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<document id="128CA5A923964A6052F0D9AB2532B8B5" ID-DOI="10.26107/RBZ-2019-0041" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576444" ID-ZooBank="AAA866A1-FBAF-486D-AB84-3AB989A269C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614778122966" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Hunyadi, András &amp; Jochum, Adrienne" docDate="2019" docId="03BF447DFF91FFA7FF77FE09FAEFFAD1" docLanguage="en" docName="RafflesBZool.67.517-535.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Tonkinospira raxajacki Pall-Gergely &amp; A. Reischutz 2019, new species" docType="treatment" docVersion="3" lastPageNumber="529" masterDocId="FF863C05FF9DFFABFFEEFA75FFE6FFC4" masterDocTitle="New Tonkinospira Jochum, Slapnik &amp; Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae)" masterLastPageNumber="535" masterPageNumber="517" pageNumber="529" updateTime="1698878161501" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="FBFEA35D57B5AF96F84EC59D366CD5C8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="230350441824C63F123CED911EDB81A7">New Tonkinospira Jochum, Slapnik &amp; Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae)</mods:title>
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<mods:name id="DA1CEE0DFF8B72FE046CFB666AC5803E" type="personal">
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<mods:namePart id="8358643D71CDC541313E85B430214E23">Páll-Gergely, Barna</mods:namePart>
</mods:name>
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<mods:namePart id="0021E9CF6615988F953E3DDBD821959A">Grego, Jozef</mods:namePart>
</mods:name>
<mods:name id="BAB2CB6FFD55CAA653241EF2416CD44B" type="personal">
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</mods:role>
<mods:namePart id="A3F6423D35D066FBA7AA8E3C826597DD">Vermeulen, Jaap J.</mods:namePart>
</mods:name>
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<mods:role id="A6AC8AFAC39725A581A1132497634984">
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<mods:namePart id="B71EFE5ED8EBF846853AAD3A9EC2B96D">Reischütz, Alexander</mods:namePart>
</mods:name>
<mods:name id="9B1E64358126C83A83B9EA511B73E0D9" type="personal">
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</mods:role>
<mods:namePart id="4F1DF857420DD9ADBC1E4343448D1F2F">Hunyadi, András</mods:namePart>
</mods:name>
<mods:name id="850EF36DBA0B4506A3BD2D0ACD955BF9" type="personal">
<mods:role id="0316260C24E1B4AD637BB3B59A4A6950">
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</mods:role>
<mods:namePart id="6B3B7663772A33806724B7B2FADD0B1B">Jochum, Adrienne</mods:namePart>
</mods:name>
<mods:typeOfResource id="D3089EB32464C74934A7CC663E5F29E6">text</mods:typeOfResource>
<mods:relatedItem id="5CED54D5DDC57B1D8543A25241AAEBDC" type="host">
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<mods:title id="1E0FD691FF21CFAF25E796B625BD79F2">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="0C6A8821D5CF42D2C8DE86CF5FF1A803">
<mods:date id="760DCC5C9449807317DB2193173C284B">2019</mods:date>
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<mods:number id="440A438B2455D16171F3902A986A81EA">2019-11-08</mods:number>
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<mods:number id="4F55872DA71CDB959DCA25951F3D5C64">67</mods:number>
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<mods:start id="46FFF2518C489D81D0F68F84CC8DF269">517</mods:start>
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<mods:identifier id="6EAD6902A433745663E26CAA3E73B4AD" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="B41BE845ED8DC35FE290E11CAA73E88E" type="Zenodo-Dep">4576444</mods:identifier>
<mods:identifier id="F915632C4D7A631C9BD836277DBD62D4" type="ZooBank">AAA866A1-FBAF-486D-AB84-3AB989A269C1</mods:identifier>
</mods:mods>
<treatment id="03BF447DFF91FFA7FF77FE09FAEFFAD1" LSID="urn:lsid:plazi:treatment:03BF447DFF91FFA7FF77FE09FAEFFAD1" httpUri="http://treatment.plazi.org/id/03BF447DFF91FFA7FF77FE09FAEFFAD1" lastPageNumber="529" pageId="12" pageNumber="529">
<subSubSection id="C30CA6E0FF91FFA7FF77FE09FDEAFB71" pageId="12" pageNumber="529" type="nomenclature">
<paragraph id="8BA9F56BFF91FFA7FF77FE09FDEAFB71" blockId="12.[153,758,1148,1237]" pageId="12" pageNumber="529">
<heading id="D0E14207FF91FFA7FF77FE09FDEAFB71" bold="true" centered="true" fontSize="10" level="2" pageId="12" pageNumber="529" reason="2">
<emphasis id="B9622979FF91FFA7FF77FE09FDEAFB71" bold="true" pageId="12" pageNumber="529">
<taxonomicName id="4C168EE8FF91FFA7FF77FE09FD17FB51" ID-CoL="7TCTZ" authority="Pall-Gergely &amp; A. Reischutz" authorityName="Pall-Gergely &amp; A. Reischutz" authorityYear="2019" box="[153,753,1148,1173]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="529" phylum="Mollusca" rank="species" species="raxajacki" status="sp. nov.">
<emphasis id="B9622979FF91FFA7FF77FE09FE78FB50" bold="true" box="[153,414,1148,1172]" italics="true" pageId="12" pageNumber="529">Tonkinospira raxajacki</emphasis>
Páll-Gergely &amp; A. Reischütz
</taxonomicName>
,
<taxonomicNameLabel id="A2519402FF91FFA7FE69FEE8FDEAFB71" box="[391,524,1181,1205]" pageId="12" pageNumber="529" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FE76FEC8FE11FB11" box="[408,503,1213,1237]" pageId="12" pageNumber="529" type="description">
<paragraph id="8BA9F56BFF91FFA7FE76FEC8FE11FB11" blockId="12.[153,758,1148,1237]" box="[408,503,1213,1237]" pageId="12" pageNumber="529">
(
<figureCitation id="132DE9EEFF91FFA7FE4EFEC8FE09FB11" box="[416,495,1213,1237]" captionStart="Fig" captionStartId="13.[113,148,2091,2113]" captionTargetBox="[131,1430,189,2067]" captionTargetId="figure-9@13.[131,1430,189,2067]" captionTargetPageId="13" captionText="Fig. 10. Tonkinospira raxajacki Páll-Gergely &amp; A. Reischütz new species, NHMW 113205 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4576478" httpUri="https://zenodo.org/record/4576478/files/figure.png" pageId="12" pageNumber="529">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FF66FE88FAA5FF31" pageId="12" pageNumber="529" type="materials_examined">
<paragraph id="8BA9F56BFF91FFA7FF66FE88FD7CFA71" blockId="12.[135,777,1277,2101]" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FF66FE88FED3FAD1" bold="true" box="[136,309,1277,1301]" pageId="12" pageNumber="529">Type material.</emphasis>
<materialsCitation id="3B7EFF36FF91FFA7FED1FE88FD7CFA71" collectingDate="2010-03" collectionCode="NHMW" collectorName="NE Thakhek &amp; Raxajack, S. &amp; Reischutz, A." country="Laos" elevation="160" latitude="17.45355" location="Thakhek" longLatPrecision="1" longitude="104.90953" pageId="12" pageNumber="529" specimenCode="NHMW 113205" specimenCount="1" stateProvince="Khammouan" typeStatus="holotype">
<typeStatus id="54AD4BC9FF91FFA7FED1FE88FE45FAD1" box="[319,419,1277,1301]" pageId="12" pageNumber="529" type="holotype">Holotype</typeStatus>
(H:
<quantity id="4CEE588EFF91FFA7FE32FE88FDD2FAD0" box="[476,564,1277,1301]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="529" unit="mm" value="1.2">1.2 mm</quantity>
, D:
<quantity id="4CEE588EFF91FFA7FD86FE88FD2BFAD0" box="[616,717,1277,1301]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="12" pageNumber="529" unit="mm" value="1.09">1.09 mm</quantity>
,
<figureCitation id="132DE9EEFF91FFA7FD32FE88FF45FAF1" captionStart="Fig" captionStartId="13.[113,148,2091,2113]" captionTargetBox="[131,1430,189,2067]" captionTargetId="figure-9@13.[131,1430,189,2067]" captionTargetPageId="13" captionText="Fig. 10. Tonkinospira raxajacki Páll-Gergely &amp; A. Reischütz new species, NHMW 113205 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4576478" httpUri="https://zenodo.org/record/4576478/files/figure.png" pageId="12" pageNumber="529">Fig. 10</figureCitation>
) (
<specimenCode id="DBB05D10FF91FFA7FF57FF68FE8BFAF1" box="[185,365,1309,1333]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="529" type="Museum">NHMW 113205</specimenCode>
),
<collectingCountry id="F301B5FBFF91FFA7FE91FF68FE53FAF0" box="[383,437,1309,1332]" name="Laos" pageId="12" pageNumber="529">Laos</collectingCountry>
,
<collectingRegion id="49D23B89FF91FFA7FE2EFF68FD59FAF1" box="[448,703,1309,1333]" country="Laos" name="Khammouan" pageId="12" pageNumber="529">Khammouane Province</collectingRegion>
, Tham Xienliab (cave),
<collectorName id="26E390BDFF91FFA7FED7FF48FE25FA91" box="[313,451,1341,1365]" pageId="12" pageNumber="529">
NE
<location id="8EC9A3B0FF91FFA7FE8DFF48FE25FA91" LSID="urn:lsid:plazi:treatment:03BF447DFF91FFA7FF77FE09FAEFFAD1:8EC9A3B0FF91FFA7FE8DFF48FE25FA91" box="[355,451,1341,1365]" country="Laos" latitude="17.45355" longLatPrecision="1" longitude="104.90953" name="Thakhek" pageId="12" pageNumber="529" stateProvince="Khammouan">Thakhek</location>
</collectorName>
, at the base of limestone rocks at the entrance of the cave, ca.
<elevation id="003B1258FF91FFA7FE06FF28FD8DFAB1" box="[488,619,1373,1397]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="529" unit="m" value="160.0">
<quantity id="4CEE588EFF91FFA7FE06FF28FDC9FAB0" box="[488,559,1373,1397]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="529" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
, approx. GPS coordinates:
<geoCoordinate id="EE2293ACFF91FFA7FEE2FF0EFE75FA50" box="[268,403,1403,1428]" degrees="17" direction="north" minutes="27.213" orientation="latitude" pageId="12" pageNumber="529" precision="1" value="17.45355">17°27.213N</geoCoordinate>
,
<geoCoordinate id="EE2293ACFF91FFA7FE72FF0EFDCDFA50" box="[412,555,1403,1428]" degrees="104" direction="east" minutes="54.572" orientation="longitude" pageId="12" pageNumber="529" precision="1" value="104.90953">104°54.572E</geoCoordinate>
(locality code: La.5), coll.
<collectorName id="26E390BDFF91FFA7FF51FFE8FEA1FA71" box="[191,327,1437,1461]" pageId="12" pageNumber="529">Raxajack, S.</collectorName>
&amp;
<collectorName id="26E390BDFF91FFA7FE83FFE8FDE4FA71" box="[365,514,1437,1461]" pageId="12" pageNumber="529">Reischütz, A.</collectorName>
,
<date id="FFA8D3ABFF91FFA7FDE1FFE8FD70FA71" box="[527,662,1437,1461]" pageId="12" pageNumber="529" value="2010-03">
<collectingDate id="EFEC2A43FF91FFA7FDE1FFE8FD70FA71" box="[527,662,1437,1461]" pageId="12" pageNumber="529" value="2010-03">March 2010</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA9F56BFF91FFA7FF66FFC8FAA5FF31" blockId="12.[135,777,1277,2101]" lastBlockId="12.[835,1475,188,245]" pageId="12" pageNumber="529">
<materialsCitation id="3B7EFF36FF91FFA7FF66FFC8FEDEF971" collectingDate="2007-11-24" collectorName="A. Abdou &amp; I. V. Muratov" country="Laos" county="Central" elevation="180" latitude="17.449284" location="Muang" longLatPrecision="1" longitude="105.113014" municipality="Thakhek" pageId="12" pageNumber="529" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
<typeStatus id="54AD4BC9FF91FFA7FF66FFC8FF1DFA11" box="[136,251,1469,1493]" pageId="12" pageNumber="529" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D103EE2FF91FFA7FEE4FFC8FE63FA11" box="[266,389,1469,1493]" pageId="12" pageNumber="529" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(MNHN-IM-2014-6406), South-
<collectingCounty id="62C88DE7FF91FFA7FF66FFA8FF3DFA31" box="[136,219,1501,1525]" pageId="12" pageNumber="529">Central</collectingCounty>
<collectingCountry id="F301B5FBFF91FFA7FF09FFA8FEC6FA30" box="[231,288,1501,1524]" name="Laos" pageId="12" pageNumber="529">Laos</collectingCountry>
,
<collectingRegion id="49D23B89FF91FFA7FEDEFFA8FDA7FA31" box="[304,577,1501,1525]" country="Laos" name="Khammouan" pageId="12" pageNumber="529">Khammouane Province</collectingRegion>
, ca.
<quantity id="4CEE588EFF91FFA7FD92FFA8FD20FA31" box="[636,710,1501,1525]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="529" unit="km" value="32.0">32 km</quantity>
ENE of
<collectingMunicipality id="6BCD6F11FF91FFA7FF45FF88FEEDF9D1" box="[171,267,1533,1557]" pageId="12" pageNumber="529">Thakhek</collectingMunicipality>
(
<location id="8EC9A3B0FF91FFA7FEF3FF88FE8BF9D1" LSID="urn:lsid:plazi:treatment:03BF447DFF91FFA7FF77FE09FAEFFAD1:8EC9A3B0FF91FFA7FEF3FF88FE8BF9D1" box="[285,365,1533,1557]" country="Laos" county="Central" latitude="17.449284" longLatPrecision="1" longitude="105.113014" municipality="Thakhek" name="Muang" pageId="12" pageNumber="529" stateProvince="Khammouan">Muang</location>
<collectingRegion id="49D23B89FF91FFA7FE96FF88FDEDF9D1" box="[376,523,1533,1557]" country="Laos" name="Khammouan" pageId="12" pageNumber="529">Khammouan</collectingRegion>
), ca.
<locationDeviation id="9A1BEC49FF91FFA7FDA2FF88FF0FF9F1" location="Mahaxai" pageId="12" pageNumber="529">
<quantity id="4CEE588EFF91FFA7FDA2FF88FD73F9D1" box="[588,661,1533,1557]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="529" unit="km" value="10.0">10 km</quantity>
WNW of Mahaxai
</locationDeviation>
, limestone, clay, black soil in limestone pockets, on and under rocks in dry secondary forest on steep N exposed slope under cliff,
<elevation id="003B1258FF91FFA7FEA2FC28FE28F9B1" box="[332,462,1629,1653]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="529" unit="m" value="180.0">
<quantity id="4CEE588EFF91FFA7FEA2FC28FE75F9B0" box="[332,403,1629,1653]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="529" unit="m" value="180.0">180 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE2293ACFF91FFA7FE33FC2EFD81F9B0" box="[477,615,1627,1652]" degrees="17" direction="north" minutes="26.957" orientation="latitude" pageId="12" pageNumber="529" precision="1" value="17.449284">17°26.957N</geoCoordinate>
,
<geoCoordinate id="EE2293ACFF91FFA7FD9BFC2EFCE1F9B0" box="[629,775,1627,1652]" degrees="105" direction="east" minutes="06.781" orientation="longitude" pageId="12" pageNumber="529" precision="1" value="105.113014">105°06.781E</geoCoordinate>
(locality code: 1L07), coll.
<collectorName id="26E390BDFF91FFA7FE56FC08FDC3F951" box="[440,549,1661,1685]" pageId="12" pageNumber="529">A. Abdou</collectorName>
&amp;
<collectorName id="26E390BDFF91FFA7FDA3FC08FD39F950" box="[589,735,1661,1684]" pageId="12" pageNumber="529">I.V. Muratov</collectorName>
,
<date id="FFA8D3ABFF91FFA7FD02FC08FEDEF971" pageId="12" pageNumber="529" value="2007-11-24">
<collectingDate id="EFEC2A43FF91FFA7FD02FC08FEDEF971" pageId="12" pageNumber="529" value="2007-11-24">24 November 2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B7EFF36FF91FFA7FEADFCE8FEA5F8F1" collectingDate="2010-03" collectorName="La. &amp; Raxajack, S. &amp; Reischutz, A." country="Laos" elevation="180" latitude="17.46145" location="East of Thakhek" longLatPrecision="1" longitude="104.91585" municipality="Tham Pha Inh" pageId="12" pageNumber="529" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
<specimenCount id="9D103EE2FF91FFA7FEADFCE8FE55F971" box="[323,435,1693,1717]" pageId="12" pageNumber="529" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(coll. RE),
<collectingCountry id="F301B5FBFF91FFA7FDDEFCE8FD80F970" box="[560,614,1693,1716]" name="Laos" pageId="12" pageNumber="529">Laos</collectingCountry>
,
<collectingRegion id="49D23B89FF91FFA7FD9FFCE8FF0CF911" country="Laos" name="Khammouan" pageId="12" pageNumber="529">Khammouane Province</collectingRegion>
,
<collectingMunicipality id="6BCD6F11FF91FFA7FF1BFCC8FE68F911" box="[245,398,1725,1749]" pageId="12" pageNumber="529">Tham Pha Inh</collectingMunicipality>
(cave),
<location id="8EC9A3B0FF91FFA7FE08FCC8FD7FF911" LSID="urn:lsid:plazi:treatment:03BF447DFF91FFA7FF77FE09FAEFFAD1:8EC9A3B0FF91FFA7FE08FCC8FD7FF911" box="[486,665,1725,1749]" country="Laos" latitude="17.46145" longLatPrecision="1" longitude="104.91585" municipality="Tham Pha Inh" name="East of Thakhek" pageId="12" pageNumber="529" stateProvince="Khammouan">East of Thakhek</location>
, ca.
<elevation id="003B1258FF91FFA7FD24FCC8FF5CF931" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="529" unit="m" value="180.0">
<quantity id="4CEE588EFF91FFA7FD24FCC8FCE1F911" box="[714,775,1725,1749]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="529" unit="m" value="180.0">180m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE2293ACFF91FFA7FF24FCAEFEB2F930" box="[202,340,1755,1780]" degrees="17" direction="north" minutes="27.687" orientation="latitude" pageId="12" pageNumber="529" precision="1" value="17.46145">17°27.687N</geoCoordinate>
,
<geoCoordinate id="EE2293ACFF91FFA7FE8CFCAEFE10F930" box="[354,502,1755,1780]" degrees="104" direction="east" minutes="54.951" orientation="longitude" pageId="12" pageNumber="529" precision="1" value="104.91585">104°54.951E</geoCoordinate>
, (locality code:
<collectorName id="26E390BDFF91FFA7FD44FCA8FD36F931" box="[682,720,1757,1781]" pageId="12" pageNumber="529">La.</collectorName>
7), at the base of limestone rocks, coll.
<collectorName id="26E390BDFF91FFA7FE00FC88FD92F8D1" box="[494,628,1789,1813]" pageId="12" pageNumber="529">Raxajack, S.</collectorName>
&amp;
<collectorName id="26E390BDFF91FFA7FD79FC88FF43F8F1" pageId="12" pageNumber="529">Reischütz, A.</collectorName>
,
<date id="FFA8D3ABFF91FFA7FF5BFD68FEA5F8F1" box="[181,323,1821,1845]" pageId="12" pageNumber="529" value="2010-03">
<collectingDate id="EFEC2A43FF91FFA7FF5BFD68FEA5F8F1" box="[181,323,1821,1845]" pageId="12" pageNumber="529" value="2010-03">March 2010</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B7EFF36FF91FFA7FEBCFD68FE2FF831" collectingDate="2007-12-09" collectorName="A. Abdou &amp; I. V. Muratov" country="Laos" elevation="160" latitude="17.405666" location="Muang" longLatPrecision="1" longitude="104.9149" municipality="South-Central" pageId="12" pageNumber="529" specimenCount="10" stateProvince="Khammouan" typeStatus="paratype">
<specimenCount id="9D103EE2FF91FFA7FEBCFD68FE04F8F1" box="[338,482,1821,1845]" pageId="12" pageNumber="529" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
(MNHN-IM-2014-6405),
<collectingMunicipality id="6BCD6F11FF91FFA7FF66FD48FEC6F891" box="[136,288,1853,1877]" pageId="12" pageNumber="529">South-Central</collectingMunicipality>
<collectingCountry id="F301B5FBFF91FFA7FEC4FD48FE87F890" box="[298,353,1853,1876]" name="Laos" pageId="12" pageNumber="529">Laos</collectingCountry>
,
<collectingRegion id="49D23B89FF91FFA7FE81FD48FD92F891" box="[367,628,1853,1877]" country="Laos" name="Khammouan" pageId="12" pageNumber="529">Khammouane Province</collectingRegion>
, ca.
<locationDeviation id="9A1BEC49FF91FFA7FD45FD48FEF9F8B1" location="Thakhek" pageId="12" pageNumber="529">
<quantity id="4CEE588EFF91FFA7FD45FD48FCE0F891" box="[683,774,1853,1877]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="12" pageNumber="529" unit="km" value="10.5">10.5 km</quantity>
E of Thakhek
</locationDeviation>
(
<location id="8EC9A3B0FF91FFA7FEDFFD28FE98F8B1" LSID="urn:lsid:plazi:treatment:03BF447DFF91FFA7FF77FE09FAEFFAD1:8EC9A3B0FF91FFA7FEDFFD28FE98F8B1" box="[305,382,1885,1909]" country="Laos" latitude="17.405666" longLatPrecision="1" longitude="104.9149" municipality="South-Central" name="Muang" pageId="12" pageNumber="529" stateProvince="Khammouan">Muang</location>
<collectingRegion id="49D23B89FF91FFA7FE66FD28FDF0F8B1" box="[392,534,1885,1909]" country="Laos" name="Khammouan" pageId="12" pageNumber="529">Khammouan</collectingRegion>
), on and under rocks, cave deposits, in secondary forest under entrance and in large cave on NE exposed steep slope, alt.
<quantity id="4CEE588EFF91FFA7FDCCFDE8FD8CF870" box="[546,618,1949,1973]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="529" unit="m" value="160.0">
<elevation id="003B1258FF91FFA7FDCCFDE8FD8CF870" box="[546,618,1949,1973]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="529" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<geoCoordinate id="EE2293ACFF91FFA7FD99FDEEFCE7F870" box="[631,769,1947,1972]" degrees="17" direction="north" minutes="24.340" orientation="latitude" pageId="12" pageNumber="529" precision="1" value="17.405666">17°24.340N</geoCoordinate>
,
<geoCoordinate id="EE2293ACFF91FFA7FF66FDCEFEFDF810" box="[136,283,1979,2004]" degrees="104" direction="east" minutes="54.894" orientation="longitude" pageId="12" pageNumber="529" precision="1" value="104.9149">104°54.894E</geoCoordinate>
(locality code: 25L07), coll.
<collectorName id="26E390BDFF91FFA7FDBCFDC8FD5AF811" box="[594,700,1981,2005]" pageId="12" pageNumber="529">A. Abdou</collectorName>
&amp;
<collectorName id="26E390BDFF91FFA7FD33FDC8FF01F830" pageId="12" pageNumber="529">I.V. Muratov</collectorName>
,
<date id="FFA8D3ABFF91FFA7FF1AFDA8FE2FF831" box="[244,457,2013,2037]" pageId="12" pageNumber="529" value="2007-12-09">
<collectingDate id="EFEC2A43FF91FFA7FF1AFDA8FE2FF831" box="[244,457,2013,2037]" pageId="12" pageNumber="529" value="2007-12-09">09 December 2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B7EFF36FF91FFA7FE39FDA8FAA6FF31" collectingDate="2017-02-17" collectionCode="JG" collectorName="J. Grego" country="Laos" county="Nong Long" latitude="18.1149" location="Spring" longLatPrecision="1" longitude="105.05154" municipality="Phou Phako" pageId="12" pageNumber="529" specimenCount="1" stateProvince="Bolikhamsay Province" typeStatus="paratype">
<specimenCount id="9D103EE2FF91FFA7FE39FDA8FDACF831" box="[471,586,2013,2037]" pageId="12" pageNumber="529" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(coll.
<collectionCode id="ED076DAEFF91FFA7FD7BFDA8FD50F831" box="[661,694,2013,2037]" pageId="12" pageNumber="529">JG</collectionCode>
),
<collectingCountry id="F301B5FBFF91FFA7FD22FDA8FCE4F830" box="[716,770,2013,2036]" name="Laos" pageId="12" pageNumber="529">Laos</collectingCountry>
,
<collectingRegion id="49D23B89FF91FFA7FF66FD88FE66F7D1" box="[136,384,2045,2069]" inferredBySuffix="true" pageId="12" pageNumber="529">Bolikhamsay Province</collectingRegion>
,
<collectingCounty id="62C88DE7FF91FFA7FE62FD88FDEEF7D1" box="[396,520,2045,2069]" pageId="12" pageNumber="529">Nong Long</collectingCounty>
<locationDeviation id="9A1BEC49FF91FFA7FDFFFD88FCE4F7D1" box="[529,770,2045,2069]" location="Lak Sao" pageId="12" pageNumber="529">
<quantity id="4CEE588EFF91FFA7FDFFFD88FDB0F7D1" box="[529,598,2045,2069]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="529" unit="km" value="10.0">10 km</quantity>
SE of Lak Sao
</locationDeviation>
,
<collectingMunicipality id="6BCD6F11FF91FFA7FF64F268FEEAF7F1" box="[138,268,2077,2101]" pageId="12" pageNumber="529">Phou Phakô</collectingMunicipality>
massif
<quantity id="4CEE588EFF91FFA7FE8CF268FE77F7F1" box="[354,401,2077,2101]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="529" unit="km" value="2.0">2km</quantity>
E of the village,
<location id="8EC9A3B0FF91FFA7FDBEF268FD71F7F1" LSID="urn:lsid:plazi:treatment:03BF447DFF91FFA7FF77FE09FAEFFAD1:8EC9A3B0FF91FFA7FDBEF268FD71F7F1" box="[592,663,2077,2101]" country="Laos" county="Nong Long" latitude="18.1149" longLatPrecision="1" longitude="105.05154" municipality="Phou Phako" name="Spring" pageId="12" pageNumber="529" stateProvince="Bolikhamsay Province">Spring</location>
and caves at NW foot of the massif, in front of the quarry,
<geoCoordinate id="EE2293ACFF91FFA7FAACFAC9FA26FF11" box="[1346,1472,188,213]" degrees="18" direction="north" minutes="6.894" orientation="latitude" pageId="12" pageNumber="529" precision="1" value="18.1149">18°6.894N</geoCoordinate>
,
<geoCoordinate id="EE2293ACFF91FFA7FCADFAA9FC2CFF31" box="[835,970,220,245]" degrees="105" direction="east" minutes="3.092" orientation="longitude" pageId="12" pageNumber="529" precision="1" value="105.05154">105°3.092E</geoCoordinate>
, coll.
<collectorName id="26E390BDFF91FFA7FBE1FAABFB8BFF31" box="[1039,1133,221,245]" pageId="12" pageNumber="529">J. Grego</collectorName>
,
<date id="FFA8D3ABFF91FFA7FB94FAA8FAA6FF31" box="[1146,1344,221,245]" pageId="12" pageNumber="529" value="2017-02-17">
<collectingDate id="EFEC2A43FF91FFA7FB94FAA8FAA6FF31" box="[1146,1344,221,245]" pageId="12" pageNumber="529" value="2017-02-17">17 February 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FCADFB68FA85FE51" pageId="12" pageNumber="529" type="diagnosis">
<paragraph id="8BA9F56BFF91FFA7FCADFB68FA85FE51" blockId="12.[834,1474,285,405]" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FCADFB68FC5EFEF1" bold="true" box="[835,952,285,309]" pageId="12" pageNumber="529">Diagnosis.</emphasis>
A small
<taxonomicName id="4C168EE8FF91FFA7FBF3FB68FB4DFEF1" authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" box="[1053,1195,285,309]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="529" phylum="Mollusca" rank="genus">
<emphasis id="B9622979FF91FFA7FBF3FB68FB4DFEF1" box="[1053,1195,285,309]" italics="true" pageId="12" pageNumber="529">Tonkinospira</emphasis>
</taxonomicName>
species with an elongateconic shell, rounded body whorl, irregularly growing whorls, conspicuously deep suture, dense spiral striation, and a peristome that is not adnate to penultimate whorl.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FCADFBC8FC71FC71" pageId="12" pageNumber="529" type="description">
<paragraph id="8BA9F56BFF91FFA7FCADFBC8FB16FC91" blockId="12.[835,1476,445,853]" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FCADFBC8FC30FE11" bold="true" box="[835,982,445,469]" pageId="12" pageNumber="529">Description.</emphasis>
Shell small for the genus; elongate-conic with rounded body whorl; 4.254.5 whorls, irregularly increasing; suture remarkably deep; protoconch consists of 1.25 whorls, finely rugose, and densely, strongly spirally striated; teleoconch reticulated with irregular, rough radial growth lines, and dense, rather regular spiral striation, ca.
<quantity id="4CEE588EFF91FFA7FCADF808FC9AFD51" box="[835,892,637,661]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="529" unit="in" value="30.0">30 in</quantity>
number in the middle shell line from apertural view; aperture oblique to shell axis from lateral view; aperture ovate-subquadrate; peristome not or very slightly expanded, not reflected; aperture not adnate to penultimate whorl and slightly separated (ventral view) from the body whorl; umbilicus deep, slightly smaller than 1/3 the diameter of the base and not covered by peristome.
</paragraph>
<paragraph id="8BA9F56BFF91FFA7FCADF908FC71FC71" blockId="12.[835,1474,893,949]" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FCADF908FBB9FC51" bold="true" box="[835,1119,893,917]" pageId="12" pageNumber="529">Measurements (in mm).</emphasis>
H = 1.141.22, D = 1.091.21 (n = 4).
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FCADF9A8FBB8FBB1" pageId="12" pageNumber="529" type="diagnosis">
<paragraph id="8BA9F56BFF91FFA7FCADF9A8FBB8FBB1" blockId="12.[835,1474,989,1141]" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FCADF9A8FB87FC31" bold="true" box="[835,1121,989,1013]" pageId="12" pageNumber="529">Differential diagnosis.</emphasis>
The most similar species is
<taxonomicName id="4C168EE8FF91FFA7FCABF988FB8DFBD1" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" box="[837,1131,1021,1045]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="529" phylum="Mollusca" rank="species" species="pauperrima">
<emphasis id="B9622979FF91FFA7FCABF988FB8DFBD1" box="[837,1131,1021,1045]" italics="true" pageId="12" pageNumber="529">Tonkinospira pauperrima</emphasis>
</taxonomicName>
, which is larger, has a more oblique aperture to the shell axis, a narrower umbilicus (the peristome partly covers the umbilicus), and has more regularly growing whorls.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FCADFEE8FAEFFAD1" pageId="12" pageNumber="529" type="etymology">
<paragraph id="8BA9F56BFF91FFA7FCADFEE8FAEFFAD1" blockId="12.[834,1474,1181,1301]" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FCADFEE8FC22FB71" bold="true" box="[835,964,1181,1205]" pageId="12" pageNumber="529">Etymology.</emphasis>
We dedicate this species to Somsamay Raxajack for enduring A. Reischütz for seven weeks during a Southeast Asia trip and in gratitude for the outstanding hospitality of his familys native village Ban Pak Nam.
</paragraph>
</subSubSection>
</treatment>
</document>