265 lines
31 KiB
XML
265 lines
31 KiB
XML
<document id="58FE807B4EE41EFAD302814B173688DE" ID-CLB-Dataset="36770" ID-DOI="10.11646/zootaxa.3878.1.7" ID-GBIF-Dataset="a2036f4e-9744-4420-88ee-f8a0cde25cc4" ID-ISSN="1175-5326" ID-Zenodo-Dep="287356" ID-ZooBank="D7F22A9C-158F-4449-BD0C-49C734AF3E8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460941702703" checkinUser="plazi" docAuthor="Löcker, Birgit" docDate="2014" docId="03C2EC72FFF1FFC4FF295639FC9BFBBF" docLanguage="en" docName="zt03878p096.pdf" docOrigin="Zootaxa 3878 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calamister obscurus Kirkaldy" docType="treatment" docVersion="7" lastPageNumber="93" masterDocId="FFFB940AFFF3FFC0FFBE5252FFF5FFA6" masterDocTitle="Taxonomic revision of Calamister Kirkaldy, 1906 (Fulgoromorpha: Cixiidae) with the description of a new species" masterLastPageNumber="96" masterPageNumber="89" pageNumber="91" updateTime="1698584400831" updateUser="ExternalLinkService">
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<mods:mods id="988A232DB10EC0881807B16C34550241" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="460E81946BBA4692643669624F7FF673">
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<mods:title id="3CC39A204CF60ABDD5C9988C373508C8">Taxonomic revision of Calamister Kirkaldy, 1906 (Fulgoromorpha: Cixiidae) with the description of a new species</mods:title>
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<mods:name id="8C2FE28B9621CF92E37D23B9EB57ECC4" type="personal">
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<mods:roleTerm id="4E2C7EAFD76B2E8844FA746FC6C334A5">Author</mods:roleTerm>
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<mods:namePart id="E74146223CE10D4314ED04ABFCDB11DC">Löcker, Birgit</mods:namePart>
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<mods:typeOfResource id="35578569C04F5A21EB3E46AAD8D74313">text</mods:typeOfResource>
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<mods:title id="B76B7067791D9683EECEC155FDCF9FE7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AFD1F4EEA11AD22D44057609EFD772F8">
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<mods:date id="67632FE54ACE2944E31554B4C1F1AC52">2014</mods:date>
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<mods:detail id="58DBD9D8D7391BD854E8D300E8DC74CF" type="volume">
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<mods:number id="F98EF088BD8454C887C8B46421A9F81C">3878</mods:number>
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</mods:detail>
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<mods:detail id="DD3CC516388E5234E2CE56A6AEBF0A67" type="issue">
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<mods:number id="317B47CB4CC59002B55E0C67FFE0E52C">1</mods:number>
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<mods:start id="2E124800AABAE4B4234168F0E96A162E">89</mods:start>
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<mods:end id="20E0DA1CF596E59F5FD40E79B2659533">96</mods:end>
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<mods:classification id="6DADF02456E6CD27140AFBF314E08274">journal article</mods:classification>
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<mods:identifier id="8AE7E0A1C8AAEA1053D07315CA133233" type="CLB-Dataset">36770</mods:identifier>
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<mods:identifier id="46A26909806337F1C973225453548C21" type="DOI">10.11646/zootaxa.3878.1.7</mods:identifier>
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<mods:identifier id="5820699CBD7853B5CBA6F5E27B1AE51B" type="GBIF-Dataset">a2036f4e-9744-4420-88ee-f8a0cde25cc4</mods:identifier>
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<mods:identifier id="D94530F240BD3AB314F859C2CED53195" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="24B0279F32D220A779E8217C38F82E0E" type="Zenodo-Dep">287356</mods:identifier>
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<treatment id="03C2EC72FFF1FFC4FF295639FC9BFBBF" ID-DOI="http://doi.org/10.5281/zenodo.6123464" ID-GBIF-Taxon="127670637" ID-Zenodo-Dep="6123464" LSID="urn:lsid:plazi:treatment:03C2EC72FFF1FFC4FF295639FC9BFBBF" httpUri="http://treatment.plazi.org/id/03C2EC72FFF1FFC4FF295639FC9BFBBF" lastPageId="4" lastPageNumber="93" pageId="2" pageNumber="91">
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<subSubSection id="C3710EEFFFF1FFC2FF295639FEA9FB01" pageId="2" pageNumber="91" type="nomenclature">
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<paragraph id="8BD45D64FFF1FFC2FF295639FDFDFB20" blockId="2.[151,520,1131,1191]" box="[151,520,1131,1158]" pageId="2" pageNumber="91">
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<heading id="D09CEA08FFF1FFC2FF295639FDFDFB20" bold="true" box="[151,520,1131,1158]" fontSize="11" level="1" pageId="2" pageNumber="91" reason="1">
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<taxonomicName id="4C6B26E7FFF1FFC2FF295639FDFDFB20" ID-CoL="PDHP" authority="Kirkaldy" authorityName="Kirkaldy" box="[151,520,1131,1158]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="obscurus">
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||
<emphasis id="B91F8176FFF1FFC2FF295639FDFDFB20" bold="true" box="[151,520,1131,1158]" pageId="2" pageNumber="91">
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<emphasis id="B91F8176FFF1FFC2FF295639FE7AFB23" bold="true" box="[151,399,1131,1157]" italics="true" pageId="2" pageNumber="91">Calamister obscurus</emphasis>
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Kirkaldy
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BD45D64FFF1FFC2FF2956DCFEA9FB01" blockId="2.[151,520,1131,1191]" box="[151,348,1166,1191]" pageId="2" pageNumber="91">
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||
(
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<figureCitation id="135041E1FFF1FFC2FF2156DCFF12FB01" box="[159,231,1166,1191]" captionStart="FIGURE 2" captionStartId="3.[151,250,2006,2028]" captionTargetBox="[151,1395,359,1975]" captionTargetId="figure@3.[151,1436,357,1986]" captionTargetPageId="3" captionText="FIGURE 2. Calamister obscurus: A habitus; B forewing; C – D head; E first and second tarsomere. Scale bar 1 mm." httpUri="https://zenodo.org/record/287358/files/figure.png" pageId="2" pageNumber="91">Figs 2</figureCitation>
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,
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<figureCitation id="135041E1FFF1FFC2FF4D56DDFEF6FB01" box="[243,259,1167,1191]" captionStart="FIGURE 4" captionStartId="6.[151,250,807,829]" captionTargetBox="[344,1242,198,782]" captionTargetId="figure@6.[338,1249,193,786]" captionTargetPageId="6" captionText="FIGURE 4. Calamister obscurus: A aedeagus ventral; B aedeagus left lateral; C – D anal tube; E – F genital styles." httpUri="https://zenodo.org/record/287360/files/figure.png" pageId="2" pageNumber="91">4</figureCitation>
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,
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<figureCitation id="135041E1FFF1FFC2FEB156DCFEE9FB01" box="[271,284,1166,1191]" captionStart="FIGURE 6" captionStartId="7.[151,250,1030,1052]" captionTargetBox="[170,1417,202,1000]" captionTargetId="figure@7.[164,1424,195,1009]" captionTargetPageId="7" captionText="FIGURE 6. Female genitalia. Calamister obscurus: A ventral; B caudal; C dorsal. Calamister cougalensis: D ventral; E caudal; F dorsal." httpUri="https://zenodo.org/record/287362/files/figure.png" pageId="2" pageNumber="91">6</figureCitation>
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A–C)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3710EEFFFF1FFC2FF295684FB3AFA38" pageId="2" pageNumber="91" type="reference_group">
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<paragraph id="8BD45D64FFF1FFC2FF295684FDB0FB4A" blockId="2.[151,581,1238,1261]" box="[151,581,1238,1261]" pageId="2" pageNumber="91">
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<treatmentCitationGroup id="AB7B7A4AFFF1FFC2FF295684FDB0FB4A" box="[151,581,1238,1261]" pageId="2" pageNumber="91">
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<taxonomicName id="4C6B26E7FFF1FFC2FF295684FDB7FB4A" ID-CoL="PDHP" authority="Kirkaldy, 1906: 403" authorityName="Kirkaldy" authorityPageNumber="403" authorityYear="1906" box="[151,578,1238,1261]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="obscurus">
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<emphasis id="B91F8176FFF1FFC2FF295684FE98FB4B" bold="true" box="[151,365,1238,1261]" italics="true" pageId="2" pageNumber="91">Calamister obscurus</emphasis>
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<treatmentCitation id="0ACA7B75FFF1FFC2FECA5684FDB7FB4A" author="Kirkaldy" box="[372,578,1238,1260]" page="403" pageId="2" pageNumber="91" year="1906">
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<bibRefCitation id="EFFA2095FFF1FFC2FECA5684FDFAFB4A" author="Kirkaldy" box="[372,527,1238,1260]" pageId="2" pageNumber="91" refString="Kirkaldy, G. W. (1906) Leafhoppers and their natural enemies. Bulletin of the Hawaiian Sugar Planter's Association Experimental Station Entomological Series, 1, 271 - 479." type="journal article" year="1906">Kirkaldy, 1906</bibRefCitation>
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: 403
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD45D64FFF1FFC2FF29574BFCACFAF0" blockId="2.[151,1437,1305,2014]" pageId="2" pageNumber="91">
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<emphasis id="B91F8176FFF1FFC2FF29574BFF16FA94" bold="true" box="[151,227,1305,1330]" pageId="2" pageNumber="91">
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<typeStatus id="54D0E3C6FFF1FFC2FF29574BFF2AFA94" box="[151,223,1305,1330]" pageId="2" pageNumber="91">Types</typeStatus>
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.
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</emphasis>
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<emphasis id="B91F8176FFF1FFC2FF4D5748FE95FA97" box="[243,352,1306,1329]" italics="true" pageId="2" pageNumber="91">
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<typeStatus id="54D0E3C6FFF1FFC2FF4D5748FE95FA97" box="[243,352,1306,1329]" pageId="2" pageNumber="91" type="lectotype">Lectotype</typeStatus>
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</emphasis>
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(designated here, examined), ♂,
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<emphasis id="B91F8176FFF1FFC2FCBF574BFC06FA94" bold="true" box="[769,1011,1305,1330]" pageId="2" pageNumber="91">
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<collectingCountry id="F37C1DF4FFF1FFC2FCBF574BFC5DFA94" box="[769,936,1305,1330]" name="Australia" pageId="2" pageNumber="91">AUSTRALIA</collectingCountry>
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, Qld:
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</emphasis>
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Bundaberg,
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<date id="FFD57BA4FFF1FFC2FB2B574BFAE9FA94" box="[1173,1308,1305,1330]" pageId="2" pageNumber="91" value="1904-09" valueMax="1904-12-31" valueMin="1904-09-00">ix.–xii.1904</date>
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(Koebele) (BPBM).
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<emphasis id="B91F8176FFF1FFC2FEBB576FFE58FAF3" box="[261,429,1341,1365]" italics="true" pageId="2" pageNumber="91">
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||
<typeStatus id="54D0E3C6FFF1FFC2FEBB576FFE58FAF3" box="[261,429,1341,1365]" pageId="2" pageNumber="91" type="paralectotype">Paralectotypes</typeStatus>
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</emphasis>
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, 2 ♀, same data as
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<typeStatus id="54D0E3C6FFF1FFC2FD36576FFD1FFAF0" box="[648,746,1341,1366]" pageId="2" pageNumber="91" type="holotype">holotype</typeStatus>
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(BPBM).
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</paragraph>
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<paragraph id="8BD45D64FFF1FFC2FF795733FB3AFA38" blockId="2.[151,1437,1305,2014]" pageId="2" pageNumber="91">
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<emphasis id="B91F8176FFF1FFC2FF795733FEE6FADC" bold="true" box="[199,275,1377,1402]" pageId="2" pageNumber="91">Notes:</emphasis>
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The original description does not mention the designation of a
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<typeStatus id="54D0E3C6FFF1FFC2FC615733FBB1FADC" box="[991,1092,1377,1402]" pageId="2" pageNumber="91" type="holotype">holotype</typeStatus>
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, therefore these specimens are regarded as
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<typeStatus id="54D0E3C6FFF1FFC2FEA057D4FE71FA38" box="[286,388,1414,1438]" pageId="2" pageNumber="91" type="syntype">syntypes</typeStatus>
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||
. A
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<typeStatus id="54D0E3C6FFF1FFC2FE1757D7FDE5FA38" box="[425,528,1413,1438]" pageId="2" pageNumber="91" type="lectotype">lectotype</typeStatus>
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is designated to provide a diagnostic reference for the species.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3710EEFFFF1FFC2FF7957FBFABBF9AC" pageId="2" pageNumber="91" type="materials_examined">
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||
<paragraph id="8BD45D64FFF1FFC2FF7957FBFABBF9AC" blockId="2.[151,1437,1305,2014]" pageId="2" pageNumber="91">
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||
<emphasis id="B91F8176FFF1FFC2FF7957FBFD1CFA64" bold="true" box="[199,745,1449,1474]" pageId="2" pageNumber="91">
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Other material examined.
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<collectingCountry id="F37C1DF4FFF1FFC2FDBF57FBFD5DFA64" box="[513,680,1449,1474]" name="Australia" pageId="2" pageNumber="91">AUSTRALIA</collectingCountry>
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, Qld:
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</emphasis>
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<specimenCount id="9D6D96EDFFF1FFC2FD4E57F8FCDDFA67" box="[752,808,1450,1474]" pageId="2" pageNumber="91" type="generic">11 ♂</specimenCount>
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, 6 ♀,
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<specimenCount id="9D6D96EDFFF1FFC2FCD457F8FC66FA67" box="[874,915,1450,1474]" pageId="2" pageNumber="91" type="generic">1 ♂</specimenCount>
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or ♀ (abdomen missing, body length indicates male), Bundaberg,
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<date id="FFD57BA4FFF1FFC2FED0579FFE03FA40" box="[366,502,1485,1510]" pageId="2" pageNumber="91" value="1919-08-20">20.viii.1919</date>
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(F. Muir) (
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<collectionCode id="ED7AC5A1FFF1FFC2FDCA579CFD3DFA40" LSID="urn:lsid:biocol.org:col:34697" box="[628,712,1486,1510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="91">BPBM</collectionCode>
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);
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<specimenCount id="9D6D96EDFFF1FFC2FD62579CFCFCFA43" box="[732,777,1486,1510]" pageId="2" pageNumber="91" type="generic">1 ♂</specimenCount>
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, 1 ♀, Coolum,
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<date id="FFD57BA4FFF1FFC2FC06579FFBDAFA40" box="[952,1071,1485,1510]" pageId="2" pageNumber="91" value="1956-09-22">22.ix.1956</date>
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(Kirkpolnik) (QM).
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<emphasis id="B91F8176FFF1FFC2FAA5579FFA97FA40" bold="true" box="[1307,1378,1485,1510]" pageId="2" pageNumber="91">
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||
<collectionCode id="ED7AC5A1FFF1FFC2FAA5579FFAABFA40" LSID="urn:lsid:biocol.org:col:15627" box="[1307,1374,1485,1510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="2" pageNumber="91">NSW</collectionCode>
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:
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</emphasis>
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2 ♀, Bullsground [state forest], Port
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<collectingCountry id="F37C1DF4FFF1FFC2FE4757A3FD81F9AC" box="[505,628,1521,1546]" name="Australia" pageId="2" pageNumber="91">Macquarie</collectingCountry>
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, beating understorey plants,
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<date id="FFD57BA4FFF1FFC2FC0A57A3FBE5F9AC" box="[948,1040,1521,1546]" pageId="2" pageNumber="91" value="2005-11">xi. 2005</date>
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(M.L. Moir MLM01135-6).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3710EEFFFF1FFC4FF795447FDEDFE99" lastPageId="4" lastPageNumber="93" pageId="2" pageNumber="91" type="diagnosis">
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||
<paragraph id="8BD45D64FFF1FFC2FF795447FDB8F9D0" blockId="2.[151,1437,1305,2014]" pageId="2" pageNumber="91">
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||
<emphasis id="B91F8176FFF1FFC2FF795447FEB4F988" bold="true" box="[199,321,1557,1582]" pageId="2" pageNumber="91">Diagnosis.</emphasis>
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This species can be distinguished from
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<taxonomicName id="4C6B26E7FFF1FFC2FCB25444FC47F98B" box="[780,946,1557,1581]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="cougalensis">
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<emphasis id="B91F8176FFF1FFC2FCB25444FC47F98B" box="[780,946,1557,1581]" italics="true" pageId="2" pageNumber="91">C. cougalensis</emphasis>
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</taxonomicName>
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by its colouration which is entirely yellow with hyaline forewings and a whitish or yellowish pterostigma. Further the phallotheca bears two spines ventrally to right-laterally, about equal in length.
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</paragraph>
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||
<paragraph id="8BD45D64FFF1FFC2FF7954D3FB45F91B" blockId="2.[151,1437,1305,2014]" pageId="2" pageNumber="91">
|
||
<emphasis id="B91F8176FFF1FFC2FF7954D3FED5F93C" bold="true" box="[199,288,1665,1690]" pageId="2" pageNumber="91">Colour.</emphasis>
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Body light brownish-yellow (testaceous), eyes and antennae darker; rarely lateral parts of mesonotum slightly darker. Forewing hyaline with testaceous veins and tubercles; pterostigma testaceous.
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</paragraph>
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<paragraph id="8BD45D64FFF1FFC2FF79549BFCAEF947" blockId="2.[151,1437,1305,2014]" box="[199,859,1737,1762]" pageId="2" pageNumber="91">
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<emphasis id="B91F8176FFF1FFC2FF79549BFE95F944" bold="true" box="[199,352,1737,1762]" pageId="2" pageNumber="91">Morphology.</emphasis>
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Body length: ♂
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<quantity id="4C93F081FFF1FFC2FDA2549BFD50F947" box="[540,677,1737,1762]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="2" pageNumber="91" unit="mm" value="4.4" valueMax="4.5" valueMin="4.3">4.3–4.5 mm</quantity>
|
||
; ♀
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<quantity id="4C93F081FFF1FFC2FD73549BFCA2F947" box="[717,855,1737,1762]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.1" metricValueMin="4.7" pageId="2" pageNumber="91" unit="mm" value="4.9" valueMax="5.1" valueMin="4.7">4.7–5.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD45D64FFF1FFC2FF7954BFFBACF830" blockId="2.[151,1437,1305,2014]" pageId="2" pageNumber="91">Head: Vertex 1.5–1.7 x wider than long; apical carina v-shaped, subapical carina u- shaped; median carina of vertex absent or if present, covering up ¼ of basal compartment of vertex; basal compartment 3.7–9.0 x longer than apical compartment. Frons 1.1–1.4 x longer than wide. Postclypeus with median carina absent or evanescent; lateral carinae absent. Anteclypeus with median and lateral carinae absent. Rostrum very short, reaching mid coxa or mid femur; apical segment of rostrum shorter than or as long as subapical segment.</paragraph>
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||
<paragraph id="8BD45D64FFF1FFC3FF7955F3FC2DFEE5" blockId="2.[151,1437,1305,2014]" lastBlockId="3.[151,1437,151,323]" lastPageId="3" lastPageNumber="92" pageId="2" pageNumber="91">
|
||
Thorax: Forewing 2.7–3.5 x longer than wide; costal margin with about 1–4 tubercles; Sc+R+M near basal cell fused, forming a minute common stem Sc+R+M; fork of ScRA+
|
||
<collectingCountry id="F37C1DF4FFF1FFC2FCC65594FC6EF878" box="[888,923,1990,2014]" name="Poland" pageId="2" pageNumber="91">RP</collectingCountry>
|
||
basad or at same level as fork CuA1+CuA2; position of r-m at same level as or slightly basad of fork MA+MP; MP apically unforked or bifid; vein delimiting subapical cell C4 more or less at same level as vein delimiting C5; subapical cell C5 distinctly longer than C4; CuA2 ending well before it reaches margin of wing or fading (reaching margin of wing with reduced thickness); 7–8 apical cells; 3 anteapical cells.
|
||
<collectingCountry id="F37C1DF4FFF0FFC3FD925354FD91FEB9" box="[556,612,262,287]" name="India" pageId="3" pageNumber="92">Hind</collectingCountry>
|
||
leg: 1st tarsomere with 7 apical teeth (rarely 6 or 8) and no platellae; 2nd tarsomere with 8 (rarely 9) apical teeth and 4 (rarely 5) platellae (
|
||
<figureCitation id="135041E1FFF0FFC3FCCA5378FC42FEE5" box="[884,951,298,323]" captionStart="FIGURE 2" captionStartId="3.[151,250,2006,2028]" captionTargetBox="[151,1395,359,1975]" captionTargetId="figure@3.[151,1436,357,1986]" captionTargetPageId="3" captionText="FIGURE 2. Calamister obscurus: A habitus; B forewing; C – D head; E first and second tarsomere. Scale bar 1 mm." httpUri="https://zenodo.org/record/287358/files/figure.png" pageId="3" pageNumber="92">Fig. 2</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
<caption id="DF140DECFFF0FFC3FF295584FAD3F84A" box="[151,1318,2006,2028]" httpUri="https://zenodo.org/record/287358/files/figure.png" pageId="3" pageNumber="92" targetBox="[151,1395,359,1975]" targetPageId="3">
|
||
<paragraph id="8BD45D64FFF0FFC3FF295584FAD3F84A" blockId="3.[151,1318,2006,2028]" box="[151,1318,2006,2028]" pageId="3" pageNumber="92">
|
||
<emphasis id="B91F8176FFF0FFC3FF295584FEE7F84D" bold="true" box="[151,274,2006,2028]" pageId="3" pageNumber="92">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6B26E7FFF0FFC3FEA75584FE1EF84A" box="[281,491,2006,2028]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B91F8176FFF0FFC3FEA75584FE1EF84A" box="[281,491,2006,2028]" italics="true" pageId="3" pageNumber="92">Calamister obscurus</emphasis>
|
||
</taxonomicName>
|
||
: A habitus; B forewing; C–D head; E first and second tarsomere. Scale bar 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD45D64FFF7FFC4FF7952C5FB66FF5E" blockId="4.[151,1437,151,1049]" pageId="4" pageNumber="93">
|
||
Male genitalia: Anal tube as in
|
||
<figureCitation id="135041E1FFF7FFC4FD9352C5FD80FF16" box="[557,629,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,807,829]" captionTargetBox="[344,1242,198,782]" captionTargetId="figure@6.[338,1249,193,786]" captionTargetPageId="6" captionText="FIGURE 4. Calamister obscurus: A aedeagus ventral; B aedeagus left lateral; C – D anal tube; E – F genital styles." httpUri="https://zenodo.org/record/287360/files/figure.png" pageId="4" pageNumber="93">Figs 4</figureCitation>
|
||
C,D. Pygofer and genital styles as in
|
||
<figureCitation id="135041E1FFF7FFC4FBA252C5FB96FF16" box="[1052,1123,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,807,829]" captionTargetBox="[344,1242,198,782]" captionTargetId="figure@6.[338,1249,193,786]" captionTargetPageId="6" captionText="FIGURE 4. Calamister obscurus: A aedeagus ventral; B aedeagus left lateral; C – D anal tube; E – F genital styles." httpUri="https://zenodo.org/record/287360/files/figure.png" pageId="4" pageNumber="93">Figs 4</figureCitation>
|
||
E,F. Aedeagus (
|
||
<figureCitation id="135041E1FFF7FFC4FAAB52C5FAAAFF16" box="[1301,1375,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,807,829]" captionTargetBox="[344,1242,198,782]" captionTargetId="figure@6.[338,1249,193,786]" captionTargetPageId="6" captionText="FIGURE 4. Calamister obscurus: A aedeagus ventral; B aedeagus left lateral; C – D anal tube; E – F genital styles." httpUri="https://zenodo.org/record/287360/files/figure.png" pageId="4" pageNumber="93">Figs 4</figureCitation>
|
||
A,B): Phallotheca ventrally to right-laterally with two large, curved spines (a, b), about equal in length; spine (a) arising near apex of aedeagus and spine (b) arising right underneath spine (a). Flagellum unarmed.
|
||
</paragraph>
|
||
<paragraph id="8BD45D64FFF7FFC4FF795356FDEDFE99" blockId="4.[151,1437,151,1049]" pageId="4" pageNumber="93">
|
||
Female genitalia: Ovipositor, wax plate and anal tube as in
|
||
<figureCitation id="135041E1FFF7FFC4FC345356FC26FEBB" box="[906,979,260,285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1030,1052]" captionTargetBox="[170,1417,202,1000]" captionTargetId="figure@7.[164,1424,195,1009]" captionTargetPageId="7" captionText="FIGURE 6. Female genitalia. Calamister obscurus: A ventral; B caudal; C dorsal. Calamister cougalensis: D ventral; E caudal; F dorsal." httpUri="https://zenodo.org/record/287362/files/figure.png" pageId="4" pageNumber="93">Figs 6</figureCitation>
|
||
A–C: wax plate oval-shaped; anal tube trapezoid, widening towards apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3710EEFFFF7FFC4FF79531CFC9BFBBF" pageId="4" pageNumber="93" type="discussion">
|
||
<paragraph id="8BD45D64FFF7FFC4FF79531CFEF8FE5A" blockId="4.[151,1437,151,1049]" pageId="4" pageNumber="93">
|
||
<emphasis id="B91F8176FFF7FFC4FF79531CFECEFEC1" bold="true" box="[199,315,334,359]" pageId="4" pageNumber="93">Remarks.</emphasis>
|
||
In general
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FE07531DFDCAFEC1" box="[441,575,335,359]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B91F8176FFF7FFC4FE07531DFDCAFEC1" box="[441,575,335,359]" italics="true" pageId="4" pageNumber="93">C. obscurus</emphasis>
|
||
</taxonomicName>
|
||
has 7 apical teeth and no platellae on the 1st tarsomere and 8 apical teeth and 4 platellae on the 2nd tarsomere. The following deviations from this formula appear to be aberrations as they only occurred on one leg of the specimen: One male from Bundaberg had only 6 and another male from the same location 8 apical teeth on the 1st tarsomere; the female from Coolum had 9 apical teeth and 5 platellae on the 2nd tarsomere.
|
||
</paragraph>
|
||
<paragraph id="8BD45D64FFF7FFC4FF79505AFC9BFBBF" blockId="4.[151,1437,151,1049]" pageId="4" pageNumber="93">
|
||
Kirkaldy’s original descriptions of the genus and species are not very specific and lack illustrations. Muir (1908) provided a dichotomous key to the genera of
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FC8C507EFC64FDE3" box="[818,913,556,581]" class="Insecta" family="Cixiidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="family">Cixiidae</taxonomicName>
|
||
which includes
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FBDC507EFB28FDE2" box="[1122,1245,556,580]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F8176FFF7FFC4FBDC507EFB28FDE2" box="[1122,1245,556,580]" italics="true" pageId="4" pageNumber="93">Calamister</emphasis>
|
||
</taxonomicName>
|
||
. Based on this identification key
|
||
<bibRefCitation id="EFFA2095FFF7FFC4FED45002FDF3FDCF" author="Jacobi" box="[362,518,592,617]" pageId="4" pageNumber="93" refString="Jacobi, A. (1928) Results of Dr E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. Rhynchota, Homoptera. 1. Fulgoridae und Cercopidae. Arkiv for Zoologi, 19 A, 1 - 50." type="journal article" year="1928">Jacobi (1928)</bibRefCitation>
|
||
identified a female specimen from the 'Blackal Range’ (this appears to be the ‘Blackall Range’ in South East Queensland) to be
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FD6F5027FCADFD2A" box="[721,856,628,652]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B91F8176FFF7FFC4FD6F5027FCADFD2A" box="[721,856,628,652]" italics="true" pageId="4" pageNumber="93">C. obscurus</emphasis>
|
||
</taxonomicName>
|
||
. Jacobi mentions features such as a minute stem of Sc+R+M; a long common stem of Sc+R and the fact that fork Sc+R and fork CuA1+CuA2 are at about the same level, situated in the basal half of the forewing; and transverse carina of vertex slightly u-shaped. These features can be found in the
|
||
<typeStatus id="54D0E3C6FFF7FFC4FEC050B3FE5BFD5F" box="[382,430,737,761]" pageId="4" pageNumber="93">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FDE850B3FD28FD5E" box="[598,733,736,760]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B91F8176FFF7FFC4FDE850B3FD28FD5E" box="[598,733,736,760]" italics="true" pageId="4" pageNumber="93">C. obscurus</emphasis>
|
||
</taxonomicName>
|
||
and/or the other material examined.
|
||
<bibRefCitation id="EFFA2095FFF7FFC4FB3950B2FAD7FD5F" author="Jacobi" box="[1159,1314,736,761]" pageId="4" pageNumber="93" refString="Jacobi, A. (1928) Results of Dr E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. Rhynchota, Homoptera. 1. Fulgoridae und Cercopidae. Arkiv for Zoologi, 19 A, 1 - 50." type="journal article" year="1928">Jacobi (1928)</bibRefCitation>
|
||
noted that apical veins (towards the margin) and crossveins were darker brown in the Blackall Range specimen. In
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FA3C5157FEEEFCE6" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="cougalensis" status="sp. nov.">
|
||
<emphasis id="B91F8176FFF7FFC4FA3C5157FEEEFCE6" italics="true" pageId="4" pageNumber="93">C. cougalensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C3C0DFFF7FFC4FE9A517BFE89FCE6" box="[292,380,809,832]" pageId="4" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the apical veins are darker but not the crossveins. In
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FC5C517BFB9CFCE6" box="[994,1129,808,832]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B91F8176FFF7FFC4FC5C517BFB9CFCE6" box="[994,1129,808,832]" italics="true" pageId="4" pageNumber="93">C. obscurus</emphasis>
|
||
</taxonomicName>
|
||
all veins are uniformously coloured throughout (
|
||
<figureCitation id="135041E1FFF7FFC4FE37511EFE25FCC3" box="[393,464,844,869]" captionStart="FIGURE 2" captionStartId="3.[151,250,2006,2028]" captionTargetBox="[151,1395,359,1975]" captionTargetId="figure@3.[151,1436,357,1986]" captionTargetPageId="3" captionText="FIGURE 2. Calamister obscurus: A habitus; B forewing; C – D head; E first and second tarsomere. Scale bar 1 mm." httpUri="https://zenodo.org/record/287358/files/figure.png" pageId="4" pageNumber="93">Figs 2</figureCitation>
|
||
A,B). Jacobi included an illustration of the forewing in his redescription of
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FAA8511FFECCFC2F" authority="Kirkaldy. The" authorityName="Kirkaldy. The" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B91F8176FFF7FFC4FAA8511FFA69FCC2" box="[1302,1436,844,868]" italics="true" pageId="4" pageNumber="93">C. obscurus</emphasis>
|
||
Kirkaldy. The
|
||
</taxonomicName>
|
||
specimen depicted varies considerably from the wing venation found in
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FB0D5122FADAFC2E" box="[1203,1327,880,904]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F8176FFF7FFC4FB0D5122FADAFC2E" box="[1203,1327,880,904]" italics="true" pageId="4" pageNumber="93">Calamister</emphasis>
|
||
</taxonomicName>
|
||
, e.g. the specimen pictured has a trifid
|
||
<collectingCountry id="F37C1DF4FFF7FFC4FE5951C7FDFCFC0A" box="[487,521,917,940]" name="Poland" pageId="4" pageNumber="93">RP</collectingCountry>
|
||
, MA and MP, however in
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FC9051C6FC5EFC0A" box="[814,939,916,940]" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91F8176FFF7FFC4FC9051C6FC5EFC0A" box="[814,939,916,940]" italics="true" pageId="4" pageNumber="93">Calamister</emphasis>
|
||
</taxonomicName>
|
||
these veins are bifid (
|
||
<figureCitation id="135041E1FFF7FFC4FB2151C6FB13FC0B" box="[1183,1254,916,941]" captionStart="FIGURE 1" captionStartId="1.[151,250,1167,1189]" captionTargetBox="[221,1366,724,1138]" captionTargetId="figure@1.[214,1372,719,1142]" captionTargetPageId="1" captionText="FIGURE 1. Calamister cougalensis: forewing." httpUri="https://zenodo.org/record/287357/files/figure.png" pageId="4" pageNumber="93">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="135041E1FFF7FFC4FB4D51C7FAF5FC0B" box="[1267,1280,917,941]" captionStart="FIGURE 3" captionStartId="5.[151,250,1951,1973]" captionTargetBox="[183,1335,193,1919]" captionTargetId="figure@5.[183,1403,193,1930]" captionTargetPageId="5" captionText="FIGURE 3. Calamister cougalensis: A habitus; B forewing with CuA 2, highlighted by arrow; C – D head. Scale bar 1 mm." httpUri="https://zenodo.org/record/287359/files/figure.png" pageId="4" pageNumber="93">3</figureCitation>
|
||
A), or even in some cases MP unforked (
|
||
<figureCitation id="135041E1FFF7FFC4FE7B51EAFDFEFC77" box="[453,523,952,977]" captionStart="FIGURE 2" captionStartId="3.[151,250,2006,2028]" captionTargetBox="[151,1395,359,1975]" captionTargetId="figure@3.[151,1436,357,1986]" captionTargetPageId="3" captionText="FIGURE 2. Calamister obscurus: A habitus; B forewing; C – D head; E first and second tarsomere. Scale bar 1 mm." httpUri="https://zenodo.org/record/287358/files/figure.png" pageId="4" pageNumber="93">Fig. 2</figureCitation>
|
||
B). Further, Jacobi’s specimen shows CuA1 and CuA2 rejoining which does not occur in any of the
|
||
<typeStatus id="54D0E3C6FFF7FFC4FECA518FFE47FC53" box="[372,434,989,1013]" pageId="4" pageNumber="93">types</typeStatus>
|
||
. This means there is considerable doubt that the specimen from the Blackall Range is
|
||
<taxonomicName id="4C6B26E7FFF7FFC4FA3C518FFF09FBBE" class="Insecta" family="Cixiidae" genus="Calamister" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="B91F8176FFF7FFC4FA3C518FFF09FBBE" italics="true" pageId="4" pageNumber="93">C. obscurus</emphasis>
|
||
</taxonomicName>
|
||
—examination of the specimen is needed to clarify this.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |