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<document id="86CAC28F796FF0773A9B28F34B4C4FC0" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A13575315FDABFD57FABDFCEA" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus ravidus Moret 2020, sp. nov." docType="treatment" docUuid="8449EDAE-130C-468E-A1F3-78EE6FB86649" docUuidSource="ZooBank" docVersion="8" lastPageNumber="33" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="32" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="B5D2A22A426F1C01217902AFD81F26FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A9DDDEF719E30F98CE6492DB558C6CEF">
<mods:title id="67F663196B90C0A981CA767114D41CBB">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="93D749722685D129CE0D65EF79D7CA56" type="personal">
<mods:role id="874B743C1FD97830564E3A42D048D57B">
<mods:roleTerm id="1C3D3A177E7A2E014BD170D7F6A58DE2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E43F2D88B0D2D44C078730FE536E7CBD">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="5794C13BEE5B9BD0A0493705DE55D3A4" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="FCFDD177B08A389709C07397C466156E">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="AAE2E98EDFA60E6E1B1DA064535A699A" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="C9997256A954DF66D81185106597AF4B" type="personal">
<mods:role id="EDD4D4D418132E6F79A329E47D8950FA">
<mods:roleTerm id="397E02B68BE0F85FFFDEBFDD1C650E57">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B0A0C44B25B39DFB887497EFA5756E09">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="1701A20110A5D9D141C51DAF4C7D4D48" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="FA9D3DB154DF64ED94F76BBE22E40F6B">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="4635762F89A07DD40D01BA9454A94401" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1685BDEB4D08DD3293DC9C950B4F35FF">text</mods:typeOfResource>
<mods:relatedItem id="93E9A3A2E169C42FDA41B48A3A45BB12" type="host">
<mods:titleInfo id="4177AA58B151B86B219255E4CD82C63D">
<mods:title id="89E1342BC49531F82AF5486D5044B531">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="51A271BC61FFF7DDC1A6BB4627460C17">
<mods:date id="FD40FE2832762803320B4D4FE6C357BF">2020</mods:date>
<mods:detail id="A9AF0DDE3BBEEF93021FF36B22243A5D" type="pubDate">
<mods:number id="4C24DCD0F226C9116B256F2E50383B52">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="0DD556BD71BDADD967CEFAE4043CCEAA" type="volume">
<mods:number id="7F7CCA9DA8ED08BFA53DCA12C9312D42">646</mods:number>
</mods:detail>
<mods:extent id="2FA7D86EDF955FEA488CFFCF622B02D0" unit="page">
<mods:start id="C60DAEDF8EF59C4045C4C6920FA8AB32">1</mods:start>
<mods:end id="05931CA2F00F8CEBF870476BBF0FB4C6">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FDA9EF6AFF55C5D5C92F58B0B6ECA1A9">journal article</mods:classification>
<mods:identifier id="431B98C7F5B64025EEF9DE9167906E24" type="CLB-Dataset">22092</mods:identifier>
<mods:identifier id="3FA217766073340FC5F9022D851936A9" type="DOI">10.5852/ejt.2020.646</mods:identifier>
<mods:identifier id="86B5B1D612427B8BC929960E75117419" type="GBIF-Dataset">c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b</mods:identifier>
<mods:identifier id="A56A249521B26DCCF71D392D672EA948" type="Zenodo-Dep">3829682</mods:identifier>
<mods:identifier id="127086C009D18B784D9D0BE025DF08E7" type="ZooBank">4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B</mods:identifier>
</mods:mods>
<treatment id="03B7726A13575315FDABFD57FABDFCEA" ID-DOI="http://doi.org/10.5281/zenodo.3848399" ID-GBIF-Taxon="164160407" ID-Zenodo-Dep="3848399" LSID="urn:lsid:zoobank.org:act:8449EDAE-130C-468E-A1F3-78EE6FB86649" httpUri="http://treatment.plazi.org/id/03B7726A13575315FDABFD57FABDFCEA" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<subSubSection id="C30490F71357532AFDABFD57FB42FD69" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BA1C37C1357532AFDABFD57FC5FFD4B" blockId="31.[368,1220,685,782]" box="[602,985,685,712]" pageId="31" pageNumber="32">
<heading id="D0E974101357532AFDABFD57FC5FFD4B" box="[602,985,685,712]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="32" reason="2">
<taxonomicName id="4C1EB8FF1357532AFDABFD57FCFEFD4B" ID-CoL="8MJX2" authority="Moret" authorityName="Moret" authorityYear="2020" box="[602,888,685,712]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="ravidus" status="sp. nov.">
<emphasis id="B96A1F6E1357532AFDABFD57FCAFFD4B" bold="true" box="[602,809,685,712]" italics="true" pageId="31" pageNumber="32">Dyscolus ravidus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151357532AFC8EFD54FC5FFD4B" box="[895,985,686,712]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C1357532AFE81FD2AFB42FD69" blockId="31.[368,1220,685,782]" box="[368,1220,720,746]" pageId="31" pageNumber="32">
<uri id="FF8FCF7E1357532AFE81FD2AFB42FD69" box="[368,1220,720,746]" pageId="31" pageNumber="32">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A91357532AFD7FFD2AFB42FD69" box="[654,1220,720,746]" pageId="31" pageNumber="32">8449EDAE-130C-468E-A1F3-78EE6FB86649</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71357532AFD29FD09FCDDFC8D" box="[728,859,755,782]" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA1C37C1357532AFD29FD09FCDDFC8D" blockId="31.[368,1220,685,782]" box="[728,859,755,782]" pageId="31" pageNumber="32">
<figureCitation id="1325DFF91357532AFD29FD09FCDDFC8D" box="[728,859,755,782]" captionStart="Figs 29–31. 29" captionStartId="30.[189,241,1924,1950]" captionTargetBox="[217,1383,358,1797]" captionTargetId="figure@30.[189,1398,335,1812]" captionTargetPageId="30" captionText="Figs 29–31. 29. Dyscolus crespoae Moret sp. nov., habitus of the female holotype (QACZ). 30– 31. D. ravidus Moret sp. nov. 30. Habitus of the male holotype (QACZ). 31. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829698" httpUri="https://zenodo.org/record/3829698/files/figure.png" pageId="31" pageNumber="32">Figs 30–31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71357532AFF4CFCCFFDA9FCFE" box="[189,559,821,893]" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BA1C37C1357532AFF4CFCCFFEC2FCCC" blockId="31.[189,324,821,847]" box="[189,324,821,847]" pageId="31" pageNumber="32">
<heading id="D0E974101357532AFF4CFCCFFEC2FCCC" bold="true" box="[189,324,821,847]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B96A1F6E1357532AFF4CFCCFFEC2FCCC" bold="true" box="[189,324,821,847]" pageId="31" pageNumber="32">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1357532AFF4CFC99FDA9FCFE" blockId="31.[189,559,867,893]" box="[189,559,867,893]" pageId="31" pageNumber="32">Latin adjective meaning ‘dark’.</paragraph>
</subSubSection>
<subSubSection id="C30490F713575315FF4CFC5FFABDFCEA" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BA1C37C1357532AFF4CFC5FFEEAFC3C" blockId="31.[189,364,933,959]" box="[189,364,933,959]" pageId="31" pageNumber="32">
<heading id="D0E974101357532AFF4CFC5FFEEAFC3C" bold="true" box="[189,364,933,959]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B96A1F6E1357532AFF4CFC5FFEEAFC3C" bold="true" box="[189,364,933,959]" pageId="31" pageNumber="32">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1357532AFF1BFC2EFEDAFC6D" blockId="31.[189,1399,980,1077]" box="[234,348,980,1006]" pageId="31" pageNumber="32">
<emphasis id="B96A1F6E1357532AFF1BFC2EFEDAFC6D" bold="true" box="[234,348,980,1006]" pageId="31" pageNumber="32">
<typeStatus id="54A57DDE1357532AFF1BFC2EFEDAFC6D" box="[234,348,980,1006]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C1357532AFF4CFC0DFBDBFBB7" blockId="31.[189,1399,980,1077]" pageId="31" pageNumber="32">
<materialsCitation id="3B76C9211357532AFF4CFC0DFBDBFBB7" ID-GBIF-Occurrence="2620303428" collectingDate="2015-03-17" collectionCode="QCAZ" collectorName="P. Moret &amp; C. Ruiz" country="Ecuador" elevation="2850" latitude="-4.116222" location="Refugio Parque Nacional Podocarpus" longLatPrecision="1" longitude="-79.17183" municipality="Cajanuma" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Loja Province" typeStatus="holotype">
<collectingCountry id="F30983EC1357532AFF4CFC0DFECDFB92" box="[189,331,1015,1041]" name="Ecuador" pageId="31" pageNumber="32">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F51357532AFE84FC0FFE0AFB92" box="[373,396,1013,1041]" pageId="31" pageNumber="32" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E1357532AFE52FC0DFDD5FB92" box="[419,595,1015,1041]" country="Ecuador" name="Loja" pageId="31" pageNumber="32">Loja Province</collectingRegion>
,
<collectingMunicipality id="6BC559061357532AFD99FC0DFD64FB92" box="[616,738,1015,1041]" pageId="31" pageNumber="32">Cajanuma</collectingMunicipality>
,
<location id="8EC195A71357532AFD07FC0DFB4AFB92" LSID="urn:lsid:plazi:treatment:03B7726A13575315FDABFD57FABDFCEA:8EC195A71357532AFD07FC0DFB4AFB92" box="[758,1228,1015,1041]" country="Ecuador" latitude="-4.116222" longLatPrecision="1" longitude="-79.17183" municipality="Cajanuma" name="Refugio Parque Nacional Podocarpus" pageId="31" pageNumber="32" stateProvince="Loja Province">Refugio Parque Nacional Podocarpus</location>
;
<geoCoordinate id="EE2AA5BB1357532AFB11FC0FFAF7FB92" box="[1248,1393,1013,1041]" degrees="4" direction="south" minutes="6" orientation="latitude" pageId="31" pageNumber="32" precision="1" seconds="58.4" value="-4.116222">4°6′58.4″ S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB1357532AFF4CFBE2FEE8FBB7" box="[189,366,1048,1076]" degrees="79" direction="west" minutes="10" orientation="longitude" pageId="31" pageNumber="32" precision="1" seconds="18.6" value="-79.17183">79°10′18.6″ W</geoCoordinate>
;
<elevation id="0033244F1357532AFE8BFBE0FD95FBB7" box="[378,531,1050,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="31" pageNumber="32" unit="m" value="2850.0">
<quantity id="4CE66E991357532AFE8BFBE0FE55FBB7" box="[378,467,1050,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="31" pageNumber="32" unit="m" value="2850.0">2850 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1357532AFDD1FBE0FD47FBB6" box="[544,705,1050,1077]" pageId="31" pageNumber="32" value="2015-03-17">
<collectingDate id="EFE41C541357532AFDD1FBE0FD47FBB6" box="[544,705,1050,1077]" pageId="31" pageNumber="32" value="2015-03-17">17 Mar. 2015</collectingDate>
</date>
;
<collectorName id="26EBA6AA1357532AFD3CFBE0FCB7FBB7" box="[717,817,1050,1076]" pageId="31" pageNumber="32">P. Moret</collectorName>
and
<collectorName id="26EBA6AA1357532AFC9BFBE0FC45FBB7" box="[874,963,1050,1076]" pageId="31" pageNumber="32">C. Ruiz</collectorName>
leg.;
<collectionCode id="ED0F5BB91357532AFBF4FBE0FBDFFBB7" box="[1029,1113,1050,1076]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="31" pageNumber="32" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C1357532AFF1BFBA1FD90FBF5" blockId="31.[189,1399,1113,1494]" box="[234,534,1113,1142]" pageId="31" pageNumber="32">
<emphasis id="B96A1F6E1357532AFF1BFBA1FEE1FBF6" bold="true" box="[234,359,1115,1141]" pageId="31" pageNumber="32">
<typeStatus id="54A57DDE1357532AFF1BFBA1FEE1FBF6" box="[234,359,1115,1141]" pageId="31" pageNumber="32" type="paratype">Paratypes</typeStatus>
</emphasis>
(7 ♂♂,
<specimenCount id="9D1808F51357532AFE3BFBA1FD8CFBF5" box="[458,522,1115,1142]" pageId="31" pageNumber="32" type="female">5 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BA1C37C1357532AFF4CFB85FC2AFA56" blockId="31.[189,1399,1113,1494]" pageId="31" pageNumber="32">
<materialsCitation id="3B76C9211357532AFF4CFB85FBEEFB1A" ID-GBIF-Occurrence="2620303367" box="[189,1128,1149,1177]" collectingDate="2015-03-17" collectionCode="CPM" collectorName="P. Moret &amp; C. Ruiz" country="Ecuador" elevation="2850" latitude="-4.116222" location="Loja Province" longLatPrecision="1" longitude="-79.17183" municipality="Cajanuma" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Loja Province">
<collectingCountry id="F30983EC1357532AFF4CFB85FECDFB1A" box="[189,331,1151,1177]" name="Ecuador" pageId="31" pageNumber="32">ECUADOR</collectingCountry>
–
<emphasis id="B96A1F6E1357532AFE98FB84FD9CFB1B" bold="true" box="[361,538,1150,1176]" pageId="31" pageNumber="32">
<collectingRegion id="49DA0D9E1357532AFE98FB84FD9CFB1B" box="[361,538,1150,1176]" country="Ecuador" name="Loja" pageId="31" pageNumber="32">Loja Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1808F51357532AFDC5FB85FDE4FB1A" box="[564,610,1149,1177]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB91357532AFBDBFB85FBEEFB1A" box="[1066,1128,1151,1177]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="31" pageNumber="32">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFB73FB85FD89FB3F" ID-GBIF-Occurrence="2620303315" collectingDate="2015-03-17" collectionCode="QCAZ" collectorName="P. Moret &amp; C. Ruiz" country="Ecuador" elevation="2850" latitude="-4.116222" location="Refugio Parque Nacional Podocarpus" longLatPrecision="1" longitude="-79.17183" municipality="Cajanuma" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFB73FB85FB36FB1A" box="[1154,1200,1149,1177]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB91357532AFE4EFB58FD89FB3F" box="[447,527,1186,1212]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="31" pageNumber="32" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFDDDFB58FBAEFB5C" ID-GBIF-Occurrence="2620303395" collectingDate="2016-08-02" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" county="Cajanuma" elevation="2840" latitude="-4.11617" location="Waypoint 166" longLatPrecision="1" longitude="-79.17204" municipality="Refugio Parque Nacional Podocarpus" pageId="31" pageNumber="32" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFDDDFB58FDDDFB3F" box="[556,603,1184,1212]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1808F51357532AFD9DFB58FD1DFB3F" box="[620,667,1186,1212]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
;
<collectingCounty id="62C0BBF01357532AFD5DFB58FCA0FB3F" box="[684,806,1186,1212]" pageId="31" pageNumber="32">Cajanuma</collectingCounty>
,
<collectingMunicipality id="6BC559061357532AFCC5FB58FB7EFB3F" box="[820,1272,1186,1212]" pageId="31" pageNumber="32">Refugio Parque Nacional Podocarpus</collectingMunicipality>
,
<location id="8EC195A71357532AFAF4FB58FF6AFB5C" LSID="urn:lsid:plazi:treatment:03B7726A13575315FDABFD57FABDFCEA:8EC195A71357532AFAF4FB58FF6AFB5C" country="Ecuador" county="Cajanuma" latitude="-4.11617" longLatPrecision="1" longitude="-79.17204" municipality="Refugio Parque Nacional Podocarpus" name="Waypoint 166" pageId="31" pageNumber="32" stateProvince="Loja">Waypoint 166</location>
;
<geoCoordinate id="EE2AA5BB1357532AFF0BFB3FFEFDFB5C" box="[250,379,1221,1247]" degrees="4.11617" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-4.11617">4.11617º S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB1357532AFE7BFB3FFDA1FB5C" box="[394,551,1221,1247]" degrees="79.17204" direction="west" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="-79.17204">79.17204º W</geoCoordinate>
;
<quantity id="4CE66E991357532AFDC5FB3FFD17FB5C" box="[564,657,1221,1247]" metricMagnitude="3" metricUnit="m" metricValue="2.84" pageId="31" pageNumber="32" unit="m" value="2840.0">
<elevation id="0033244F1357532AFDC5FB3FFD17FB5C" box="[564,657,1221,1247]" metricMagnitude="3" metricUnit="m" metricValue="2.84" pageId="31" pageNumber="32" unit="m" value="2840.0">2840 m</elevation>
</quantity>
;
<date id="FFA0E5BC1357532AFD6FFB3FFCB2FB5C" box="[670,820,1221,1247]" pageId="31" pageNumber="32" value="2016-08-02">
<collectingDate id="EFE41C541357532AFD6FFB3FFCB2FB5C" box="[670,820,1221,1247]" pageId="31" pageNumber="32" value="2016-08-02">2 Aug. 2016</collectingDate>
</date>
;
<collectorName id="26EBA6AA1357532AFCB0FB3FFC20FB5C" box="[833,934,1221,1247]" pageId="31" pageNumber="32">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB91357532AFC1BFB3FFBAEFB5C" box="[1002,1064,1221,1247]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="31" pageNumber="32">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFBB2FB3FFD9CFA80" ID-GBIF-Occurrence="2620303364" collectingDate="2016-08-02" collectionCode="MNHN" collectorName="P. Moret" country="Ecuador" county="Cajanuma" elevation="2840" latitude="-4.11617" location="Waypoint" longLatPrecision="1" longitude="-79.17204" municipality="Refugio Parque Nacional Podocarpus" pageId="31" pageNumber="32" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFBB2FB3FFBF6FB5C" box="[1091,1136,1219,1247]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1808F51357532AFB71FB3FFB28FB5C" box="[1152,1198,1221,1247]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0F5BB91357532AFE31FB13FD9CFA80" box="[448,538,1257,1283]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="31" pageNumber="32">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFDC3FB12FE43FAA6" ID-GBIF-Occurrence="2620303346" collectingDate="2001-01-06" collectionCode="QCAZ" collectorName="I. G. Tapia" country="Ecuador" county="Valladolid" location="Podocarpus" municipality="Limite del Parque Jocotoco" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFDC3FB12FDD9FA81" box="[562,607,1254,1282]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
;
<collectingCounty id="62C0BBF01357532AFD9DFB12FD6EFA81" box="[620,744,1256,1282]" pageId="31" pageNumber="32">Valladolid</collectingCounty>
,
<collectingMunicipality id="6BC559061357532AFD02FB12FBB2FA80" box="[755,1076,1256,1283]" pageId="31" pageNumber="32">Límite del Parque Jocotoco</collectingMunicipality>
y
<location id="8EC195A71357532AFBA0FB12FB58FA81" LSID="urn:lsid:plazi:treatment:03B7726A13575315FDABFD57FABDFCEA:8EC195A71357532AFBA0FB12FB58FA81" box="[1105,1246,1256,1282]" country="Ecuador" county="Valladolid" municipality="Limite del Parque Jocotoco" name="Podocarpus" pageId="31" pageNumber="32" stateProvince="Loja">Podocarpus</location>
;
<date id="FFA0E5BC1357532AFB18FB12FAF4FA81" box="[1257,1394,1256,1283]" pageId="31" pageNumber="32" value="2001-01-06">
<collectingDate id="EFE41C541357532AFB18FB12FAF4FA81" box="[1257,1394,1256,1283]" pageId="31" pageNumber="32" value="2001-01-06">6 Jan. 2001</collectingDate>
</date>
;
<collectorName id="26EBA6AA1357532AFF4CFAF1FEB5FAA6" box="[189,307,1291,1317]" pageId="31" pageNumber="32">I.G. Tapia</collectorName>
leg.;
<collectionCode id="ED0F5BB91357532AFE84FAF1FE43FAA6" box="[373,453,1291,1317]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="31" pageNumber="32" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFE2FFAF1FE8BFACA" ID-GBIF-Occurrence="2620303361" collectingDate="1994-02-12" collectionCode="QCAZ" collectorName="G. Onore" country="Ecuador" elevation="2800" location="Parque Nacional Podocarpus" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFE2FFAF1FD8DFAA5" box="[478,523,1291,1318]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
;
<location id="8EC195A71357532AFDEBFAF6FCF7FAA6" LSID="urn:lsid:plazi:treatment:03B7726A13575315FDABFD57FABDFCEA:8EC195A71357532AFDEBFAF6FCF7FAA6" box="[538,881,1291,1318]" country="Ecuador" name="Parque Nacional Podocarpus" pageId="31" pageNumber="32" stateProvince="Loja">Parque Nacional Podocarpus</location>
;
<elevation id="0033244F1357532AFC8CFAF1FB90FAA6" box="[893,1046,1291,1318]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="31" pageNumber="32" unit="m" value="2800.0">
<quantity id="4CE66E991357532AFC8CFAF1FC50FAA5" box="[893,982,1291,1318]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="31" pageNumber="32" unit="m" value="2800.0">2800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1357532AFBD5FAF1FB44FAA6" box="[1060,1218,1291,1317]" pageId="31" pageNumber="32" value="1994-02-12">
<collectingDate id="EFE41C541357532AFBD5FAF1FB44FAA6" box="[1060,1218,1291,1317]" pageId="31" pageNumber="32" value="1994-02-12">12 Feb. 1994</collectingDate>
</date>
;
<collectorName id="26EBA6AA1357532AFB3FFAF1FABDFAA6" box="[1230,1339,1291,1317]" pageId="31" pageNumber="32">G. Onore</collectorName>
leg.;
<collectionCode id="ED0F5BB91357532AFF4CFAD5FE8BFACA" box="[189,269,1327,1353]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="31" pageNumber="32" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFED6FAD5FBDAFACA" ID-GBIF-Occurrence="2620303415" box="[295,1116,1326,1353]" collectingDate="1994-12-28" collectionCode="CISEC" collectorName="G. Onore" country="Ecuador" elevation="2700" location="Mamanuma" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFED6FAD5FED3FACA" box="[295,341,1327,1353]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
;
<location id="8EC195A71357532AFE95FAD5FE72FACA" LSID="urn:lsid:plazi:treatment:03B7726A13575315FDABFD57FABDFCEA:8EC195A71357532AFE95FAD5FE72FACA" box="[356,500,1327,1353]" country="Ecuador" name="Mamanuma" pageId="31" pageNumber="32" stateProvince="Loja">Mamanuma</location>
;
<elevation id="0033244F1357532AFDF1FAD5FD1DFACB" box="[512,667,1326,1353]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="31" pageNumber="32" unit="m" value="2700.0">
<quantity id="4CE66E991357532AFDF1FAD5FDDCFACA" box="[512,602,1327,1353]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="31" pageNumber="32" unit="m" value="2700.0">2700 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1357532AFD58FAD5FCCDFACA" box="[681,843,1327,1353]" pageId="31" pageNumber="32" value="1994-12-28">
<collectingDate id="EFE41C541357532AFD58FAD5FCCDFACA" box="[681,843,1327,1353]" pageId="31" pageNumber="32" value="1994-12-28">28 Dec. 1994</collectingDate>
</date>
;
<collectorName id="26EBA6AA1357532AFCA6FAD5FC43FACA" box="[855,965,1327,1353]" pageId="31" pageNumber="32">G. Onore</collectorName>
leg.;
<collectionCode id="ED0F5BB91357532AFBF8FAD5FBDAFACA" box="[1033,1116,1327,1353]" collectionName="CISEC" pageId="31" pageNumber="32">CISEC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFB87FAD5FBE1FA0C" ID-GBIF-Occurrence="2620303433" collectingDate="2015-03-17" collectionCode="CPM" collectorName="P. Moret &amp; C. Ruiz" country="Ecuador" county="Parque Nacional Podocarpus" elevation="2970" latitude="-79.16841" location="Waypoint 397" municipality="Cajanuma" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFB87FAD5FB22FACA" box="[1142,1188,1325,1353]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
;
<collectingCounty id="62C0BBF01357532AFB42FAD5FECFFAEF" pageId="31" pageNumber="32">Parque Nacional Podocarpus</collectingCounty>
,
<collectingMunicipality id="6BC559061357532AFEA4FAA8FE49FAEF" box="[341,463,1362,1388]" pageId="31" pageNumber="32">Cajanuma</collectingMunicipality>
,
<location id="8EC195A71357532AFE2BFAA8FD05FAEF" LSID="urn:lsid:plazi:treatment:03B7726A13575315FDABFD57FABDFCEA:8EC195A71357532AFE2BFAA8FD05FAEF" box="[474,643,1362,1388]" country="Ecuador" county="Parque Nacional Podocarpus" latitude="-79.16841" municipality="Cajanuma" name="Waypoint 397" pageId="31" pageNumber="32" stateProvince="Loja">Waypoint 397</location>
;
<geoCoordinate id="EE2AA5BB1357532AFD61FAAAFC8CFAEF" box="[656,778,1360,1388]" degrees="4" direction="south" minutes="7" orientation="latitude" pageId="31" pageNumber="32" precision="1" seconds="2.4" value="-4.1173334">4°7′2.4″ S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB1357532AFCE6FAAAFC36FAEF" box="[791,944,1360,1388]" degrees="79" direction="south" minutes="10" orientation="latitude" pageId="31" pageNumber="32" precision="1" seconds="6.3" value="-79.16841">79°10′6.3″ S</geoCoordinate>
;
<elevation id="0033244F1357532AFC4CFAA8FBD1FAEF" box="[957,1111,1362,1388]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" pageId="31" pageNumber="32" unit="m" value="2970.0">
<quantity id="4CE66E991357532AFC4CFAA8FB90FAEF" box="[957,1046,1362,1388]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" pageId="31" pageNumber="32" unit="m" value="2970.0">2970 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1357532AFB95FAA8FA80FAEF" box="[1124,1286,1361,1388]" pageId="31" pageNumber="32" value="2015-03-17">
<collectingDate id="EFE41C541357532AFB95FAA8FA80FAEF" box="[1124,1286,1361,1388]" pageId="31" pageNumber="32" value="2015-03-17">17 Mar. 2015</collectingDate>
</date>
;
<collectorName id="26EBA6AA1357532AFAE2FAA8FAF1FAEF" box="[1299,1399,1362,1388]" pageId="31" pageNumber="32">P. Moret</collectorName>
and
<collectorName id="26EBA6AA1357532AFF1EFA8FFECFFA0C" box="[239,329,1397,1423]" pageId="31" pageNumber="32">C. Ruiz</collectorName>
leg.; COI voucher PM397-02, BOLD sequence SUM058-18;
<collectionCode id="ED0F5BB91357532AFBD8FA8FFBE1FA0C" box="[1065,1127,1397,1423]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="31" pageNumber="32">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFB73FA8FFB23FA31" ID-GBIF-Occurrence="2620303301" collectingDate="2015-03-17" collectionCode="CPM" collectorName="P. Moret &amp; C. Ruiz" country="Ecuador" county="Parque Nacional Podocarpus" elevation="2970" latitude="-79.16841" location="Waypoint" municipality="Cajanuma" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFB73FA8FFB29FA0C" box="[1154,1199,1395,1423]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
; same collection data as for preceding; COI voucher PM397-03, BOLD sequence SUM249-18;
<collectionCode id="ED0F5BB91357532AFB96FA62FB23FA31" box="[1127,1189,1432,1458]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="31" pageNumber="32">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211357532AFB31FA62FC2EFA56" ID-GBIF-Occurrence="2620303360" collectingDate="1988-07-21" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" elevation="2850" location="Saraguro-Loja" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Loja">
<specimenCount id="9D1808F51357532AFB31FA62FB69FA30" box="[1216,1263,1432,1459]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
;
<locationDeviation id="9A13DA5E1357532AFB0FFA62FEE9FA56" pageId="31" pageNumber="32">km 10 vía Saraguro–Loja</locationDeviation>
;
<elevation id="0033244F1357532AFE8AFA41FD92FA56" box="[379,532,1467,1494]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="31" pageNumber="32" unit="m" value="2850.0">
<quantity id="4CE66E991357532AFE8AFA41FE52FA55" box="[379,468,1467,1494]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="31" pageNumber="32" unit="m" value="2850.0">2850 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1357532AFDD0FA41FD33FA56" box="[545,693,1467,1493]" pageId="31" pageNumber="32" value="1988-07-21">
<collectingDate id="EFE41C541357532AFDD0FA41FD33FA56" box="[545,693,1467,1493]" pageId="31" pageNumber="32" value="1988-07-21">21 Jul. 1988</collectingDate>
</date>
;
<collectorName id="26EBA6AA1357532AFD30FA46FCA2FA55" box="[705,804,1468,1494]" pageId="31" pageNumber="32">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB91357532AFC97FA41FC2EFA56" box="[870,936,1467,1493]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="31" pageNumber="32">CPM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1C37C1357532AFF4CFA06FE52F995" blockId="31.[189,468,1532,1558]" box="[189,468,1532,1558]" pageId="31" pageNumber="32">
<heading id="D0E974101357532AFF4CFA06FE52F995" bold="true" box="[189,468,1532,1558]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B96A1F6E1357532AFF4CFA06FE52F995" bold="true" box="[189,468,1532,1558]" pageId="31" pageNumber="32">Diagnostic description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1357532AFF4CF9D1FEB1F826" blockId="31.[189,1399,1579,1957]" pageId="31" pageNumber="32">
Habitus:
<figureCitation id="1325DFF91357532AFED9F9D1FEF8F9C6" box="[296,382,1579,1606]" captionStart="Figs 29–31. 29" captionStartId="30.[189,241,1924,1950]" captionTargetBox="[217,1383,358,1797]" captionTargetId="figure@30.[189,1398,335,1812]" captionTargetPageId="30" captionText="Figs 29–31. 29. Dyscolus crespoae Moret sp. nov., habitus of the female holotype (QACZ). 30– 31. D. ravidus Moret sp. nov. 30. Habitus of the male holotype (QACZ). 31. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829698" httpUri="https://zenodo.org/record/3829698/files/figure.png" pageId="31" pageNumber="32">Fig. 30</figureCitation>
. Wingless. Body length:
<quantity id="4CE66E991357532AFD52F9D1FCC4F9C5" box="[675,834,1579,1606]" metricMagnitude="-2" metricUnit="m" metricValue="1.015" metricValueMax="1.08" metricValueMin="0.95" pageId="31" pageNumber="32" unit="mm" value="10.15" valueMax="10.8" valueMin="9.5">9.5–10.8 mm</quantity>
. Body colour from reddish brown to brunneous; legs, antennae and mouthparts reddish brown. Elytral microsculpture transverse. Head elongate, markedly constricted basally, eyes moderately convex, genae flat. Pronotum subcordiform; sides arcuate anterad, sinuate posterad before the hind angles which are sharp and right angled. Elytra slightly convex; striae entire but narrow and shallowly impressed, intervals flat; third interval without discal setae. Last visible abdominal ventrite with one pair (♂) or two pairs (
<specimenCount id="9D1808F51357532AFC73F921FC13F976" box="[898,917,1755,1781]" pageId="31" pageNumber="32" type="female">♀</specimenCount>
) of setae along its apical margin. Legs: tarsi smooth dorsally, metatarsomeres 1–3 bisulcate, fourth metatarsomere with one pair of dorsolateral subapical setae, apical lobes moderately long, the external lobe&gt; 2 times longer than the inner lobe; fifth metatarsomere asetose ventrally.
<emphasis id="B96A1F6E1357532AFD8BF8BFFCB6F8DC" box="[634,816,1861,1887]" italics="true" pageId="31" pageNumber="32">Male genitalia:</emphasis>
<figureCitation id="1325DFF91357532AFCC6F8BFFC0AF8DC" box="[823,908,1861,1887]" captionStart="Figs 29–31. 29" captionStartId="30.[189,241,1924,1950]" captionTargetBox="[217,1383,358,1797]" captionTargetId="figure@30.[189,1398,335,1812]" captionTargetPageId="30" captionText="Figs 29–31. 29. Dyscolus crespoae Moret sp. nov., habitus of the female holotype (QACZ). 30– 31. D. ravidus Moret sp. nov. 30. Habitus of the male holotype (QACZ). 31. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829698" httpUri="https://zenodo.org/record/3829698/files/figure.png" pageId="31" pageNumber="32">Fig. 31</figureCitation>
. Ventral side of the median lobe bisinuate in lateral view; apical blade elongate; endophallus with a dense squamose area.
<emphasis id="B96A1F6E1357532AFB53F892FAE8F801" box="[1186,1390,1896,1922]" italics="true" pageId="31" pageNumber="32">Female genitalia</emphasis>
: unstudied.
</paragraph>
<paragraph id="8BA1C37C13685315FF4CFEF1FEE5FEA6" blockId="32.[189,355,267,293]" box="[189,355,267,293]" pageId="32" pageNumber="33">
<heading id="D0E9741013685315FF4CFEF1FEE5FEA6" bold="true" box="[189,355,267,293]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B96A1F6E13685315FF4CFEF1FEE5FEA6" bold="true" box="[189,355,267,293]" pageId="32" pageNumber="33">Comparisons</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13685315FF4CFEC3FE66FDED" blockId="32.[189,1399,313,622]" pageId="32" pageNumber="33">
According to the currently available molecular data,
<taxonomicName id="4C1EB8FF13685315FCCDFEC3FBDDFED7" authority="Moret" authorityName="Moret" authorityYear="2020" box="[828,1115,313,340]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ravidus" status="sp. nov.">
<emphasis id="B96A1F6E13685315FCCDFEC3FB8FFED0" box="[828,1033,313,339]" italics="true" pageId="32" pageNumber="33">Dyscolus ravidus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513685315FB94FEC0FB45FED7" box="[1125,1219,314,340]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the same clade as
<taxonomicName id="4C1EB8FF13685315FE90FEA7FDBDFEF4" authority="Moret" authorityName="Moret" authorityYear="2020" box="[353,571,349,375]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="crespoae" status="sp. nov.">
<emphasis id="B96A1F6E13685315FE90FEA7FE6BFEF5" box="[353,493,349,374]" italics="true" pageId="32" pageNumber="33">D. crespoae</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513685315FDB1FEA7FD1CFEF4" box="[576,666,349,375]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
(see above in the description of the latter for the morphological differences between them). However, by its external characters,
<taxonomicName id="4C1EB8FF13685315FC45FE7BFB07FE19" authority="Moret" authorityName="Moret" authorityYear="2020" box="[948,1153,384,410]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ravidus" status="sp. nov.">
<emphasis id="B96A1F6E13685315FC45FE7BFBB7FE19" box="[948,1073,384,410]" italics="true" pageId="32" pageNumber="33">D. ravidus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513685315FB78FE7AFB62FE19" box="[1161,1252,384,410]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
has a closer resemblance to
<taxonomicName id="4C1EB8FF13685315FE89FE5EFDBBFE3D" authority="Moret" authorityName="Moret" authorityYear="2020" box="[376,573,420,446]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="marini" status="sp. nov.">
<emphasis id="B96A1F6E13685315FE89FE5EFE6AFE3E" box="[376,492,420,445]" italics="true" pageId="32" pageNumber="33">D. marini</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513685315FDB7FE5EFD23FE3D" box="[582,677,420,446]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
, which belongs to a different clade.
<taxonomicName id="4C1EB8FF13685315FBA8FE59FAF1FE3D" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1113,1399,419,446]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ravidus" status="sp. nov.">
<emphasis id="B96A1F6E13685315FBA8FE59FAA0FE3E" box="[1113,1318,419,445]" italics="true" pageId="32" pageNumber="33">Dyscolus ravidus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513685315FF4CFE3DFE9FFE62" box="[189,281,455,481]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C1EB8FF13685315FE47FE3DFDFCFE62" authority="Moret" authorityName="Moret" authorityYear="2020" box="[438,634,455,481]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="marini" status="sp. nov.">
<emphasis id="B96A1F6E13685315FE47FE3DFDAFFE63" box="[438,553,455,480]" italics="true" pageId="32" pageNumber="33">D. marini</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513685315FD73FE3DFD58FE62" box="[642,734,455,481]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
by the following characters: elytra brownish in contrast with a black or piceous black pronotum; eyes less convex; hind angles of the pronotum sharper, right angled; elytral striae narrower and more shallowly impressed; third interval without discal setae; elytral microsculpture transverse; male genitalia: apical blade of the median lobe two times as long as in
<taxonomicName id="4C1EB8FF13685315FF4CFDAEFEF8FDED" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,382,596,622]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="marini" status="sp. nov.">
<emphasis id="B96A1F6E13685315FF4CFDAEFEA9FDEE" box="[189,303,596,621]" italics="true" pageId="32" pageNumber="33">D. marini</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513685315FE74FDAEFE66FDED" box="[389,480,596,622]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BA1C37C13685315FF4CFD77FE9BFD24" blockId="32.[189,285,653,679]" box="[189,285,653,679]" pageId="32" pageNumber="33">
<heading id="D0E9741013685315FF4CFD77FE9BFD24" bold="true" box="[189,285,653,679]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B96A1F6E13685315FF4CFD77FE9BFD24" bold="true" box="[189,285,653,679]" pageId="32" pageNumber="33">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13685315FF4CFD46FB6DFD7A" blockId="32.[189,1399,699,761]" pageId="32" pageNumber="33">
Upper montane forest on the Eastern slope of the
<taxonomicName id="4C1EB8FF13685315FCF9FD41FCD0FD56" authorityName="Stal" authorityYear="1866" box="[776,854,699,725]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="54" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
, between
<date id="FFA0E5BC13685315FC3DFD46FBFAFD55" box="[972,1148,699,726]" pageId="32" pageNumber="33" value="2700" valueMax="2900">
2700 and
<quantity id="4CE66E9913685315FBB0FD46FBFAFD55" box="[1089,1148,700,726]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="32" pageNumber="33" unit="m" value="2900.0">2900</quantity>
</date>
m a.s.l. Active at the beginning of the night (7.30–9 pm) on the ground and in leaf axils of epiphytic bromeliads.
</paragraph>
<paragraph id="8BA1C37C13685315FF4CFCDAFE6DFCB9" blockId="32.[189,491,800,826]" box="[189,491,800,826]" pageId="32" pageNumber="33">
<heading id="D0E9741013685315FF4CFCDAFE6DFCB9" bold="true" box="[189,491,800,826]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B96A1F6E13685315FF4CFCDAFE6DFCB9" bold="true" box="[189,491,800,826]" pageId="32" pageNumber="33">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13685315FF4CFCB5FABDFCEA" blockId="32.[189,1339,847,873]" box="[189,1339,847,873]" pageId="32" pageNumber="33">
Only known from the Parque Nacional Podocarpus in Southern
<collectingCountry id="F30983EC13685315FC5AFCB5FB89FCEA" box="[939,1039,847,873]" name="Ecuador" pageId="32" pageNumber="33">Ecuador</collectingCountry>
. Probably microendemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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