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<document ID-DOI="http://doi.org/10.5281/zenodo.5193614" ID-GBIF-Dataset="a02f182b-32e4-4649-afe0-983cb21ceb9e" ID-GBIF-Taxon="160690014" ID-Zenodo-Dep="5193614" approvalRequired="5" approvalRequired_for_illustrations="3" approvalRequired_for_treatments="2" checkinTime="1576077498408" checkinUser="jeremy" docAuthor="M. A. Lieftinck" docDate="1940" docId="03B487FFFF8EFFE6FF07BCEEFC2FAD79" docLanguage="en" docName="Lieftinck - 1940 - Descriptions and records of South-east Asiatic Odonata (II) ABBYYed.pdf.imf" docOrigin="Treubia 17 (4)" docStyle="DocumentStyle{}" docTitle="Coeliccia arcuata Lieftinck, 1940, sp. n." docType="treatment" docVersion="13" lastPageId="18" lastPageNumber="354" masterDocId="FF8DFF87FF9EFFF4FFF0BE36FF92AF52" masterDocTitle="Descriptions and records of South-East Asiatic Odonata (II)" masterLastPageNumber="392" masterPageNumber="337" pageId="16" pageNumber="353" updateTime="1632324461921" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Descriptions and records of South-East Asiatic Odonata (II)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>M. A. Lieftinck</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Treubia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>1940</mods:date>
<mods:detail type="pubDate">
<mods:number>1940-01-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>17</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>337</mods:start>
<mods:end>392</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3571116</mods:identifier>
<mods:identifier type="GBIF-Dataset">a02f182b-32e4-4649-afe0-983cb21ceb9e</mods:identifier>
<mods:identifier type="Zenodo-Dep">3571116</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5193614" ID-GBIF-Taxon="160690014" ID-Zenodo-Dep="5193614" LSID="urn:lsid:plazi:treatment:03B487FFFF8EFFE6FF07BCEEFC2FAD79" httpUri="http://treatment.plazi.org/id/03B487FFFF8EFFE6FF07BCEEFC2FAD79" lastPageId="18" lastPageNumber="354" pageId="16" pageNumber="353">
<subSubSection box="[247,945,728,764]" pageId="16" pageNumber="353" type="nomenclature">
<paragraph blockId="16.[0,1933,0,2949]" box="[247,945,728,764]" pageId="16" pageNumber="353">
<emphasis bold="true" box="[247,601,728,764]" pageId="16" pageNumber="353">
<taxonomicName authorityName="M.A. Lieftinck" authorityYear="1940" box="[247,592,728,764]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="arcuata" status="sp. n.">Coeliccia arcuata</taxonomicName>
,
</emphasis>
<taxonomicNameLabel box="[627,719,728,764]" pageId="16" pageNumber="353" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation box="[756,919,728,764]" captionStart="Fig" captionStart-0="Fig" captionStart-1="Fig" captionStartId="17.[225,275,1195,1223]" captionStartId-0="17.[225,275,1195,1223]" captionStartId-1="17.[324,386,2232,2264]" captionTargetBox="[228,1073,591,1173]" captionTargetBox-0="[228,1073,591,1173]" captionTargetBox-1="[322,1601,1617,2220]" captionTargetPageId="17" captionTargetPageId-0="17" captionTargetPageId-1="17" captionText="Fig . 7 . Coeliccia arcuata , sp . n . ♂ . Anal ap ­ pendages , dorsal view and right side ." captionText-0="Fig. 7. Coeliccia arcuata, sp. n. ♂. Anal ap ­ pendages, dorsal view and right side ." captionText-1="Fig. 8 8.. Coeliccia arcuata, sp. n. ♀ ♀.. Prothorax, dorsal view and right side ." figureDoi="http://doi.org/10.5281/zenodo.3571133" figureDoi-0="http://doi.org/10.5281/zenodo.3571133" figureDoi-1="http://doi.org/10.5281/zenodo.3571137" httpUri="https://zenodo.org/record/3571133/files/figure.png" httpUri-0="https://zenodo.org/record/3571133/files/figure.png" httpUri-1="https://zenodo.org/record/3571137/files/figure.png" pageId="16" pageNumber="353">fig. 7 - 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="353" type="materials_examined">
<paragraph blockId="16.[0,1933,0,2949]" pageId="16" pageNumber="353">
<materialsCitation ID-GBIF-Occurrence="2467684757" collectingDateMax="1937-06" collectingDateMin="1934-04" collectorName="M. E. Walsh" country="Malaysia" location="E. Borneo, N. Koetai, Sangkoelirang distr., Kariorang, Batau Besi and Babi Djoeton" pageId="16" pageNumber="353" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="E. Borneo">
Material studied: —
<specimenCount box="[747,896,785,821]" pageId="16" pageNumber="353" type="male">2 males</specimenCount>
,
<specimenCount box="[925,1109,785,821]" pageId="16" pageNumber="353" type="female">3 females</specimenCount>
, E.
<collectingRegion box="[1201,1366,785,821]" pageId="16" pageNumber="353">Borneo</collectingRegion>
,
<collectingMunicipality box="[1418,1610,785,821]" pageId="16" pageNumber="353">N. Koetai</collectingMunicipality>
,
<collectingRegion pageId="16" pageNumber="353">Sang- koelirang distr</collectingRegion>
.,
<location LSID="urn:lsid:plazi:treatment:03B487FFFF8EFFE6FF07BCEEFC2FAD79:8EC26032FF8EFFE4FDC7BD70FD67AC38" box="[567,757,838,874]" country="Borneo" municipality="N. Koetai" name="Kariorang" pageId="16" pageNumber="353" stateProvince="Borneo">Kariorang</location>
,
<location LSID="urn:lsid:plazi:treatment:03B487FFFF8EFFE6FF07BCEEFC2FAD79:8EC26032FF8EFFE4FCE3BD70FACCAC38" box="[787,1374,838,874]" country="Borneo" municipality="N. Koetai" name="Batau Besi and Babi Djoeton" pageId="16" pageNumber="353" stateProvince="Borneo">Batau Besi and Babi Djoeton</location>
,
<date box="[1405,1610,838,874]" pageId="16" pageNumber="353" value="1934-04">
<collectingDate box="[1405,1610,838,874]" pageId="16" pageNumber="353" value="1934-04">April 1934</collectingDate>
</date>
,
<collectingDate pageId="16" pageNumber="353">May- June</collectingDate>
,
<collectingDate box="[367,450,894,928]" pageId="16" pageNumber="353">1937</collectingDate>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2467684747" box="[479,1209,892,928]" collectingCode="Buitenzorg Museum" collectingDateMax="1937-06" collectingDateMin="1937-05" collectorName="M. E. Walsh" country="Malaysia" location="Batau Besi" pageId="16" pageNumber="353" specimenCount="1" specimenCount-male="1" stateProvince="E. Borneo" typeStatus="holotype">
<collectorName box="[479,731,892,928]" pageId="16" pageNumber="353">M. E. Walsh</collectorName>
leg.
<typeStatus box="[835,1068,892,928]" pageId="16" pageNumber="353">Holotype</typeStatus>
<specimenCount box="[1079,1107,892,928]" pageId="16" pageNumber="353" type="male">♂</specimenCount>
and
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2467684722" collectingCode="Buitenzorg Museum" collectingDateMax="1937-06" collectingDateMin="1937-05" collectorName="M. E. Walsh" country="Malaysia" location="Batau Besi" pageId="16" pageNumber="353" specimenCount="1" specimenCount-female="1" stateProvince="E. Borneo" typeStatus="allotype">
<typeStatus box="[1233,1441,892,928]" pageId="16" pageNumber="353">allotype</typeStatus>
<specimenCount box="[1471,1494,892,928]" pageId="16" pageNumber="353" type="female">♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03B487FFFF8EFFE6FF07BCEEFC2FAD79:8EC26032FF8EFFE4FA05BD4AF95AACF2" box="[1525,1736,892,928]" country="Borneo" municipality="N. Koetai" name="Batau Besi" pageId="16" pageNumber="353" stateProvince="Borneo">Batau Besi</location>
,
<collectingDate box="[249,440,945,981]" pageId="16" pageNumber="353">May-June</collectingDate>
,
<collectingDate box="[474,557,947,981]" pageId="16" pageNumber="353">1937</collectingDate>
, in the Buitenzorg Museum
</materialsCitation>
.
</paragraph>
<paragraph blockId="16.[0,1933,0,2949]" pageId="16" pageNumber="353">
<emphasis box="[331,423,1000,1036]" italics="true" pageId="16" pageNumber="353">Male</emphasis>
(semiad.). — Labium pale orange-yellow. Labrum wholly black. Mandible-bases, genae and anteclypeus bright blue. Postclypeus, frons and vertex black, postclypeus very shining, base and apex of second joint of antennae and a pair of elongate transverse marks just posterior to the antennae, on either side connecting each lateral ocellus with the margin of compound eye, blue. These head-marks are irregular in shape, rather expanded mesially, narrower and linear laterally. Remainder of head black, save for a pair of very small, transverse, blue postocular spots, restricted to the middle of the epicranial lobes and just visible in dorsal view.
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="355" pageId="16" pageNumber="353" type="description">
<paragraph blockId="16.[0,1933,0,2949]" pageId="16" pageNumber="353">Prothorax with anterior and posterior lobes black, middle lobe blue with a fine median black line. Transverse lateral suture black. Posterior lobe simple, its free margin with a very shallow concavity so as to form a minute obtuseangulate projection on either side of it.</paragraph>
<paragraph blockId="16.[0,1933,0,2949]" pageId="16" pageNumber="353">Synthorax bronzy-black above, light orange-yellow at the sides and underneath; antehumeral stripes broad, complete, gently tapering to a point dorsally and ceasing about 0.5 mm before ante-alar triangles, which are black; at the top of either shoulder and outdistancing the stripes, lies an oval yellow spot. On the side of the thorax a black line, narrowly connected dorsally with the black of the mesepimeron, on the second lateral suture, stopping short 0.5 mm before the spiracle.</paragraph>
<paragraph blockId="16.[0,1933,0,2949]" pageId="16" pageNumber="353">Legs with the coxae and trochanters pale orange-yellow. Femora yellow with fine blackish exterior line and with the knees also obscured; tibiae and tarsi dirty yellowish, tibiae finely black interiorly and with black apices, distal half of last tarsal joint and claws black. Spines brown.</paragraph>
<paragraph blockId="16.[0,1933,0,2949]" pageId="16" pageNumber="353">
Wings hyaline. Neuration very similar to
<emphasis box="[1163,1386,2287,2323]" italics="true" pageId="16" pageNumber="353">
<taxonomicName authorityName="Laidlaw" authorityYear="1918" box="[1163,1379,2287,2323]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="flavostriata">flavostriata</taxonomicName>
,
</emphasis>
L aid. Costal side of
<emphasis box="[306,327,2342,2378]" italics="true" pageId="16" pageNumber="353">q</emphasis>
a trifle longer in both pairs of wings, and hence distal side slightly less oblique than in that species. Pterostigma blackish-brown, shaped much as in
<taxonomicName authorityName="Laidlaw" authorityYear="1918" box="[246,457,2449,2485]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="flavostriata">
<emphasis box="[246,457,2449,2485]" italics="true" pageId="16" pageNumber="353">flavostriata</emphasis>
</taxonomicName>
but a trifle longer, and costal side very slightly shorter than anal side. 14 postnodals in fore wing, 13 in hind wing.
</paragraph>
<paragraph blockId="16.[0,1933,0,2949]" lastBlockId="17.[221,1718,309,559]" lastPageId="17" lastPageNumber="354" pageId="16" pageNumber="353">Abdomen slender. Segm. 1 bright yellow, with a thick median line of black, 2 yellow-brown, paler below, with narrow black apical ring; 3 - 6 each with an indistinct, transverse basal yellow ring, interrupted mid-dorsally (most conspicuous on 3). and narrow black apical rings. The remaining segments progressively darker, 9 10 black; 9 with a large blue dorsal patch, bluntly pointed basallv, and 10 with a mid-dorsal point of the same colour, at extreme base. Articulation between 9 and 10 likewise blue.</paragraph>
<paragraph blockId="17.[224,1723,204,258]" box="[224,1275,206,243]" pageId="17" pageNumber="354">
<pageNumber ambiguity="4" box="[224,285,207,243]" fuzzyness="0" pageId="17" pageNumber="354" score="264.0" value="354">354</pageNumber>
Treubia Deel 17, 1940, Afl. 4.
</paragraph>
<paragraph blockId="17.[1081,1723,577,1312]" pageId="17" pageNumber="354">
Anal appendages black, superior pair longer than segm. 10, inferiors a trifle longer than the superiors. Superior pair cylindrical, rather stout, very slightly incurved in dorsal view, evenly but- strongly downbent in side-view; each carries a short, acute, internal tooth (not visible in dorsal view) at the end of its basal fifth, which is directed ventrad and slightly basad. Inferior appendages slender, cylindrical, incurved apically (
<figureCitation box="[1097,1193,1274,1312]" captionStart="Fig" captionStartId="17.[225,275,1195,1223]" captionTargetBox="[228,1073,591,1173]" captionTargetPageId="17" captionText="Fig. 7. Coeliccia arcuata, sp. n. ♂. Anal ap ­ pendages, dorsal view and right side ." figureDoi="http://doi.org/10.5281/zenodo.3571133" httpUri="https://zenodo.org/record/3571133/files/figure.png" pageId="17" pageNumber="354">fig. 7</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3571133" ID-Zenodo-Dep="3571133" httpUri="https://zenodo.org/record/3571133/files/figure.png" pageId="17" pageNumber="354" startId="17.[225,275,1195,1223]" targetBox="[228,1073,591,1173]" targetPageId="17">
<paragraph pageId="17" pageNumber="354">
Fig. 7.
<taxonomicName authorityName="M.A. Lieftinck" authorityYear="1940" box="[378,676,1192,1224]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="353" phylum="Arthropoda" rank="species" species="arcuata" status="sp. n.">
<emphasis box="[378,525,1192,1224]" italics="true" pageId="17" pageNumber="354">Coeliccia</emphasis>
<emphasis box="[546,676,1192,1224]" italics="true" pageId="17" pageNumber="354">arcuata</emphasis>
</taxonomicName>
<emphasis box="[676,682,1192,1224]" italics="true" pageId="17" pageNumber="354">,</emphasis>
sp. n. ♂. Anal ap ­ pendages, dorsal view and right side
</paragraph>
<paragraph box="[941,946,1231,1259]" pageId="17" pageNumber="354">.</paragraph>
</caption>
<paragraph blockId="17.[218,1715,1327,1526]" pageId="17" pageNumber="354">
Female (ad). — Differs from the male only in details of coloration and in the shape of the prothorax. Posterior lobe of prothorax shaped as shown in
<figureCitation box="[277,374,1434,1472]" captionStart="Fig" captionStartId="17.[324,386,2232,2264]" captionTargetBox="[322,1601,1617,2220]" captionTargetPageId="17" captionText="Fig . 8 8 . . Coeliccia arcuata , sp . n . ♀ ♀ . . Prothorax , dorsal view and right side ." figureDoi="http://doi.org/10.5281/zenodo.3571137" httpUri="https://zenodo.org/record/3571137/files/figure.png" pageId="17" pageNumber="354">fig. 8</figureCitation>
, the median division of the lobe depressed. Antehumeral stripes and metepisternum blue.
</paragraph>
<paragraph lastBlockId="17.[213,1712,2290,2645]" pageId="17" pageNumber="354">Wings with 12-14/11-13 postnodals. Abdomen coloured as in male. Distal 2/3 to 1/3 of segm. 8 entirely, and a thick stripe bordering the tergal margin, bright blue; 9 and 10 and ann] appendages wholly black. Valves very long, with slightly convex ventral border, projecting beyond apex of segun. 10 for almost twice t-he length of this segment.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3571137" ID-Zenodo-Dep="3571137" box="[324,1590,2231,2264]" httpUri="https://zenodo.org/record/3571137/files/figure.png" pageId="17" pageNumber="354" startId="17.[324,386,2232,2264]" targetBox="[322,1601,1617,2220]" targetPageId="17">
<paragraph box="[324,1585,2231,2264]" pageId="17" pageNumber="354">
Fig. 8 8..
<taxonomicName authorityName="M.A. Lieftinck" authorityYear="1940" box="[471,762,2232,2264]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="353" phylum="Arthropoda" rank="species" species="arcuata" status="sp. n.">
<emphasis bold="true" box="[471,615,2232,2264]" italics="true" pageId="17" pageNumber="354">Coeliccia</emphasis>
<emphasis bold="true" box="[633,762,2232,2264]" italics="true" pageId="17" pageNumber="354">arcuata</emphasis>
</taxonomicName>
<emphasis bold="true" box="[762,770,2232,2264]" italics="true" pageId="17" pageNumber="354">,</emphasis>
sp. n. ♀ ♀.. Prothorax, dorsal view and right side
</paragraph>
<paragraph box="[1585,1590,2232,2264]" pageId="17" pageNumber="354">.</paragraph>
</caption>
<paragraph blockId="17.[213,1712,2290,2645]" lastBlockId="18.[249,1746,302,555]" lastPageId="18" lastPageNumber="355" pageId="17" pageNumber="354">
Length: ♂ abd. + app. 35 35.5, hw. 20.5 21; ♀ 32 - 365 (incl. valves). 21 - 24 mm. This species falls in Laidlaw ’s Group 3, of
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[1232,1471,307,343]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="membranipes">
<emphasis box="[1232,1471,307,343]" italics="true" pageId="18" pageNumber="355">membranipes</emphasis>
</taxonomicName>
(Ramb.), and seems to find its nearest allies in
<taxonomicName authorityName="Laidlaw" authorityYear="1918" box="[909,1121,360,396]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="flavostriata">
<emphasis box="[909,1121,360,396]" italics="true" pageId="18" pageNumber="355">flavostriata</emphasis>
</taxonomicName>
Laid.,
<taxonomicName authorityName="Laidlaw" authorityYear="1918" box="[1270,1436,360,396]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="campioni">
<emphasis box="[1270,1436,360,396]" italics="true" pageId="18" pageNumber="355">campioni</emphasis>
</taxonomicName>
Laid.,
<taxonomicName authorityName="Laidlaw" authorityYear="1932" box="[1583,1745,360,396]" class="Insecta" family="Platycnemididae" genus="Coeliccia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis box="[1583,1745,360,396]" italics="true" pageId="18" pageNumber="355">lieftincki</emphasis>
</taxonomicName>
Laid., and the species next to be described as
<emphasis box="[1155,1332,414,450]" italics="true" pageId="18" pageNumber="355">
<taxonomicName authorityName="M.A. Lieftinck" authorityYear="1940" box="[1155,1325,414,450]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="coomansi" status="sp. n.">coomansi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel box="[1353,1446,414,450]" pageId="18" pageNumber="355" rank="species">sp. n.</taxonomicNameLabel>
1). It is easily distinguished from all these by the different shape of the♂ anal appendages and the structure of the ♀ prothorax.
</paragraph>
</subSubSection>
</treatment>
</document>
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