281 lines
42 KiB
XML
281 lines
42 KiB
XML
<document id="6080960CAD92398E5B2BD3919F42C150" ID-CLB-Dataset="28158" ID-DOI="10.11646/zootaxa.4563.1.1" ID-GBIF-Dataset="3e3068c1-c158-4d97-9aec-31fc80551edb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2600858" ID-ZooBank="A44D4C11-ADA1-48A8-9F00-B46BDF7B2A49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553170725873" checkinUser="plazi" docAuthor="Seeman, Owen D." docDate="2019" docId="03B63A508D59015AFF6010E302CA4241" docLanguage="en" docName="zootaxa.4563.1.1.pdf" docOrigin="Zootaxa 4563 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megisthanidae" docType="key" docVersion="8" lastPageNumber="38" masterDocId="FF8F42288D7D017FFFF7166307564009" masterDocTitle="The megisthanid mites (Mesostigmata: Megisthanidae) of Australia" masterLastPageNumber="40" masterPageNumber="1" pageNumber="37" updateTime="1698702915705" updateUser="ExternalLinkService">
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<mods:title id="FFD0E13249645F2B901CC66DD4670D77">The megisthanid mites (Mesostigmata: Megisthanidae) of Australia</mods:title>
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<mods:namePart id="D19B755A5DAFA7DA379905117B556C48">Seeman, Owen D.</mods:namePart>
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<mods:title id="43AF7F7673F5D8BCFDA930E4357DD53F">Zootaxa</mods:title>
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<mods:date id="CEDDA407648E62ADB8DE69D58D772E75">2019</mods:date>
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<mods:number id="8512BC63A461416408AD16AAEB516CC1">2019-02-28</mods:number>
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<treatment id="03B63A508D59015AFF6010E302CA4241" ID-DOI="http://doi.org/10.5281/zenodo.5924279" ID-GBIF-Taxon="154474393" ID-Zenodo-Dep="5924279" LSID="urn:lsid:plazi:treatment:03B63A508D59015AFF6010E302CA4241" httpUri="http://treatment.plazi.org/id/03B63A508D59015AFF6010E302CA4241" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
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<subSubSection id="C305D8CD8D59015BFF6010E303EC4693" box="[151,1210,1664,1690]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BA08B468D59015BFF6010E303EC4693" blockId="36.[151,1210,1664,1690]" box="[151,1210,1664,1690]" pageId="36" pageNumber="37">
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<heading id="D0E83C2A8D59015BFF6010E303EC4693" bold="true" box="[151,1210,1664,1690]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
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<emphasis id="B96B57548D59015BFF6010E303EC4693" bold="true" box="[151,1210,1664,1690]" pageId="36" pageNumber="37">
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Key to species of Australian
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<taxonomicName id="4C1FF0C58D59015BFE0C10E305E54693" ID-CoL="CJ4" ID-ENA="48836" baseAuthorityName="Womersley" baseAuthorityYear="1937" box="[507,691,1664,1690]" class="Arachnida" family="Megisthanidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="family">Megisthanidae</taxonomicName>
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(females required; males supplementary)
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</emphasis>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C305D8CD8D59015AFF6010A502CA4241" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="key">
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<keyStep id="30EB93E38D59015BFF6010A502CD473A" pageId="36" pageNumber="37">
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<paragraph id="8BA08B468D59015BFF6010A502CD46F0" blockId="36.[151,1436,1734,2019]" pageId="36" pageNumber="37">
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<keyLead id="30EE28738D59015BFF6010A502CD46F0" pageId="36" pageNumber="37">
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1. Dorsal shield lyriform (
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<figureCitation id="132497C38D59015BFE3F10A5054E46D2" box="[456,536,1734,1755]" captionStart="FIGURE 6" captionStartId="14.[151,250,1168,1190]" captionTargetBox="[155,1429,184,1151]" captionTargetId="figure@14.[151,1436,181,1156]" captionTargetPageId="14" captionText="FIGURE 6. Megisthanus manonae sp. nov., female, photomicrographs. (A) dorsum; (B) venter. Scale bar = 200 µm." httpUri="https://zenodo.org/record/2600870/files/figure.png" pageId="36" pageNumber="37">Fig. 6A</figureCitation>
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); posterior dorsal shield setae thick, short, with strong barbs (
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<figureCitation id="132497C38D59015BFB8010A503E346D2" box="[1143,1205,1734,1755]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1701]" captionTargetBox="[186,1408,405,1653]" captionTargetId="figure@15.[151,1436,373,1658]" captionTargetPageId="15" captionText="FIGURE 7. Megisthanus manonae sp. nov., female. (A) sternogenital region; (B) internal genitalia, fused mesogynal and latigynal shields (mg+lg), position of sternogynal setae indicated by dotted circles; (C) vaginal heads (vh) and vaginal arms (va), which articulate with sternogynal processes (sgp) arising from thickened sternogynal margins (sgm); (D) posterior dorsal shield setae. Megisthanus leviathanicus Seeman, 2017, female. (E) posterior dorsal shield setae." httpUri="https://zenodo.org/record/2600872/files/figure.png" pageId="36" pageNumber="37">Figs 7</figureCitation>
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D–E); internal genitalia with well-developed latigynal element (
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<figureCitation id="132497C38D59015BFDA9108705FF46F0" box="[606,681,1764,1785]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1701]" captionTargetBox="[186,1408,405,1653]" captionTargetId="figure@15.[151,1436,373,1658]" captionTargetPageId="15" captionText="FIGURE 7. Megisthanus manonae sp. nov., female. (A) sternogenital region; (B) internal genitalia, fused mesogynal and latigynal shields (mg+lg), position of sternogynal setae indicated by dotted circles; (C) vaginal heads (vh) and vaginal arms (va), which articulate with sternogynal processes (sgp) arising from thickened sternogynal margins (sgm); (D) posterior dorsal shield setae. Megisthanus leviathanicus Seeman, 2017, female. (E) posterior dorsal shield setae." httpUri="https://zenodo.org/record/2600872/files/figure.png" pageId="36" pageNumber="37">Fig. 7B</figureCitation>
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)............................................................. 2
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</keyLead>
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</paragraph>
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<paragraph id="8BA08B468D59015BFF60116202CD473A" blockId="36.[151,1436,1734,2019]" pageId="36" pageNumber="37">
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<keyLead id="30EE28738D59015BFF60116202CD473A" pageId="36" pageNumber="37">
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- Dorsal shield ovate (
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<figureCitation id="132497C38D59015BFE50116206A5471F" box="[423,499,1793,1814]" captionStart="FIGURE 1" captionStartId="6.[151,250,1918,1940]" captionTargetBox="[167,1421,199,1889]" captionTargetId="figure@6.[161,1425,195,1894]" captionTargetPageId="6" captionText="FIGURE 1. Megisthanus modestus Berlese, 1910, female, photomicrographs. (A) dorsal shield (secondarily sclerotised), specimen from Lamington NP; (B) dorsal shield (lacking secondary sclerotisation), specimen from Sydney; (C) venter, specimen from Lamington NP; (D) venter, specimen from Sydney. Scale bars = 200 µm." httpUri="https://zenodo.org/record/2600860/files/figure.png" pageId="36" pageNumber="37">Fig. 1A</figureCitation>
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); posterior dorsal setae slender, usually longer, smooth or with fine barbs (
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<figureCitation id="132497C38D59015BFB4B1162025A471F" box="[1212,1292,1793,1814]" captionStart="FIGURE 3" captionStartId="8.[151,250,1850,1872]" captionTargetBox="[159,1420,203,1800]" captionTargetId="figure@8.[151,1436,193,1828]" captionTargetPageId="8" captionText="FIGURE 3. Megisthanus modestus Berlese, 1910, specimens from Lamington NP. (A) female, sternogenital region; (B) internal genitalia of same specimen (shields and thickened margins not shown); (C) internal genitalia, different specimen (shield and thickened margins shown); (D) female, posterior dorsal shield and setae; (E) female, femur IV; (F) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600864/files/figure.png" pageId="36" pageNumber="37">Figs 3D</figureCitation>
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,
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<figureCitation id="132497C38D59015BFAE111620217471F" box="[1302,1345,1793,1814]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="36" pageNumber="37">17C</figureCitation>
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); internal genitalia lacking well-developed latigynal element (
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<figureCitation id="132497C38D59015BFD25117D045A473A" box="[722,780,1822,1843]" captionStart="FIGURE 3" captionStartId="8.[151,250,1850,1872]" captionTargetBox="[159,1420,203,1800]" captionTargetId="figure@8.[151,1436,193,1828]" captionTargetPageId="8" captionText="FIGURE 3. Megisthanus modestus Berlese, 1910, specimens from Lamington NP. (A) female, sternogenital region; (B) internal genitalia of same specimen (shields and thickened margins not shown); (C) internal genitalia, different specimen (shield and thickened margins shown); (D) female, posterior dorsal shield and setae; (E) female, femur IV; (F) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600864/files/figure.png" pageId="36" pageNumber="37">Figs 3</figureCitation>
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B–C)................................................. 3
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="30EB93E38D59015BFF60115F02CA4767" pageId="36" pageNumber="37">
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||
<paragraph id="8BA08B468D59015BFF60115F02CA4758" blockId="36.[151,1436,1734,2019]" box="[151,1436,1851,1873]" pageId="36" pageNumber="37">
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||
<keyLead id="30EE28738D59015BFF60115F02CA4758" box="[151,1436,1851,1873]" pageId="36" pageNumber="37">
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2. Posterior dorsal shield setae each with 0–2 barbs (
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||
<figureCitation id="132497C38D59015BFD37115F045D4758" box="[704,779,1852,1873]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1701]" captionTargetBox="[186,1408,405,1653]" captionTargetId="figure@15.[151,1436,373,1658]" captionTargetPageId="15" captionText="FIGURE 7. Megisthanus manonae sp. nov., female. (A) sternogenital region; (B) internal genitalia, fused mesogynal and latigynal shields (mg+lg), position of sternogynal setae indicated by dotted circles; (C) vaginal heads (vh) and vaginal arms (va), which articulate with sternogynal processes (sgp) arising from thickened sternogynal margins (sgm); (D) posterior dorsal shield setae. Megisthanus leviathanicus Seeman, 2017, female. (E) posterior dorsal shield setae." httpUri="https://zenodo.org/record/2600872/files/figure.png" pageId="36" pageNumber="37">Fig. 7D</figureCitation>
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).........................................
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<taxonomicName id="4C1FF0C58D59015BFB0B115F02CA4758" authorityName="Seeman" authorityYear="2017" box="[1276,1436,1851,1873]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="leviathanicus">
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||
<emphasis id="B96B57548D59015BFB0B115F02464758" box="[1276,1296,1852,1873]" italics="true" pageId="36" pageNumber="37">M</emphasis>
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||
.
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<emphasis id="B96B57548D59015BFAEB115802CA4758" box="[1308,1436,1851,1873]" italics="true" pageId="36" pageNumber="37">leviathanicus</emphasis>
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</taxonomicName>
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</keyLead>
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||
</paragraph>
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<paragraph id="8BA08B468D59015BFF60113A02CA4767" blockId="36.[151,1436,1734,2019]" box="[151,1436,1880,1902]" pageId="36" pageNumber="37">
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||
<keyLead id="30EE28738D59015BFF60113A02CA4767" box="[151,1436,1880,1902]" pageId="36" pageNumber="37">
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- Posterior dorsal shield setae with many (ca. 10) barbs (
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<figureCitation id="132497C38D59015BFD18113A046F4767" box="[751,825,1881,1902]" captionStart="FIGURE 7" captionStartId="15.[151,250,1679,1701]" captionTargetBox="[186,1408,405,1653]" captionTargetId="figure@15.[151,1436,373,1658]" captionTargetPageId="15" captionText="FIGURE 7. Megisthanus manonae sp. nov., female. (A) sternogenital region; (B) internal genitalia, fused mesogynal and latigynal shields (mg+lg), position of sternogynal setae indicated by dotted circles; (C) vaginal heads (vh) and vaginal arms (va), which articulate with sternogynal processes (sgp) arising from thickened sternogynal margins (sgm); (D) posterior dorsal shield setae. Megisthanus leviathanicus Seeman, 2017, female. (E) posterior dorsal shield setae." httpUri="https://zenodo.org/record/2600872/files/figure.png" pageId="36" pageNumber="37">Fig. 7E</figureCitation>
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).................................
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<taxonomicName id="4C1FF0C58D59015BFB26113A021F4767" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[1233,1353,1881,1902]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="manonae" status="sp. nov.">
|
||
<emphasis id="B96B57548D59015BFB26113A03B34767" box="[1233,1253,1881,1902]" italics="true" pageId="36" pageNumber="37">M</emphasis>
|
||
.
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||
<emphasis id="B96B57548D59015BFB06113A021F4767" box="[1265,1353,1881,1902]" italics="true" pageId="36" pageNumber="37">manonae</emphasis>
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</taxonomicName>
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||
<emphasis id="B96B57548D59015BFAB8113B02CA4767" bold="true" box="[1359,1436,1880,1902]" pageId="36" pageNumber="37">
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||
<taxonomicNameLabel id="A258EA2F8D59015BFAB8113B02CA4767" box="[1359,1436,1880,1902]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
|
||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="30EB93E38D59015BFF60111502CA47EA" pageId="36" pageNumber="37">
|
||
<paragraph id="8BA08B468D59015BFF60111502CA47A0" blockId="36.[151,1436,1734,2019]" pageId="36" pageNumber="37">
|
||
<keyLead id="30EE28738D59015BFF60111502CA47A0" pageId="36" pageNumber="37">
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||
3. Seta
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<emphasis id="B96B57548D59015BFEE7111606624782" box="[272,308,1909,1931]" italics="true" pageId="36" pageNumber="37">pd2</emphasis>
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on femur IV on large projection (
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||
<figureCitation id="132497C38D59015BFD8D111505854782" box="[634,723,1910,1931]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="36" pageNumber="37">Fig. 17D</figureCitation>
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||
); dorsal shield densely covered with short fine setae (
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||
<figureCitation id="132497C38D59015BFB21111502794782" box="[1238,1327,1910,1931]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="36" pageNumber="37">Fig. 16A</figureCitation>
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); sternogynal shields widely separated (by about width of sternogynal shield) (
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<figureCitation id="132497C38D59015BFC8711F7049147A0" box="[880,967,1940,1961]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="36" pageNumber="37">Fig. 17A</figureCitation>
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)...................
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<taxonomicName id="4C1FF0C58D59015BFB4C11F7021F47A0" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[1211,1353,1939,1961]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="womersleyi" status="sp. nov.">
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||
<emphasis id="B96B57548D59015BFB4C11F7039947A0" box="[1211,1231,1940,1961]" italics="true" pageId="36" pageNumber="37">M</emphasis>
|
||
.
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||
<emphasis id="B96B57548D59015BFB2B11F0021F47A0" box="[1244,1353,1939,1961]" italics="true" pageId="36" pageNumber="37">womersleyi</emphasis>
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</taxonomicName>
|
||
<emphasis id="B96B57548D59015BFAB811F002CA47A0" bold="true" box="[1359,1436,1939,1961]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="A258EA2F8D59015BFAB811F002CA47A0" box="[1359,1436,1939,1961]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D59015BFF6011D202CA47EA" blockId="36.[151,1436,1734,2019]" pageId="36" pageNumber="37">
|
||
<keyLead id="30EE28738D59015BFF6011D202CA47EA" pageId="36" pageNumber="37">
|
||
- Seta
|
||
<emphasis id="B96B57548D59015BFEE711D3066547CF" box="[272,307,1968,1990]" italics="true" pageId="36" pageNumber="37">pd2</emphasis>
|
||
on femur IV not on projection (
|
||
<figureCitation id="132497C38D59015BFD9311D205F847CF" box="[612,686,1969,1990]" captionStart="FIGURE 3" captionStartId="8.[151,250,1850,1872]" captionTargetBox="[159,1420,203,1800]" captionTargetId="figure@8.[151,1436,193,1828]" captionTargetPageId="8" captionText="FIGURE 3. Megisthanus modestus Berlese, 1910, specimens from Lamington NP. (A) female, sternogenital region; (B) internal genitalia of same specimen (shields and thickened margins not shown); (C) internal genitalia, different specimen (shield and thickened margins shown); (D) female, posterior dorsal shield and setae; (E) female, femur IV; (F) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600864/files/figure.png" pageId="36" pageNumber="37">Fig. 3E</figureCitation>
|
||
); dorsal shield less hypertrichous, with at least some long setae (
|
||
<figureCitation id="132497C38D59015BFAE411D2023447CF" box="[1299,1378,1969,1990]" captionStart="FIGURE 1" captionStartId="6.[151,250,1918,1940]" captionTargetBox="[167,1421,199,1889]" captionTargetId="figure@6.[161,1425,195,1894]" captionTargetPageId="6" captionText="FIGURE 1. Megisthanus modestus Berlese, 1910, female, photomicrographs. (A) dorsal shield (secondarily sclerotised), specimen from Lamington NP; (B) dorsal shield (lacking secondary sclerotisation), specimen from Sydney; (C) venter, specimen from Lamington NP; (D) venter, specimen from Sydney. Scale bars = 200 µm." httpUri="https://zenodo.org/record/2600860/files/figure.png" pageId="36" pageNumber="37">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D59015BFA9B11D202C147CF" box="[1388,1431,1969,1990]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="36" pageNumber="37">12A</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D59015BFF1611AD065B47EA" box="[225,269,1998,2019]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="36" pageNumber="37">21A</figureCitation>
|
||
); sternogynal shields not widely separated (by much less than width of sternogynal shield) (
|
||
<figureCitation id="132497C38D59015BFB8911AD039A47EA" box="[1150,1228,1998,2019]" captionStart="FIGURE 3" captionStartId="8.[151,250,1850,1872]" captionTargetBox="[159,1420,203,1800]" captionTargetId="figure@8.[151,1436,193,1828]" captionTargetPageId="8" captionText="FIGURE 3. Megisthanus modestus Berlese, 1910, specimens from Lamington NP. (A) female, sternogenital region; (B) internal genitalia of same specimen (shields and thickened margins not shown); (C) internal genitalia, different specimen (shield and thickened margins shown); (D) female, posterior dorsal shield and setae; (E) female, femur IV; (F) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600864/files/figure.png" pageId="36" pageNumber="37">Fig. 3A</figureCitation>
|
||
)................ 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB93E38D58015AFF6016F402CD410D" pageId="37" pageNumber="38">
|
||
<paragraph id="8BA08B468D58015AFF6016F402CD40C3" blockId="37.[151,1436,151,584]" pageId="37" pageNumber="38">
|
||
<keyLead id="30EE28738D58015AFF6016F402CD40C3" pageId="37" pageNumber="38">
|
||
4. Medial dorsal shield with two distinct
|
||
<typeStatus id="54A435E48D58015AFDA416F405D040A5" box="[595,646,151,172]" pageId="37" pageNumber="38">types</typeStatus>
|
||
of setae: a short, heavily barbed form and a long, smooth or sparsely barbed form (
|
||
<figureCitation id="132497C38D58015AFF1016D6067B40C3" box="[231,301,181,202]" captionStart="FIGURE 11" captionStartId="19.[151,250,1876,1898]" captionTargetBox="[159,1428,1009,1844]" captionTargetId="figure@19.[151,1436,996,1854]" captionTargetPageId="19" captionText="FIGURE 11. Mid-opisthonotal setae of: (A) Megisthanus simoneae sp. nov.; (B) Megisthanus thorelli Womersley, 1937." httpUri="https://zenodo.org/record/2600880/files/figure.png" pageId="37" pageNumber="38">Figs 11</figureCitation>
|
||
A–B)......................................................................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D58015AFF6016B102CD410D" blockId="37.[151,1436,151,584]" pageId="37" pageNumber="38">
|
||
<keyLead id="30EE28738D58015AFF6016B102CD410D" pageId="37" pageNumber="38">
|
||
- Medial dorsal shield with one
|
||
<typeStatus id="54A435E48D58015AFDF616B1057C40EE" box="[513,554,210,231]" pageId="37" pageNumber="38">type</typeStatus>
|
||
of seta, a long, smooth, barbed form (some long setae may be present on margins) (
|
||
<figureCitation id="132497C38D58015AFABF16B102C140EE" box="[1352,1431,210,231]" captionStart="FIGURE 1" captionStartId="6.[151,250,1918,1940]" captionTargetBox="[167,1421,199,1889]" captionTargetId="figure@6.[161,1425,195,1894]" captionTargetPageId="6" captionText="FIGURE 1. Megisthanus modestus Berlese, 1910, female, photomicrographs. (A) dorsal shield (secondarily sclerotised), specimen from Lamington NP; (B) dorsal shield (lacking secondary sclerotisation), specimen from Sydney; (C) venter, specimen from Lamington NP; (D) venter, specimen from Sydney. Scale bars = 200 µm." httpUri="https://zenodo.org/record/2600860/files/figure.png" pageId="37" pageNumber="38">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D58015AFF16168C065A410D" box="[225,268,239,260]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="37" pageNumber="38">12A</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D58015AFEE1168C0617410D" box="[278,321,239,260]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="37" pageNumber="38">21A</figureCitation>
|
||
)........................................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB93E38D58015AFF60176E02CA4154" pageId="37" pageNumber="38">
|
||
<paragraph id="8BA08B468D58015AFF60176E02CA412B" blockId="37.[151,1436,151,584]" box="[151,1436,268,290]" pageId="37" pageNumber="38">
|
||
<keyLead id="30EE28738D58015AFF60176E02CA412B" box="[151,1436,268,290]" pageId="37" pageNumber="38">
|
||
5. Medial opisthosoma with distinct patch of short pilose setae (
|
||
<figureCitation id="132497C38D58015AFCDA176E04D4412B" box="[813,898,269,290]" captionStart="FIGURE 11" captionStartId="19.[151,250,1876,1898]" captionTargetBox="[159,1428,1009,1844]" captionTargetId="figure@19.[151,1436,996,1854]" captionTargetPageId="19" captionText="FIGURE 11. Mid-opisthonotal setae of: (A) Megisthanus simoneae sp. nov.; (B) Megisthanus thorelli Womersley, 1937." httpUri="https://zenodo.org/record/2600880/files/figure.png" pageId="37" pageNumber="38">Fig. 11B</figureCitation>
|
||
)...................................
|
||
<taxonomicName id="4C1FF0C58D58015AFAC1176E02CA412B" authorityName="Womersley" authorityYear="1937" box="[1334,1436,268,290]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="thorelli">
|
||
<emphasis id="B96B57548D58015AFAC1176E021C412B" box="[1334,1354,269,290]" italics="true" pageId="37" pageNumber="38">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D58015AFAA1176F02CA412B" box="[1366,1436,268,290]" italics="true" pageId="37" pageNumber="38">thorelli</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D58015AFF60174902CA4154" blockId="37.[151,1436,151,584]" pageId="37" pageNumber="38">
|
||
<keyLead id="30EE28738D58015AFF60174902CA4154" pageId="37" pageNumber="38">
|
||
- Medial opisthosoma without distinct patch of short pilose setae (
|
||
<figureCitation id="132497C38D58015AFCA4174904FA4136" box="[851,940,298,319]" captionStart="FIGURE 11" captionStartId="19.[151,250,1876,1898]" captionTargetBox="[159,1428,1009,1844]" captionTargetId="figure@19.[151,1436,996,1854]" captionTargetPageId="19" captionText="FIGURE 11. Mid-opisthonotal setae of: (A) Megisthanus simoneae sp. nov.; (B) Megisthanus thorelli Womersley, 1937." httpUri="https://zenodo.org/record/2600880/files/figure.png" pageId="37" pageNumber="38">Fig. 11A</figureCitation>
|
||
), differentiation between long and short setal
|
||
<typeStatus id="54A435E48D58015AFA9E174902CA4136" box="[1385,1436,298,319]" pageId="37" pageNumber="38">types</typeStatus>
|
||
less distinct than on podosoma...........................................................
|
||
<taxonomicName id="4C1FF0C58D58015AFB3A172B021E4154" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[1229,1352,327,349]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="simoneae" status="sp. nov.">
|
||
<emphasis id="B96B57548D58015AFB3A172B03B74154" box="[1229,1249,328,349]" italics="true" pageId="37" pageNumber="38">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D58015AFB19172B021E4154" box="[1262,1352,328,349]" italics="true" pageId="37" pageNumber="38">simoneae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D58015AFAB8172402CA4154" bold="true" box="[1359,1436,327,349]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A258EA2F8D58015AFAB8172402CA4154" box="[1359,1436,327,349]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB93E38D58015AFF60170602CD419E" pageId="37" pageNumber="38">
|
||
<paragraph id="8BA08B468D58015AFF60170602CA4173" blockId="37.[151,1436,151,584]" box="[151,1436,356,378]" pageId="37" pageNumber="38">
|
||
<keyLead id="30EE28738D58015AFF60170602CA4173" box="[151,1436,356,378]" pageId="37" pageNumber="38">
|
||
6. Setae st4 present (
|
||
<figureCitation id="132497C38D58015AFE781706068A4173" box="[399,476,357,378]" captionStart="FIGURE 3" captionStartId="8.[151,250,1850,1872]" captionTargetBox="[159,1420,203,1800]" captionTargetId="figure@8.[151,1436,193,1828]" captionTargetPageId="8" captionText="FIGURE 3. Megisthanus modestus Berlese, 1910, specimens from Lamington NP. (A) female, sternogenital region; (B) internal genitalia of same specimen (shields and thickened margins not shown); (C) internal genitalia, different specimen (shield and thickened margins shown); (D) female, posterior dorsal shield and setae; (E) female, femur IV; (F) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600864/files/figure.png" pageId="37" pageNumber="38">Fig. 3A</figureCitation>
|
||
); host
|
||
<taxonomicName id="4C1FF0C58D58015AFDEB170705C64173" authorityName="Kaup" authorityYear="1868" box="[540,656,356,378]" class="Insecta" family="Passallidae" genus="Pharochilus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D58015AFDEB170705C64173" box="[540,656,356,378]" italics="true" pageId="37" pageNumber="38">Pharochilus</emphasis>
|
||
</taxonomicName>
|
||
spp.............................................
|
||
<taxonomicName id="4C1FF0C58D58015AFB271706021F4173" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[1232,1353,356,378]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="modestus" status="sp. nov.">
|
||
<emphasis id="B96B57548D58015AFB27170603B24173" box="[1232,1252,357,378]" italics="true" pageId="37" pageNumber="38">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D58015AFB071707021F4173" box="[1264,1353,356,378]" italics="true" pageId="37" pageNumber="38">modestus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D58015AFAB8170702CA4173" bold="true" box="[1359,1436,356,378]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A258EA2F8D58015AFAB8170702CA4173" box="[1359,1436,356,378]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D58015AFF6017E102CD419E" blockId="37.[151,1436,151,584]" box="[151,1435,385,407]" pageId="37" pageNumber="38">
|
||
<keyLead id="30EE28738D58015AFF6017E102CD419E" box="[151,1435,385,407]" pageId="37" pageNumber="38">
|
||
- Setae st4 absent (
|
||
<figureCitation id="132497C38D58015AFE7117E106B4419E" box="[390,482,386,407]" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="37" pageNumber="38">Figs 14A</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D58015AFE1B17E10541419E" box="[492,535,386,407]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="37" pageNumber="38">22B</figureCitation>
|
||
); host
|
||
<taxonomicName id="4C1FF0C58D58015AFDA017E20581419E" authorityName="Kaup" authorityYear="1868" box="[599,727,385,407]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D58015AFDA017E20581419E" box="[599,727,385,407]" italics="true" pageId="37" pageNumber="38">Aulacocyclus</emphasis>
|
||
</taxonomicName>
|
||
spp....................................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB93E38D58015AFF6017FC02CA4241" pageId="37" pageNumber="38">
|
||
<paragraph id="8BA08B468D58015AFF6017FC02CA41F9" blockId="37.[151,1436,151,584]" pageId="37" pageNumber="38">
|
||
<keyLead id="30EE28738D58015AFF6017FC02CA41F9" pageId="37" pageNumber="38">
|
||
7. Podonotum and opisthonotum densely setose (
|
||
<figureCitation id="132497C38D58015AFD5717FC05AC41BD" box="[672,762,415,436]" captionStart="FIGURE 12" captionStartId="22.[151,250,1822,1844]" captionTargetBox="[157,1428,207,1794]" captionTargetId="figure@22.[151,1436,193,1801]" captionTargetPageId="22" captionText="FIGURE 12. Megisthanus southcotti sp. nov. photomicrographs. (A) female, dorsal shield; (B) female, venter; (C) major male, venter; (D) major male, sternoventral shield. Scale bars: 200 µm (A–C), 100 µm (D)." httpUri="https://zenodo.org/record/2600882/files/figure.png" pageId="37" pageNumber="38">Fig. 12A</figureCitation>
|
||
); genital setae long, 60–110; with 5–11 setae behind margin of genital shield (
|
||
<figureCitation id="132497C38D58015AFEDD17DE06D141DB" box="[298,391,445,466]" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="37" pageNumber="38">Fig. 14A</figureCitation>
|
||
); male sternoventral shield highly hypertrichous around genital opening (approximately 50-60 setae;
|
||
<figureCitation id="132497C38D58015AFA8217DE065C41E6" captionStart="FIGURE 14" captionStartId="24.[151,250,1705,1727]" captionTargetBox="[180,1394,196,1617]" captionTargetId="figure@24.[151,1436,193,1684]" captionTargetPageId="24" captionText="FIGURE 14. Megisthanus southcotti sp. nov. (A) female, sternogenital region; (B) internal genitalia; (C) female, femur IV; (D) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600886/files/figure.png" pageId="37" pageNumber="38">Fig. 14D</figureCitation>
|
||
)...............................................................................
|
||
<taxonomicName id="4C1FF0C58D58015AFB3D17B9021C41E6" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[1226,1354,473,495]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="southcotti" status="sp. nov.">
|
||
<emphasis id="B96B57548D58015AFB3D17B9038841E6" box="[1226,1246,474,495]" italics="true" pageId="37" pageNumber="38">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D58015AFB1C17BA021C41E6" box="[1259,1354,473,495]" italics="true" pageId="37" pageNumber="38">southcotti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D58015AFAB817B902CA41F9" bold="true" box="[1359,1436,474,496]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A258EA2F8D58015AFAB817B902CA41F9" box="[1359,1436,474,496]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D58015AFF60179402CA4241" blockId="37.[151,1436,151,584]" pageId="37" pageNumber="38">
|
||
<keyLead id="30EE28738D58015AFF60179402CA4241" pageId="37" pageNumber="38">
|
||
- Podonotum more setose than opisthonotum (
|
||
<figureCitation id="132497C38D58015AFD6F179405A24205" box="[664,756,503,524]" captionStart="FIGURE 21" captionStartId="33.[151,250,1957,1979]" captionTargetBox="[170,1410,633,1925]" captionTargetId="figure@33.[161,1425,625,1936]" captionTargetPageId="33" captionText="FIGURE 21. Megisthanus zachariei sp. nov. photomicrographs. (A) female dorsal shield; (B) female venter; (C) female posterior dorsal shield; (D) major male ventrianal shield. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600900/files/figure.png" pageId="37" pageNumber="38">Fig. 21A</figureCitation>
|
||
); genital setae short, 20–35; with 0–2 setae behind margin of genital shield (
|
||
<figureCitation id="132497C38D58015AFED0147606D74223" box="[295,385,533,554]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="37" pageNumber="38">Fig. 22A</figureCitation>
|
||
); male sternoventral shield much less hypertrichous around genital opening (at most 20 setae;
|
||
<figureCitation id="132497C38D58015AFAFE1476020A4223" box="[1289,1372,533,554]" captionStart="FIGURE 22" captionStartId="34.[151,250,1933,1955]" captionTargetBox="[199,1380,210,1889]" captionTargetId="figure@34.[186,1410,193,1915]" captionTargetPageId="34" captionText="FIGURE 22. Megisthanus zachariei sp. nov. (A) female, posterior dorsal shield; (B) female, sternogenital region; (C) internal genitalia; (D) female, femur IV; (E) major male, sternogenital region. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600902/files/figure.png" pageId="37" pageNumber="38">Fig. 22E</figureCitation>
|
||
)........................................................................................
|
||
<taxonomicName id="4C1FF0C58D58015AFB271451021F424E" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[1232,1353,561,583]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="zachariei" status="sp. nov.">
|
||
<emphasis id="B96B57548D58015AFB27145103B2424E" box="[1232,1252,562,583]" italics="true" pageId="37" pageNumber="38">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D58015AFB071452021F424E" box="[1264,1353,561,583]" italics="true" pageId="37" pageNumber="38">zachariei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D58015AFAB8145102CA4241" bold="true" box="[1359,1436,562,584]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A258EA2F8D58015AFAB8145102CA4241" box="[1359,1436,562,584]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |