treatments-xml/data/03/A8/D7/03A8D74FFF9BFFF1C9C2FC8D4588F86F.xml

<document id="664BAB617B025F623AC8CB1F8009BDEC" ID-CLB-Dataset="29235" ID-DOI="10.11646/zootaxa.4497.1.1" ID-GBIF-Dataset="d5b94fdb-db02-4530-85aa-12d9757e3313" ID-ISSN="1175-5326" ID-Zenodo-Dep="1451056" ID-ZooBank="890E26F1-1B03-4F13-9299-A5E8BED8439D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538973324849" checkinUser="plazi" docAuthor="Grehan, John R. &amp; Mielke, Os. G. C." docDate="2018" docId="03A8D74FFF9BFFF1C9C2FC8D4588F86F" docLanguage="en" docName="zootaxa.4497.1.1.pdf" docOrigin="Zootaxa 4497 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kozloviella viazmenskyi ♂, Grehan &amp; C. Mielke 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="FF91AF37FF98FFF4C955FF924474FFBB" masterDocTitle="New species of Dugdaleiella, gen. nov., Kozloviella, gen. nov., and Pfitzneriella Viette from upper elevation Andes of Ecuador and Peru (Lepidoptera: Hepialidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="4" updateTime="1698567105610" updateUser="ExternalLinkService">
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<mods:title id="196332E8609DB11D83C7194C146D8789">New species of Dugdaleiella, gen. nov., Kozloviella, gen. nov., and Pfitzneriella Viette from upper elevation Andes of Ecuador and Peru (Lepidoptera: Hepialidae)</mods:title>
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<mods:namePart id="AAF836CE0BC2D0FC8C60243761A8EECD">Grehan, John R.</mods:namePart>
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<mods:namePart id="2064F44E5AC2A51027DDA29F4AEB186C">Mielke, Os. G. C.</mods:namePart>
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<mods:date id="9EF99A088DDC0E4006D34D08C1C6E645">2018</mods:date>
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<treatment id="03A8D74FFF9BFFF1C9C2FC8D4588F86F" ID-DOI="http://doi.org/10.5281/zenodo.5957831" ID-GBIF-Taxon="148651435" ID-Zenodo-Dep="5957831" LSID="urn:lsid:plazi:treatment:03A8D74FFF9BFFF1C9C2FC8D4588F86F" httpUri="http://treatment.plazi.org/id/03A8D74FFF9BFFF1C9C2FC8D4588F86F" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<subSubSection id="C31B35D2FF9BFFF7C9C2FC8D46BFFCE0" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BBE6659FF9BFFF7C9C2FC8D4674FC81" blockId="3.[151,715,799,859]" box="[151,512,799,826]" pageId="3" pageNumber="4">
<heading id="D0F6D135FF9BFFF7C9C2FC8D4674FC81" bold="true" box="[151,512,799,826]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="B975BA4BFF9BFFF7C9C2FC8D4674FC81" bold="true" box="[151,512,799,826]" pageId="3" pageNumber="4">
<taxonomicName id="4C011DDAFF9BFFF7C9C2FC8D45C3FC82" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[151,439,799,825]" class="Insecta" family="Hepialidae" genus="Kozloviella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="viazmenskyi" status="sp. nov.">
<emphasis id="B975BA4BFF9BFFF7C9C2FC8D45C3FC82" bold="true" box="[151,439,799,825]" italics="true" pageId="3" pageNumber="4">Kozloviella viazmenskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9BFFF7C8E8FCB24674FC81" box="[445,512,800,826]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE6659FF9BFFF7C9C2FCD046BFFCE0" blockId="3.[151,715,799,859]" box="[151,715,834,859]" pageId="3" pageNumber="4">
(
<figureCitation id="133A7ADCFF9BFFF7C9CAFCD04569FCE0" box="[159,285,834,859]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="3" pageNumber="4">Figs 1a, 1b</figureCitation>
, 
<figureCitation id="133A7ADCFF9BFFF7C87FFCD14533FCE0" box="[298,327,835,859]" captionStart="FIGURES 8–9" captionStartId="5.[151,264,1506,1529]" captionTargetBox="[151,1435,193,1486]" captionTargetId="figure@5.[151,1435,193,1486]" captionTargetPageId="5" captionText="FIGURES 8–9. Voucher specimens. 8, Dugdaleiella monticola male. 8a, LT (ZMHB); 8b, Cotopaxi, Ecuador (CMNH); 8c, Volcán Chiles, Ecuador (CMNH); 8d, male, Volcán Chiles, Ecuador (CMNH). 9, Labial palpi (male unless otherwise specified), 9a, Kozloviella viazmenskyi sp. n.; 9b, Pfitzneriella antonkozlovi sp. n.; 9c, P. rawlinsi sp. n.; 9d, P. titarenkoi sp. n.; 9e, P. yuliyakovalevae sp. n.; 9f, P. yuliyakovalevae sp. n. (female); 9g, Dugdaleiella monticola. Photos: Jane Hyland (Fig. 8), John Grehan (Figs 9a–b, d–h), and Carlos Mielke (Fig. 9c). Scale 1 mm (when present)." httpUri="https://zenodo.org/record/1451060/files/figure.png" pageId="3" pageNumber="4">9a</figureCitation>
, 
<figureCitation id="133A7ADCFF9BFFF7C807FCD14509FCE0" box="[338,381,835,859]" captionStart="FIGURE 10" captionStartId="6.[151,250,1649,1671]" captionTargetBox="[152,1437,195,1610]" captionTargetId="figure@6.[152,1437,195,1610]" captionTargetPageId="6" captionText="FIGURE 10. Wing venation (male unless otherwise specified). 10a, Kozloviella viazmenskyi sp. n.; 10b, P. antonkozlovi sp. n.; 10c, P. olafi sp. n.; 10d, P. rawlinsi sp. n.; 10e, P. remota (from photo only, venation incomplete due to damaged condition of HT); 10f, P. yuliyakovalevae sp. n.; 10g, P. yuliyakovalevae sp. n. (female); 10h, Dugdaleiella monticola." httpUri="https://zenodo.org/record/1451062/files/figure.png" pageId="3" pageNumber="4">10a</figureCitation>
, 
<figureCitation id="133A7ADCFF9BFFF7C8DCFCD145C6FCE0" box="[393,434,835,859]" captionStart="FIGURE 11" captionStartId="7.[151,250,1429,1451]" captionTargetBox="[150,1436,810,1407]" captionTargetId="figure@7.[150,1436,810,1407]" captionTargetPageId="7" captionText="FIGURE 11. Wing markings. 11a; Kozloviella viazmenskyi sp. n., 11b; Pfitzneriella antonkozlovi sp. n.; 11c, P. olafi sp. n.; 11d, P. rawlinsi sp. n.; 11e, P. remota; 11f, P. yuliyakovalevae sp. n. Blue line—edge of white banding." httpUri="https://zenodo.org/record/1451064/files/figure.png" pageId="3" pageNumber="4">11a</figureCitation>
, 
<figureCitation id="133A7ADCFF9BFFF7C8EBFCD1459DFCE0" box="[446,489,835,859]" captionStart="FIGURES 12–14. 12" captionStartId="9.[151,264,1172,1195]" captionTargetBox="[150,1436,193,1151]" captionTargetId="figure@9.[150,1436,193,1151]" captionTargetPageId="9" captionText="FIGURES 12–14. 12, Antenna. Pfitzneriella rawlinsi sp. n. (unscaled). 13, Legs. 13a, Kozloviella viazmenskyi sp. n.; 13b, P. olafi sp. n.; 13c, P. rawlinsi sp. n.; 13d, P. titarenkoi sp. n.; 13e, P. yuliyakovalevae sp. n. 14, Metatibia. 14a, P. antonkozlovi sp. n.; 14b, P. rawlinsi sp. n.; 14c, P. titarenkoi; 14d, P. yuliyakovalevae sp. n.; 14e, P. antonkozlovi sp. n.; 14f, Dugdaleiella monticola. Photos: John Grehan (Figs 12, 14e, 14f), Carlos Mielke (Figs 13,14a–d). Scale 1 mm." httpUri="https://zenodo.org/record/1451066/files/figure.png" pageId="3" pageNumber="4">13a</figureCitation>
, 
<figureCitation id="133A7ADCFF9BFFF7C8A0FCD04622FCE0" box="[501,598,834,859]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[197,1386,199,1851]" captionTargetId="figure@10.[197,1386,199,1851]" captionTargetPageId="10" captionText="FIGURES 15–16. Tergosternal connection. 15a, Kozloviella viazmenskyi sp. n.; 15b, Pfitzneriella yuliyakovalevae sp. n.; 15c, P. rawlinsi sp. n.; 15d, Dugdaleiella monticola. 16, Abdomen (males unless otherwise specified). 16a, K. viazmenskyi sp. n.; 16b, P. antonkozlovi sp. n.; 16c, P. olafi sp. n.; 16d, P. rawlinsi sp. n.; 16e, P. titarenkoi sp. n.; 16f, P. yuliyakovalevae sp. n.; 16g, P. yuliyakovalevae sp. n. (female); 16h–16j, Dugdaleiella monticola. 16h, tergum II; 16i, sternum II; 16j, tergum and sternum VIII. Photos: John Grehan (Figs 15d, 16h–16j), Carlos Mielke (Figs 15a–15c, 16a–16g)." httpUri="https://zenodo.org/record/1451068/files/figure.png" pageId="3" pageNumber="4">15a, 16a</figureCitation>
, 
<figureCitation id="133A7ADCFF9BFFF7CB34FCD146B6FCE0" box="[609,706,834,859]" captionStart="FIGURES 17–25" captionStartId="11.[151,264,1644,1667]" captionTargetBox="[151,1435,193,1629]" captionTargetId="figure@11.[151,1435,193,1629]" captionTargetPageId="11" captionText="FIGURES 17–25. Genitalia (male unless otherwise specified). 17, Kozloviella viazmenskyi sp. n. HT. 17a, ventral; 17b, lateral. 18, Pfitzneriella antonkozlovi sp. n. HT, ventral. 19, P. olafi sp. n. HT, ventral. 20, P. rawlinsi sp. n. HT, ventral. 21, P. remota HT (redrawn from Viette (1951a)). 22, P. titarenkoi sp. n. HT, ventral. 23, P. yuliyakovalevae sp. n. 23a, ventral; 23b, lateral. 24, P. yuliyakovalevae sp. n. PT female. 24a, posterior; 24b, lateral. 25, Dugdaleiella monticola. 25a, LT (ZMHB); 25b, Cotopaxi, Parque Nacional Cotopaxi. Dissection JRG 136 (CMNH). sa = saccus, fs = fultura superior, fi = fultura inferior, as = apodemal suture. Scale 1 mm (when present). Photos: Carlos Mielke (Figs 17–20, 22–25b)." httpUri="https://zenodo.org/record/1451070/files/figure.png" pageId="3" pageNumber="4">17a, 17b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF9BFFF7C9C2FC1845F5FB88" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BBE6659FF9BFFF7C9C2FC1845F5FB88" blockId="3.[151,1437,906,1975]" pageId="3" pageNumber="4">
<emphasis id="B975BA4BFF9BFFF7C9C2FC18457EFC18" bold="true" box="[151,266,906,931]" pageId="3" pageNumber="4">Diagnosis</emphasis>
. Similar in general color and FW texture to many species of the cibyrine clade (sensu Grehan 2012), but contrasting with narrower forewings and slightly concave costal margin, and particularly a narrow and oblique, black V-shaped line that appears to be unique to this species. The yellowish orange color of this species contrasts with the grayish brown of 
<taxonomicName id="4C011DDAFF9BFFF7C8B4FC644607FBB4" authority="Grehan &amp; Mielke, 2018" authorityName="Grehan &amp; Mielke" authorityYear="2018" box="[481,627,1014,1039]" class="Insecta" family="Hepialidae" genus="Dugdaleiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975BA4BFF9BFFF7C8B4FC644607FBB4" box="[481,627,1014,1039]" italics="true" pageId="3" pageNumber="4">Dugdaleiella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9BFFF7CBD4FC6446A3FBB4" box="[641,727,1014,1039]" pageId="3" pageNumber="4" rank="genus">
<emphasis id="B975BA4BFF9BFFF7CBD4FC6446A3FBB4" bold="true" box="[641,727,1014,1039]" pageId="3" pageNumber="4">gen. n.</emphasis>
</taxonomicNameLabel>
and the yellowish brown to grayish brown color tones of 
<taxonomicName id="4C011DDAFF9BFFF7C9C2FB884556FB88" authorityName="Grehan &amp; Mielke" authorityYear="2018" box="[151,290,1050,1075]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B975BA4BFF9BFFF7C9C2FB884556FB88" box="[151,290,1050,1075]" italics="true" pageId="3" pageNumber="4">Pfitzneriella</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF9BFFF7C992FBAC45E4F8F0" pageId="3" pageNumber="4" type="description">
<paragraph id="8BBE6659FF9BFFF7C992FBAC466BFBC0" blockId="3.[151,1437,906,1975]" pageId="3" pageNumber="4">
<emphasis id="B975BA4BFF9BFFF7C992FBAC4525FBEC" bold="true" box="[199,337,1086,1111]" pageId="3" pageNumber="4">Description</emphasis>
. Male (
<figureCitation id="133A7ADCFF9BFFF7C8F2FBAC4652FBEC" box="[423,550,1086,1111]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="3" pageNumber="4">Figs 1a, 1b</figureCitation>
): Wingspan: 
<quantity id="4CF9CBBCFF9BFFF7CBEEFBAD4779FBED" box="[699,781,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="4" unit="mm" value="34.0">34 mm</quantity>
; FW length: 
<quantity id="4CF9CBBCFF9BFFF7CACBFBAD479AFBED" box="[926,1006,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="4" unit="mm" value="17.0">17 mm</quantity>
, width: 
<quantity id="4CF9CBBCFF9BFFF7CD12FBAD40FDFBED" box="[1095,1161,1087,1110]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="4" unit="mm" value="7.0">7 mm</quantity>
, ratio 2.4: 1; HW length: 
<quantity id="4CF9CBBCFF9BFFF7C9C2FBF1449CFBC1" box="[151,232,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="4" unit="mm" value="14.0">14 mm</quantity>
, width: 
<quantity id="4CF9CBBCFF9BFFF7C817FBF145E6FBC1" box="[322,402,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="4" unit="mm" value="11.0">11 mm</quantity>
, ratio 1.3: 1.
</paragraph>
<paragraph id="8BBE6659FF9BFFF7C992FB144124FA94" blockId="3.[151,1437,906,1975]" pageId="3" pageNumber="4">
<emphasis id="B975BA4BFF9BFFF7C992FB14457FFB24" box="[199,267,1158,1183]" italics="true" pageId="3" pageNumber="4">Head.</emphasis>
Scales covered with dark brown tipped yellowish brown piliform scales, as also are the frons and vertex. Interocular-antennal antennal scales piliform, extending over the eye. Antenna filiform with 39 flagellomeres, annuli slightly compressed, laterally covered with numerous sensillae chaetica, the basal 3-4 annuli covered with some lamellar scales; scape barrel shaped, covered with lamellar scales. Labial palpi long (longer than surrounding scales); distal palpomere digitiform, 2.3 times length of basal segment, subapical sensory pit present (
<figureCitation id="133A7ADCFF9BFFF7CDA6FA844137FA94" box="[1267,1347,1302,1327]" captionStart="FIGURES 8–9" captionStartId="5.[151,264,1506,1529]" captionTargetBox="[151,1435,193,1486]" captionTargetId="figure@5.[151,1435,193,1486]" captionTargetPageId="5" captionText="FIGURES 8–9. Voucher specimens. 8, Dugdaleiella monticola male. 8a, LT (ZMHB); 8b, Cotopaxi, Ecuador (CMNH); 8c, Volcán Chiles, Ecuador (CMNH); 8d, male, Volcán Chiles, Ecuador (CMNH). 9, Labial palpi (male unless otherwise specified), 9a, Kozloviella viazmenskyi sp. n.; 9b, Pfitzneriella antonkozlovi sp. n.; 9c, P. rawlinsi sp. n.; 9d, P. titarenkoi sp. n.; 9e, P. yuliyakovalevae sp. n.; 9f, P. yuliyakovalevae sp. n. (female); 9g, Dugdaleiella monticola. Photos: Jane Hyland (Fig. 8), John Grehan (Figs 9a–b, d–h), and Carlos Mielke (Fig. 9c). Scale 1 mm (when present)." httpUri="https://zenodo.org/record/1451060/files/figure.png" pageId="3" pageNumber="4">Fig. 9a</figureCitation>
).
</paragraph>
<paragraph id="8BBE6659FF9BFFF7C992FAA84738F92C" blockId="3.[151,1437,906,1975]" pageId="3" pageNumber="4">
<emphasis id="B975BA4BFF9BFFF7C992FAA8456CFAE8" box="[199,280,1338,1363]" italics="true" pageId="3" pageNumber="4">Thorax</emphasis>
. Dorsal scales greyish brown, ventral yellowish brown with darker brown at apex of scale. Legs (
<figureCitation id="133A7ADCFF9BFFF7CC3AFAA844B4FACC" captionStart="FIGURES 12–14. 12" captionStartId="9.[151,264,1172,1195]" captionTargetBox="[150,1436,193,1151]" captionTargetId="figure@9.[150,1436,193,1151]" captionTargetPageId="9" captionText="FIGURES 12–14. 12, Antenna. Pfitzneriella rawlinsi sp. n. (unscaled). 13, Legs. 13a, Kozloviella viazmenskyi sp. n.; 13b, P. olafi sp. n.; 13c, P. rawlinsi sp. n.; 13d, P. titarenkoi sp. n.; 13e, P. yuliyakovalevae sp. n. 14, Metatibia. 14a, P. antonkozlovi sp. n.; 14b, P. rawlinsi sp. n.; 14c, P. titarenkoi; 14d, P. yuliyakovalevae sp. n.; 14e, P. antonkozlovi sp. n.; 14f, Dugdaleiella monticola. Photos: John Grehan (Figs 12, 14e, 14f), Carlos Mielke (Figs 13,14a–d). Scale 1 mm." httpUri="https://zenodo.org/record/1451066/files/figure.png" pageId="3" pageNumber="4">Fig. 13a</figureCitation>
) coloured as the thorax: pro-, meso-, meta- legs length ratio 1.3:1.5:1; epiphysis present; tibial and tarsal scales long, but not obscuring tarsomeres, dorsal scales of tarsi 1–4 about 1/3 longer than tarsomeres, ventral tarsal scales short; metaleg pale orange-brown, femoral and tibial scales piliform. Wings narrow, slightly wider distally. FW distinctly narrow compared to HW, mid-costal margin concave, outer margin convex, curving gradually to anal margin; venation hepialine (
<bibRefCitation id="EF901BA8FF9BFFF7C88FFA7C46D4F9BC" box="[474,672,1518,1543]" pageId="3" pageNumber="4" refId="ref14701" refString="Dumbleton, L. J. (1966) Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera). New Zealand Journal of Science, 9, 920 - 981." type="journal article">Dumbleton 1966</bibRefCitation>
), base of FW anal vein strongly curved anteriorly, distally curved posteriorly (
<figureCitation id="133A7ADCFF9BFFF7C876F98045F2F990" box="[291,390,1554,1579]" captionStart="FIGURE 10" captionStartId="6.[151,250,1649,1671]" captionTargetBox="[152,1437,195,1610]" captionTargetId="figure@6.[152,1437,195,1610]" captionTargetPageId="6" captionText="FIGURE 10. Wing venation (male unless otherwise specified). 10a, Kozloviella viazmenskyi sp. n.; 10b, P. antonkozlovi sp. n.; 10c, P. olafi sp. n.; 10d, P. rawlinsi sp. n.; 10e, P. remota (from photo only, venation incomplete due to damaged condition of HT); 10f, P. yuliyakovalevae sp. n.; 10g, P. yuliyakovalevae sp. n. (female); 10h, Dugdaleiella monticola." httpUri="https://zenodo.org/record/1451062/files/figure.png" pageId="3" pageNumber="4">Fig. 10a</figureCitation>
); dorsal ground colour pale yellowish orange with areas of darker orange-brown and dark brown shading and scattered black markings; oblique black line forming a V-shape with outer line extending between Cu and apex (
<figureCitation id="133A7ADCFF9BFFF7C8F5F9C84675F9C8" box="[416,513,1626,1651]" captionStart="FIGURE 11" captionStartId="7.[151,250,1429,1451]" captionTargetBox="[150,1436,810,1407]" captionTargetId="figure@7.[150,1436,810,1407]" captionTargetPageId="7" captionText="FIGURE 11. Wing markings. 11a; Kozloviella viazmenskyi sp. n., 11b; Pfitzneriella antonkozlovi sp. n.; 11c, P. olafi sp. n.; 11d, P. rawlinsi sp. n.; 11e, P. remota; 11f, P. yuliyakovalevae sp. n. Blue line—edge of white banding." httpUri="https://zenodo.org/record/1451064/files/figure.png" pageId="3" pageNumber="4">Fig. 11a</figureCitation>
); orange-brown shading predominant over outer discal cells and near apex. HW greyish brown with pale reddish brown towards outer margin.
</paragraph>
<paragraph id="8BBE6659FF9BFFF7C992F93046E6F964" blockId="3.[151,1437,906,1975]" pageId="3" pageNumber="4">
<emphasis id="B975BA4BFF9BFFF7C992F9304545F900" box="[199,305,1698,1723]" italics="true" pageId="3" pageNumber="4">Abdomen</emphasis>
. Greyish brown, narrow, distally wider around genitalic region. Tergum VIII rectangular, wider than long, sternum VII wider than long (
<figureCitation id="133A7ADCFF9BFFF7CB70F95446F1F964" box="[549,645,1734,1759]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[197,1386,199,1851]" captionTargetId="figure@10.[197,1386,199,1851]" captionTargetPageId="10" captionText="FIGURES 15–16. Tergosternal connection. 15a, Kozloviella viazmenskyi sp. n.; 15b, Pfitzneriella yuliyakovalevae sp. n.; 15c, P. rawlinsi sp. n.; 15d, Dugdaleiella monticola. 16, Abdomen (males unless otherwise specified). 16a, K. viazmenskyi sp. n.; 16b, P. antonkozlovi sp. n.; 16c, P. olafi sp. n.; 16d, P. rawlinsi sp. n.; 16e, P. titarenkoi sp. n.; 16f, P. yuliyakovalevae sp. n.; 16g, P. yuliyakovalevae sp. n. (female); 16h–16j, Dugdaleiella monticola. 16h, tergum II; 16i, sternum II; 16j, tergum and sternum VIII. Photos: John Grehan (Figs 15d, 16h–16j), Carlos Mielke (Figs 15a–15c, 16a–16g)." httpUri="https://zenodo.org/record/1451068/files/figure.png" pageId="3" pageNumber="4">Fig. 16a</figureCitation>
).
</paragraph>
<paragraph id="8BBE6659FF9BFFF7C992F9784758F89D" blockId="3.[151,1437,906,1975]" pageId="3" pageNumber="4">
<emphasis id="B975BA4BFF9BFFF7C992F9784504F8B8" box="[199,368,1770,1795]" italics="true" pageId="3" pageNumber="4">Male genitalia</emphasis>
. See description of the genus. Ventral surface of saccus forming two subrectangular lobes, separated medially. Phallus membranous without cornutus.
</paragraph>
<paragraph id="8BBE6659FF9BFFF7C992F8A045E4F8F0" blockId="3.[151,1437,906,1975]" box="[199,400,1842,1867]" pageId="3" pageNumber="4">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF9BFFF1C992F8C44772F90F" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="distribution">
<paragraph id="8BBE6659FF9BFFF7C992F8C444A6F80C" blockId="3.[151,1437,906,1975]" pageId="3" pageNumber="4">
<emphasis id="B975BA4BFF9BFFF7C992F8C4452DF8D4" bold="true" box="[199,345,1878,1903]" pageId="3" pageNumber="4">Distribution</emphasis>
. 
<collectingRegion id="49C5A8BBFF9BFFF7C832F8C445B8F8D4" box="[359,460,1878,1903]" country="Sri Lanka" name="Northern" pageId="3" pageNumber="4">Northern</collectingRegion>
<collectingCountry id="F31626C9FF9BFFF7C886F8C5467DF8D5" box="[467,521,1879,1902]" name="Peru" pageId="3" pageNumber="4">Peru</collectingCountry>
, known from the 
<typeStatus id="54BAD8FBFF9BFFF7CB9BF8C5468AF8D4" box="[718,766,1879,1903]" pageId="3" pageNumber="4">type</typeStatus>
locality only (
<figureCitation id="133A7ADCFF9BFFF7CAF7F8C44059F8D4" box="[930,1069,1878,1903]" captionStart="FIGURES 27–29" captionStartId="20.[151,264,1710,1733]" captionTargetBox="[156,1435,363,1685]" captionTargetId="figure@20.[151,1436,363,1690]" captionTargetPageId="20" captionText="FIGURES 27–29. Distribution. 27, Dugdaleiella gen. n., Kozloviella gen. n., and Pfitzneriella locality records. 27a, Ecuador and Peru. Tectonic boundaries approximate; 27b, Dugdaleiella gen. n. and P. lucicola in Ecuador. 28, Locality records of Peruvian Pfitzneriella in relation to eastern Andean montane forests (modified from Young &amp; León (2000) with permission by Kenneth Young). 29, Principal groups of Hepialidae in Central and South America. Phassus/Schausiana/Phthius (solid line), cibyrine genera of Hepialidae (dotted line), Dugdaleiella gen. n./Kozloviella gen. n./Pfitzneriella (pale green), Uiridigigas (red), and southern Andean genera (blue)." httpUri="https://zenodo.org/record/1451074/files/figure.png" pageId="3" pageNumber="4">Figs 27a, 29</figureCitation>
<emphasis id="B975BA4BFF9BFFF7CD79F8C4404AF8D4" bold="true" box="[1068,1086,1878,1903]" pageId="3" pageNumber="4">).</emphasis>
Collected in a mountain valley in the eastern Andes adjacent to disturbed forest with local clearing and farming activities at 
<quantity id="4CF9CBBCFF9BFFF7CD96F8E84155F829" box="[1219,1313,1914,1939]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="3" pageNumber="4" unit="m" value="2050.0">2,050 m</quantity>
(
<figureCitation id="133A7ADCFF9BFFF7CC67F8E844B1F80C" captionStart="FIGURE 26" captionStartId="14.[151,250,1674,1696]" captionTargetBox="[152,1435,266,1652]" captionTargetId="figure@14.[152,1435,195,1652]" captionTargetPageId="14" captionText="FIGURE 26. Collection localities. 26a–26b, Kozloviella viazmenskyi sp. n. Peru, Cerro Machete, 2,050 m; 26c, Pfitzneriella antonkozlovi sp. n., Peru, Marañon Province, 3,270 m; 26d, P. titarenkoi, sp. n. and P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 3,380 m; 26e–26f, P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 2,960 m. With permission by Anton Kozlov and Yuliya Kovaleva, foreground 26b, 26e, Anton Kozlov and 26f, Yuliya Kovaleva. Photos: Anton Kozlov." httpUri="https://zenodo.org/record/1451072/files/figure.png" pageId="3" pageNumber="4">Figs 26a, 26b</figureCitation>
).
</paragraph>
<caption id="DF7E36D1FF9CFFF0C9C2F9394743F8E0" httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,264,1707,1730]" targetBox="[151,1436,195,1660]" targetPageId="4">
<paragraph id="8BBE6659FF9CFFF0C9C2F9394743F8E0" blockId="4.[151,1436,1707,1883]" pageId="4" pageNumber="5">
<emphasis id="B975BA4BFF9CFFF0C9C2F9394542F97A" bold="true" box="[151,310,1707,1730]" pageId="4" pageNumber="5">FIGURES 1–7</emphasis>
. Voucher specimens. 1, 
<taxonomicName id="4C011DDAFF9CFFF0CB66F93E475DF979" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[563,809,1708,1730]" class="Insecta" family="Hepialidae" genus="Kozloviella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="viazmenskyi" status="gen. nov.">
<emphasis id="B975BA4BFF9CFFF0CB66F93E475DF979" box="[563,809,1708,1730]" italics="true" pageId="4" pageNumber="5">Kozloviella viazmenskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9CFFF0CA64F93E47B6F979" box="[817,962,1708,1730]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B975BA4BFF9CFFF0CA64F93E4723F97A" bold="true" box="[817,855,1708,1729]" pageId="4" pageNumber="5">gen</emphasis>
. 
<emphasis id="B975BA4BFF9CFFF0CA30F93E4707F97A" bold="true" box="[869,883,1708,1729]" pageId="4" pageNumber="5">n</emphasis>
., 
<emphasis id="B975BA4BFF9CFFF0CADDF93E47D4F97A" bold="true" box="[904,928,1708,1729]" pageId="4" pageNumber="5">sp</emphasis>
. 
<emphasis id="B975BA4BFF9CFFF0CAFBF93E47C8F97A" bold="true" box="[942,956,1708,1729]" pageId="4" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
male HT (CMNH). 1a, dorsal; 1b, ventral. 2, 
<taxonomicName id="4C011DDAFF9CFFF0C9C2F95945E9F95A" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[151,413,1739,1761]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="antonkozlovi" status="sp. nov.">
<emphasis id="B975BA4BFF9CFFF0C9C2F95945E9F95A" box="[151,413,1739,1761]" italics="true" pageId="4" pageNumber="5">Pfitzneriella antonkozlovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9CFFF0C8F6F95945A8F95A" box="[419,476,1739,1761]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B975BA4BFF9CFFF0C8F6F95945CFF95B" bold="true" box="[419,443,1739,1760]" pageId="4" pageNumber="5">sp</emphasis>
. 
<emphasis id="B975BA4BFF9CFFF0C89DF95945A2F95B" bold="true" box="[456,470,1739,1760]" pageId="4" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
male HT (CMNH), 2a, dorsal; 2b, ventral. 3, 
<taxonomicName id="4C011DDAFF9CFFF0CAE4F95E478DF95A" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[945,1017,1739,1761]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="olafi" status="sp. nov.">
<emphasis id="B975BA4BFF9CFFF0CAE4F95E478DF95A" box="[945,1017,1739,1761]" italics="true" pageId="4" pageNumber="5">P. olafi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9CFFF0CD55F959404DF95A" box="[1024,1081,1739,1761]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B975BA4BFF9CFFF0CD55F959406CF95B" bold="true" box="[1024,1048,1739,1760]" pageId="4" pageNumber="5">sp</emphasis>
. 
<emphasis id="B975BA4BFF9CFFF0CD70F9594047F95B" bold="true" box="[1061,1075,1739,1760]" pageId="4" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
male HT (MNHN), 3a, dorsal; 3b, ventral. 4, 
<taxonomicName id="4C011DDAFF9CFFF0C856F978451BF944" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[259,367,1769,1791]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rawlinsi" status="sp. nov.">
<emphasis id="B975BA4BFF9CFFF0C856F978451BF944" box="[259,367,1769,1791]" italics="true" pageId="4" pageNumber="5">P. rawlinsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9CFFF0C820F97845DAF944" box="[373,430,1770,1791]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B975BA4BFF9CFFF0C820F97845F9F944" bold="true" box="[373,397,1770,1791]" pageId="4" pageNumber="5">sp</emphasis>
. 
<emphasis id="B975BA4BFF9CFFF0C8CCF97845D3F944" bold="true" box="[409,423,1770,1791]" pageId="4" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, 
<taxonomicName id="4C011DDAFF9CFFF0CDCDF978408CF944" baseAuthorityName="Pfitzner" baseAuthorityYear="1906" box="[1176,1272,1770,1791]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="B975BA4BFF9CFFF0CDCDF978408CF944" box="[1176,1272,1770,1791]" italics="true" pageId="4" pageNumber="5">P. remota</emphasis>
</taxonomicName>
male HT dorsal (MNHN). 6, 
<taxonomicName id="4C011DDAFF9CFFF0C849F89B45E3F8A5" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[284,407,1800,1822]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="titarenkoi" status="sp. nov.">
<emphasis id="B975BA4BFF9CFFF0C849F89B45E3F8A5" box="[284,407,1800,1822]" italics="true" pageId="4" pageNumber="5">P. titarenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9CFFF0C8CBF89A45A3F8A5" box="[414,471,1800,1822]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B975BA4BFF9CFFF0C8CBF89A45C2F8A6" bold="true" box="[414,438,1800,1821]" pageId="4" pageNumber="5">sp</emphasis>
. 
<emphasis id="B975BA4BFF9CFFF0C896F89A45A5F8A6" bold="true" box="[451,465,1800,1821]" pageId="4" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
male HT (CMNH), 6a, dorsal; 6b, ventral. 7, 
<taxonomicName id="4C011DDAFF9CFFF0CAFAF89B401FF8A5" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[943,1131,1800,1822]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="yuliyakovalevae" status="sp. nov.">
<emphasis id="B975BA4BFF9CFFF0CAFAF89B401FF8A5" box="[943,1131,1800,1822]" italics="true" pageId="4" pageNumber="5">P. yuliyakovalevae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9CFFF0CD27F89A40DFF8A5" box="[1138,1195,1800,1822]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B975BA4BFF9CFFF0CD27F89A40FEF8A6" bold="true" box="[1138,1162,1800,1821]" pageId="4" pageNumber="5">sp</emphasis>
. 
<emphasis id="B975BA4BFF9CFFF0CDC2F89A40D1F8A6" bold="true" box="[1175,1189,1800,1821]" pageId="4" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c).
</paragraph>
</caption>
<caption id="DF7E36D1FF9DFFF1C9C2FA704066F9CF" httpUri="https://zenodo.org/record/1451060/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,264,1506,1529]" targetBox="[151,1435,193,1486]" targetPageId="5">
<paragraph id="8BBE6659FF9DFFF1C9C2FA704066F9CF" blockId="5.[151,1436,1506,1652]" pageId="5" pageNumber="6">
<emphasis id="B975BA4BFF9DFFF1C9C2FA704541FA43" bold="true" box="[151,309,1506,1529]" pageId="5" pageNumber="6">FIGURES 8–9</emphasis>
. Voucher specimens. 8
<taxonomicName id="4C011DDAFF9DFFF1CB73FA714751FA42" box="[550,805,1507,1529]" pageId="5" pageNumber="6" rank="subSpecies" subSpecies="monticola">
, 
<emphasis id="B975BA4BFF9DFFF1CB61FA714751FA42" box="[564,805,1507,1529]" italics="true" pageId="5" pageNumber="6">Dugdaleiella monticola</emphasis>
</taxonomicName>
male. 8a, LT (ZMHB); 8b, Cotopaxi, Ecuador (CMNH); 8c, Volcán Chiles, Ecuador (CMNH); 8d, male, Volcán Chiles, Ecuador (CMNH). 9, Labial palpi (male unless otherwise specified), 9a, 
<taxonomicName id="4C011DDAFF9DFFF1C87EF9B2466AF98D" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[299,542,1568,1590]" class="Insecta" family="Hepialidae" genus="Kozloviella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="viazmenskyi" status="sp. nov.">
<emphasis id="B975BA4BFF9DFFF1C87EF9B2466AF98D" box="[299,542,1568,1590]" italics="true" pageId="5" pageNumber="6">Kozloviella viazmenskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9DFFF1CB70F9B34629F98D" box="[549,605,1569,1590]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B975BA4BFF9DFFF1CB70F9B34649F98D" bold="true" box="[549,573,1569,1590]" pageId="5" pageNumber="6">sp</emphasis>
. 
<emphasis id="B975BA4BFF9DFFF1CB1CF9B34622F98D" bold="true" box="[585,598,1569,1590]" pageId="5" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
; 9b, 
<taxonomicName id="4C011DDAFF9DFFF1CBC5F9B247E5F98D" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[656,913,1568,1590]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="antonkozlovi" status="sp. nov.">
<emphasis id="B975BA4BFF9DFFF1CBC5F9B247E5F98D" box="[656,913,1568,1590]" italics="true" pageId="5" pageNumber="6">Pfitzneriella antonkozlovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9DFFF1CAC2F9B347A4F98D" box="[919,976,1569,1590]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B975BA4BFF9DFFF1CAC2F9B347DBF98D" bold="true" box="[919,943,1569,1590]" pageId="5" pageNumber="6">sp</emphasis>
. 
<emphasis id="B975BA4BFF9DFFF1CAE9F9B347BEF98D" bold="true" box="[956,970,1569,1590]" pageId="5" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
; 9c, 
<taxonomicName id="4C011DDAFF9DFFF1CD57F9B0401FF98D" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[1026,1131,1568,1591]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rawlinsi" status="sp. nov.">
<emphasis id="B975BA4BFF9DFFF1CD57F9B0401FF98D" box="[1026,1131,1568,1591]" italics="true" pageId="5" pageNumber="6">P. rawlinsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9DFFF1CD24F9B340DDF98D" box="[1137,1193,1569,1590]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B975BA4BFF9DFFF1CD24F9B340FDF98D" bold="true" box="[1137,1161,1569,1590]" pageId="5" pageNumber="6">sp</emphasis>
. 
<emphasis id="B975BA4BFF9DFFF1CDC0F9B340D7F98D" bold="true" box="[1173,1187,1569,1590]" pageId="5" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
; 9d, 
<taxonomicName id="4C011DDAFF9DFFF1CD89F9B04122F98D" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[1244,1366,1568,1591]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="titarenkoi" status="sp. nov.">
<emphasis id="B975BA4BFF9DFFF1CD89F9B04122F98D" box="[1244,1366,1568,1591]" italics="true" pageId="5" pageNumber="6">P. titarenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9DFFF1CC09F9B341E0F98D" box="[1372,1428,1569,1590]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B975BA4BFF9DFFF1CC09F9B34100F98D" bold="true" box="[1372,1396,1569,1590]" pageId="5" pageNumber="6">sp</emphasis>
. 
<emphasis id="B975BA4BFF9DFFF1CCD5F9B341F9F98D" bold="true" box="[1408,1421,1569,1590]" pageId="5" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
; 9e, 
<taxonomicName id="4C011DDAFF9DFFF1C9E9F9D2450CF9EE" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[188,376,1599,1621]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yuliyakovalevae" status="sp. nov.">
<emphasis id="B975BA4BFF9DFFF1C9E9F9D2450CF9EE" box="[188,376,1599,1621]" italics="true" pageId="5" pageNumber="6">P. yuliyakovalevae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9DFFF1C82AF9AD45CDF9EE" box="[383,441,1599,1621]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B975BA4BFF9DFFF1C82AF9AD45E3F9EF" bold="true" box="[383,407,1599,1620]" pageId="5" pageNumber="6">sp</emphasis>
. 
<emphasis id="B975BA4BFF9DFFF1C8F1F9AD45C6F9EF" bold="true" box="[420,434,1599,1620]" pageId="5" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
; 9f, 
<taxonomicName id="4C011DDAFF9DFFF1C8BCF9D246D2F9EE" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[489,678,1599,1621]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yuliyakovalevae" status="sp. nov.">
<emphasis id="B975BA4BFF9DFFF1C8BCF9D246D2F9EE" box="[489,678,1599,1621]" italics="true" pageId="5" pageNumber="6">P. yuliyakovalevae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9DFFF1CBF8F9AD4692F9EE" box="[685,742,1599,1621]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B975BA4BFF9DFFF1CBF8F9AD46B1F9EF" bold="true" box="[685,709,1599,1620]" pageId="5" pageNumber="6">sp</emphasis>
. 
<emphasis id="B975BA4BFF9DFFF1CB87F9AD4694F9EF" bold="true" box="[722,736,1599,1620]" pageId="5" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
(female); 9g
<taxonomicName id="4C011DDAFF9DFFF1CA39F9AD4010F9EE" box="[876,1124,1599,1621]" pageId="5" pageNumber="6" rank="subSpecies" subSpecies="monticola">
, 
<emphasis id="B975BA4BFF9DFFF1CA2DF9AD4010F9EE" box="[888,1124,1599,1621]" italics="true" pageId="5" pageNumber="6">Dugdaleiella monticola</emphasis>
</taxonomicName>
. Photos: Jane Hyland (Fig. 8), John Grehan (Figs 9a–b, d–h), and Carlos Mielke (Fig. 9c). Scale 1 mm (when present).
</paragraph>
</caption>
<paragraph id="8BBE6659FF9DFFF1C992F9094772F90F" blockId="5.[151,1437,1691,2004]" box="[199,774,1691,1716]" pageId="5" pageNumber="6">
<emphasis id="B975BA4BFF9DFFF1C992F9094545F90F" bold="true" box="[199,305,1691,1716]" pageId="5" pageNumber="6">Ethology</emphasis>
. Attracted to UV-light at collecting sheet.
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF9DFFF1C992F92D4740F963" box="[199,820,1727,1752]" pageId="5" pageNumber="6" type="biology_ecology">
<paragraph id="8BBE6659FF9DFFF1C992F92D4740F963" blockId="5.[151,1437,1691,2004]" box="[199,820,1727,1752]" pageId="5" pageNumber="6">
<emphasis id="B975BA4BFF9DFFF1C992F92D4530F963" bold="true" box="[199,324,1727,1752]" pageId="5" pageNumber="6">Host plant</emphasis>
. Larval habits and host plants are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF9DFFF1C992F9714486F8FF" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BBE6659FF9DFFF1C992F9714486F8FF" blockId="5.[151,1437,1691,2004]" pageId="5" pageNumber="6">
<emphasis id="B975BA4BFF9DFFF1C992F9714532F947" bold="true" box="[199,326,1763,1788]" pageId="5" pageNumber="6">Etymology</emphasis>
. This species is named after Alexander Lvovich Viazmensky (
<collectingRegion id="49C5A8BBFF9DFFF1CD68F9714174F947" box="[1085,1280,1763,1788]" country="Russia" name="Saint Petersburg" pageId="5" pageNumber="6">Saint Petersburg</collectingRegion>
, 
<collectingCountry id="F31626C9FF9DFFF1CC44F9714114F947" box="[1297,1376,1763,1788]" name="Russia" pageId="5" pageNumber="6">Russia</collectingCountry>
), the famous Russian aquarellist and the passionate fan of mushrooms, at the special request of Anton Kozlov (
<collectingRegion id="49C5A8BBFF9DFFF1CC60F89541ECF8A4" box="[1333,1432,1799,1823]" country="Russia" name="Moscow" pageId="5" pageNumber="6">Moscow</collectingRegion>
, 
<collectingCountry id="F31626C9FF9DFFF1C9C2F8B94493F8FF" box="[151,231,1835,1860]" name="Russia" pageId="5" pageNumber="6">Russia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF9DFFF1C992F8DD4588F86F" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BBE6659FF9DFFF1C992F8DD4588F86F" blockId="5.[151,1437,1691,2004]" pageId="5" pageNumber="6">
<emphasis id="B975BA4BFF9DFFF1C992F8DD451BF8D3" bold="true" box="[199,367,1871,1896]" pageId="5" pageNumber="6">
<typeStatus id="54BAD8FBFF9DFFF1C992F8DD4576F8D3" box="[199,258,1871,1896]" pageId="5" pageNumber="6">Type</typeStatus>
material
</emphasis>
. 
<typeStatus id="54BAD8FBFF9DFFF1C82BF8DD4591F8D3" box="[382,485,1871,1896]" pageId="5" pageNumber="6" type="holotype">
<emphasis id="B975BA4BFF9DFFF1C82BF8DD4591F8D3" box="[382,485,1871,1896]" italics="true" pageId="5" pageNumber="6">Holotype</emphasis>
</typeStatus>
male (with the following labels separated by forward slashes): /S. America, 
<collectionCode id="ED10FE9CFF9DFFF1CC1AF8C241ECF8DC" box="[1359,1432,1872,1895]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14398" name="Universit<69> di Perugia" pageId="5" pageNumber="6">
<collectingCountry id="F31626C9FF9DFFF1CC1AF8C241ECF8DC" box="[1359,1432,1872,1895]" name="Peru" pageId="5" pageNumber="6">PERU</collectingCountry>
</collectionCode>
, 
<collectingRegion id="49C5A8BBFF9DFFF1C9C2F8E144A5F837" box="[151,209,1907,1932]" country="Peru" name="Piura" pageId="5" pageNumber="6">Piura</collectingRegion>
reg. Huancabamba prov. Cerro Machete. 
<geoCoordinate id="EE35009EFF9DFFF1CBEEF8E14745F837" box="[699,817,1907,1932]" direction="south" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="-0.084640004">0 5.0784 S</geoCoordinate>
, 0 
<geoCoordinate id="EE35009EFF9DFFF1CA1AF8E147A4F830" box="[847,976,1907,1932]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="-79.3362">79.3362 W</geoCoordinate>
. h= 
<quantity id="4CF9CBBCFF9DFFF1CAA9F8E1402CF830" box="[1020,1112,1907,1932]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="5" pageNumber="6" unit="m" value="2050.0">2050 m</quantity>
, 
<date id="FFBF4099FF9DFFF1CD3DF8E640ABF837" box="[1128,1247,1908,1932]" pageId="5" pageNumber="6" value="2017-02-10">10/II/2017</date>
. A. Kozlov, Yu. Kovaleva, R. Gortovannyi leg. / 
<typeStatus id="54BAD8FBFF9DFFF1CB54F805460CF80B" box="[513,632,1943,1968]" pageId="5" pageNumber="6" type="holotype">Holotypus</typeStatus>
, 
<taxonomicName id="4C011DDAFF9DFFF1CBD1F80540D3F80B" authority="♂, Grehan &amp; C. Mielke" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[644,1191,1943,1968]" class="Insecta" family="Hepialidae" genus="Kozloviella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="viazmenskyi">
<emphasis id="B975BA4BFF9DFFF1CBD1F80547E7F80B" box="[644,915,1943,1968]" italics="true" pageId="5" pageNumber="6">Kozloviella viazmenskyi</emphasis>
<specimenCount id="9D07ADD0FF9DFFF1CACEF80A47C6F814" box="[923,946,1944,1967]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
, Grehan &amp; C. Mielke
</taxonomicName>
det. 2017/ Dissection JRG 258/ (
<collectionCode id="ED10FE9CFF9DFFF1C85BF82E4513F86F" box="[270,359,1980,2004]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="5" pageNumber="6">CMNH</collectionCode>
). 
<figureCitation id="133A7ADCFF9DFFF1C82CF8294583F86F" box="[377,503,1979,2004]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="5" pageNumber="6">Figs 1a, 1b</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>