751,842,1958,1982]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="317" unit="mm" value="0.7">0.7 mm</quantity>
). PreOc 1/1, tall and narrow; no PreSubOc. PosOc 2/ 2, the upper one larger. Tem 1+2 on each side. Mental width
<quantity id="4C956189DB1B5B7FCD37F8088E92F812" box="[864,954,1997,2021]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="317" unit="mm" value="1.9">1.9 mm</quantity>
, mental length
<quantity id="4C956189DB1B5B7FCA39F80889E2F812" box="[1134,1226,1997,2021]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="317" unit="mm" value="1.2">1.2 mm</quantity>
. IL 8/8, 1st pair in contact behind mental, IL
<quantity id="4C956189DB185B7CCFECFF5D8F3EFF46" box="[443,534,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="2" pageNumber="318" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1 to 4 in</quantity>
contact with anterior chin shields. First pair of chin shields distinctly longer than 2nd pair.
</paragraph>
<paragraph id="8BD2CC6CDB185B7CCE90FF278E3EFED4" blockId="2.[151,1436,151,290]" pageId="2" pageNumber="318">
Dentition. Thirteen MT, curved backwards, sharp, the two posterior ones enlarged, kukri-like. Hemipenes. The hemipenes
<emphasis id="B919107EDB185B7CCC53FECF8F61FED7" box="[516,585,266,289]" italics="true" pageId="2" pageNumber="318">in situ</emphasis>
reach the 18th SC.
</paragraph>
<caption id="DF129CE4DB185B7CCEC0FB078EF9FB2E" box="[151,977,1218,1240]" httpUri="https://zenodo.org/record/844790/files/figure.png" pageId="2" pageNumber="318" targetBox="[151,1436,342,1196]" targetPageId="2">
<paragraph id="8BD2CC6CDB185B7CCEC0FB078EF9FB2E" blockId="2.[151,977,1218,1240]" box="[151,977,1218,1240]" pageId="2" pageNumber="318">
<emphasis id="B919107EDB185B7CCEC0FB078C3BFB21" bold="true" box="[151,275,1218,1240]" pageId="2" pageNumber="318">FIGURE 1.</emphasis>
Live holotype of
<taxonomicName id="4C6DB7EFDB185B7CCF9FFB078F47FB2E" authority="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels, 2017" authorityName="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels" authorityYear="2017" box="[456,623,1218,1240]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="318" phylum="Chordata" rank="species" species="saiyok" status="sp. nov.">
<emphasis id="B919107EDB185B7CCF9FFB078F47FB2E" box="[456,623,1218,1240]" italics="true" pageId="2" pageNumber="318">Oligodon saiyok</emphasis>
</taxonomicName>
<emphasis id="B919107EDB185B7CCC22FB078FEDFB21" bold="true" box="[629,709,1218,1239]" pageId="2" pageNumber="318">
<taxonomicNameLabel id="A22AAD05DB185B7CCC22FB078FEDFB21" box="[629,709,1218,1239]" pageId="2" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Photograph by K. Kunya.
</paragraph>
</caption>
<paragraph id="8BD2CC6CDB185B7CCE90FB3A8859F8E5" blockId="2.[151,1437,1279,2030]" pageId="2" pageNumber="318">Coloration in life. Dorsal surface of head grayish brown as the dorsum, but paler on the sides of the snout and rostral. Four chevron marks pointing frontwards on the head and neck. Chevrons blackish brown, somewhat paler grayish brown in their center, edged by darker pigmentation. Anteriormost chevron with its apex at the level of the posterior edge of the rostral, covering part of the internasals, prefrontals and frontal, and descending through the eye to cover part of the 5th and 6th SL. Second chevron with its tip in the middle of the frontal, covering part of the frontal, parietals, temporals and descending behind the mouth angle to the ventrals. Third chevron with its anterior tip on the parietals, covering part of the parietals and the nape. Fourth chevron smallest, located midway between the third chevron and the first dorsal blotch. Dorsum surface grayish brown, with all dorsal scales finely but densely dotted with blackish brown. A series of 21 blackish brown blotches on dorsum, somewhat paler grayish brown in their center, edged by darker pigmentation, 2 or 3 DSR long and 5 or 6 DSR broad, i.e. covering the vertebral scale row and 2 or 3 adjacent rows. First blotch located at 14 scales behind parietals. An irregular blackish transversal zig-zag separates chevrons. Some irregular blackish marks on lower flanks under each chevron. Five similarly colored blotches above tail. No dorsal stripes. Ventral surface of head uniform cream. Belly cream with numerous subrectangular dark grayish brown blotches increasing in density posteriorly. Underside of tail cream with some subrectangular dark grayish brown blotches anteriorly. Iris light golden brown, speckled with black.</paragraph>
</subSubSection>
<subSubSection id="C3779FE7DB185B7DCE90F8DB8CE7FEB6" lastPageId="3" lastPageNumber="319" pageId="2" pageNumber="318" type="discussion">
<paragraph id="8BD2CC6CDB185B7DCE90F8DB8CE7FEB6" blockId="2.[151,1437,1279,2030]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="319" pageId="2" pageNumber="318">
<emphasis id="B919107EDB185B7CCE90F8DB8C14F8C1" bold="true" box="[199,316,1822,1847]" pageId="2" pageNumber="318">Variation.</emphasis>
<tableCitation id="C6EFF9D7DB185B7CCF11F8DB8CB1F8C1" box="[326,409,1822,1847]" captionStart="TABLE 1" captionStartId="6.[151,239,1150,1173]" captionTargetBox="[159,1402,1297,1401]" captionText="TABLE 1. Meristic and morphometric (in mm) data for the holotype and paratype of Oligodon saiyok sp. nov. Paired meristic characters are given left / right. Paired measurements are given for the right side. Supralabial numbers are followed in brackets by the ones contacting the orbit. Infralabial numbers are followed in brackets by how many contact the anterior chin shields. A = anal plate; S = single. For the other abbreviations, see Material and methods." httpUri="http://table.plazi.org/id/DF129CE4DB1C5B78CEC0FBBB882EFB03" pageId="2" pageNumber="318" tableUuid="DF129CE4DB1C5B78CEC0FBBB882EFB03">Table 1</tableCitation>
presents the main meristic and morphometric characters for the holotype and the paratype. The largest known specimen is the holotype. The relative tail length (TaL/TL) for the female paratype is 11.6 %, i.e., slightly shorter than in the male. The paratypes frontal is 1.4 times as long as wide (frontal length
<quantity id="4C956189DB185B7CCB69F8A288BFF888" box="[1342,1431,1895,1919]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="318" unit="mm" value="4.2">4.2 mm</quantity>
, frontal width
<quantity id="4C956189DB185B7CCF67F8488CA3F853" box="[304,395,1933,1957]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="318" unit="mm" value="2.9">2.9 mm</quantity>
). In the paratype, the DSR reduction from 17 to 15 occurs above the 131st VEN by fusion of rows 4 and 5 on the left side and fusion of rows 3 and 4 on the right side. The most noticeable difference between the holotype and the paratype is the dorsum color pattern (see
<figureCitation id="1356D0E9DB185B7CCD33F8108E9AF818" box="[868,946,2005,2030]" captionStart="FIGURE 5" captionStartId="5.[151,250,1029,1051]" captionTargetBox="[172,1416,193,1006]" captionTargetId="figure@5.[172,1416,193,1006]" captionTargetPageId="5" captionText="FIGURE 5. Live paratype of Oligodon saiyok sp. nov. Photograph by K. Kunya." httpUri="https://zenodo.org/record/844798/files/figure.png" pageId="2" pageNumber="318">Figs. 5</figureCitation>
and
<figureCitation id="1356D0E9DB185B7CCDBBF8108ED4F81B" box="[1004,1020,2005,2029]" captionStart="FIGURE 7" captionStartId="6.[151,250,1089,1111]" captionTargetBox="[151,1436,193,1067]" captionTargetId="figure@6.[151,1436,193,1067]" captionTargetPageId="6" captionText="FIGURE 7. Left view of the head of the preserved paratype of Oligodon saiyok sp. nov. Photograph by M. Sumontha." httpUri="https://zenodo.org/record/844802/files/figure.png" pageId="2" pageNumber="318">7</figureCitation>
). Contrary to the male holotype, the female paratype does not show dorsal dark blotches but well-defined white rings bordered by a black line, in a number that is comparable to the number of blotches in the male holotype. Excluding the similarly colored chevron on the nape, the dorsum displays 22 such rings and the tail 4. These rings do not fully encircle the body as they do not reach the ventrals. The posterior rings are less contrasted than the anterior ones. The color of the iris in life is the same as in the holotype.
</paragraph>
</subSubSection>
<caption id="DF129CE4DB195B7DCEC0FA8B89DCFA92" box="[151,1268,1358,1380]" httpUri="https://zenodo.org/record/844792/files/figure.png" pageId="3" pageNumber="319" targetBox="[151,1436,373,1337]" targetPageId="3">
<paragraph id="8BD2CC6CDB195B7DCEC0FA8B89DCFA92" blockId="3.[151,1268,1358,1380]" box="[151,1268,1358,1380]" pageId="3" pageNumber="319">
<emphasis id="B919107EDB195B7DCEC0FA8B8C3AFA95" bold="true" box="[151,274,1358,1380]" pageId="3" pageNumber="319">FIGURE 2.</emphasis>
Ventral view of the preserved holotype of
<taxonomicName id="4C6DB7EFDB195B7DCC93FA8B8E43FA92" authority="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels, 2017" authorityName="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels" authorityYear="2017" box="[708,875,1358,1380]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="saiyok" status="sp. nov.">
<emphasis id="B919107EDB195B7DCC93FA8B8E43FA92" box="[708,875,1358,1380]" italics="true" pageId="3" pageNumber="319">Oligodon saiyok</emphasis>
</taxonomicName>
<emphasis id="B919107EDB195B7DCD27FA8A8EE9FA92" bold="true" box="[880,961,1359,1380]" pageId="3" pageNumber="319">
<taxonomicNameLabel id="A22AAD05DB195B7DCD27FA8A8EE9FA92" box="[880,961,1359,1380]" pageId="3" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Photograph by M. Sumontha.
</paragraph>
</caption>
<paragraph id="8BD2CC6CDB195B7DCE90FA54880CFA04" blockId="3.[151,1436,1425,1990]" pageId="3" pageNumber="319">
<emphasis id="B919107EDB195B7DCE90FA548F78FA5C" bold="true" box="[199,592,1425,1450]" pageId="3" pageNumber="319">Distribution and natural history.</emphasis>
<taxonomicName id="4C6DB7EFDB195B7DCC0FFA548E39FA5C" authority="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels, 2017" authorityName="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels" authorityYear="2017" box="[600,785,1425,1450]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="saiyok" status="sp. nov.">
<emphasis id="B919107EDB195B7DCC0FFA548E39FA5C" box="[600,785,1425,1450]" italics="true" pageId="3" pageNumber="319">Oligodon saiyok</emphasis>
</taxonomicName>
<emphasis id="B919107EDB195B7DCD4EFA548E5CFA5C" bold="true" box="[793,884,1425,1450]" pageId="3" pageNumber="319">
<taxonomicNameLabel id="A22AAD05DB195B7DCD4EFA548E5CFA5C" box="[793,884,1425,1450]" pageId="3" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently known only from its
<typeStatus id="54D672CEDB195B7DCAA5FA57880AFA5C" box="[1266,1314,1426,1450]" pageId="3" pageNumber="319">type</typeStatus>
locality in Sai Yok District (
<figureCitation id="1356D0E9DB195B7DCF37FA708C87FA38" box="[352,431,1461,1486]" captionStart="FIGURE 8" captionStartId="7.[151,250,1460,1482]" captionTargetBox="[343,1247,266,1438]" captionTargetId="figure@7.[340,1247,265,1439]" captionTargetPageId="7" captionText="FIGURE 8. Geographical location of the type-locality (red star) of Oligodon saiyok sp. nov. in Kanchanaburi Province, western Thailand. Map by W. Sodoab." httpUri="https://zenodo.org/record/844804/files/figure.png" pageId="3" pageNumber="319">Figs. 8</figureCitation>
and
<figureCitation id="1356D0E9DB195B7DCFBDFA738CD2FA38" box="[490,506,1462,1486]" captionStart="FIGURE 9" captionStartId="8.[151,250,1723,1745]" captionTargetBox="[151,1436,736,1700]" captionTargetId="figure@8.[151,1436,736,1700]" captionTargetPageId="8" captionText="FIGURE 9. Biotope of Oligodon saiyok sp. nov. at the type locality. Photograph by M. Sumontha." httpUri="https://zenodo.org/record/844806/files/figure.png" pageId="3" pageNumber="319">9</figureCitation>
). Both individuals were found active at night (18.00 and 20.00 respectively) in a Buddhist temple at the entrance of a cave on a limestone hill. They were rather aggressive when caught.
</paragraph>
<paragraph id="8BD2CC6CDB195B7DCE90FA388E41F8AC" blockId="3.[151,1436,1425,1990]" pageId="3" pageNumber="319">
Other reptiles we found in syntopy at the
<typeStatus id="54D672CEDB195B7DCCCCFA3B8FE3F9E0" box="[667,715,1534,1558]" pageId="3" pageNumber="319">type</typeStatus>
locality include
<taxonomicName id="4C6DB7EFDB195B7DCDD0FA388864F9E0" authority="(Boulenger)" authorityName="Boulenger" box="[903,1356,1533,1558]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B919107EDB195B7DCDD0FA388994F9E3" box="[903,1212,1533,1558]" italics="true" pageId="3" pageNumber="319">Cyrtodactylus cf. peguensis</emphasis>
(Boulenger)
</taxonomicName>
and
<taxonomicName id="4C6DB7EFDB195B7DCBD5FA3B8F89F9CF" authority="Bauer, Sumontha &amp; Pauwels, 2003" authorityName="Bauer, Sumontha &amp; Pauwels" authorityYear="2003" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="tigroides">
<emphasis id="B919107EDB195B7DCBD5FA3B8DD2F9CC" italics="true" pageId="3" pageNumber="319">C. tigroides</emphasis>
<bibRefCitation id="EFFCB19DDB195B7DCF53F9E78F89F9CF" author="Bauer" box="[260,673,1569,1594]" pageId="3" pageNumber="319" refString="Bauer, A. M., Sumontha, M. &amp; Pauwels, O. S. G. (2003) Two new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Thailand. Zootaxa, 376 (1), 1 - 18." type="journal article" year="2003">Bauer, Sumontha &amp; Pauwels, 2003</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6DB7EFDB195B7DCCE7F9E78C0BF9A8" authority="Bauer, Sumontha, Grossmann, Pauwels &amp; Vogel, 2004" authorityName="Bauer, Sumontha, Grossmann, Pauwels &amp; Vogel" authorityYear="2004" class="Reptilia" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="hangseesom">
<emphasis id="B919107EDB195B7DCCE7F9E78E81F9CC" box="[688,937,1569,1594]" italics="true" pageId="3" pageNumber="319">Dixonius hangseesom</emphasis>
<bibRefCitation id="EFFCB19DDB195B7DCDE3F9E78C0BF9A8" author="Bauer" pageId="3" pageNumber="319" refString="Bauer, A. M., Sumontha, M., Grossmann, W., Pauwels, O. S. G. &amp; Vogel, G. (2004) A new species of Dixonius (Squamata: Gekkonidae) from Kanchanaburi Province, western Thailand. Current Herpetology, 23 (1), 17 - 26. https: // doi. org / 10.5358 / hsj. 23.17" type="journal article" year="2004">Bauer, Sumontha, Grossmann, Pauwels &amp; Vogel, 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6DB7EFDB195B7DCF66F9808F6EF9A8" authority="(Linnaeus)" authorityName="Linnaeus" box="[305,582,1605,1630]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="gecko">
<emphasis id="B919107EDB195B7DCF66F9808CECF9A8" box="[305,452,1605,1630]" italics="true" pageId="3" pageNumber="319">Gekko gecko</emphasis>
(Linnaeus)
</taxonomicName>
and
<taxonomicName id="4C6DB7EFDB195B7DCCD5F983884EF9A8" authority="Bauer, Sumontha &amp; Pauwels, 2008" authorityName="Bauer, Sumontha &amp; Pauwels" authorityYear="2008" box="[642,1382,1605,1630]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="nutaphandi">
<emphasis id="B919107EDB195B7DCCD5F9838E0AF9A8" box="[642,802,1605,1630]" italics="true" pageId="3" pageNumber="319">G. nutaphandi</emphasis>
<bibRefCitation id="EFFCB19DDB195B7DCD7BF9838997F9A8" author="Bauer" box="[812,1215,1605,1630]" pageId="3" pageNumber="319" refString="Bauer, A. M., Sumontha, M. &amp; Pauwels, O. S. G. (2008) A new red-eyed Gekko (Reptilia: Gekkonidae) from Kanchanaburi Province, Thailand. Zootaxa, 1750, 32 - 42." type="journal article" year="2008">Bauer, Sumontha &amp; Pauwels, 2008</bibRefCitation>
(Gekkonidae)
</taxonomicName>
, and
<taxonomicName id="4C6DB7EFDB195B7DCEC0F9AF8F94F974" authority="Smith" authorityName="Smith" authorityYear="1943" box="[151,700,1641,1666]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="kanburiensis">
<emphasis id="B919107EDB195B7DCEC0F9AF8CE6F974" box="[151,462,1641,1666]" italics="true" pageId="3" pageNumber="319">Trimeresurus kanburiensis</emphasis>
Smith (Viperidae)
</taxonomicName>
. Syntopic amphibians we observed included
<taxonomicName id="4C6DB7EFDB195B7DCB55F9AC8F63F950" authority="(Schneider)" authorityName="Schneider" class="Amphibia" family="Bufonidae" genus="Duttaphrynus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="melanostictus">
<emphasis id="B919107EDB195B7DCB55F9AC8C19F950" italics="true" pageId="3" pageNumber="319">Duttaphrynus melanostictus</emphasis>
(Schneider) (Bufonidae)
</taxonomicName>
,
<taxonomicName id="4C6DB7EFDB195B7DCC01F9488E63F950" authority="Gray" authorityName="Gray" authorityYear="1831" box="[598,843,1677,1702]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="pulchra">
<emphasis id="B919107EDB195B7DCC01F9488E26F950" box="[598,782,1677,1702]" italics="true" pageId="3" pageNumber="319">Kaloula pulchra</emphasis>
Gray
</taxonomicName>
,
<taxonomicName id="4C6DB7EFDB195B7DCD0EF94888B3F950" authority="(Taylor)" authorityName="Taylor" box="[857,1435,1677,1702]" class="Amphibia" family="Microhylidae" genus="Micryletta" kingdom="Animalia" order="Anura" pageId="3" pageNumber="319" phylum="Chordata" rank="subSpecies" species="inornata" subSpecies="lineata">
<emphasis id="B919107EDB195B7DCD0EF94889A2F950" box="[857,1162,1677,1702]" italics="true" pageId="3" pageNumber="319">Micryletta inornata lineata</emphasis>
(Taylor) (Microhylidae)
</taxonomicName>
and
<taxonomicName id="4C6DB7EFDB195B7DCE99F9748E7AF93C" authority="Hallowell" authorityName="Hallowell" authorityYear="1861" box="[206,850,1713,1738]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="megacephalus">
<emphasis id="B919107EDB195B7DCE99F9748F2DF93C" box="[206,517,1713,1738]" italics="true" pageId="3" pageNumber="319">Polypedates megacephalus</emphasis>
Hallowell (Rhacophoridae)
</taxonomicName>
. A color pattern dimorphism similar to the one observed in
<taxonomicName id="4C6DB7EFDB195B7DCF77F9108CF2F918" authority="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels, 2017" authorityName="Sumontha &amp; Kunya &amp; Dangsri &amp; Pauwels" authorityYear="2017" box="[288,474,1749,1774]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="saiyok" status="sp. nov.">
<emphasis id="B919107EDB195B7DCF77F9108CF2F918" box="[288,474,1749,1774]" italics="true" pageId="3" pageNumber="319">Oligodon saiyok</emphasis>
</taxonomicName>
<emphasis id="B919107EDB195B7DCFB7F9108F12F918" bold="true" box="[480,570,1749,1774]" pageId="3" pageNumber="319">
<taxonomicNameLabel id="A22AAD05DB195B7DCFB7F9108F12F918" box="[480,570,1749,1774]" pageId="3" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has been documented for
<taxonomicName id="4C6DB7EFDB195B7DCD34F9138936F91B" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[867,1054,1750,1773]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="species" species="purpurascens">
<emphasis id="B919107EDB195B7DCD34F9138936F91B" box="[867,1054,1750,1773]" italics="true" pageId="3" pageNumber="319">O. purpurascens</emphasis>
</taxonomicName>
by van
<bibRefCitation id="EFFCB19DDB195B7DCA2FF910884DF918" box="[1144,1381,1749,1774]" pageId="3" pageNumber="319" refString="van Rooijen, J., Wood, P. L., Grismer, J. L., Grismer, L. L. &amp; Grossmann, W. (2011) Color pattern dimorphism in the colubrid snake Oligodon purpurascens (Schlegel, 1837) (Reptilia: Squamata). Russian Journal of Herpetology, 18 (3), 215 - 220." type="journal article">
Rooijen
<emphasis id="B919107EDB195B7DCA8EF9138822F918" box="[1241,1290,1749,1774]" italics="true" pageId="3" pageNumber="319">et al</emphasis>
. (2011)
</bibRefCitation>
. The blotched pattern provides a cryptic coloration in the leaf litter, while the conspicuous ringed pattern suggests a mimicry of an aposematic signal, as noted by van
<bibRefCitation id="EFFCB19DDB195B7DCC8FF8D88EE1F8C0" box="[728,969,1821,1846]" pageId="3" pageNumber="319" refString="van Rooijen, J., Wood, P. L., Grismer, J. L., Grismer, L. L. &amp; Grossmann, W. (2011) Color pattern dimorphism in the colubrid snake Oligodon purpurascens (Schlegel, 1837) (Reptilia: Squamata). Russian Journal of Herpetology, 18 (3), 215 - 220." type="journal article">
Rooijen
<emphasis id="B919107EDB195B7DCD6EF8DB8E43F8C0" box="[825,875,1821,1846]" italics="true" pageId="3" pageNumber="319">et al</emphasis>
. (2011)
</bibRefCitation>
; these latter authors did not mention any link between the pattern observed and the sex of the individuals.
</paragraph>
<subSubSection id="C3779FE7DB195B7DCE90F8A08E5FF830" pageId="3" pageNumber="319" type="etymology">
<paragraph id="8BD2CC6CDB195B7DCE90F8A08E5FF830" blockId="3.[151,1436,1425,1990]" pageId="3" pageNumber="319">
<emphasis id="B919107EDB195B7DCE90F8A08C66F888" bold="true" box="[199,334,1893,1918]" pageId="3" pageNumber="319">Etymology.</emphasis>
The specific epithet is an invariable noun in honor of the administrative district where the typelocality lies. We suggest the following common names:
<emphasis id="B919107EDB195B7DCD59F84F8EF4F854" box="[782,988,1929,1954]" italics="true" pageId="3" pageNumber="319">Ngu Ngod Sai Yok</emphasis>
(Thai), Sai Yok Kukri Snake (English),
<emphasis id="B919107EDB195B7DCEC0F8688C53F830" box="[151,379,1965,1990]" italics="true" pageId="3" pageNumber="319">
<taxonomicName id="4C6DB7EFDB195B7DCEC0F8688C29F830" authorityName="Boie in Fitzinger" authorityYear="1826" box="[151,257,1965,1990]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="319" phylum="Chordata" rank="genus">Oligodon</taxonomicName>
de SaÏ Yok
</emphasis>
(French), and
<emphasis id="B919107EDB195B7DCC77F86B8FD4F830" box="[544,764,1965,1990]" italics="true" pageId="3" pageNumber="319">Sai Yok Kukrinatter</emphasis>
(German).
</paragraph>
</subSubSection>
</treatment>
</document>