treatments-xml/data/03/A8/87/03A887EBFFDBFFE2FEFBFEC3B3437950.xml

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<mods:title id="C9574C97FBF2DB4798C125EAC6FEAE5C">New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America</mods:title>
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<mods:namePart id="184DB3D810AD6A0D0FBAFE29D42FFB30">Kohlmann, Bert</mods:namePart>
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<mods:namePart id="F2DFE8EF1E3B59146DDD88BCE785C114">Delgado, Enio Cano And Leonardo</mods:namePart>
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<heading id="D0F68191FFDBFFE2FEFBFEC3B23C7D0D" bold="true" box="[264,1108,283,310]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B975EAEFFFDBFFE2FEFBFEC3B23C7D0D" bold="true" box="[264,1108,283,310]" pageId="11" pageNumber="12">
Key to the species of the 
<taxonomicName id="4C014D7EFFDBFFE2FDCFFEC4B4F57D0E" ID-CoL="7ZMLY" box="[572,669,284,310]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="remotus">
<emphasis id="B975EAEFFFDBFFE2FDCFFEC4B4F57D0E" bold="true" box="[572,669,284,310]" italics="true" pageId="11" pageNumber="12">remotus</emphasis>
</taxonomicName>
complex based on the major males
</emphasis>
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<keyLead id="30F095C8FFDBFFE2FEFBFEBFB3437D91" pageId="11" pageNumber="12">
1. Head unarmed having only a gibbosity, clypeus tridentate; thorax unarmed. 
<collectingCountry id="F316766DFFDBFFE2FB54FEBFB34E7DB9" box="[1191,1318,359,385]" name="Costa Rica" pageId="11" pageNumber="12">Costa Rica</collectingCountry>
<emphasis id="B975EAEFFFDBFFE2FEB1FE57B2207D90" box="[322,1096,399,424]" italics="true" pageId="11" pageNumber="12">
........................................................................................ 
<taxonomicName id="4C014D7EFFDBFFE2FC3AFE57B2207D90" ID-CoL="7ZMN8" authority="Solis &amp; Kohlmann" authorityName="Solis &amp; Kohlmann" box="[969,1096,399,424]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tridentatus">tridentatus</taxonomicName>
</emphasis>
Solís &amp; Kohlmann
</keyLead>
</paragraph>
<paragraph id="8BBE36FDFFDBFFE2FEFBFE6FB3457DE9" blockId="11.[264,1325,359,1384]" box="[264,1325,439,465]" pageId="11" pageNumber="12">
<keyLead id="30F095C8FFDBFFE2FEFBFE6FB3457DE9" box="[264,1325,439,465]" pageId="11" pageNumber="12">­ Head armed with a horn, clypeus bidentate; thorax armed .......................................... 2</keyLead>
</paragraph>
</keyStep>
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2. Pygidial margin incomplete, its inner border completely effaced ventrally. Chiapas, 
<collectingCountry id="F316766DFFDBFFE2FECBFDDFB7D27E19" box="[312,442,519,545]" name="Guatemala" pageId="11" pageNumber="12">Guatemala</collectingCountry>
, 
<collectingCountry id="F316766DFFDBFFE2FE36FDDFB42D7E19" box="[453,581,519,545]" name="Costa Rica" pageId="11" pageNumber="12">Costa Rica</collectingCountry>
and 
<collectingCountry id="F316766DFFDBFFE2FD8DFDDFB4B37E19" box="[638,731,519,545]" name="Panama" pageId="11" pageNumber="12">Panamá</collectingCountry>
.............................................. 
<emphasis id="B975EAEFFFDBFFE2FBCFFDDFB2BE7E18" box="[1084,1238,519,544]" italics="true" pageId="11" pageNumber="12">costaricencis</emphasis>
Gahan
</keyLead>
</paragraph>
<paragraph id="8BBE36FDFFDBFFE2FEFBFDF7B3457E71" blockId="11.[264,1325,359,1384]" box="[264,1325,559,585]" pageId="11" pageNumber="12">
<keyLead id="30F095C8FFDBFFE2FEFBFDF7B3457E71" box="[264,1325,559,585]" pageId="11" pageNumber="12">­ Pygidial margin complete, its inner border entirely engraved ...................................... 3</keyLead>
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<paragraph id="8BBE36FDFFDBFFE2FEFBFD8FB3437EA1" blockId="11.[264,1325,359,1384]" pageId="11" pageNumber="12">
<keyLead id="30F095C8FFDBFFE2FEFBFD8FB3437EA1" pageId="11" pageNumber="12">3. Eighth elytral stria incomplete; frontal pronotal declivity with a carina running through its middle ....................................................................................................... 4</keyLead>
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<keyLead id="30F095C8FFDBFFE2FEFBFD7FB3437ED1" pageId="11" pageNumber="12">­ Eighth elytral stria complete; frontal pronotal declivity without a carina running through its middle ....................................................................................................... 5</keyLead>
</paragraph>
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<keyLead id="30F095C8FFDBFFE2FEFBFD2FB3437F59" pageId="11" pageNumber="12">
4. Antennae dark­brown; male anterolateral pronotal angles not forming median point, but presenting an excavation behind these angles (
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), median pronotal prominences diverging (
<figureCitation id="133A2A78FFDBFFE2FDFCFC9FB43C7F59" box="[527,596,839,865]" captionStart="FIGURE. 1" captionStartId="6.[264,369,1766,1790]" captionTargetBox="[452,1164,396,1670]" captionTargetId="figure@6.[365,1223,361,1708]" captionTargetPageId="6" captionText="FIGURE. 1. Dorsal view of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype)." httpUri="https://zenodo.org/record/168711/files/figure.png" pageId="11" pageNumber="12">Fig. 1</figureCitation>
). 
<collectingCountry id="F316766DFFDBFFE2FD9EFC9FB4847F59" box="[621,748,839,865]" name="Guatemala" pageId="11" pageNumber="12">Guatemala</collectingCountry>
. 
<taxonomicName id="4C014D7EFFDBFFE2FD0CFC9FB2887F59" ID-CoL="7ZMFW" authority="Kohlmann, Cano &amp; Delgado" authorityName="Kohlmann, Cano &amp; Delgado" box="[767,1248,839,865]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="caliginosus" status="sp. nov.">
<emphasis id="B975EAEFFFDBFFE2FD0CFC9FB5ED7F58" box="[767,901,839,864]" italics="true" pageId="11" pageNumber="12">caliginosus</emphasis>
Kohlmann, Cano &amp; Delgado
</taxonomicName>
, 
<taxonomicNameLabel id="A2465794FFDBFFE2FB1FFC9FB3437F59" box="[1260,1323,839,865]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE36FDFFDBFFE2FEFBFCB7B3437839" blockId="11.[264,1325,359,1384]" pageId="11" pageNumber="12">
<keyLead id="30F095C8FFDBFFE2FEFBFCB7B3437839" pageId="11" pageNumber="12">
­ Antennae reddish­brown; male anterolateral pronotal angles forming a small downwardly directed median point and separated from the angles by a more or less level area (
<figureCitation id="133A2A78FFDBFFE2FE8BFC67B7A87FE1" box="[376,448,959,985]" captionStart="FIGURE. 4" captionStartId="9.[264,370,1708,1732]" captionTargetBox="[525,1064,323,1678]" captionTargetId="figure@9.[533,1053,793,1144]" captionTargetPageId="9" captionText="FIGURE. 4. Lateral views of head and pronotum of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype, top), Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype, middle) and Copris sallei (bottom)." httpUri="https://zenodo.org/record/168713/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
), median pronotal prominences strongly diverging (
<figureCitation id="133A2A78FFDBFFE2FBEDFC67B20D7FE1" box="[1054,1125,959,985]" captionStart="FIGURE. 2" captionStartId="7.[264,370,1696,1720]" captionTargetBox="[391,1181,414,1621]" captionTargetId="figure@7.[365,1223,326,1673]" captionTargetPageId="7" captionText="FIGURE. 2. Dorsal view of Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype)." httpUri="https://zenodo.org/record/168712/files/figure.png" pageId="11" pageNumber="12">Fig. 2</figureCitation>
). 
<collectingCountry id="F316766DFFDBFFE2FB88FC67B2927FE1" box="[1147,1274,959,985]" name="Guatemala" pageId="11" pageNumber="12">Guatemala</collectingCountry>
and 
<collectingCountry id="F316766DFFDBFFE2FECBFC3FB7C27839" box="[312,426,999,1025]" name="Honduras" pageId="11" pageNumber="12">Honduras</collectingCountry>
<emphasis id="B975EAEFFFDBFFE2FE44FC3FB51A7838" box="[439,882,999,1024]" italics="true" pageId="11" pageNumber="12">
............................................. 
<taxonomicName id="4C014D7EFFDBFFE2FCF5FC3FB51A7838" ID-CoL="7ZML8" authority="Kohlmann, Cano and Delgado" authorityName="Kohlmann, Cano and Delgado" box="[774,882,999,1024]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nubilosus" status="sp. nov.">nubilosus</taxonomicName>
</emphasis>
Kohlmann, Cano and Delgado, 
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</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F52E58FFDBFFE2FEFBFBD7B3437840" pageId="11" pageNumber="12">
<paragraph id="8BBE36FDFFDBFFE2FEFBFBD7B3427869" blockId="11.[264,1325,359,1384]" pageId="11" pageNumber="12">
<keyLead id="30F095C8FFDBFFE2FEFBFBD7B3427869" pageId="11" pageNumber="12">
5. Protibial apical spur abruptly bent inward very near apex. Oklahoma, Texas, Coahuila, Nuevo León and Tamaulipas .............................................................. 
<taxonomicName id="4C014D7EFFDBFFE2FB92FBEFB3427869" ID-CoL="7ZMLY" authority="LeConte" authorityName="LeConte" box="[1121,1322,1079,1105]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="remotus">
<emphasis id="B975EAEFFFDBFFE2FB92FBEFB2D57868" box="[1121,1213,1079,1104]" italics="true" pageId="11" pageNumber="12">remotus</emphasis>
LeConte
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBE36FDFFDBFFE2FEFBFB86B3437840" blockId="11.[264,1325,359,1384]" box="[264,1323,1118,1144]" pageId="11" pageNumber="12">
<keyLead id="30F095C8FFDBFFE2FEFBFB86B3437840" box="[264,1323,1118,1144]" pageId="11" pageNumber="12">
­ Protibial apical spur curved or slightly curved inward (
<figureCitation id="133A2A78FFDBFFE2FC4AFB86B5977840" box="[953,1023,1118,1144]" captionStart="FIGURE. 3" captionStartId="8.[264,370,1696,1720]" captionText="FIGURE. 3. Dorsal view of C. sallei Harold." pageId="11" pageNumber="12">Fig. 3</figureCitation>
) ................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F52E58FFDBFFE2FEFBFB5EB3437950" pageId="11" pageNumber="12">
<paragraph id="8BBE36FDFFDBFFE2FEFBFB5EB34278C8" blockId="11.[264,1325,359,1384]" pageId="11" pageNumber="12">
<keyLead id="30F095C8FFDBFFE2FEFBFB5EB34278C8" pageId="11" pageNumber="12">
6. Anterior pronotal margin with a minute, acute median tooth (
<figureCitation id="133A2A78FFDBFFE2FBF2FB5EB2207898" box="[1025,1096,1158,1184]" captionStart="FIGURE. 3" captionStartId="8.[264,370,1696,1720]" captionText="FIGURE. 3. Dorsal view of C. sallei Harold." pageId="11" pageNumber="12">Fig. 3</figureCitation>
); elytral striae with large, less transverse punctures, distinctly crenating the margins of the strial channels; protibial apical spurs blunt (
<figureCitation id="133A2A78FFDBFFE2FD71FB0EB4A178C8" box="[642,713,1238,1264]" captionStart="FIGURE. 3" captionStartId="8.[264,370,1696,1720]" captionText="FIGURE. 3. Dorsal view of C. sallei Harold." pageId="11" pageNumber="12">Fig. 3</figureCitation>
). Veracruz to Chiapas ........................ 
<taxonomicName id="4C014D7EFFDBFFE2FB61FB0FB34278C8" ID-CoL="7ZMM8" authority="Harold" authorityName="Harold" box="[1170,1322,1238,1264]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sallei">
<emphasis id="B975EAEFFFDBFFE2FB61FB0FB2BA78C8" box="[1170,1234,1239,1264]" italics="true" pageId="11" pageNumber="12">sallei</emphasis>
Harold
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBE36FDFFDBFFE2FEFBFB26B3437950" blockId="11.[264,1325,359,1384]" pageId="11" pageNumber="12">
<keyLead id="30F095C8FFDBFFE2FEFBFB26B3437950" pageId="11" pageNumber="12">
­ Anterior pronotal margin not forming any median point or angle; elytral striae with fine, very transverse punctures which at most only slightly crenate the edges of the channel; protibial apical spurs pointed. Michoacán 
<emphasis id="B975EAEFFFDBFFE2FC7FFA97B2737950" box="[908,1051,1359,1384]" italics="true" pageId="11" pageNumber="12">
.. 
<taxonomicName id="4C014D7EFFDBFFE2FC6CFA97B2737950" ID-CoL="7ZMKN" authority="Matthews and Halffter" authorityName="Matthews and Halffter" box="[927,1051,1359,1384]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mexicanus">mexicanus</taxonomicName>
</emphasis>
Matthews and Halffter
</keyLead>
</paragraph>
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