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<document id="1726F9A889467ED4DB019CE8D92F650B" ID-CLB-Dataset="22881" ID-DOI="10.11646/zootaxa.4763.4.5" ID-GBIF-Dataset="17770afd-8c50-46f3-8b81-e5bc2b93d545" ID-ISSN="1175-5334" ID-Zenodo-Dep="3762144" ID-ZooBank="urn:lsid:zoobank.org:pub:3B0E1BAD-72EE-4E68-ABF2-6EE3F9792A94" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587571907482" checkinUser="carolina" docAuthor="Salata, Sebastian, Khalili-Moghadam, Arsalan &amp; Borowiec, Lech" docDate="2020" docId="03BC9E52FB4C5B01FF3DD1CCFB5AE684" docLanguage="en" docName="zootaxa.4763.4.5.pdf.imf" docOrigin="Zootaxa 4763 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Camponotus samius Forel 1889" docType="treatment" docVersion="7" lastPageNumber="558" masterDocId="FF85E62AFB455B0CFFAAD458FF98E67F" masterDocTitle="Review of the Camponotus samius complex (Hymenoptera: Formicidae) in the Turano-Balkan region, with the description of a new species from Iran" masterLastPageNumber="562" masterPageNumber="545" pageNumber="554" updateTime="1698835525197" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0A27F849004C3978D3314CB4811866FA">Review of the Camponotus samius complex (Hymenoptera: Formicidae) in the Turano-Balkan region, with the description of a new species from Iran</mods:title>
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<mods:namePart id="6DD9044229ED4C3C8B645916B54715C8">Salata, Sebastian</mods:namePart>
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<mods:namePart id="6FAC387D4EBC8B25D18A946DCD8A60CA">Khalili-Moghadam, Arsalan</mods:namePart>
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<mods:namePart id="0C429B6241EBE16C8F1B5428A0A37559">Borowiec, Lech</mods:namePart>
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<mods:title id="A033D52A696B2E5ABBFD2EF0BAA612C3">Zootaxa</mods:title>
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<mods:date id="B26FC3071F5D4D98C70A17FFDE3C73A5">2020</mods:date>
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<mods:number id="728CBE918A9AF6AF3735262798379FDA">4763</mods:number>
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<mods:identifier id="082B20DFF20C7C7556C99710C1E10653" type="DOI">10.11646/zootaxa.4763.4.5</mods:identifier>
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<treatment id="03BC9E52FB4C5B01FF3DD1CCFB5AE684" ID-DOI="http://doi.org/10.5281/zenodo.3804810" ID-GBIF-Taxon="163732304" ID-Zenodo-Dep="3804810" LSID="urn:lsid:plazi:treatment:03BC9E52FB4C5B01FF3DD1CCFB5AE684" httpUri="http://treatment.plazi.org/id/03BC9E52FB4C5B01FF3DD1CCFB5AE684" lastPageId="13" lastPageNumber="558" pageId="9" pageNumber="554">
<subSubSection id="C30F7CCFFB4C5B05FF3DD1CCFDB8E3D1" box="[151,544,1428,1454]" pageId="9" pageNumber="554" type="nomenclature">
<paragraph id="8BAA2F44FB4C5B05FF3DD1CCFDB8E3D1" blockId="9.[151,544,1428,1490]" box="[151,544,1428,1454]" pageId="9" pageNumber="554">
<heading id="D0E29828FB4C5B05FF3DD1CCFDB8E3D1" bold="true" box="[151,544,1428,1454]" fontSize="10" level="2" pageId="9" pageNumber="554" reason="2">
<taxonomicName id="4C1554C7FB4C5B05FF3DD1CCFDB8E3D1" ID-CoL="68WKF" authority="Forel, 1889" authorityName="Forel" authorityYear="1889" box="[151,544,1428,1454]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="samius">
<emphasis id="B961F356FB4C5B05FF3DD1CCFDB8E3D1" bold="true" box="[151,544,1428,1454]" pageId="9" pageNumber="554">
<emphasis id="B961F356FB4C5B05FF3DD1CCFE15E3D1" bold="true" box="[151,397,1428,1454]" italics="true" pageId="9" pageNumber="554">Camponotus samius</emphasis>
<bibRefCitation id="EF8452B5FB4C5B05FE3ED1CCFDB8E3D1" author="Forel, A." box="[404,544,1428,1454]" pageId="9" pageNumber="554" pagination="255 - 265" refId="ref6995" refString="Forel, A. (1889) Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen. Berliner Entomologische Zeitschrift, 32, 255 - 265. [1888] https: // doi. org / 10.1002 / mmnd. 47918880404" type="journal article" year="1889">Forel, 1889</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F7CCFFB4C5B05FF3DD1EFFEF0E3AD" box="[151,360,1463,1490]" pageId="9" pageNumber="554" type="description">
<paragraph id="8BAA2F44FB4C5B05FF3DD1EFFEF0E3AD" blockId="9.[151,544,1428,1490]" box="[151,360,1463,1490]" pageId="9" pageNumber="554">
<figureCitation id="132E33C1FB4C5B05FF3DD1EFFE8EE3AD" box="[151,278,1463,1490]" captionStart="FIGURES 15–16" captionStartId="12.[151,264,1999,2024]" captionTargetBox="[151,1436,417,1975]" captionTargetId="figure@12.[151,1436,417,1975]" captionTargetPageId="12" captionText="FIGURES 15–16. Camponotus samius Forel: 15—major worker lateral, 16—minor worker lateral (scale bar = 2mm)." figureDoi="http://doi.org/10.5281/zenodo.3762160" httpUri="https://zenodo.org/record/3762160/files/figure.png" pageId="9" pageNumber="554">Figs 15–16</figureCitation>
,
<figureCitation id="132E33C1FB4C5B05FE88D1EFFEF0E3AD" box="[290,360,1463,1490]" captionStart="FIGURES 22–25" captionStartId="15.[151,264,1896,1921]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure@15.[151,1436,181,1870]" captionTargetPageId="15" captionText="FIGURES 22–25. Head in full-face view: 22, 23 Camponotus laconicus Emery: 22—major worker, 23—minor worker; 24, 25 Camponotus samius Forel: 24—major worker, 25—minor worker (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3762166" httpUri="https://zenodo.org/record/3762166/files/figure.png" pageId="9" pageNumber="554">24–25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30F7CCFFB4C5B05FF3DD1A4FAB9E1DC" pageId="9" pageNumber="554" type="reference_group">
<paragraph id="8BAA2F44FB4C5B05FF3DD1A4FE30E02E" blockId="9.[151,1437,1531,1955]" pageId="9" pageNumber="554">
<treatmentCitationGroup id="AB05086AFB4C5B05FF3DD1A4FE30E02E" pageId="9" pageNumber="554">
<treatmentCitation id="0AB40955FB4C5B05FF3DD1A4FD1FE06B" author="Forel, A." box="[151,647,1531,1556]" page="262" pageId="9" pageNumber="554" year="1889">
<taxonomicName id="4C1554C7FB4C5B05FF3DD1A4FD1FE06B" ID-CoL="68WKF" authority="Forel, 1889: 262" authorityName="Forel" authorityPageNumber="262" authorityYear="1889" box="[151,647,1531,1556]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="subSpecies" species="rubripes" subSpecies="samius">
<emphasis id="B961F356FB4C5B05FF3DD1A4FEECE06B" box="[151,372,1532,1556]" italics="true" pageId="9" pageNumber="554">Camponotus rubripes</emphasis>
r.
<emphasis id="B961F356FB4C5B05FE3AD1A5FE4EE06B" box="[400,470,1533,1556]" italics="true" pageId="9" pageNumber="554">samius</emphasis>
<bibRefCitation id="EF8452B5FB4C5B05FE77D1A3FD1FE06B" author="Forel, A." box="[477,647,1531,1556]" pageId="9" pageNumber="554" pagination="255 - 265" refId="ref6995" refString="Forel, A. (1889) Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen. Berliner Entomologische Zeitschrift, 32, 255 - 265. [1888] https: // doi. org / 10.1002 / mmnd. 47918880404" type="journal article" year="1889">Forel, 1889: 262</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(s.w.). Raised to species:
<treatmentCitation id="0AB40955FB4C5B05FC38D1A3FBE6E06B" author="Dalla Torre, K. W." box="[914,1150,1531,1556]" page="251" pageId="9" pageNumber="554" year="1893">
<bibRefCitation id="EF8452B5FB4C5B05FC38D1A3FBE6E06B" author="Dalla Torre, K. W." box="[914,1150,1531,1556]" pageId="9" pageNumber="554" refId="ref6652" refString="Dalla Torre, K. W. (1893) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae (Heterogyna). W. Engelmann, Leipzig, 289 pp. https: // doi. org / 10.5962 / bhl. title. 8794" type="book" year="1893">Dalla Torre, 1893: 251</bibRefCitation>
</treatmentCitation>
. Subspecies of
<treatmentCitation id="0AB40955FB4C5B05FAB6D1A4FE6BE04C" author="Emery, C." page="370" pageId="9" pageNumber="554" year="1896">
<taxonomicName id="4C1554C7FB4C5B05FAB6D1A4FE6BE04C" ID-CoL="QDYV" authority=": Emery, 1896: 370" authorityName="Emery" authorityPageNumber="370" authorityYear="1896" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B961F356FB4C5B05FAB6D1A4FEB7E04D" italics="true" pageId="9" pageNumber="554">Camponotus maculatus</emphasis>
:
<bibRefCitation id="EF8452B5FB4C5B05FE96D243FE6BE04C" author="Emery, C." box="[316,499,1562,1587]" pageId="9" pageNumber="554" pagination="363 - 382" refId="ref6770" refString="Emery, C. (1896) Saggio di un catalogo sistematico dei generi Camponotus, Polyrhachis e affini. Memorie della Reale Accademia delle Scienze dell'Istituto di Bologna, 5, 363 - 382." type="journal article" year="1896">Emery, 1896: 370</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB40955FB4C5B05FE56D243FD3EE04C" author="Emery, C." box="[508,678,1562,1587]" page="35" pageId="9" pageNumber="554" year="1905">
<bibRefCitation id="EF8452B5FB4C5B05FE56D243FD3EE04C" author="Emery, C." box="[508,678,1562,1587]" pageId="9" pageNumber="554" pagination="27 - 44" refId="ref6828" refString="Emery, C. (1905) Le forme paleartiche del Camponotus maculatus F. Rendiconti delle Sessioni della Reale Accademia delle Scienze dell'Istituto di Bologna, 9, 27 - 44." type="journal article" year="1905">Emery, 1905: 35</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB40955FB4C5B05FD1AD242FCC2E04C" author="Forel, A." box="[688,858,1562,1587]" page="187" pageId="9" pageNumber="554" year="1906">
<bibRefCitation id="EF8452B5FB4C5B05FD1AD242FCC2E04C" author="Forel, A." box="[688,858,1562,1587]" pageId="9" pageNumber="554" pagination="187 - 190" refId="ref7044" refString="Forel, A. (1906) Fourmis d'Asie mineure et de la Dobrudscha recoltees par M. le Dr. Oscar Vogt et Mme Cecile Vogt, Dr. med. Annales de la Societe Entomologique de Belgique, 50, 187 - 190." type="journal article" year="1906">Forel, 1906: 187</bibRefCitation>
</treatmentCitation>
. Revived status as species:
<treatmentCitation id="0AB40955FB4C5B05FBC4D243FA92E04C" author="Emery, C." box="[1134,1290,1562,1587]" page="6" pageId="9" pageNumber="554" year="1920">
<bibRefCitation id="EF8452B5FB4C5B05FBC4D243FA92E04C" author="Emery, C." box="[1134,1290,1562,1587]" pageId="9" pageNumber="554" pagination="3 - 48" refId="ref6899" refString="Emery, C. (1920) Studi sui Camponotus. Bullettino della Societa Entomologica Italiana, 52, 3 - 48." type="journal article" year="1920">Emery, 1920: 6</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB40955FB4C5B05FABED243FF68E02E" author="Emery, C." page="100" pageId="9" pageNumber="554" year="1925">
<bibRefCitation id="EF8452B5FB4C5B05FABED243FF68E02E" author="Emery, C." pageId="9" pageNumber="554" pagination="1 - 302" refId="ref6922" refString="Emery, C. (1925) Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum, 183, 1 - 302." type="journal article" year="1925">Emery, 1925: 100</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB40955FB4C5B05FF51D260FE3CE02E" author="Finzi, B." box="[251,420,1592,1617]" page="318" pageId="9" pageNumber="554" year="1930">
<bibRefCitation id="EF8452B5FB4C5B05FF51D260FE3CE02E" author="Finzi, B." box="[251,420,1592,1617]" pageId="9" pageNumber="554" pagination="309 - 319" refId="ref6948" refString="Finzi, B. (1930) Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes. XII. Teil. Die Ameisen der Jonischen Inseln. Sitzungsberichte der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse, Abteilung I, 139, 309 - 319." type="journal article" year="1930">Finzi, 1930: 318</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA2F44FB4C5B05FF3DD200FAA3E0F0" blockId="9.[151,1437,1531,1955]" pageId="9" pageNumber="554">
<typeStatus id="54AE91E6FB4C5B05FF3DD200FF72E00F" box="[151,234,1624,1648]" pageId="9" pageNumber="554" type="syntype">Syntype</typeStatus>
major and minor workers, Marathokampos, Samos,
<collectingCountry id="F3026FD4FB4C5B05FD56D200FCDCE00F" box="[764,836,1624,1648]" name="Greece" pageId="9" pageNumber="554">Greece</collectingCountry>
(MSNG) [
<typeStatus id="54AE91E6FB4C5B05FC05D200FB9DE00F" box="[943,1029,1624,1648]" pageId="9" pageNumber="554" type="syntype">Syntype</typeStatus>
major and minor workers images examined, AntWeb, CASENT0910260 and CASENT0910259, photos by Zach Lieberman, available on AntWeb.org].
</paragraph>
<paragraph id="8BAA2F44FB4C5B05FF3DD2CEFF66E0B3" blockId="9.[151,1437,1531,1955]" pageId="9" pageNumber="554">
<treatmentCitationGroup id="AB05086AFB4C5B05FF3DD2CEFF66E0B3" pageId="9" pageNumber="554">
=
<treatmentCitation id="0AB40955FB4C5B05FF0FD2CEFD5CE0D2" author="Emery, C." box="[165,708,1684,1710]" page="6" pageId="9" pageNumber="554" year="1920">
<taxonomicName id="4C1554C7FB4C5B05FF0FD2CEFD5CE0D2" authority="Emery, 1920: 6" authorityName="Emery" authorityPageNumber="6" authorityYear="1920" box="[165,708,1684,1710]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="variety" species="samius" variety="spagnolinii">
<emphasis id="B961F356FB4C5B05FF0FD2CEFEECE0D2" box="[165,372,1686,1709]" italics="true" pageId="9" pageNumber="554">Camponotus samius</emphasis>
var.
<emphasis id="B961F356FB4C5B05FE00D2CDFD84E0D2" box="[426,540,1685,1709]" italics="true" pageId="9" pageNumber="554">spagnolinii</emphasis>
<bibRefCitation id="EF8452B5FB4C5B05FD8ED2CEFD5CE0D2" author="Emery, C." box="[548,708,1684,1710]" pageId="9" pageNumber="554" pagination="3 - 48" refId="ref6899" refString="Emery, C. (1920) Studi sui Camponotus. Bullettino della Societa Entomologica Italiana, 52, 3 - 48." type="journal article" year="1920">Emery, 1920: 6</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(w.);
<treatmentCitation id="0AB40955FB4C5B05FCABD2CCFBC2E0D2" author="Agosti, D. &amp; Collingwood, C. A." box="[769,1114,1684,1709]" page="59" pageId="9" pageNumber="554" year="1987">
<bibRefCitation id="EF8452B5FB4C5B05FCABD2CCFBC2E0D2" author="Agosti, D. &amp; Collingwood, C. A." box="[769,1114,1684,1709]" pageId="9" pageNumber="554" pagination="51 - 62" refId="ref6381" refString="Agosti, D. &amp; Collingwood, C. A. (1987) A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60, 51 - 62." type="journal article" year="1987">Agosti &amp; Collingwood, 1987: 59</bibRefCitation>
</treatmentCitation>
, as a junior synonym of
<emphasis id="B961F356FB4C5B05FAF4D2CEFF66E0B3" italics="true" pageId="9" pageNumber="554">
<taxonomicName id="4C1554C7FB4C5B05FAF4D2CEFF62E0B3" ID-CoL="68WKF" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="samius">C. samius</taxonomicName>
.
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA2F44FB4C5B05FF3DD28BFBE4E176" blockId="9.[151,1437,1531,1955]" pageId="9" pageNumber="554">
<typeStatus id="54AE91E6FB4C5B05FF3DD28BFF72E094" box="[151,234,1747,1771]" pageId="9" pageNumber="554" type="syntype">Syntype</typeStatus>
major and minor workers, Büyükada,
<collectingCountry id="F3026FD4FB4C5B05FDC4D28AFD2EE094" box="[622,694,1746,1771]" name="Turkey" pageId="9" pageNumber="554">Turkey</collectingCountry>
(MHNG) [
<typeStatus id="54AE91E6FB4C5B05FC8CD28BFCE4E094" box="[806,892,1747,1771]" pageId="9" pageNumber="554" type="syntype">Syntype</typeStatus>
major and minor workers images examined, AntWeb, CASENT0905291 and CASENT0905290, photos by Will Ericson, available on AntWeb.org].
</paragraph>
<paragraph id="8BAA2F44FB4C5B05FF3DD348FD9CE11A" blockId="9.[151,1437,1531,1955]" pageId="9" pageNumber="554">
<treatmentCitationGroup id="AB05086AFB4C5B05FF3DD348FD9CE11A" pageId="9" pageNumber="554">
<emphasis id="B961F356FB4C5B05FF3DD348FF3DE158" box="[151,165,1808,1831]" italics="true" pageId="9" pageNumber="554">=</emphasis>
<treatmentCitation id="0AB40955FB4C5B05FF0FD348FCD3E157" author="Forel, A." box="[165,843,1807,1832]" page="187" pageId="9" pageNumber="554" year="1906">
<taxonomicName id="4C1554C7FB4C5B05FF0FD348FCD3E157" authority="Forel, 1906: 187" authorityName="Forel" authorityPageNumber="187" authorityYear="1906" box="[165,843,1807,1832]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="variety" species="maculatus" subSpecies="samius" variety="bosphoricus">
<emphasis id="B961F356FB4C5B05FF0FD348FE0EE157" box="[165,406,1808,1832]" italics="true" pageId="9" pageNumber="554">Camponotus maculatus</emphasis>
r.
<emphasis id="B961F356FB4C5B05FE04D349FE6CE157" box="[430,500,1809,1832]" italics="true" pageId="9" pageNumber="554">samius</emphasis>
var.
<emphasis id="B961F356FB4C5B05FD8ED348FD38E157" box="[548,672,1808,1832]" italics="true" pageId="9" pageNumber="554">bosphoricus</emphasis>
<bibRefCitation id="EF8452B5FB4C5B05FD0FD357FCD3E157" author="Forel, A." box="[677,843,1807,1832]" pageId="9" pageNumber="554" pagination="187 - 190" refId="ref7044" refString="Forel, A. (1906) Fourmis d'Asie mineure et de la Dobrudscha recoltees par M. le Dr. Oscar Vogt et Mme Cecile Vogt, Dr. med. Annales de la Societe Entomologique de Belgique, 50, 187 - 190." type="journal article" year="1906">Forel, 1906: 187</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(w.)
<collectingCountry id="F3026FD4FB4C5B05FCD7D357FC51E157" box="[893,969,1807,1832]" name="Turkey" pageId="9" pageNumber="554">Turkey</collectingCountry>
; unavailable (infrasubspecific) name. Material referred to
<emphasis id="B961F356FB4C5B05FE9CD376FE30E139" box="[310,424,1838,1862]" italics="true" pageId="9" pageNumber="554">spagnolinii</emphasis>
by
<treatmentCitation id="0AB40955FB4C5B05FE61D377FD19E138" author="Emery, C." box="[459,641,1838,1863]" page="100" pageId="9" pageNumber="554" year="1925">
<bibRefCitation id="EF8452B5FB4C5B05FE61D377FD19E138" author="Emery, C." box="[459,641,1838,1863]" pageId="9" pageNumber="554" pagination="1 - 302" refId="ref6922" refString="Emery, C. (1925) Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum, 183, 1 - 302." type="journal article" year="1925">Emery, 1925: 100</bibRefCitation>
</treatmentCitation>
. Material referred to
<taxonomicName id="4C1554C7FB4C5B05FCF9D377FC01E139" ID-CoL="68WKF" box="[851,921,1839,1862]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="samius">
<emphasis id="B961F356FB4C5B05FCF9D377FC01E139" box="[851,921,1839,1862]" italics="true" pageId="9" pageNumber="554">samius</emphasis>
</taxonomicName>
by
<treatmentCitation id="0AB40955FB4C5B05FC16D377FA91E138" author="Atanassov, N. &amp; Dlussky, G. M." box="[956,1289,1838,1863]" page="219" pageId="9" pageNumber="554" year="1992">
<bibRefCitation id="EF8452B5FB4C5B05FC16D377FA91E138" author="Atanassov, N. &amp; Dlussky, G. M." box="[956,1289,1838,1863]" pageId="9" pageNumber="554" pagination="1 - 310" refId="ref6432" refString="Atanassov, N. &amp; Dlussky, G. M. (1992) Fauna of Bulgaria. Hymenoptera, Formicidae. Fauna na Bulgariya, 22, 1 - 310. [in Bulgarian]" type="journal article" year="1992">Atanassov &amp; Dlussky, 1992: 219</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB40955FB4C5B05FAB8D376FF7BE11A" author="Bolton, B." page="89" pageId="9" pageNumber="554" year="1995">
<bibRefCitation id="EF8452B5FB4C5B05FAB8D376FF7BE11A" author="Bolton, B." pageId="9" pageNumber="554" refId="ref6468" refString="Bolton, B. (1995) A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, 504 pp." type="book" year="1995">Bolton, 1995: 89</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB40955FB4C5B05FF44D314FD99E11A" author="Kiran, K. &amp; Karaman, C." box="[238,513,1868,1893]" page="8" pageId="9" pageNumber="554" year="2012">
<bibRefCitation id="EF8452B5FB4C5B05FF44D314FD99E11A" author="Kiran, K. &amp; Karaman, C." box="[238,513,1868,1893]" pageId="9" pageNumber="554" pagination="1 - 38" refId="ref7258" refString="Kiran, K. &amp; Karaman, C. (2012) First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa, 3548 (1), 1 - 38. http: // doi. org / 10.11646 / zootaxa. 3548.1.1" type="journal article" year="2012">
Kiran &amp;
<collectingRegion id="49D1E1A6FB4C5B05FEE2D316FE3FE11A" box="[328,423,1870,1893]" country="Turkey" name="Karaman" pageId="9" pageNumber="554">Karaman</collectingRegion>
, 2012: 8
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA2F44FB4C5B05FF3DD334FAB9E1DC" blockId="9.[151,1437,1531,1955]" pageId="9" pageNumber="554">
<treatmentCitationGroup id="AB05086AFB4C5B05FF3DD334FAB9E1DC" pageId="9" pageNumber="554">
<emphasis id="B961F356FB4C5B05FF3DD334FF3DE1FC" box="[151,165,1900,1923]" italics="true" pageId="9" pageNumber="554">=</emphasis>
<treatmentCitation id="0AB40955FB4C5B05FF0FD334FC3EE1FB" author="Forel, A." box="[165,934,1899,1924]" page="255" pageId="9" pageNumber="554" year="1889">
<taxonomicName id="4C1554C7FB4C5B05FF0FD334FC3EE1FB" authority="Forel, 1889: 255" authorityName="Forel" authorityPageNumber="255" authorityYear="1889" box="[165,934,1899,1924]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="variety" species="rubripes" subSpecies="cognatus" variety="cognatopilicornis">
<emphasis id="B961F356FB4C5B05FF0FD334FE10E1FB" box="[165,392,1900,1924]" italics="true" pageId="9" pageNumber="554">Camponotus rubripes</emphasis>
r.
<emphasis id="B961F356FB4C5B05FE0DD335FD9AE1FB" box="[423,514,1901,1924]" italics="true" pageId="9" pageNumber="554">cognatus</emphasis>
var.
<emphasis id="B961F356FB4C5B05FD90D334FD75E1FB" box="[570,749,1900,1924]" italics="true" pageId="9" pageNumber="554">cognatopilicornis</emphasis>
<bibRefCitation id="EF8452B5FB4C5B05FD5CD333FC3EE1FB" author="Forel, A." box="[758,934,1899,1924]" pageId="9" pageNumber="554" pagination="255 - 265" refId="ref6995" refString="Forel, A. (1889) Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen. Berliner Entomologische Zeitschrift, 32, 255 - 265. [1888] https: // doi. org / 10.1002 / mmnd. 47918880404" type="journal article" year="1889">Forel, 1889: 255</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 262 (w.)
<collectingCountry id="F3026FD4FB4C5B05FBBED334FBC7E1FB" box="[1044,1119,1900,1924]" name="Greece" pageId="9" pageNumber="554">Greece</collectingCountry>
; unavailable (infrasubspecific) name. Material referred to
<taxonomicName id="4C1554C7FB4C5B05FE7DD3D3FD8DE1DD" ID-CoL="QDVV" authorityName="Emery" authorityYear="1920" box="[471,533,1931,1954]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="ionius">
<emphasis id="B961F356FB4C5B05FE7DD3D3FD8DE1DD" box="[471,533,1931,1954]" italics="true" pageId="9" pageNumber="554">ionius</emphasis>
</taxonomicName>
by
<treatmentCitation id="0AB40955FB4C5B05FD91D3D3FD6BE1DC" author="Emery, C." box="[571,755,1930,1955]" page="100" pageId="9" pageNumber="554" year="1925">
<bibRefCitation id="EF8452B5FB4C5B05FD91D3D3FD6BE1DC" author="Emery, C." box="[571,755,1930,1955]" pageId="9" pageNumber="554" pagination="1 - 302" refId="ref6922" refString="Emery, C. (1925) Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum, 183, 1 - 302." type="journal article" year="1925">Emery, 1925: 100</bibRefCitation>
</treatmentCitation>
. Material referred to
<taxonomicName id="4C1554C7FB4C5B05FC63D3D3FB97E1DD" ID-CoL="68WKF" box="[969,1039,1931,1954]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="samius">
<emphasis id="B961F356FB4C5B05FC63D3D3FB97E1DD" box="[969,1039,1931,1954]" italics="true" pageId="9" pageNumber="554">samius</emphasis>
</taxonomicName>
by
<treatmentCitation id="0AB40955FB4C5B05FB9CD3D2FA85E1DC" author="Radchenko, A. G." box="[1078,1309,1930,1955]" page="809" pageId="9" pageNumber="554" year="1997">
<bibRefCitation id="EF8452B5FB4C5B05FB9CD3D2FA85E1DC" author="Radchenko, A. G." box="[1078,1309,1930,1955]" pageId="9" pageNumber="554" pagination="806 - 815" refId="ref7450" refString="Radchenko, A. G. (1997) Review of ants of the subgenera Tanaemyrmex, Colobopsis, Myrmamblis, Myrmosericus, Orthonotomyrmex and Paramyrmamblis of the genus Camponotus (Hymenoptera, Formicidae) in the Asian Palearctic. Zoologicheskii Zhurnal, 76, 806 - 815. [in Russian]" type="journal article" year="1997">Radchenko, 1997: 809</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF6A7FCCFB4F5B06FF3DD393FAD3E19B" ID-DOI="http://doi.org/10.5281/zenodo.3762156" ID-Zenodo-Dep="3762156" box="[151,1355,1995,2020]" httpUri="https://zenodo.org/record/3762156/files/figure.png" pageId="10" pageNumber="555" startId="10.[151,264,1995,2020]" targetBox="[163,1425,181,1969]" targetPageId="10">
<paragraph id="8BAA2F44FB4F5B06FF3DD393FAD3E19B" blockId="10.[151,1355,1995,2020]" box="[151,1355,1995,2020]" pageId="10" pageNumber="555">
<emphasis id="B961F356FB4F5B06FF3DD393FED4E19B" bold="true" box="[151,332,1995,2020]" pageId="10" pageNumber="555">FIGURES 11–12</emphasis>
.
<taxonomicName id="4C1554C7FB4F5B06FEF3D394FDF3E19B" authority="Emery" authorityName="Emery" authorityYear="1920" box="[345,619,1996,2020]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="555" phylum="Arthropoda" rank="species" species="ionius">
<emphasis id="B961F356FB4F5B06FEF3D394FD85E19C" box="[345,541,1996,2019]" italics="true" pageId="10" pageNumber="555">Camponotus ionius</emphasis>
Emery
</taxonomicName>
: 11—
<taxonomicName id="4C1554C7FB4F5B06FD09D393FD79E19B" box="[675,737,1995,2020]" pageId="10" pageNumber="555" rank="species" species="major">major</taxonomicName>
worker lateral, 12—
<taxonomicName id="4C1554C7FB4F5B06FC19D393FC69E19B" box="[947,1009,1995,2020]" pageId="10" pageNumber="555" rank="species" species="minor">minor</taxonomicName>
worker lateral (scale bar = 2mm).
</paragraph>
</caption>
<caption id="DF6A7FCCFB4E5B07FF3DD391FAE9E19D" ID-DOI="http://doi.org/10.5281/zenodo.3762158" ID-Zenodo-Dep="3762158" box="[151,1393,1993,2018]" httpUri="https://zenodo.org/record/3762158/files/figure.png" pageId="11" pageNumber="556" startId="11.[151,264,1993,2018]" targetBox="[199,1388,184,1967]" targetPageId="11">
<paragraph id="8BAA2F44FB4E5B07FF3DD391FAE9E19D" blockId="11.[151,1393,1993,2018]" box="[151,1393,1993,2018]" pageId="11" pageNumber="556">
<emphasis id="B961F356FB4E5B07FF3DD391FECCE19D" bold="true" box="[151,340,1993,2018]" pageId="11" pageNumber="556">FIGURES 13–14.</emphasis>
<taxonomicName id="4C1554C7FB4E5B07FEF0D392FD08E19D" authority="Emery" authorityName="Emery" box="[346,656,1994,2018]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="laconicus">
<emphasis id="B961F356FB4E5B07FEF0D392FE42E19E" box="[346,474,1994,2017]" italics="true" pageId="11" pageNumber="556">Camponotus</emphasis>
<emphasis id="B961F356FB4E5B07FE4BD392FDDAE19D" box="[481,578,1994,2018]" italics="true" pageId="11" pageNumber="556">laconicus</emphasis>
Emery
</taxonomicName>
: 13—
<taxonomicName id="4C1554C7FB4E5B07FD62D391FC9EE19D" box="[712,774,1993,2018]" pageId="11" pageNumber="556" rank="species" species="major">major</taxonomicName>
worker lateral, 14—
<taxonomicName id="4C1554C7FB4E5B07FC72D391FB8EE19D" box="[984,1046,1993,2018]" pageId="11" pageNumber="556" rank="species" species="minor">minor</taxonomicName>
worker lateral (scale bar = 2mm).
</paragraph>
</caption>
<subSubSection id="C30F7CCFFB495B00FF3DD4C0FD71E760" pageId="12" pageNumber="557" type="diagnosis">
<paragraph id="8BAA2F44FB495B00FF3DD4C0FD71E760" blockId="12.[151,1436,152,396]" pageId="12" pageNumber="557">
<emphasis id="B961F356FB495B00FF3DD4C0FE8AE6CC" bold="true" box="[151,274,152,179]" pageId="12" pageNumber="557">Diagnosis.</emphasis>
Hindtibia and scape with decumbent to suberect pilosity but without additional erect setae; gena with numerous erect setae; minor and major workers with head uniformly black to dark brown, mesosoma and legs with variable colouration from yellow to reddish-brown, gaster in major and minor workers always with distinctly brighter basal part; pilosity on gaster sparse and thin.
</paragraph>
</subSubSection>
<subSubSection id="C30F7CCFFB495B00FF6DD570FB63E7F4" pageId="12" pageNumber="557" type="distribution">
<paragraph id="8BAA2F44FB495B00FF6DD570FB63E7F4" blockId="12.[151,1436,152,396]" pageId="12" pageNumber="557">
<emphasis id="B961F356FB495B00FF6DD570FEF9E73C" bold="true" box="[199,353,296,323]" pageId="12" pageNumber="557">Distribution.</emphasis>
<collectingCountry id="F3026FD4FB495B00FECDD571FE2EE73C" box="[359,438,297,323]" name="Greece" pageId="12" pageNumber="557">Greece</collectingCountry>
(
<bibRefCitation id="EF8452B5FB495B00FE6FD571FD47E73C" author="Salata, S. &amp; Borowiec, L." box="[453,735,297,323]" pageId="12" pageNumber="557" pagination="1 - 51" refId="ref7503" refString="Salata, S. &amp; Borowiec, L. (2018) Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology, 27, 1 - 51. [online 008] http: // doi. org / 10.5281 / zenodo. 2199191" type="journal article" year="2018">Salata &amp; Borowiec 2018</bibRefCitation>
),
<collectingCountry id="F3026FD4FB495B00FD58D571FCDAE73C" box="[754,834,297,323]" name="Turkey" pageId="12" pageNumber="557">Turkey</collectingCountry>
(
<bibRefCitation id="EF8452B5FB495B00FCFAD571FBC7E73C" author="Kiran, K. &amp; Karaman, C." box="[848,1119,297,323]" pageId="12" pageNumber="557" pagination="1 - 38" refId="ref7258" refString="Kiran, K. &amp; Karaman, C. (2012) First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa, 3548 (1), 1 - 38. http: // doi. org / 10.11646 / zootaxa. 3548.1.1" type="journal article" year="2012">
Kiran &amp;
<collectingRegion id="49D1E1A6FB495B00FC1CD571FB85E73C" box="[950,1053,297,323]" country="Turkey" name="Karaman" pageId="12" pageNumber="557">Karaman</collectingRegion>
2012
</bibRefCitation>
),
<collectingCountry id="F3026FD4FB495B00FBD9D571FB40E73C" box="[1139,1240,297,323]" name="Romania" pageId="12" pageNumber="557">Romania</collectingCountry>
(
<bibRefCitation id="EF8452B5FB495B00FB4CD571FEC6E718" author="Tausan, I. &amp; Lapeva-Gjonova, A." pageId="12" pageNumber="557" pagination="352 - 354" refId="ref7631" refString="Tausan, I. &amp; Lapeva-Gjonova, A. (2017) Camponotus samius Forel, 1889 (Hymenoptera: Formicidae) - at the north edge of its European distribution. North-Western Journal of Zoology, 13 (2), 352 - 354." type="journal article" year="2017">Tausan &amp; Lapeva-Gjonova 2017</bibRefCitation>
),
<collectingCountry id="F3026FD4FB495B00FEDBD515FE49E718" box="[369,465,333,359]" name="Bulgaria" pageId="12" pageNumber="557">Bulgaria</collectingCountry>
(
<bibRefCitation id="EF8452B5FB495B00FE75D515FC86E718" author="Lapeva-Gjonova, L. &amp; Antonova, V. &amp; Radchenko, A. G. &amp; Atanasova, M." box="[479,798,333,359]" pageId="12" pageNumber="557" pagination="1 - 124" refId="ref7308" refString="Lapeva-Gjonova, L., Antonova, V., Radchenko, A. G. &amp; Atanasova, M. (2010) Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria. ZooKeys, 62, 1 - 124. https: // doi. org / 10.3897 / zookeys. 62.430" type="journal article" year="2010">
Lapeva-Gjonova
<emphasis id="B961F356FB495B00FD09D516FD43E718" box="[675,731,333,359]" italics="true" pageId="12" pageNumber="557">et al.</emphasis>
2010
</bibRefCitation>
). We consider the record from
<collectingCountry id="F3026FD4FB495B00FBDBD515FB62E718" box="[1137,1274,333,359]" name="Afghanistan" pageId="12" pageNumber="557">Afghanistan</collectingCountry>
(
<bibRefCitation id="EF8452B5FB495B00FAA2D515FF48E7F3" author="Collingwood, C. A." pageId="12" pageNumber="557" pagination="51 - 79" refId="ref6608" refString="Collingwood, C. A. (1961 [1960]) The third Danish Expedition to Central Asia. Zoological Results 27. Formicidae (Insecta) from Afghanistan. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 123, 51 - 79." type="journal article" year="1961">Collingwood 1961</bibRefCitation>
) as doubtful due to the remote locality of this place from the known distribution of the species.
</paragraph>
</subSubSection>
<caption id="DF6A7FCCFB495B00FF3DD397FAD0E197" ID-DOI="http://doi.org/10.5281/zenodo.3762160" ID-Zenodo-Dep="3762160" box="[151,1352,1999,2024]" httpUri="https://zenodo.org/record/3762160/files/figure.png" pageId="12" pageNumber="557" startId="12.[151,264,1999,2024]" targetBox="[151,1436,417,1975]" targetPageId="12">
<paragraph id="8BAA2F44FB495B00FF3DD397FAD0E197" blockId="12.[151,1352,1999,2024]" box="[151,1352,1999,2024]" pageId="12" pageNumber="557">
<emphasis id="B961F356FB495B00FF3DD397FECCE197" bold="true" box="[151,340,1999,2024]" pageId="12" pageNumber="557">FIGURES 15–16.</emphasis>
<taxonomicName id="4C1554C7FB495B00FEF0D389FDFEE197" authority="Forel" authorityName="Forel" box="[346,614,1999,2024]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="samius">
<emphasis id="B961F356FB495B00FEF0D389FE42E197" box="[346,474,2001,2024]" italics="true" pageId="12" pageNumber="557">Camponotus</emphasis>
<emphasis id="B961F356FB495B00FE4BD389FDBFE197" box="[481,551,2001,2024]" italics="true" pageId="12" pageNumber="557">samius</emphasis>
Forel
</taxonomicName>
: 15—
<taxonomicName id="4C1554C7FB495B00FD0AD397FD46E197" box="[672,734,1999,2024]" pageId="12" pageNumber="557" rank="species" species="major">major</taxonomicName>
worker lateral, 16—
<taxonomicName id="4C1554C7FB495B00FC05D397FC75E197" box="[943,1005,1999,2024]" pageId="12" pageNumber="557" rank="species" species="minor">minor</taxonomicName>
worker lateral (scale bar = 2mm).
</paragraph>
</caption>
<subSubSection id="C30F7CCFFB485B01FF6DD4C0FB5AE684" pageId="13" pageNumber="558" type="biology_ecology">
<paragraph id="8BAA2F44FB485B01FF6DD4C0FB5AE684" blockId="13.[151,1436,152,251]" pageId="13" pageNumber="558">
<emphasis id="B961F356FB485B01FF6DD4C0FEBFE6CC" bold="true" box="[199,295,152,179]" pageId="13" pageNumber="558">Biology.</emphasis>
<taxonomicName id="4C1554C7FB485B01FE9BD4C1FD8EE6CC" box="[305,534,153,179]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="558" phylum="Arthropoda" rank="species" species="samius">
<emphasis id="B961F356FB485B01FE9BD4C1FD8EE6CC" box="[305,534,153,179]" italics="true" pageId="13" pageNumber="558">Camponotus samius</emphasis>
</taxonomicName>
is a thermophile species most often recorded from pine forests. It also occurs in stream valleys with deciduous trees, suburban areas with maquis, pastures with shrubs or from dry hills in the coastal area. Nests are located under large stones. Workers are active usually at dawn and dusk.
</paragraph>
</subSubSection>
</treatment>
</document>
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