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<document id="CD940908C9CAE09A431ADC641DE4CFF4" ID-DOI="10.11646/zootaxa.5016.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222210" ID-ZooBank="6E3BF843-16C1-4B91-AB72-D5C1D556384A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629379786491" checkinUser="plazi" docAuthor="Dick, Matthew H., Grischenko, Andrei V., Gordon, Dennis P. &amp; Ostrovsky, Andrew N." docDate="2021" docId="03BCE061A16EFFC3B790FA035A59FD81" docLanguage="en" docName="zootaxa.5016.3.2.pdf" docOrigin="Zootaxa 5016 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Juxtacribrilina pushkini Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky 2021, n. sp." docType="treatment" docUuid="CDED1A27-C2BC-4867-9F62-E395036D70F5" docUuidSource="ZooBank" docVersion="4" lastPageNumber="352" masterDocId="FF859819A161FFD0B707FFA55E06FFE0" masterDocTitle="The “ Cribrilina annulata &quot; problem and new species of Juxtacribrilina (Bryozoa Cheilostomata: Cribrilinidae) from the North Pacific" masterLastPageNumber="364" masterPageNumber="333" pageNumber="348" updateTime="1699070475915" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B413447D1F18678D81B8E1CAF2860220">The “ Cribrilina annulata &quot; problem and new species of Juxtacribrilina (Bryozoa Cheilostomata: Cribrilinidae) from the North Pacific</mods:title>
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<mods:namePart id="7B4E670F79669A2BADB60DA450091953">Dick, Matthew H.</mods:namePart>
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<mods:affiliation id="16E5BBF4453558700761C9311B7AC31C">Department of Biological Sciences, Faculty of Science, Hokkaido University, N 10 W 8, Kita-ku, Sapporo 060 - 0810, Japan. dickmatthewh @ gmail. com; https: // orcid. org / 0000 - 0001 - 5125 - 5159</mods:affiliation>
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<mods:namePart id="7613F2B84EFD035F2970630526CEAF56">Grischenko, Andrei V.</mods:namePart>
<mods:affiliation id="DCE375171415042D708CF2201820F760">Department of Invertebrate Zoology and Aquatic Ecology, Biological Faculty, Perm State National Research University, Bukirev Street 15, GSP, Perm 614990, Russia. &amp; A. V. Zhirmunsky National Scientific Center of Marine Biology, Far East Branch, Russian Academy of Sciences, Palchevskogo Street 17, Vladivostok 690041, Russia.</mods:affiliation>
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<mods:namePart id="F3A48C209BFE4E4524394009B3F2857D">Gordon, Dennis P.</mods:namePart>
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<mods:affiliation id="3FCDC9A0DB4B3E22BA16C8AC7C916A80">National Institute of Water and Atmospheric Research, Private Bag 14901 Kilbirnie, Wellington 6022, New Zealand. dennis. gordon @ niwa. co. nz; http: // orcid. org / 0000 - 0001 - 9398 - 996 X</mods:affiliation>
<mods:nameIdentifier id="BC298D5D3737157F68491982024BFFA6" type="email">dennis.gordon@niwa.co.nz</mods:nameIdentifier>
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<mods:namePart id="3FC75717A71ADEF3C28E639B1E21FB8F">Ostrovsky, Andrew N.</mods:namePart>
<mods:affiliation id="FD6FEBF4D177417C0997F5C8C9AAEAD9">Department of Invertebrate Zoology, Faculty of Biology, Saint Petersburg State University, Universitetskaja nab. 7 / 9, Saint Petersburg 199034, Russia. &amp; Department of Palaeontology, Faculty of Earth Sciences, Geography and Astronomy, University of Vienna, Althanstr. 14, Vienna 1090, Austria.</mods:affiliation>
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<mods:date id="7BA5067201408BBFC7BD58397670C5F7">2021</mods:date>
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<heading id="D0E2E61BA16EFFDFB790FA035C0FFA21" bold="true" box="[151,521,1446,1473]" fontSize="11" level="1" pageId="15" pageNumber="348" reason="1">
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<taxonomicName id="4C152AF4A16EFFDFB790FA035FB9FA21" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[151,447,1446,1473]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="348" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A16EFFDFB790FA035FB9FA21" bold="true" box="[151,447,1446,1473]" italics="true" pageId="15" pageNumber="348">Juxtacribrilina pushkini</emphasis>
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<paragraph id="8BAA5177A16EFFDFB790FA6E5F7CFA05" blockId="15.[151,931,1446,1544]" box="[151,378,1483,1509]" pageId="15" pageNumber="348">
(
<figureCitation id="132E4DF2A16EFFDFB798FA6E5EEEFA05" box="[159,232,1483,1509]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="15" pageNumber="348">Figs 9</figureCitation>
,
<figureCitation id="132E4DF2A16EFFDFB7F3FA6E5F15FA05" box="[244,275,1483,1509]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="15" pageNumber="348">10</figureCitation>
;
<tableCitation id="C69764CCA16EFFDFB619FA6E5F77FA05" box="[286,369,1483,1509]" captionStart="TABLE 3" captionStartId="16.[151,239,476,500]" captionText="TABLE 3. Measurements in micrometers and counts (presented as average and range of values) for Juxtacribrilina pushkini n. sp.; intertidal specimens from Ketchikan, Alaska; square brackets indicate sample sizes other than given at top." pageId="15" pageNumber="348">Table 3</tableCitation>
)
</paragraph>
<paragraph id="8BAA5177A16EFFDFB790FA555DA5F9E8" blockId="15.[151,931,1446,1544]" box="[151,931,1520,1544]" pageId="15" pageNumber="348">
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<paragraph id="8BAA5177A16EFFDFB790F9965DE4F9AB" blockId="15.[151,994,1587,1612]" box="[151,994,1587,1612]" pageId="15" pageNumber="348">
<treatmentCitationGroup id="AB057659A16EFFDFB790F9965DE4F9AB" box="[151,994,1587,1612]" pageId="15" pageNumber="348">
<treatmentCitation id="0AB47766A16EFFDFB790F9965C6CF9AB" author="Dick, M. H. &amp; Grischenko, A. V. &amp; Mawatari, S. F." box="[151,618,1587,1611]" page="3718" pageId="15" pageNumber="348" year="2005">
<taxonomicName id="4C152AF4A16EFFDFB790F9965C6CF9AB" ID-CoL="84Z6S" authority=": Dick et al. 2005, p. 3718" authorityName="Dick" authorityPageNumber="3718" authorityYear="2005" box="[151,618,1587,1611]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="348" phylum="Bryozoa" rank="species" species="annulata">
<emphasis id="B9618D65A16EFFDFB790F9965F58F9AB" box="[151,350,1587,1611]" italics="true" pageId="15" pageNumber="348">Cribrilina annulata</emphasis>
:
<bibRefCitation id="EF842C86A16EFFDFB66BF9965C17F9AB" author="Dick, M. H. &amp; Grischenko, A. V. &amp; Mawatari, S. F." box="[364,529,1587,1611]" pageId="15" pageNumber="348" pagination="3687 - 3784" refId="ref21456" refString="Dick, M. H., Grischenko, A. V. &amp; Mawatari, S. F. (2005) Intertidal Bryozoa (Cheilostomata) of Ketchikan, Alaska. Journal of Natural History, 39 (43), 3687 - 3784. https: // doi. org / 10.1080 / 00222930500415195" type="journal article" year="2005">
Dick
<emphasis id="B9618D65A16EFFDFB6A4F9915FD1F9AB" box="[419,471,1587,1611]" italics="true" pageId="15" pageNumber="348">et al.</emphasis>
2005
</bibRefCitation>
, p. 3718
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, fig. 7A, B;
<bibRefCitation id="EF842C86A16EFFDFB5E1F9915D89F9AB" author="Yang, H. J. &amp; Seo, J. &amp; Min, B. S. &amp; Grischenko, A. V. &amp; Gordon, D. P." box="[742,911,1587,1612]" pageId="15" pageNumber="348" pagination="216 - 234" refId="ref23909" refString="Yang, H. J., Seo, J. E, Min, B. S., Grischenko, A. V. &amp; Gordon, D. P. (2018) Cribrilinidae (Bryozoa: Cheilostomata) of Korea. Zootaxa, 4377 (2), 216 - 234. https: // doi. org / 10.11646 / zootaxa. 4377.2.4" type="journal article" year="2018">
Yang
<emphasis id="B9618D65A16EFFDFB426F9915D52F9AB" box="[801,852,1587,1611]" italics="true" pageId="15" pageNumber="348">et al.</emphasis>
2018
</bibRefCitation>
, fig. 13.
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</paragraph>
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<subSubSection id="C30F02FCA16EFFDFB790F9D75CF1F869" pageId="15" pageNumber="348" type="diagnosis">
<paragraph id="8BAA5177A16EFFDFB790F9D75CF1F869" blockId="15.[151,1437,1650,2037]" pageId="15" pageNumber="348">
<emphasis id="B9618D65A16EFFDFB790F9D75F14F96C" bold="true" box="[151,274,1650,1676]" pageId="15" pageNumber="348">Diagnosis.</emphasis>
Non-ovicellate zooids with 1419 costae, 35 oral spines (usually 3 or 4); intercostal lacunae small,
<quantity id="4CEDFC92A16EFFDFB790F9325EE9F951" box="[151,239,1687,1713]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.54" metricValueMin="1.524" pageId="15" pageNumber="348" unit="in" value="8.0" valueMax="10.0" valueMin="6.0">610 in</quantity>
transverse series. One or two frontal pore chambers on proximal gymnocyst of most zooids. Ovicellate zooids (including ooecium) roughly same size as non-ovicellate zooids occurring but uncommon in basal layer. Reduced or dwarf marginal ovicellate zooids occurring; frontal dwarfs often abundant in colony center. Frontal shield of dwarf zooids as large as or larger than ooecial complex, with 5 or 6 costae. Ooecium in zooids in basal layer moderately reduced, cap-like, with several irregular pseudopores; ooecium in dwarf zooids typically with 2 transversely arranged, oval or irregular pseudopores, with additional pair of smaller pseudopores sometimes present. Ancestrula cribriform, with 4 or 5 oral spines.
</paragraph>
</subSubSection>
<subSubSection id="C30F02FCA16EFFDFB7C0F8365D71F815" pageId="15" pageNumber="348" type="etymology">
<paragraph id="8BAA5177A16EFFDFB7C0F8365D71F815" blockId="15.[151,1437,1650,2037]" pageId="15" pageNumber="348">
<emphasis id="B9618D65A16EFFDFB7C0F8365F48F84D" bold="true" box="[199,334,1939,1965]" pageId="15" pageNumber="348">Etymology.</emphasis>
Named for Dr. Alexander Fedorovitch Pushkin (19382019), research associate at the Zoological Institute, Russian Academy of Sciences, zoologist and hydrobiologist, working mostly on Arctic and Antarctic benthic invertebrates and a leading world expert on Pycnogonida.
</paragraph>
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<emphasis id="B9618D65A171FFC0B7C0FF335FB7FF51" bold="true" box="[199,433,150,177]" pageId="16" pageNumber="349">Material examined.</emphasis>
<materialsCitation id="3B7D5B2AA171FFC0B6BEFF325B1DFF35" collectingDate="2003-09-09" collectingDateMax="2003-09-12" collectingDateMin="2003-09-09" collectionCode="MLWL" elevation="5" location="Ketchikan" pageId="16" pageNumber="349" specimenCount="1" stateProvince="Alaska" typeStatus="holotype">
<emphasis id="B9618D65A171FFC0B6BEFF325C26FF51" box="[441,544,151,177]" italics="true" pageId="16" pageNumber="349">
<typeStatus id="54AEEFD5A171FFC0B6BEFF325C26FF51" box="[441,544,151,177]" pageId="16" pageNumber="349" type="holotype">Holotype</typeStatus>
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: YPM-IZ-100361, vicinity of
<location id="8ECA07ACA171FFC0B47FFF325DE8FF51" LSID="urn:lsid:plazi:treatment:03BCE061A16EFFC3B790FA035A59FD81:8ECA07ACA171FFC0B47FFF325DE8FF51" box="[888,1006,151,177]" name="Ketchikan" pageId="16" pageNumber="349" stateProvince="Alaska">Ketchikan</location>
,
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,
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<collectingDate id="EFEF8E5FA171FFC0B351FF325B56FF51" box="[1110,1360,151,177]" pageId="16" pageNumber="349" value="2003-09-09" valueMax="2003-09-12" valueMin="2003-09-09">912 September 2003</collectingDate>
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, lower rocky intertidal, -0.46 to -
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<elevation id="0038B644A171FFC0B6ADFF1E5FF1FF35" box="[426,503,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.9" pageId="16" pageNumber="349" unit="m" value="0.49">0.49 m</elevation>
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relative to mean low water level (
<collectionCode id="ED04C9B2A171FFC0B458FF1E5DBFFF35" box="[863,953,187,213]" pageId="16" pageNumber="349">MLWL</collectionCode>
); on rock, mounted on SEM stub
</materialsCitation>
.
<materialsCitation id="3B7D5B2AA171FFC0B224FF1E5A1FFE85" collectingDate="2003-09-09" collectingDateMax="2003-09-12" collectingDateMin="2003-09-09" collectionCode="MLWL" county="East Tongass Narrows" elevation="5" latitude="55.51161" location="Settlers Cove" longLatPrecision="1" longitude="-131.72517" municipality="Ketchikan" pageId="16" pageNumber="349" specimenCount="2" stateProvince="Alaska" typeStatus="paratype">
<emphasis id="B9618D65A171FFC0B224FF1E5B93FF34" box="[1315,1429,187,212]" italics="true" pageId="16" pageNumber="349">
<typeStatus id="54AEEFD5A171FFC0B224FF1E5B93FF34" box="[1315,1429,187,212]" pageId="16" pageNumber="349" type="paratype">Paratypes</typeStatus>
</emphasis>
: YPM-IZ-106544, YPM-IZ-106545, YPM-IZ-106546,
<collectingCounty id="62CB29FBA171FFC0B40DFF7A5A14FF19" box="[778,1042,223,249]" pageId="16" pageNumber="349">East Tongass Narrows</collectingCounty>
,
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,
<collectingRegion id="49D19F95A171FFC0B3AEFF7A5AF1FF19" box="[1193,1271,223,249]" country="United States of America" name="Alaska" pageId="16" pageNumber="349">Alaska</collectingRegion>
(
<geoCoordinate id="EE2137B0A171FFC0B20CFF7B5B9EFF19" box="[1291,1432,222,249]" degrees="55.31657" direction="north" orientation="latitude" pageId="16" pageNumber="349" precision="1" value="55.31657">55.31657°N</geoCoordinate>
,
<geoCoordinate id="EE2137B0A171FFC0B790FEA75F3EFEFD" box="[151,312,258,285]" degrees="131.51990" direction="west" orientation="longitude" pageId="16" pageNumber="349" precision="1" value="-131.5199">131.51990°W</geoCoordinate>
),
<date id="FFAB77B7A171FFC0B657FEA65C59FEFD" box="[336,607,259,285]" pageId="16" pageNumber="349" value="2003-09-11" valueMax="2003-09-12" valueMin="2003-09-11">
<collectingDate id="EFEF8E5FA171FFC0B657FEA65C59FEFD" box="[336,607,259,285]" pageId="16" pageNumber="349" value="2003-09-11" valueMax="2003-09-12" valueMin="2003-09-11">1112 September 2003</collectingDate>
</date>
, lower rocky intertidal; YPM-IZ-106547, collection information as for
<typeStatus id="54AEEFD5A171FFC0B790FE825EFAFEA1" box="[151,252,295,321]" pageId="16" pageNumber="349" type="holotype">holotype</typeStatus>
, on SEM stub; YPM-IZ-100481,
<location id="8ECA07ACA171FFC0B568FE825D01FEA1" LSID="urn:lsid:plazi:treatment:03BCE061A16EFFC3B790FA035A59FD81:8ECA07ACA171FFC0B568FE825D01FEA1" box="[623,775,295,321]" county="East Tongass Narrows" latitude="55.51161" longLatPrecision="1" longitude="-131.72517" municipality="Ketchikan" name="Settlers Cove" pageId="16" pageNumber="349" stateProvince="Alaska">Settlers Cove</location>
,
<location id="8ECA07ACA171FFC0B416FE825D82FEA1" LSID="urn:lsid:plazi:treatment:03BCE061A16EFFC3B790FA035A59FD81:8ECA07ACA171FFC0B416FE825D82FEA1" box="[785,900,295,321]" county="East Tongass Narrows" latitude="55.51161" longLatPrecision="1" longitude="-131.72517" municipality="Ketchikan" name="Ketchikan" pageId="16" pageNumber="349" stateProvince="Alaska">Ketchikan</location>
vicinity,
<collectingRegion id="49D19F95A171FFC0B4EEFE825A31FEA1" box="[1001,1079,295,321]" country="United States of America" name="Alaska" pageId="16" pageNumber="349">Alaska</collectingRegion>
(
<geoCoordinate id="EE2137B0A171FFC0B342FE835AD7FEA1" box="[1093,1233,294,321]" degrees="55.51161" direction="north" orientation="latitude" pageId="16" pageNumber="349" precision="1" value="55.51161">55.51161°N</geoCoordinate>
,
<geoCoordinate id="EE2137B0A171FFC0B3DCFE835B7AFEA1" box="[1243,1404,294,321]" degrees="131.72517" direction="west" orientation="longitude" pageId="16" pageNumber="349" precision="1" value="-131.72517">131.72517°W</geoCoordinate>
),
<date id="FFAB77B7A171FFC0B289FE825F57FE85" pageId="16" pageNumber="349" value="2003-09-09">
<collectingDate id="EFEF8E5FA171FFC0B289FE825F57FE85" pageId="16" pageNumber="349" value="2003-09-09">9 September 2003</collectingDate>
</date>
, lower rocky intertidal, -
<quantity id="4CEDFC92A171FFC0B565FEEE5CB7FE85" box="[610,689,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="4.6" pageId="16" pageNumber="349" unit="m" value="0.46">
<elevation id="0038B644A171FFC0B565FEEE5CB7FE85" box="[610,689,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="4.6" pageId="16" pageNumber="349" unit="m" value="0.46">0.46 m</elevation>
</quantity>
from
<collectionCode id="ED04C9B2A171FFC0B5F2FEEE5D4AFE85" box="[757,844,331,357]" pageId="16" pageNumber="349">MLWL</collectionCode>
; all on SEM stubs
</materialsCitation>
.
<emphasis id="B9618D65A171FFC0B324FEEE5ACCFE85" box="[1059,1226,331,357]" italics="true" pageId="16" pageNumber="349">Other material</emphasis>
: YPM-IZ-106548, collection information as for
<typeStatus id="54AEEFD5A171FFC0B6D9FECA5C42FE69" box="[478,580,367,393]" pageId="16" pageNumber="349" type="holotype">holotype</typeStatus>
; on SEM stub. All specimens collected by M.H. Dick.
</paragraph>
</subSubSection>
<subSubSection id="C30F02FCA171FFC2B7C0FE365D2CFD11" lastPageId="18" lastPageNumber="351" pageId="16" pageNumber="349" type="description">
<paragraph id="8BAA5177A171FFC0B7C0FE365C0BFE4D" blockId="16.[151,1437,150,429]" box="[199,525,403,429]" pageId="16" pageNumber="349">
<emphasis id="B9618D65A171FFC0B7C0FE365F78FE4D" bold="true" box="[199,382,403,429]" pageId="16" pageNumber="349">Measurements.</emphasis>
See
<tableCitation id="C69764CCA171FFC0B6B4FE365C0EFE4D" box="[435,520,403,429]" captionStart="TABLE 3" captionStartId="16.[151,239,476,500]" captionText="TABLE 3. Measurements in micrometers and counts (presented as average and range of values) for Juxtacribrilina pushkini n. sp.; intertidal specimens from Ketchikan, Alaska; square brackets indicate sample sizes other than given at top." pageId="16" pageNumber="349">Table 3</tableCitation>
.
</paragraph>
<caption id="DF6A01FFA171FFC0B790FE795EC7FDDC" pageId="16" pageNumber="349" startId="16.[151,239,476,500]">
<paragraph id="8BAA5177A171FFC0B790FE795EC7FDDC" blockId="16.[151,1436,476,572]" pageId="16" pageNumber="349">
<emphasis id="B9618D65A171FFC0B790FE795F08FE14" bold="true" box="[151,270,476,500]" pageId="16" pageNumber="349">TABLE 3.</emphasis>
Measurements in micrometers and counts (presented as average and range of values) for
<taxonomicName id="4C152AF4A171FFC0B3FBFE795EF7FDF8" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="349" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A171FFC0B3FBFE795EF7FDF8" italics="true" pageId="16" pageNumber="349">Juxtacribrilina pushkini</emphasis>
</taxonomicName>
<emphasis id="B9618D65A171FFC0B7FDFDA45F31FDF9" bold="true" box="[250,311,513,537]" pageId="16" pageNumber="349">
<taxonomicNameLabel id="A252301EA171FFC0B7FDFDA45F31FDF9" box="[250,311,513,537]" pageId="16" pageNumber="349" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; intertidal specimens from Ketchikan, Alaska; square brackets indicate sample sizes other than given at top.
</paragraph>
</caption>
<paragraph id="8BAA5177A171FFC0B50BFDE85B57F8F3" pageId="16" pageNumber="349">
<table id="F915A3D7A171002FB7A1FDE85B8FF8F4" box="[166,1417,589,1812]" gridcols="5" gridrows="29" pageId="16" pageNumber="349">
<tr id="35255335A171002FB7A1FDE85B8FFD69" box="[166,1417,589,649]" gridrow="0" pageId="16" pageNumber="349" rowspan-0="1">
<th id="76F43A49A171002FB50BFDE85CBFFD69" box="[524,697,589,649]" gridcol="1" gridrow="0" pageId="16" pageNumber="349">YPM-IZ-100361 (Holotype)</th>
<th id="76F43A49A171002FB5E8FDE85D9CFD69" box="[751,922,589,649]" gridcol="2" gridrow="0" pageId="16" pageNumber="349">YPM-IZ-106545 (Paratype)</th>
<th id="76F43A49A171002FB4D4FDE85A8EFD69" box="[979,1160,589,649]" gridcol="3" gridrow="0" pageId="16" pageNumber="349">YPM-IZ-10048 (Paratype)</th>
<th id="76F43A49A171002FB3BCFDE85B8FFD69" box="[1211,1417,589,649]" gridcol="4" gridrow="0" pageId="16" pageNumber="349">YPM-IZ-106546 (Paratype)</th>
</tr>
<tr id="35255335A171002FB7A1FD3C5B8FFD51" box="[166,1417,665,689]" gridrow="1" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FD3C5FB8FD51" box="[166,446,665,689]" gridcol="0" gridrow="1" pageId="16" pageNumber="349">Non-ovicellate zooids</th>
<td id="76F43A49A171002FB50BFD3C5CBFFD51" box="[524,697,665,689]" gridcol="1" gridrow="1" pageId="16" pageNumber="349">n = 15, 1</td>
<td id="76F43A49A171002FB5E8FD3C5D9CFD51" box="[751,922,665,689]" gridcol="2" gridrow="1" pageId="16" pageNumber="349">n = 15, 1</td>
<td id="76F43A49A171002FB4D4FD3C5A8EFD51" box="[979,1160,665,689]" gridcol="3" gridrow="1" pageId="16" pageNumber="349">n = 15, 1</td>
<td id="76F43A49A171002FB3BCFD3C5B8FFD51" box="[1211,1417,665,689]" gridcol="4" gridrow="1" pageId="16" pageNumber="349">n = 15, 1</td>
</tr>
<tr id="35255335A171002FB7A1FD645B8FFD39" box="[166,1417,705,729]" gridrow="2" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FD645FB8FD39" box="[166,446,705,729]" gridcol="0" gridrow="2" pageId="16" pageNumber="349">Zooid length</th>
<td id="76F43A49A171002FB50BFD645CBFFD39" box="[524,697,705,729]" gridcol="1" gridrow="2" pageId="16" pageNumber="349">622 (555715)</td>
<td id="76F43A49A171002FB5E8FD645D9CFD39" box="[751,922,705,729]" gridcol="2" gridrow="2" pageId="16" pageNumber="349">559 (461665)</td>
<td id="76F43A49A171002FB4D4FD645A8EFD39" box="[979,1160,705,729]" gridcol="3" gridrow="2" pageId="16" pageNumber="349">619 (543681)</td>
<td id="76F43A49A171002FB3BCFD645B8FFD39" box="[1211,1417,705,729]" gridcol="4" gridrow="2" pageId="16" pageNumber="349">587 (500683)</td>
</tr>
<tr id="35255335A171002FB7A1FD4C5B8FFCE2" box="[166,1417,745,770]" gridrow="3" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FD4C5FB8FCE2" box="[166,446,745,770]" gridcol="0" gridrow="3" pageId="16" pageNumber="349">Zooid width</th>
<td id="76F43A49A171002FB50BFD4C5CBFFCE2" box="[524,697,745,770]" gridcol="1" gridrow="3" pageId="16" pageNumber="349">360 (321393)</td>
<td id="76F43A49A171002FB5E8FD4C5D9CFCE2" box="[751,922,745,770]" gridcol="2" gridrow="3" pageId="16" pageNumber="349">362 (312398)</td>
<td id="76F43A49A171002FB4D4FD4C5A8EFCE2" box="[979,1160,745,770]" gridcol="3" gridrow="3" pageId="16" pageNumber="349">342 (298393)</td>
<td id="76F43A49A171002FB3BCFD4C5B8FFCE2" box="[1211,1417,745,770]" gridcol="4" gridrow="3" pageId="16" pageNumber="349">369 (404427)</td>
</tr>
<tr id="35255335A171002FB7A1FCB45B8FFCCA" box="[166,1417,785,810]" gridrow="4" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FCB45FB8FCCA" box="[166,446,785,810]" gridcol="0" gridrow="4" pageId="16" pageNumber="349">Orifice length</th>
<td id="76F43A49A171002FB50BFCB45CBFFCCA" box="[524,697,785,810]" gridcol="1" gridrow="4" pageId="16" pageNumber="349">108 (94123)</td>
<td id="76F43A49A171002FB5E8FCB45D9CFCCA" box="[751,922,785,810]" gridcol="2" gridrow="4" pageId="16" pageNumber="349">106 (88143)</td>
<td id="76F43A49A171002FB4D4FCB45A8EFCCA" box="[979,1160,785,810]" gridcol="3" gridrow="4" pageId="16" pageNumber="349">121 (108165)</td>
<td id="76F43A49A171002FB3BCFCB45B8FFCCA" box="[1211,1417,785,810]" gridcol="4" gridrow="4" pageId="16" pageNumber="349">110 (87127)</td>
</tr>
<tr id="35255335A171002FB7A1FC9F5B8FFCB2" box="[166,1417,826,850]" gridrow="5" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FC9F5FB8FCB2" box="[166,446,826,850]" gridcol="0" gridrow="5" pageId="16" pageNumber="349">Orifice width</th>
<td id="76F43A49A171002FB50BFC9F5CBFFCB2" box="[524,697,826,850]" gridcol="1" gridrow="5" pageId="16" pageNumber="349">173 (156189)</td>
<td id="76F43A49A171002FB5E8FC9F5D9CFCB2" box="[751,922,826,850]" gridcol="2" gridrow="5" pageId="16" pageNumber="349">170 (153210)</td>
<td id="76F43A49A171002FB4D4FC9F5A8EFCB2" box="[979,1160,826,850]" gridcol="3" gridrow="5" pageId="16" pageNumber="349">162 (121177)</td>
<td id="76F43A49A171002FB3BCFC9F5B8FFCB2" box="[1211,1417,826,850]" gridcol="4" gridrow="5" pageId="16" pageNumber="349">174 (153186)</td>
</tr>
<tr id="35255335A171002FB7A1FCC75B8FFC9A" box="[166,1417,866,890]" gridrow="6" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FCC75FB8FC9A" box="[166,446,866,890]" gridcol="0" gridrow="6" pageId="16" pageNumber="349">Number of costae</th>
<td id="76F43A49A171002FB50BFCC75CBFFC9A" box="[524,697,866,890]" gridcol="1" gridrow="6" pageId="16" pageNumber="349">16.1 (1418)</td>
<td id="76F43A49A171002FB5E8FCC75D9CFC9A" box="[751,922,866,890]" gridcol="2" gridrow="6" pageId="16" pageNumber="349">15.1 (1417)</td>
<td id="76F43A49A171002FB4D4FCC75A8EFC9A" box="[979,1160,866,890]" gridcol="3" gridrow="6" pageId="16" pageNumber="349">17.1 (1417)</td>
<td id="76F43A49A171002FB3BCFCC75B8FFC9A" box="[1211,1417,866,890]" gridcol="4" gridrow="6" pageId="16" pageNumber="349">16.3 (1419)</td>
</tr>
<tr id="35255335A171002FB7A1FC2F5B8FFC43" box="[166,1417,906,931]" gridrow="7" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FC2F5FB8FC43" box="[166,446,906,931]" gridcol="0" gridrow="7" pageId="16" pageNumber="349">No. oral spines</th>
<td id="76F43A49A171002FB50BFC2F5CBFFC43" box="[524,697,906,931]" gridcol="1" gridrow="7" pageId="16" pageNumber="349">3.1 (34)</td>
<td id="76F43A49A171002FB5E8FC2F5D9CFC43" box="[751,922,906,931]" gridcol="2" gridrow="7" pageId="16" pageNumber="349">3.2 (34)</td>
<td id="76F43A49A171002FB4D4FC2F5A8EFC43" box="[979,1160,906,931]" gridcol="3" gridrow="7" pageId="16" pageNumber="349">3.7 (35) [n = 42]</td>
<td id="76F43A49A171002FB3BCFC2F5B8FFC43" box="[1211,1417,906,931]" gridcol="4" gridrow="7" pageId="16" pageNumber="349">3.5 (34) [n = 42, 1]</td>
</tr>
<tr id="35255335A171002FB7A1FC175B8FFC0F" box="[166,1417,946,1007]" gridrow="8" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FC175FB8FC0F" box="[166,446,946,1007]" gridcol="0" gridrow="8" pageId="16" pageNumber="349">No, intercostal lacunae1</th>
<td id="76F43A49A171002FB50BFC175CBFFC0F" box="[524,697,946,1007]" gridcol="1" gridrow="8" pageId="16" pageNumber="349">8.6 (710) [n = 20, 1]</td>
<td id="76F43A49A171002FB5E8FC175D9CFC0F" box="[751,922,946,1007]" gridcol="2" gridrow="8" pageId="16" pageNumber="349">9.0 (810) [n = 20, 1]</td>
<td id="76F43A49A171002FB4D4FC175A8EFC0F" box="[979,1160,946,1007]" gridcol="3" gridrow="8" pageId="16" pageNumber="349">8.7 (710) [n = 20, 1]</td>
<td id="76F43A49A171002FB3BCFC175B8FFC0F" box="[1211,1417,946,1007]" gridcol="4" gridrow="8" pageId="16" pageNumber="349">8.2 (610) [n = 20, 1]</td>
</tr>
<tr id="35255335A171002FB7A1FC5A5B8FFBF7" box="[166,1417,1023,1047]" gridrow="9" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FC5A5FB8FBF7" box="[166,446,1023,1047]" gridcol="0" gridrow="9" pageId="16" pageNumber="349">Ovicellate zooids, basal</th>
<td id="76F43A49A171002FB50BFC5A5CBFFBF7" box="[524,697,1023,1047]" gridcol="1" gridrow="9" pageId="16" pageNumber="349">n = 5, 1</td>
<td id="76F43A49A171002FB5E8FC5A5D9CFBF7" box="[751,922,1023,1047]" gridcol="2" gridrow="9" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FC5A5A8EFBF7" box="[979,1160,1023,1047]" gridcol="3" gridrow="9" pageId="16" pageNumber="349">n = 6, 1</td>
<td id="76F43A49A171002FB3BCFC5A5B8FFBF7" box="[1211,1417,1023,1047]" gridcol="4" gridrow="9" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FB835B8FFBDF" box="[166,1417,1062,1087]" gridrow="10" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FB835FB8FBDF" box="[166,446,1062,1087]" gridcol="0" gridrow="10" pageId="16" pageNumber="349">Zooid length2</th>
<td id="76F43A49A171002FB50BFB835CBFFBDF" box="[524,697,1062,1087]" gridcol="1" gridrow="10" pageId="16" pageNumber="349">567 (513647)</td>
<td id="76F43A49A171002FB5E8FB835D9CFBDF" box="[751,922,1062,1087]" gridcol="2" gridrow="10" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FB835A8EFBDF" box="[979,1160,1062,1087]" gridcol="3" gridrow="10" pageId="16" pageNumber="349">607 (550659)</td>
<td id="76F43A49A171002FB3BCFB835B8FFBDF" box="[1211,1417,1062,1087]" gridcol="4" gridrow="10" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FBEA5B8FFB87" box="[166,1417,1103,1127]" gridrow="11" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FBEA5FB8FB87" box="[166,446,1103,1127]" gridcol="0" gridrow="11" pageId="16" pageNumber="349">Zooid width</th>
<td id="76F43A49A171002FB50BFBEA5CBFFB87" box="[524,697,1103,1127]" gridcol="1" gridrow="11" pageId="16" pageNumber="349">340 (309380)</td>
<td id="76F43A49A171002FB5E8FBEA5D9CFB87" box="[751,922,1103,1127]" gridcol="2" gridrow="11" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FBEA5A8EFB87" box="[979,1160,1103,1127]" gridcol="3" gridrow="11" pageId="16" pageNumber="349">301 (256381)</td>
<td id="76F43A49A171002FB3BCFBEA5B8FFB87" box="[1211,1417,1103,1127]" gridcol="4" gridrow="11" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FBDD5B8FFB70" box="[166,1417,1144,1168]" gridrow="12" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FBDD5FB8FB70" box="[166,446,1144,1168]" gridcol="0" gridrow="12" pageId="16" pageNumber="349">Orifice length</th>
<td id="76F43A49A171002FB50BFBDD5CBFFB70" box="[524,697,1144,1168]" gridcol="1" gridrow="12" pageId="16" pageNumber="349">86 (63104)</td>
<td id="76F43A49A171002FB5E8FBDD5D9CFB70" box="[751,922,1144,1168]" gridcol="2" gridrow="12" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FBDD5A8EFB70" box="[979,1160,1144,1168]" gridcol="3" gridrow="12" pageId="16" pageNumber="349">81 (7984)</td>
<td id="76F43A49A171002FB3BCFBDD5B8FFB70" box="[1211,1417,1144,1168]" gridcol="4" gridrow="12" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FB055B8FFB58" box="[166,1417,1184,1208]" gridrow="13" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FB055FB8FB58" box="[166,446,1184,1208]" gridcol="0" gridrow="13" pageId="16" pageNumber="349">Orifice width</th>
<td id="76F43A49A171002FB50BFB055CBFFB58" box="[524,697,1184,1208]" gridcol="1" gridrow="13" pageId="16" pageNumber="349">198 (185212)</td>
<td id="76F43A49A171002FB5E8FB055D9CFB58" box="[751,922,1184,1208]" gridcol="2" gridrow="13" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FB055A8EFB58" box="[979,1160,1184,1208]" gridcol="3" gridrow="13" pageId="16" pageNumber="349">167 (156188)</td>
<td id="76F43A49A171002FB3BCFB055B8FFB58" box="[1211,1417,1184,1208]" gridcol="4" gridrow="13" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FB6D5B8FFB00" box="[166,1417,1224,1248]" gridrow="14" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FB6D5FB8FB00" box="[166,446,1224,1248]" gridcol="0" gridrow="14" pageId="16" pageNumber="349">Ooecial complex length</th>
<td id="76F43A49A171002FB50BFB6D5CBFFB00" box="[524,697,1224,1248]" gridcol="1" gridrow="14" pageId="16" pageNumber="349">161 (148186)</td>
<td id="76F43A49A171002FB5E8FB6D5D9CFB00" box="[751,922,1224,1248]" gridcol="2" gridrow="14" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FB6D5A8EFB00" box="[979,1160,1224,1248]" gridcol="3" gridrow="14" pageId="16" pageNumber="349">152 (129185)</td>
<td id="76F43A49A171002FB3BCFB6D5B8FFB00" box="[1211,1417,1224,1248]" gridcol="4" gridrow="14" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FB555B8FFAE8" box="[166,1417,1264,1288]" gridrow="15" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FB555FB8FAE8" box="[166,446,1264,1288]" gridcol="0" gridrow="15" pageId="16" pageNumber="349">Ooecial complex width</th>
<td id="76F43A49A171002FB50BFB555CBFFAE8" box="[524,697,1264,1288]" gridcol="1" gridrow="15" pageId="16" pageNumber="349">246 (238251)</td>
<td id="76F43A49A171002FB5E8FB555D9CFAE8" box="[751,922,1264,1288]" gridcol="2" gridrow="15" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FB555A8EFAE8" box="[979,1160,1264,1288]" gridcol="3" gridrow="15" pageId="16" pageNumber="349">241 (218285)</td>
<td id="76F43A49A171002FB3BCFB555B8FFAE8" box="[1211,1417,1264,1288]" gridcol="4" gridrow="15" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FABD5B8FFAD1" box="[166,1417,1304,1329]" gridrow="16" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FABD5FB8FAD1" box="[166,446,1304,1329]" gridcol="0" gridrow="16" pageId="16" pageNumber="349">Ooecium length</th>
<td id="76F43A49A171002FB50BFABD5CBFFAD1" box="[524,697,1304,1329]" gridcol="1" gridrow="16" pageId="16" pageNumber="349">83 (68107)</td>
<td id="76F43A49A171002FB5E8FABD5D9CFAD1" box="[751,922,1304,1329]" gridcol="2" gridrow="16" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FABD5A8EFAD1" box="[979,1160,1304,1329]" gridcol="3" gridrow="16" pageId="16" pageNumber="349">75 (5694)</td>
<td id="76F43A49A171002FB3BCFABD5B8FFAD1" box="[1211,1417,1304,1329]" gridcol="4" gridrow="16" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FAE45B8FFAB9" box="[166,1417,1345,1369]" gridrow="17" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FAE45FB8FAB9" box="[166,446,1345,1369]" gridcol="0" gridrow="17" pageId="16" pageNumber="349">Ooecium width</th>
<td id="76F43A49A171002FB50BFAE45CBFFAB9" box="[524,697,1345,1369]" gridcol="1" gridrow="17" pageId="16" pageNumber="349">181 (142205)</td>
<td id="76F43A49A171002FB5E8FAE45D9CFAB9" box="[751,922,1345,1369]" gridcol="2" gridrow="17" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FAE45A8EFAB9" box="[979,1160,1345,1369]" gridcol="3" gridrow="17" pageId="16" pageNumber="349">147 (123182)</td>
<td id="76F43A49A171002FB3BCFAE45B8FFAB9" box="[1211,1417,1345,1369]" gridcol="4" gridrow="17" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FACC5B8FFA61" box="[166,1417,1385,1409]" gridrow="18" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FACC5FB8FA61" box="[166,446,1385,1409]" gridcol="0" gridrow="18" pageId="16" pageNumber="349">Number of costae</th>
<td id="76F43A49A171002FB50BFACC5CBFFA61" box="[524,697,1385,1409]" gridcol="1" gridrow="18" pageId="16" pageNumber="349">13.0 (1114)</td>
<td id="76F43A49A171002FB5E8FACC5D9CFA61" box="[751,922,1385,1409]" gridcol="2" gridrow="18" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FACC5A8EFA61" box="[979,1160,1385,1409]" gridcol="3" gridrow="18" pageId="16" pageNumber="349">13.7 (1315)</td>
<td id="76F43A49A171002FB3BCFACC5B8FFA61" box="[1211,1417,1385,1409]" gridcol="4" gridrow="18" pageId="16" pageNumber="349"></td>
</tr>
<tr id="35255335A171002FB7A1FA345B8FFA49" box="[166,1417,1425,1449]" gridrow="19" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FA345FB8FA49" box="[166,446,1425,1449]" gridcol="0" gridrow="19" pageId="16" pageNumber="349">Frontal dwarf zooids</th>
<td id="76F43A49A171002FB50BFA345CBFFA49" box="[524,697,1425,1449]" gridcol="1" gridrow="19" pageId="16" pageNumber="349">n = 15, 1</td>
<td id="76F43A49A171002FB5E8FA345D9CFA49" box="[751,922,1425,1449]" gridcol="2" gridrow="19" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FA345A8EFA49" box="[979,1160,1425,1449]" gridcol="3" gridrow="19" pageId="16" pageNumber="349">n = 1</td>
<td id="76F43A49A171002FB3BCFA345B8FFA49" box="[1211,1417,1425,1449]" gridcol="4" gridrow="19" pageId="16" pageNumber="349">n = 9, 1</td>
</tr>
<tr id="35255335A171002FB7A1FA1D5B8FFA32" box="[166,1417,1464,1490]" gridrow="20" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FA1D5FB8FA32" box="[166,446,1464,1490]" gridcol="0" gridrow="20" pageId="16" pageNumber="349">Zooid length2</th>
<td id="76F43A49A171002FB50BFA1D5CBFFA32" box="[524,697,1464,1490]" gridcol="1" gridrow="20" pageId="16" pageNumber="349">344 (314385)</td>
<td id="76F43A49A171002FB5E8FA1D5D9CFA32" box="[751,922,1464,1490]" gridcol="2" gridrow="20" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FA1D5A8EFA32" box="[979,1160,1464,1490]" gridcol="3" gridrow="20" pageId="16" pageNumber="349">357</td>
<td id="76F43A49A171002FB3BCFA1D5B8FFA32" box="[1211,1417,1464,1490]" gridcol="4" gridrow="20" pageId="16" pageNumber="349">334 (290370)</td>
</tr>
<tr id="35255335A171002FB7A1FA475B8FFA1A" box="[166,1417,1506,1530]" gridrow="21" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1FA475FB8FA1A" box="[166,446,1506,1530]" gridcol="0" gridrow="21" pageId="16" pageNumber="349">Zooid width</th>
<td id="76F43A49A171002FB50BFA475CBFFA1A" box="[524,697,1506,1530]" gridcol="1" gridrow="21" pageId="16" pageNumber="349">269 (230304)</td>
<td id="76F43A49A171002FB5E8FA475D9CFA1A" box="[751,922,1506,1530]" gridcol="2" gridrow="21" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4FA475A8EFA1A" box="[979,1160,1506,1530]" gridcol="3" gridrow="21" pageId="16" pageNumber="349">215</td>
<td id="76F43A49A171002FB3BCFA475B8FFA1A" box="[1211,1417,1506,1530]" gridcol="4" gridrow="21" pageId="16" pageNumber="349">263 (249283)</td>
</tr>
<tr id="35255335A171002FB7A1F9AF5B8FF9C2" box="[166,1417,1546,1570]" gridrow="22" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1F9AF5FB8F9C2" box="[166,446,1546,1570]" gridcol="0" gridrow="22" pageId="16" pageNumber="349">Orifice length</th>
<td id="76F43A49A171002FB50BF9AF5CBFF9C2" box="[524,697,1546,1570]" gridcol="1" gridrow="22" pageId="16" pageNumber="349">64 (5380)</td>
<td id="76F43A49A171002FB5E8F9AF5D9CF9C2" box="[751,922,1546,1570]" gridcol="2" gridrow="22" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4F9AF5A8EF9C2" box="[979,1160,1546,1570]" gridcol="3" gridrow="22" pageId="16" pageNumber="349">59</td>
<td id="76F43A49A171002FB3BCF9AF5B8FF9C2" box="[1211,1417,1546,1570]" gridcol="4" gridrow="22" pageId="16" pageNumber="349">64 (5373)</td>
</tr>
<tr id="35255335A171002FB7A1F9975B8FF9AA" box="[166,1417,1586,1610]" gridrow="23" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1F9975FB8F9AA" box="[166,446,1586,1610]" gridcol="0" gridrow="23" pageId="16" pageNumber="349">Orifice width</th>
<td id="76F43A49A171002FB50BF9975CBFF9AA" box="[524,697,1586,1610]" gridcol="1" gridrow="23" pageId="16" pageNumber="349">156 (116179)</td>
<td id="76F43A49A171002FB5E8F9975D9CF9AA" box="[751,922,1586,1610]" gridcol="2" gridrow="23" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4F9975A8EF9AA" box="[979,1160,1586,1610]" gridcol="3" gridrow="23" pageId="16" pageNumber="349">140</td>
<td id="76F43A49A171002FB3BCF9975B8FF9AA" box="[1211,1417,1586,1610]" gridcol="4" gridrow="23" pageId="16" pageNumber="349">158 (136171)</td>
</tr>
<tr id="35255335A171002FB7A1F9FF5B8FF993" box="[166,1417,1626,1651]" gridrow="24" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1F9FF5FB8F993" box="[166,446,1626,1651]" gridcol="0" gridrow="24" pageId="16" pageNumber="349">Ooecial complex length</th>
<td id="76F43A49A171002FB50BF9FF5CBFF993" box="[524,697,1626,1651]" gridcol="1" gridrow="24" pageId="16" pageNumber="349">146 (121177)</td>
<td id="76F43A49A171002FB5E8F9FF5D9CF993" box="[751,922,1626,1651]" gridcol="2" gridrow="24" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4F9FF5A8EF993" box="[979,1160,1626,1651]" gridcol="3" gridrow="24" pageId="16" pageNumber="349">146</td>
<td id="76F43A49A171002FB3BCF9FF5B8FF993" box="[1211,1417,1626,1651]" gridcol="4" gridrow="24" pageId="16" pageNumber="349">148 (130172)</td>
</tr>
<tr id="35255335A171002FB7A1F9265B8FF97B" box="[166,1417,1667,1691]" gridrow="25" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1F9265FB8F97B" box="[166,446,1667,1691]" gridcol="0" gridrow="25" pageId="16" pageNumber="349">Ooecial complex width</th>
<td id="76F43A49A171002FB50BF9265CBFF97B" box="[524,697,1667,1691]" gridcol="1" gridrow="25" pageId="16" pageNumber="349">209 (189231)</td>
<td id="76F43A49A171002FB5E8F9265D9CF97B" box="[751,922,1667,1691]" gridcol="2" gridrow="25" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4F9265A8EF97B" box="[979,1160,1667,1691]" gridcol="3" gridrow="25" pageId="16" pageNumber="349">185</td>
<td id="76F43A49A171002FB3BCF9265B8FF97B" box="[1211,1417,1667,1691]" gridcol="4" gridrow="25" pageId="16" pageNumber="349">223 (208241)</td>
</tr>
<tr id="35255335A171002FB7A1F90E5B8FF923" box="[166,1417,1707,1731]" gridrow="26" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1F90E5FB8F923" box="[166,446,1707,1731]" gridcol="0" gridrow="26" pageId="16" pageNumber="349">Ooecium length</th>
<td id="76F43A49A171002FB50BF90E5CBFF923" box="[524,697,1707,1731]" gridcol="1" gridrow="26" pageId="16" pageNumber="349">65 (5092)</td>
<td id="76F43A49A171002FB5E8F90E5D9CF923" box="[751,922,1707,1731]" gridcol="2" gridrow="26" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4F90E5A8EF923" box="[979,1160,1707,1731]" gridcol="3" gridrow="26" pageId="16" pageNumber="349">70</td>
<td id="76F43A49A171002FB3BCF90E5B8FF923" box="[1211,1417,1707,1731]" gridcol="4" gridrow="26" pageId="16" pageNumber="349">75 (5897)</td>
</tr>
<tr id="35255335A171002FB7A1F9765B8FF90B" box="[166,1417,1747,1771]" gridrow="27" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1F9765FB8F90B" box="[166,446,1747,1771]" gridcol="0" gridrow="27" pageId="16" pageNumber="349">Ooecium width</th>
<td id="76F43A49A171002FB50BF9765CBFF90B" box="[524,697,1747,1771]" gridcol="1" gridrow="27" pageId="16" pageNumber="349">126 (106149)</td>
<td id="76F43A49A171002FB5E8F9765D9CF90B" box="[751,922,1747,1771]" gridcol="2" gridrow="27" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4F9765A8EF90B" box="[979,1160,1747,1771]" gridcol="3" gridrow="27" pageId="16" pageNumber="349">116</td>
<td id="76F43A49A171002FB3BCF9765B8FF90B" box="[1211,1417,1747,1771]" gridcol="4" gridrow="27" pageId="16" pageNumber="349">141 (130151)</td>
</tr>
<tr id="35255335A171002FB7A1F95E5B8FF8F4" box="[166,1417,1787,1812]" gridrow="28" pageId="16" pageNumber="349">
<th id="76F43A49A171002FB7A1F95E5FB8F8F4" box="[166,446,1787,1812]" gridcol="0" gridrow="28" pageId="16" pageNumber="349">Number of costae</th>
<td id="76F43A49A171002FB50BF95E5CBFF8F4" box="[524,697,1787,1812]" gridcol="1" gridrow="28" pageId="16" pageNumber="349">5.7 (56)</td>
<td id="76F43A49A171002FB5E8F95E5D9CF8F4" box="[751,922,1787,1812]" gridcol="2" gridrow="28" pageId="16" pageNumber="349"></td>
<td id="76F43A49A171002FB4D4F95E5A8EF8F4" box="[979,1160,1787,1812]" gridcol="3" gridrow="28" pageId="16" pageNumber="349">8</td>
<td id="76F43A49A171002FB3BCF95E5B8FF8F4" box="[1211,1417,1787,1812]" gridcol="4" gridrow="28" pageId="16" pageNumber="349">5.7 (56)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAA5177A171FFC0B790F8BA5B7DF8D9" blockId="16.[151,1403,1823,1849]" box="[151,1403,1823,1849]" pageId="16" pageNumber="349">
<tableNote id="76F350F9A171FFC0B790F8BA5B7DF8D9" box="[151,1403,1823,1849]" pageId="16" pageNumber="349" targetBox="[166,1417,589,1812]" targetPageId="16">
<superScript id="7C60FC3FA171FFC0B790F8BA5E99F8CD" attach="right" box="[151,159,1823,1837]" fontSize="6" pageId="16" pageNumber="349">1</superScript>
Number of lacunae in transverse series between second and third pairs of costae from orifice.
<superScript id="7C60FC3FA171FFC0B38FF8BA5A96F8CD" attach="right" box="[1160,1168,1823,1837]" fontSize="6" pageId="16" pageNumber="349">2</superScript>
Includes the ooecium.
</tableNote>
</paragraph>
<paragraph id="8BAA5177A171FFC1B7C0F8C25D13FE4D" blockId="16.[151,1437,1895,2030]" lastBlockId="17.[151,1437,151,573]" lastPageId="17" lastPageNumber="350" pageId="16" pageNumber="349">
<emphasis id="B9618D65A171FFC0B7C0F8C25F5FF861" bold="true" box="[199,345,1895,1921]" pageId="16" pageNumber="349">Description.</emphasis>
Colonies encrusting, multiserial, unilaminar except for frontal dwarf zooids (
<figureCitation id="132E4DF2A171FFC0B3ECF8C25B4CF861" box="[1259,1354,1895,1921]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="16" pageNumber="349">Fig. 9D</figureCitation>
). Nonovicellate zooids (
<figureCitation id="132E4DF2A171FFC0B658F82E5FBDF845" box="[351,443,1931,1957]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="16" pageNumber="349">Figs 9A</figureCitation>
,
<figureCitation id="132E4DF2A171FFC0B6C3F82E5C12F846" box="[452,532,1931,1958]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="16" pageNumber="349">10C, D</figureCitation>
) typically oval in outline, widest in middle, closely packed; gymnocyst negligible, sometimes wider in proximal corners or tapering proximally as long caudal zone. Ovicellate and non-ovicellate zooids in basal layer each bearing 1 or 2 (exceptionally 3) oval, triangular, or quadrangular frontal pore chambers on proximal gymnocyst (
<figureCitation id="132E4DF2A170FFC1B6B6FF325C2BFF51" box="[433,557,151,177]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="17" pageNumber="350">Fig. 9AC</figureCitation>
); frontal pore chambers multiporous. Frontal shield convex, tumid, sometimes with slight keel. Costae oval or somewhat flattened in cross section,
<quantity id="4CEDFC92A170FFC1B4AFFF1E5A0BFF35" box="[936,1037,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.826" metricValueMin="3.556" pageId="17" pageNumber="350" unit="in" value="16.5" valueMax="19.0" valueMin="14.0">1419 in</quantity>
number (average per colony 15.1 17.1) in non-ovicellate zooids, suboral pair slightly swollen but not wider than rest, sometimes with small, fused tips extending fronto-distally (
<figureCitation id="132E4DF2A170FFC1B6EBFEA65C56FEFD" box="[492,592,259,285]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="17" pageNumber="350">Fig. 10C</figureCitation>
) to form slight mucro; each costa with minute lumen pseudopore at or near tip. Intercostal lacunae small; circular, oval, or subcircular; 610 (usually 8 or 9) in transverse series (average per colony 8.29.0). Secondary orifice (
<figureCitation id="132E4DF2A170FFC1B536FEEE5C96FE85" box="[561,656,331,357]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="17" pageNumber="350">Figs 9A</figureCitation>
,
<figureCitation id="132E4DF2A170FFC1B59AFEEE5CCBFE85" box="[669,717,331,357]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="17" pageNumber="350">10C</figureCitation>
) roughly semicircular, markedly wider than long; oral spines 3 or 4 (rarely 5), lateral pair often longer and heavier than those in between; median spine(s) flattened, basally wide, tapering, single or paired, in latter case sometimes fused.
</paragraph>
<paragraph id="8BAA5177A170FFC1B7C0FE125D5EFDDD" blockId="17.[151,1437,151,573]" pageId="17" pageNumber="350">
Ovicellate zooids uncommonly present in basal layer (
<figureCitation id="132E4DF2A170FFC1B412FE125D8EFE31" box="[789,904,439,465]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="17" pageNumber="350">Fig. 9B, C</figureCitation>
), scattered singly and in small groups.Ovicellate zooids in colony interior as large (including ooecium) as non-ovicellate zooids, with 1115 costae; suboral pair of costae wider, thicker than rest, with part of fused tips often extending fronto-distally to form sharp or blunt, often flattened mucro (
<figureCitation id="132E4DF2A170FFC1B650FD865FABFDDD" box="[343,429,547,573]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="17" pageNumber="350">Fig. 9B</figureCitation>
) proximal to and overhanging orifice.
</paragraph>
<caption id="DF6A01FFA170FFC1B790FA7E5D98F947" ID-DOI="http://doi.org/10.5281/zenodo.5222230" ID-Zenodo-Dep="5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="17" pageNumber="350" startId="17.[151,250,1499,1523]" targetBox="[151,1436,597,1475]" targetPageId="17">
<paragraph id="8BAA5177A170FFC1B790FA7E5D98F947" blockId="17.[151,1437,1499,1703]" pageId="17" pageNumber="350">
<emphasis id="B9618D65A170FFC1B790FA7E5F13FA13" bold="true" box="[151,277,1499,1523]" pageId="17" pageNumber="350">FIGURE 9.</emphasis>
<taxonomicName id="4C152AF4A170FFC1B619FA7E5C13FA13" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[286,533,1499,1523]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="350" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A170FFC1B619FA7E5C13FA13" box="[286,533,1499,1523]" italics="true" pageId="17" pageNumber="350">Juxtacribrilina pushkini</emphasis>
</taxonomicName>
<emphasis id="B9618D65A170FFC1B51AFA7E5C5FFA13" bold="true" box="[541,601,1499,1523]" pageId="17" pageNumber="350">
<taxonomicNameLabel id="A252301EA170FFC1B51AFA7E5C5FFA13" box="[541,601,1499,1523]" pageId="17" pageNumber="350" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm.
</paragraph>
</caption>
<paragraph id="8BAA5177A170FFC2B7C0F9705CB3FEFD" blockId="17.[151,1436,1749,2028]" lastBlockId="18.[151,1437,151,969]" lastPageId="18" lastPageNumber="351" pageId="17" pageNumber="350">
Ooecial complex (
<figureCitation id="132E4DF2A170FFC1B693F9705C0DF90F" box="[404,523,1749,1775]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="17" pageNumber="350">Fig. 9B, C</figureCitation>
) in zooids in basal layer comprising ooecium and pair of widened, flattened, nonarticulated, modified latero-oral spines meeting at midline to form proximal margin; spines tapering and twisted in distal half to two-thirds to form concave proximal margin of ooecial complex, and sometimes raised and thickened in midline to form low mucro limited by conspicuous &quot;fold&quot; near middle of modified spine on each side (small arrowheads,
<figureCitation id="132E4DF2A170FFC1B622F8C05F7DF89F" box="[293,379,1893,1919]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="17" pageNumber="350">Fig. 9B</figureCitation>
); each spine bearing lumen pseudopore near truncate tip at midline. Proximal margin of ooecial complex and proximal margin of orifice both labiate in appearance. Ooecium moderately reduced, cap-like, roughly twice as wide as long, with 23 moderately large, circular to irregular pseudopores; ooecial cap abutting rather than passing underneath pair of modified spines forming proximal margin of complex; cap sometimes with thickened or folded proximal margin (abutting modified pair of spines) bearing additional 2 or 3 small pseudopores (
<figureCitation id="132E4DF2A173FFC2B268FF325EBDFF35" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="18" pageNumber="351">Fig. 9C</figureCitation>
). Ooecium likely kenozooidal, arising from roof of distal pore chamber, which buds next-distal zooid (
<figureCitation id="132E4DF2A173FFC2B247FF1E5B91FF35" box="[1344,1431,187,213]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="18" pageNumber="351">Fig. 9B</figureCitation>
, arrow) or fails to bud, remaining as circular opening (
<figureCitation id="132E4DF2A173FFC2B5FEFF7A5D54FF19" box="[761,850,223,249]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="18" pageNumber="351">Fig. 9B</figureCitation>
, arrowheads); basal ovicellate zooids thus often but not always in columnar series with other zooids.
</paragraph>
<paragraph id="8BAA5177A173FFC2B7C0FE825AF0FD65" blockId="18.[151,1437,151,969]" pageId="18" pageNumber="351">
Dwarf ovicellate zooids typically present in center of colony (
<figureCitation id="132E4DF2A173FFC2B493FE825DF7FEA1" box="[916,1009,295,321]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="18" pageNumber="351">Fig. 9D</figureCitation>
), often densely packed, budded from proximal frontal pore chambers of zooids in basal layer; frontal shield (
<figureCitation id="132E4DF2A173FFC2B4AAFEEE5A37FE85" box="[941,1073,331,357]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="18" pageNumber="351">Fig. 10A, B</figureCitation>
) conspicuous, occupying half of zooid length; costae usually 5 or 6 (rarely, 7 or 8); suboral pair thicker than others, similar to suboral pair in basal ovicellate zooids, often forming median mucro; other costae centripetally directed toward midline, forming tight shield, with small, circular intercostal lacunae. Ooecial complex as described above for basal ovicellate zooids; ooecium small, cap-like (
<figureCitation id="132E4DF2A173FFC2B6C7FE7E5C2CFE15" box="[448,554,475,501]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="18" pageNumber="351">Fig. 10B</figureCitation>
), but kenozooidal, terminal, budded from maternal zooid; typically bearing two moderately large, circular or oval, transversely arranged pseudopores; proximal margin of ooecium (abutting modified pair of latero-oral spines) sometimes thickened and bearing additional smaller pseudopores (
<figureCitation id="132E4DF2A173FFC2B228FD865B9EFDDD" box="[1327,1432,547,573]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="18" pageNumber="351">Fig. 10B</figureCitation>
, arrowheads). Dwarf zooids sometimes with up to three basal pore chambers in distal (ooecial), disto-lateral, or lateral positions (
<figureCitation id="132E4DF2A173FFC2B65EFDCE5FC7FD65" box="[345,449,619,645]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="18" pageNumber="351">Fig. 10A</figureCitation>
). Dwarf zooids occasionally present marginally in basal layer (
<figureCitation id="132E4DF2A173FFC2B385FDCE5AEFFD65" box="[1154,1257,619,645]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="18" pageNumber="351">Fig. 10C</figureCitation>
).
</paragraph>
<paragraph id="8BAA5177A173FFC2B7C0FD2A5D2CFD11" blockId="18.[151,1437,151,969]" pageId="18" pageNumber="351">
Ancestrula (
<figureCitation id="132E4DF2A173FFC2B648FD2A5FB3FD49" box="[335,437,655,681]" captionStart="FIGURE 10" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1436,993,1871]" captionTargetId="figure-396@18.[151,1436,993,1871]" captionTargetPageId="18" captionText="FIGURE 10. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm." figureDoi="http://doi.org/10.5281/zenodo.5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="18" pageNumber="351">Fig. 10D</figureCitation>
) cribriform, similar to but smaller than subsequent zooids, with 10
<emphasis id="B9618D65A173FFC2B39CFD2A5AAFFD49" bold="true" box="[1179,1193,655,681]" pageId="18" pageNumber="351"></emphasis>
13 costae, surrounded by 6 periancestrular zooids (1 distal, 2 disto-lateral, 2 proximolateral, 1 proximal); 4 (
<emphasis id="B9618D65A173FFC2B357FD165A58FD2C" box="[1104,1118,691,716]" italics="true" pageId="18" pageNumber="351">n</emphasis>
= 2) or 5 (
<emphasis id="B9618D65A173FFC2B3D3FD165AE4FD2C" box="[1236,1250,691,716]" italics="true" pageId="18" pageNumber="351">n</emphasis>
= 5) oral spines; if 5, distalmost, median spine thinner and shorter than rest.
</paragraph>
</subSubSection>
<subSubSection id="C30F02FCA173FFC3B7C0FD5E5A59FD81" lastPageId="19" lastPageNumber="352" pageId="18" pageNumber="351" type="discussion">
<paragraph id="8BAA5177A173FFC2B7C0FD5E5B9BFC29" blockId="18.[151,1437,151,969]" pageId="18" pageNumber="351">
<emphasis id="B9618D65A173FFC2B7C0FD5E5F3DFCF5" bold="true" box="[199,315,763,789]" pageId="18" pageNumber="351">Remarks.</emphasis>
<bibRefCitation id="EF842C86A173FFC2B645FD5E5C0EFCF5" author="Dick, M. H. &amp; Grischenko, A. V. &amp; Mawatari, S. F." box="[322,520,762,789]" pageId="18" pageNumber="351" pagination="3687 - 3784" refId="ref21456" refString="Dick, M. H., Grischenko, A. V. &amp; Mawatari, S. F. (2005) Intertidal Bryozoa (Cheilostomata) of Ketchikan, Alaska. Journal of Natural History, 39 (43), 3687 - 3784. https: // doi. org / 10.1080 / 00222930500415195" type="journal article" year="2005">
Dick
<emphasis id="B9618D65A173FFC2B678FD5E5FB1FCF5" box="[383,439,763,789]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
(2005)
</bibRefCitation>
described this species (as
<taxonomicName id="4C152AF4A173FFC2B437FD5E5A0AFCF5" box="[816,1036,763,789]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="351" phylum="Bryozoa" rank="species" species="annulata">
<emphasis id="B9618D65A173FFC2B437FD5E5A0AFCF5" box="[816,1036,763,789]" italics="true" pageId="18" pageNumber="351">Cribrilina annulata</emphasis>
</taxonomicName>
) from the same locality, Ketchikan,
<collectingRegion id="49D19F95A173FFC2B790FCBA5EEEFCD9" box="[151,232,799,825]" country="United States of America" name="Alaska" pageId="18" pageNumber="351">Alaska</collectingRegion>
. The zooidal measurements differ somewhat between that study and this, possibly due to different methods of measurement (light microscopy using an ocular micrometer in the previous study; measurements from SEM images here). The costal count for non-ovicellate zooids also differs (12
<emphasis id="B9618D65A173FFC2B4B2FCC25DC5FC61" bold="true" box="[949,963,871,897]" pageId="18" pageNumber="351"></emphasis>
17 previously; 14
<emphasis id="B9618D65A173FFC2B383FCC25A94FC61" bold="true" box="[1156,1170,871,897]" pageId="18" pageNumber="351"></emphasis>
19 here), with the lower previous numbers possibly due to inclusion of some zooids in the zone of astogenetic change and a lower sample size. The number of oral spines in the ancestrula was previously given as four, but ancestrulae more often have five spines.
</paragraph>
<caption id="DF6A01FFA173FFC2B790F8C25A8BF80B" ID-DOI="http://doi.org/10.5281/zenodo.5222232" ID-Zenodo-Dep="5222232" httpUri="https://zenodo.org/record/5222232/files/figure.png" pageId="18" pageNumber="351" startId="18.[151,250,1895,1919]" targetBox="[151,1436,993,1871]" targetPageId="18">
<paragraph id="8BAA5177A173FFC2B790F8C25A8BF80B" blockId="18.[151,1437,1895,2027]" pageId="18" pageNumber="351">
<emphasis id="B9618D65A173FFC2B790F8C25F22F89F" bold="true" box="[151,292,1895,1919]" pageId="18" pageNumber="351">FIGURE 10.</emphasis>
<taxonomicName id="4C152AF4A173FFC2B629F8C25C21F89F" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[302,551,1895,1919]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="351" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A173FFC2B629F8C25C21F89F" box="[302,551,1895,1919]" italics="true" pageId="18" pageNumber="351">Juxtacribrilina pushkini</emphasis>
</taxonomicName>
<emphasis id="B9618D65A173FFC2B536F8C25C68F89F" bold="true" box="[561,622,1895,1919]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A252301EA173FFC2B536F8C25C68F89F" box="[561,622,1895,1919]" pageId="18" pageNumber="351" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Ketchikan, Alaska; AC, holotype, YPM-IZ-100361; D, paratype, YPM-IZ- 100481. A. Dwarf zooids overlying non-ovicellate basal zooids. B. Dwarf zooid; arrowheads indicate thickened proximal margin of ooecium bearing pair of small, additional pseudopores. C. Dwarf zooid budded from basal pore chamber at colony margin. D. Ancestrula and periancestrular zooids. Scale bars: A, C, 500 µm; B, 100 µm; D, 250 µm.
</paragraph>
</caption>
<paragraph id="8BAA5177A172FFC3B7C9FF325A54FDDD" blockId="19.[151,1437,151,609]" pageId="19" pageNumber="352">
<taxonomicName id="4C152AF4A172FFC3B7C9FF325FDDFF51" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[206,475,151,177]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B7C9FF325FDDFF51" box="[206,475,151,177]" italics="true" pageId="19" pageNumber="352">Juxtacribrilina pushkini</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B6E6FF325C26FF51" bold="true" box="[481,544,151,177]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B6E6FF325C26FF51" box="[481,544,151,177]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C152AF4A172FFC3B5B5FF325D36FF51" box="[690,816,151,177]" class="Gymnolaemata" family="Cribrilinidae" genus="Cribrilina" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="annulata">
<emphasis id="B9618D65A172FFC3B5B5FF325D36FF51" box="[690,816,151,177]" italics="true" pageId="19" pageNumber="352">J. annulata</emphasis>
</taxonomicName>
in the same ways as does
<taxonomicName id="4C152AF4A172FFC3B354FF325ACCFF50" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[1107,1226,151,176]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="ezoensis" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B354FF325ACCFF50" box="[1107,1226,151,176]" italics="true" pageId="19" pageNumber="352">J. ezoensis</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B3D6FF325B09FF51" bold="true" box="[1233,1295,151,177]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B3D6FF325B09FF51" box="[1233,1295,151,177]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(see above).
<taxonomicName id="4C152AF4A172FFC3B790FF1E5FA2FF35" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[151,420,187,213]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B790FF1E5FA2FF35" box="[151,420,187,213]" italics="true" pageId="19" pageNumber="352">Juxtacribrilina pushkini</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B6ADFF1E5FEEFF35" bold="true" box="[426,488,187,213]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B6ADFF1E5FEEFF35" box="[426,488,187,213]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C152AF4A172FFC3B51AFF1E5C95FF34" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[541,659,187,212]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="ezoensis" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B51AFF1E5C95FF34" box="[541,659,187,212]" italics="true" pageId="19" pageNumber="352">J. ezoensis</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B59EFF1E5CD1FF35" bold="true" box="[665,727,187,213]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B59EFF1E5CD1FF35" box="[665,727,187,213]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are quite similar in most aspects, including zooidal dimensions, the number of oral spines, and the morphology of ovicellate zooids in the basal layer. However, non-ovicellate zooids in
<taxonomicName id="4C152AF4A172FFC3B604FEA65F7AFEFD" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[259,380,259,285]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B604FEA65F7AFEFD" box="[259,380,259,285]" italics="true" pageId="19" pageNumber="352">J. pushkini</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B685FEA65FC6FEFD" bold="true" box="[386,448,259,285]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B685FEA65FC6FEFD" box="[386,448,259,285]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
tend to have more costae (1419, overall average 16.2) than in
<taxonomicName id="4C152AF4A172FFC3B384FEA65AFCFEFC" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[1155,1274,259,284]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="ezoensis" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B384FEA65AFCFEFC" box="[1155,1274,259,284]" italics="true" pageId="19" pageNumber="352">J. ezoensis</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B206FEA65B39FEFD" bold="true" box="[1281,1343,259,285]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B206FEA65B39FEFD" box="[1281,1343,259,285]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(1218, overall average 14.8); the intercostal lacunae in basal zooids are smaller and more numerous (8 or
<quantity id="4CEDFC92A172FFC3B3F0FE825B25FEA1" box="[1271,1315,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="19" pageNumber="352" unit="in" value="9.0">9 in</quantity>
transverse series) than in
<taxonomicName id="4C152AF4A172FFC3B63BFEEE5FB5FE84" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[316,435,331,356]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="ezoensis" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B63BFEEE5FB5FE84" box="[316,435,331,356]" italics="true" pageId="19" pageNumber="352">J. ezoensis</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B6BCFEEE5FFCFE85" bold="true" box="[443,506,331,357]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B6BCFEEE5FFCFE85" box="[443,506,331,357]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(usually 68); the costal shield in dwarf zooids is larger, tighter, and more convex, bearing 5 or 6 costae, compared to
<taxonomicName id="4C152AF4A172FFC3B51EFECA5C96FE68" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[537,656,367,392]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="ezoensis" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B51EFECA5C96FE68" box="[537,656,367,392]" italics="true" pageId="19" pageNumber="352">J. ezoensis</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B591FECA5CD2FE69" bold="true" box="[662,724,367,393]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B591FECA5CD2FE69" box="[662,724,367,393]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(smaller and flatter, typically bearing 3 or 4 costae); and there is a conspicuous angular &quot;fold&quot; (
<figureCitation id="132E4DF2A172FFC3B6EFFE365C39FE4D" box="[488,575,403,429]" captionStart="FIGURE 9" captionStartId="17.[151,250,1499,1523]" captionTargetBox="[151,1436,597,1475]" captionTargetId="figure-208@17.[151,1436,597,1475]" captionTargetPageId="17" captionText="FIGURE 9. Juxtacribrilina pushkini n. sp., Ketchikan, Alaska; A, D, holotype, YPM-IZ-100361; B, C, paratype, YPM-IZ- 100481. A. Non-ovicellate zooids in basal layer; every zooid bearing one or two frontal pore chambers on proximal gymnocyst. B. Non-ovicellate and three ovicellate zooids in basal layer; large arrowheads, zooids with non-budding distal pore chamber; small arrowheads, “folds” seen in modified latero-oral spines contributing to ooecial complex; arrow, zooid with distal pore chamber contributing to next zooid. C. Reduced ovicellate zooid (center) at colony margin. D. Part of colony, showing densely packed dwarf zooids in colony center. Scale bars: AC, 250 µm; D, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.5222230" httpUri="https://zenodo.org/record/5222230/files/figure.png" pageId="19" pageNumber="352">Fig. 9B</figureCitation>
, small arrowheads) near the middle of each of the modified latero-oral spines of ovicellate zooids in the basal layer, whereas this fold is lacking in
<taxonomicName id="4C152AF4A172FFC3B493FE125A0DFE30" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[916,1035,439,464]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="ezoensis" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B493FE125A0DFE30" box="[916,1035,439,464]" italics="true" pageId="19" pageNumber="352">J. ezoensis</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B315FE125A56FE31" bold="true" box="[1042,1104,439,465]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B315FE125A56FE31" box="[1042,1104,439,465]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. The ancestrula in
<taxonomicName id="4C152AF4A172FFC3B224FE125B9AFE31" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[1315,1436,439,465]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B224FE125B9AFE31" box="[1315,1436,439,465]" italics="true" pageId="19" pageNumber="352">J. pushkini</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B790FE7E5ED3FE15" bold="true" box="[151,213,475,501]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B790FE7E5ED3FE15" box="[151,213,475,501]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has four or five oral spines but more commonly five; in
<taxonomicName id="4C152AF4A172FFC3B444FE7E5DBFFE14" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[835,953,475,500]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="ezoensis" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B444FE7E5DBFFE14" box="[835,953,475,500]" italics="true" pageId="19" pageNumber="352">J. ezoensis</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B4B8FE7E5DFBFE15" bold="true" box="[959,1021,475,501]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B4B8FE7E5DFBFE15" box="[959,1021,475,501]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, only four oral spines were observed. The two species are most readily distinguished by the size and number of the intercostal lacunae in zooids in the basal layer, and by the number of costae and size of the costal shield in dwarf zooids.
</paragraph>
<paragraph id="8BAA5177A172FFC3B7C0FDE35A59FD81" blockId="19.[151,1437,151,609]" box="[199,1119,582,609]" pageId="19" pageNumber="352">
<emphasis id="B9618D65A172FFC3B7C0FDE35F5CFD80" bold="true" box="[199,346,582,608]" pageId="19" pageNumber="352">Occurrence.</emphasis>
At present,
<taxonomicName id="4C152AF4A172FFC3B6E6FDE25C5CFD81" authority="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky, 2021" authorityName="Dick &amp; Grischenko &amp; Gordon &amp; Ostrovsky" authorityYear="2021" box="[481,602,583,609]" class="Gymnolaemata" family="Cribrilinidae" genus="Juxtacribrilina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="352" phylum="Bryozoa" rank="species" species="pushkini" status="sp. nov.">
<emphasis id="B9618D65A172FFC3B6E6FDE25C5CFD81" box="[481,602,583,609]" italics="true" pageId="19" pageNumber="352">J. pushkini</emphasis>
</taxonomicName>
<emphasis id="B9618D65A172FFC3B566FDE25CA6FD81" bold="true" box="[609,672,583,609]" pageId="19" pageNumber="352">
<taxonomicNameLabel id="A252301EA172FFC3B566FDE25CA6FD81" box="[609,672,583,609]" pageId="19" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is known only from Ketchikan,
<collectingRegion id="49D19F95A172FFC3B30DFDE25A5DFD81" box="[1034,1115,583,609]" country="United States of America" name="Alaska" pageId="19" pageNumber="352">Alaska</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>